144 lines
20 KiB
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144 lines
20 KiB
XML
<document id="5FE287C5ADF0804A32AA09B828581C4D" ID-DOI="10.11646/zootaxa.2135.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5321522" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629624871390" checkinUser="felipe" docAuthor="Stiller, M." docDate="2009" docId="03BD0C4BFF99D15BFF46F92FFAB82EF3" docLanguage="en" docName="zt02135p056.pdf" docOrigin="Zootaxa 2135 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2135.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Theronus Stiller, 2009, gen.n." docType="treatment" docVersion="5" lastPageNumber="34" masterDocId="FF847433FFB9D17AFFD1FFE6FFD02D45" masterDocTitle="Revision of Elginus Theron (Hemiptera: Cicadellidae: Deltocephalinae) with the description of two new genera and comments on the grassland leafhopper fauna in South Africa" masterLastPageNumber="56" masterPageNumber="1" pageNumber="33" updateTime="1699085644247" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="372A35CF924114230EC6A4FBCE813909">Revision of Elginus Theron (Hemiptera: Cicadellidae: Deltocephalinae) with the description of two new genera and comments on the grassland leafhopper fauna in South Africa</mods:title>
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<mods:namePart id="7A95FE3BE45154DC6E55625ACE32B3D8">Stiller, M.</mods:namePart>
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<mods:title id="A0753AF36C66D8E8E04FC9D666521787">Zootaxa</mods:title>
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<mods:part id="FDB395C7EC3B8DAE15F9F2CD424D5598">
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<mods:date id="551AACD11ECE0BBE4B54E4A0725DC3D9">2009</mods:date>
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<mods:number id="19881FC7415E25C07731197171082B2A">2009-06-18</mods:number>
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<mods:number id="D37A282F2809B04C146DD315A9D9D429">2135</mods:number>
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<mods:url id="96A2D6825CCE7461838432C9A9601772">https://biotaxa.org/Zootaxa/article/view/zootaxa.2135.1.1</mods:url>
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<mods:identifier id="BCEE328457CD160F301DACC5F05F1F87" type="DOI">10.11646/zootaxa.2135.1.1</mods:identifier>
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<mods:identifier id="18BE61A73781AC005748BBBBD1DE7D7F" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BD0C4BFF99D15BFF46F92FFAB82EF3" ID-DOI="http://doi.org/10.5281/zenodo.5334114" ID-Zenodo-Dep="5334114" LSID="urn:lsid:plazi:treatment:03BD0C4BFF99D15BFF46F92FFAB82EF3" httpUri="http://treatment.plazi.org/id/03BD0C4BFF99D15BFF46F92FFAB82EF3" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
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<subSubSection id="C30EEED6FF99D15AFF46F92FFE8E2BA7" box="[151,350,1736,1763]" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="8BABBD5DFF99D15AFF46F92FFE8E2BA7" blockId="32.[151,350,1736,1763]" box="[151,350,1736,1763]" pageId="32" pageNumber="33">
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<heading id="D0E30A31FF99D15AFF46F92FFE8E2BA7" bold="true" box="[151,350,1736,1763]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
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<emphasis id="B960614FFF99D15AFF46F92FFE8E2BA7" bold="true" box="[151,350,1736,1763]" pageId="32" pageNumber="33">
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<taxonomicName id="4C14C6DEFF99D15AFF46F92FFEDC2BA6" ID-CoL="8RGTJ" authorityName="Stiller" authorityYear="2009" box="[151,268,1737,1763]" class="Insecta" family="Cicadellidae" genus="Theronus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B960614FFF99D15AFF46F92FFEDC2BA6" bold="true" box="[151,268,1737,1763]" italics="true" pageId="32" pageNumber="33">Theronus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A253DC34FF99D15AFEC2F92EFE8E2BA7" box="[275,350,1736,1762]" pageId="32" pageNumber="33" rank="genus">gen.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30EEED6FF99D15AFF46F8F4FE8F2A0C" pageId="32" pageNumber="33" type="type_taxon">
