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<document id="A2B871E951A0B204681DE2099CBC0BBB" ID-CLB-Dataset="27980" ID-DOI="10.11646/zootaxa.4514.4.4" ID-GBIF-Dataset="7e5b5624-cb34-457f-b588-2f20485c3de5" ID-ISSN="1175-5326" ID-Zenodo-Dep="2609336" ID-ZooBank="E5B09509-BF8A-49A5-8CF1-C7B8F411E039" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553622768940" checkinUser="plazi" docAuthor="Shobharani, M., C. A. Viraktamath &amp; Webb, M. D." docDate="2018" docId="03B9270D8B2BEC65B5EF4433FE4D4059" docLanguage="en" docName="zootaxa.4514.4.4.pdf" docOrigin="Zootaxa 4514 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tambila variabilis" docType="treatment" docVersion="8" lastPageNumber="506" masterDocId="FF805F758B2FEC60B578436CFFB5440F" masterDocTitle="Revision of the penthimiine leafhopper genus Tambila Distant (Hemiptera: Cicadellidae: Deltocephalinae) of the Indian subcontinent" masterLastPageNumber="515" masterPageNumber="501" pageNumber="505" updateTime="1698713540995" updateUser="ExternalLinkService">
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<mods:title id="119B4A7230D891D85334793247E9AC66">Revision of the penthimiine leafhopper genus Tambila Distant (Hemiptera: Cicadellidae: Deltocephalinae) of the Indian subcontinent</mods:title>
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<mods:namePart id="598A4D692C540ECA6A1A19FA7AE5D975">C. A. Viraktamath</mods:namePart>
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<mods:date id="D4774ADCA68A8B962F1ABD641F59880E">2018</mods:date>
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<treatment id="03B9270D8B2BEC65B5EF4433FE4D4059" ID-DOI="http://doi.org/10.5281/zenodo.5967615" ID-GBIF-Taxon="154498435" ID-Zenodo-Dep="5967615" LSID="urn:lsid:plazi:treatment:03B9270D8B2BEC65B5EF4433FE4D4059" httpUri="http://treatment.plazi.org/id/03B9270D8B2BEC65B5EF4433FE4D4059" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505">
<subSubSection id="C30AC5908B2BEC64B5EF4433FE524376" box="[151,487,1887,1913]" pageId="4" pageNumber="505" type="nomenclature">
<paragraph id="8BAF961B8B2BEC64B5EF4433FE524376" blockId="4.[151,487,1887,1946]" box="[151,487,1887,1913]" pageId="4" pageNumber="505">
<heading id="D0E721778B2BEC64B5EF4433FE524376" bold="true" box="[151,487,1887,1913]" fontSize="11" level="1" pageId="4" pageNumber="505" reason="1">
<taxonomicName id="4C10ED988B2BEC64B5EF4433FE524376" ID-CoL="8JZHY" authority="(Distant)" baseAuthorityName="Distant" box="[151,487,1887,1913]" class="Insecta" family="Cicadellidae" genus="Tambila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B9644A098B2BEC64B5EF4433FE524376" bold="true" box="[151,487,1887,1913]" pageId="4" pageNumber="505">
<emphasis id="B9644A098B2BEC64B5EF4433FEC54376" bold="true" box="[151,368,1887,1913]" italics="true" pageId="4" pageNumber="505">Tambila variabilis</emphasis>
(Distant)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30AC5908B2BEC64B5EF44EDFEC54395" box="[151,368,1921,1946]" pageId="4" pageNumber="505" type="description">
<paragraph id="8BAF961B8B2BEC64B5EF44EDFEC54395" blockId="4.[151,487,1887,1946]" box="[151,368,1921,1946]" pageId="4" pageNumber="505">
<figureCitation id="132B8A9E8B2BEC64B5EF44EDFEA24395" box="[151,279,1921,1946]" captionStart="FIGURES 1222" captionStartId="7.[151,264,1907,1930]" captionTargetBox="[249,1385,219,1797]" captionTargetId="figure@7.[151,1436,193,1906]" captionTargetPageId="7" captionText="FIGURES 1222. Species of Tambila dorsal view. 1220, T. majuscula. 1213: holotype male (12) and topotype female (13); 14, T. gravelyi, lectotype male dorsal view; 1520, colour variation in T. majuscula: specimens from Dharmasthala female (15), Meppadi male (16), Jog