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<paragraph id="8BABBD5DFF99D15AFF46F8F4FCFB2A6C" blockId="32.[151,811,1810,1865]" box="[151,811,1810,1834]" pageId="32" pageNumber="33">
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<emphasis id="B960614FFF99D15AFF46F8F4FEF32A6F" bold="true" box="[151,291,1810,1834]" pageId="32" pageNumber="33">
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<typeStatus id="54AF03FFFF99D15AFF46F8F4FF1F2A6F" box="[151,207,1810,1834]" pageId="32" pageNumber="33">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="4C14C6DEFF99D15AFEE3F8F4FE3D2A6C" ID-CoL="8RP3H" box="[306,493,1810,1833]" class="Insecta" family="Cicadellidae" genus="Theronus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="priapus" status="sp. nov.">
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<emphasis id="B960614FFF99D15AFEE3F8F4FE3D2A6C" box="[306,493,1810,1833]" italics="true" pageId="32" pageNumber="33">Theronus priapus</emphasis>
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</taxonomicName>
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<emphasis id="B960614FFF99D15AFE25F8F4FDF92A6F" bold="true" box="[500,553,1810,1834]" pageId="32" pageNumber="33">
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<taxonomicNameLabel id="A253DC34FF99D15AFE25F8F4FDF92A6F" box="[500,553,1810,1834]" pageId="32" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="132FA1D8FF99D15AFDE9F8F4FD622A6C" box="[568,690,1810,1833]" captionStart="FIGURES 47–50" captionStartId="44.[151,272,1235,1259]" captionTargetBox="[165,1429,198,1170]" captionTargetId="figure-62@44.[132,1462,148,1243]" captionTargetPageId="44" captionText="FIGURES 47–50. Theronus priapus and Lecacis species, habitus views. Fig. 47, T. priapus sp.n., male; Fig. 48, T. priapus, female brachypter; Fig. 49, T. priapus, female macropter; Fig. 50, L. platypennis Theron, male." figureDoi="http://doi.org/10.5281/zenodo.5321540" httpUri="https://zenodo.org/record/5321540/files/figure.png" pageId="32" pageNumber="33">Figs 47–49</figureCitation>
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,
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<figureCitation id="132FA1D8FF99D15AFD6FF8F4FCF02A6C" box="[702,800,1810,1833]" captionStart="FIGURES 284–290" captionStartId="55.[151,271,1874,1898]" captionTargetBox="[260,1328,188,1820]" captionTargetId="figure-65@55.[208,1377,138,1856]" captionTargetPageId="55" captionText="FIGURES 284–290. Theronus priapus. Fig. 284, aedeagus, ventral view; Fig. 285, aedeagus, lateral view; Fig. 286, plate and valve; Fig. 287, apex of abdomen, dorsal view, depicting position of genitalia; Fig. 288, style; Fig. 289, sternite VII, female; Fig. 290, apex of abdomen, lateral view, depicting position of genitalia." figureDoi="http://doi.org/10.5281/zenodo.5363535" httpUri="https://zenodo.org/record/5363535/files/figure.png" pageId="32" pageNumber="33">284–290</figureCitation>
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).
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</paragraph>
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<paragraph id="8BABBD5DFF99D15AFF46F8D4FE8F2A0C" blockId="32.[151,811,1810,1865]" box="[151,351,1842,1865]" pageId="32" pageNumber="33">Gender masculine.</paragraph>
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</subSubSection>
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<subSubSection id="C30EEED6FF99D15BFF46F89EFAB82EF3" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="etymology">
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<paragraph id="8BABBD5DFF99D15AFF46F89EFEF22AFF" blockId="32.[151,1436,1912,2016]" pageId="32" pageNumber="33">
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<emphasis id="B960614FFF99D15AFF46F89EFECF2AD7" bold="true" box="[151,287,1912,1938]" pageId="32" pageNumber="33">Etymology</emphasis>
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: Named for the collector, J.G. Theron, who published taxonomic papers on leafhoppers between 1970–1988.
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</paragraph>
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<paragraph id="8BABBD5DFF99D15BFF14F820FC7D2C36" blockId="32.[151,1436,1912,2016]" lastBlockId="33.[151,1436,152,951]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
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<emphasis id="B960614FFF99D15AFF14F820FE022AA5" bold="true" box="[197,466,1990,2016]" pageId="32" pageNumber="33">External morphology</emphasis>
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. Vertex with disc smooth, anterior margin shagreened. Clypeus light brown with ochraceous horizontal arcs. Clypellus broad, 1.3 times as long as wide, tapering apically. Pronotum without carina. Submacropter with tegmina with three anteapical cells, four narrow apical cells, reduced appendix; hind wing about half as long as tegmina, jugal lobe present, folded over anal area. Macropters rare, with hind wing as long as tegmina. Profemur with short, stout setae in row AV, about 8 long, fine intercalary setae, one AM1 seta. Protibial setal formula 1+4; mesotibial setal formula 4+4; metafemur setal formula 2+2+1, with median pair consisting of one robust, long seta, one fine, short seta.
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</paragraph>
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<paragraph id="8BABBD5DFF98D15BFF14FE99FEDE2F71" blockId="33.[151,1436,152,951]" pageId="33" pageNumber="34">
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<emphasis id="B960614FFF98D15BFF14FE99FE562CDC" bold="true" box="[197,390,383,409]" pageId="33" pageNumber="34">Genital capsule</emphasis>
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.
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<emphasis id="B960614FFF98D15BFE44FE99FE042CDC" bold="true" box="[405,468,383,409]" pageId="33" pageNumber="34">Male</emphasis>
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. Aedeagus extremely long, narrow, flattened laterally; extending into tergite VIII, sometimes into tergite VII; gonopore subapical, dorsal (
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<figureCitation id="132FA1D8FF98D15BFCE4FE40FC302C85" box="[821,992,422,448]" captionStart="FIGURES 284–290" captionStartId="55.[151,271,1874,1898]" captionTargetBox="[260,1328,188,1820]" captionTargetId="figure-65@55.[208,1377,138,1856]" captionTargetPageId="55" captionText="FIGURES 284–290. Theronus priapus. Fig. 284, aedeagus, ventral view; Fig. 285, aedeagus, lateral view; Fig. 286, plate and valve; Fig. 287, apex of abdomen, dorsal view, depicting position of genitalia; Fig. 288, style; Fig. 289, sternite VII, female; Fig. 290, apex of abdomen, lateral view, depicting position of genitalia." figureDoi="http://doi.org/10.5281/zenodo.5363535" httpUri="https://zenodo.org/record/5363535/files/figure.png" pageId="33" pageNumber="34">Figs 284, 285</figureCitation>
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). Connective linear, arms fused (
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<figureCitation id="132FA1D8FF98D15BFABDFE40FF182CA2" captionStart="FIGURES 284–290" captionStartId="55.[151,271,1874,1898]" captionTargetBox="[260,1328,188,1820]" captionTargetId="figure-65@55.[208,1377,138,1856]" captionTargetPageId="55" captionText="FIGURES 284–290. Theronus priapus. Fig. 284, aedeagus, ventral view; Fig. 285, aedeagus, lateral view; Fig. 286, plate and valve; Fig. 287, apex of abdomen, dorsal view, depicting position of genitalia; Fig. 288, style; Fig. 289, sternite VII, female; Fig. 290, apex of abdomen, lateral view, depicting position of genitalia." figureDoi="http://doi.org/10.5281/zenodo.5363535" httpUri="https://zenodo.org/record/5363535/files/figure.png" pageId="33" pageNumber="34">Fig. 284</figureCitation>
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). Plate about half length of pygofer; apex denticulate (
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<figureCitation id="132FA1D8FF98D15BFC89FE2BFC122CA2" box="[856,962,461,487]" captionStart="FIGURES 284–290" captionStartId="55.[151,271,1874,1898]" captionTargetBox="[260,1328,188,1820]" captionTargetId="figure-65@55.[208,1377,138,1856]" captionTargetPageId="55" captionText="FIGURES 284–290. Theronus priapus. Fig. 284, aedeagus, ventral view; Fig. 285, aedeagus, lateral view; Fig. 286, plate and valve; Fig. 287, apex of abdomen, dorsal view, depicting position of genitalia; Fig. 288, style; Fig. 289, sternite VII, female; Fig. 290, apex of abdomen, lateral view, depicting position of genitalia." figureDoi="http://doi.org/10.5281/zenodo.5363535" httpUri="https://zenodo.org/record/5363535/files/figure.png" pageId="33" pageNumber="34">Fig. 286</figureCitation>
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). Anal tube superficial, not invaginated into pygofer. Pygofer lobe setose, posterior margin uniformly rounded; in dorsal view acutely triangular (
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<figureCitation id="132FA1D8FF98D15BFAB9FE12FED22F71" captionStart="FIGURES 284–290" captionStartId="55.[151,271,1874,1898]" captionTargetBox="[260,1328,188,1820]" captionTargetId="figure-65@55.[208,1377,138,1856]" captionTargetPageId="55" captionText="FIGURES 284–290. Theronus priapus. Fig. 284, aedeagus, ventral view; Fig. 285, aedeagus, lateral view; Fig. 286, plate and valve; Fig. 287, apex of abdomen, dorsal view, depicting position of genitalia; Fig. 288, style; Fig. 289, sternite VII, female; Fig. 290, apex of abdomen, lateral view, depicting position of genitalia." figureDoi="http://doi.org/10.5281/zenodo.5363535" httpUri="https://zenodo.org/record/5363535/files/figure.png" pageId="33" pageNumber="34">Figs 287, 290</figureCitation>
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).
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</paragraph>
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<paragraph id="8BABBD5DFF98D15BFF14FDA6FAD52F1E" blockId="33.[151,1436,152,951]" box="[197,1285,576,603]" pageId="33" pageNumber="34">
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<emphasis id="B960614FFF98D15BFF14FDA6FE562F1F" bold="true" box="[197,390,576,602]" pageId="33" pageNumber="34">Genital capsule</emphasis>
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.
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<emphasis id="B960614FFF98D15BFE45FDA6FE3F2F1F" bold="true" box="[404,495,576,602]" pageId="33" pageNumber="34">Female</emphasis>
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. Pygofer with numerous macrosetae, ovipositor as long as pygofer.
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</paragraph>
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<paragraph id="8BABBD5DFF98D15BFF14FD81FAB82EF3" blockId="33.[151,1436,152,951]" pageId="33" pageNumber="34">
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<emphasis id="B960614FFF98D15BFF14FD81FEA12FC4" bold="true" box="[197,369,615,641]" pageId="33" pageNumber="34">Relationships</emphasis>
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. This genus is a typical member of the tribe
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<taxonomicName id="4C14C6DEFF98D15BFC5CFD8EFBEC2FC7" authorityName="Dallas" authorityYear="1870" box="[909,1084,616,642]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="tribe" tribe="Deltocephalini">Deltocephalini</taxonomicName>
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that is defined by having the connective fused to the aedeagus (
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<bibRefCitation id="EF85C0ACFF98D15BFDFAFD68FCB72FED" author="Zahniser & Dietrich" box="[555,871,654,680]" firstAuthor="Zahniser" pageId="33" pageNumber="34" pagination="1 - 24" refId="ref38641" refString="Zahniser, J. N. & Dietrich, C. H. (2008) Phylogeny of the leafhopper subfamily Deltocephalinae (Insecta: Auchenorrhyncha: Cicadellidae) and related subfamilies based on morphology. Systematics and Biodiversity, 6 (1), 1 - 24." type="journal article" year="2008">Zahniser & Dietrich, 2008</bibRefCitation>
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;
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<bibRefCitation id="EF85C0ACFF98D15BFCA5FD68FBB52FED" author="Ossiannilsson" box="[884,1125,654,680]" firstAuthor="Ossiannilsson" pageId="33" pageNumber="34" pagination="594 - 978" refId="ref38021" refString="Ossiannilsson, F. (1983) The Auchenorrhyncha (Homoptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, 7 (3), 594 - 978." type="journal article" year="1983">Ossiannilsson, 1983</bibRefCitation>
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). It is characterized by the male genitalia that are so large that they extend into abdominal segment VIII or even VII, and have the aedeagus blade-like and curved ventroposteriorly. This genus is not closely related to any Afrotropical genus in this tribe such as
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<taxonomicName id="4C14C6DEFF98D15BFEAFFCE4FD362E59" authority="Edwards. Occasionally" authorityName="Edwards. Occasionally" authorityYear="1922" box="[382,742,770,796]" class="Insecta" family="Cicadellidae" genus="Recilia" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis id="B960614FFF98D15BFEAFFCE4FE012E5E" box="[382,465,770,795]" italics="true" pageId="33" pageNumber="34">Recilia</emphasis>
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Edwards. Occasionally
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</taxonomicName>
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, however, similar colouration is found in pale specimens of species of genera such as
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<taxonomicName id="4C14C6DEFF98D15BFE0BFCCFFDB42E07" authorityName="Theron (Theron" authorityYear="1975" box="[474,612,809,834]" class="Insecta" family="Cicadellidae" genus="Pravistylus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis id="B960614FFF98D15BFE0BFCCFFDB42E07" box="[474,612,809,834]" italics="true" pageId="33" pageNumber="34">Pravistylus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C14C6DEFF98D15BFD73FCCFFCD82E07" authorityName="Theron. The" authorityYear="1975" box="[674,776,809,834]" class="Insecta" family="Cicadellidae" genus="Vilargus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis id="B960614FFF98D15BFD73FCCFFCD82E07" box="[674,776,809,834]" italics="true" pageId="33" pageNumber="34">Vilargus</emphasis>
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</taxonomicName>
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that are in the tribe
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<taxonomicName id="4C14C6DEFF98D15BFBD8FCCFFB4D2E06" authorityName="Distant" authorityYear="1908" box="[1033,1181,809,835]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="tribe" tribe="Paralimnini">Paralimnini</taxonomicName>
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. No genera from the Palaearctic Region bearing any genitalic resemblance were listed by
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<bibRefCitation id="EF85C0ACFF98D15BFC1AFCB6FB162E2F" author="Ossiannilsson" box="[971,1222,848,874]" firstAuthor="Ossiannilsson" pageId="33" pageNumber="34" pagination="594 - 978" refId="ref38021" refString="Ossiannilsson, F. (1983) The Auchenorrhyncha (Homoptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, 7 (3), 594 - 978." type="journal article" year="1983">Ossiannilsson (1983)</bibRefCitation>
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or
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<bibRefCitation id="EF85C0ACFF98D15BFB3EFCB6FA472E2F" author="Ribaut" box="[1263,1431,848,874]" firstAuthor="Ribaut" pageId="33" pageNumber="34" pagination="1 - 474" refId="ref38051" refString="Ribaut, H. (1952) Homopteres Auchenorhynques. II (Jassidae). Faune de France, 57, 1 - 474." type="journal article" year="1952">Ribaut (1952)</bibRefCitation>
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, although some bear similar external features, such as colour and shape.
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||
<bibRefCitation id="EF85C0ACFF98D15BFC0BFC90FB602ED5" author="Linnavuori" box="[986,1200,886,912]" firstAuthor="Linnavuori" pageId="33" pageNumber="34" pagination="1 - 370" refId="ref37750" refString="Linnavuori, R. (1959) Revision of the Neotropical Deltocephalinae and some related subfamilies (Homoptera). Annales Zoologici Societatis Zoologicae Botanicae Fennicae ' Vanamo', 20 (1), 1 - 370." type="journal article" year="1959">Linnavuori (1959)</bibRefCitation>
|
||
described a number of genera from the Nearctic Region in the
|
||
<taxonomicName id="4C14C6DEFF98D15BFD56FC7BFCE62EF2" authorityName="Dallas" authorityYear="1870" box="[647,822,925,951]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="tribe" tribe="Deltocephalini">Deltocephalini</taxonomicName>
|
||
, but none show any relationship with
|
||
<emphasis id="B960614FFF98D15BFB22FC7BFAB82EF3" box="[1267,1384,925,950]" italics="true" pageId="33" pageNumber="34">
|
||
<taxonomicName id="4C14C6DEFF98D15BFB22FC7BFAB42EF3" authorityName="Stiller" authorityYear="2009" box="[1267,1380,925,950]" class="Insecta" family="Cicadellidae" genus="Theronus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Theronus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |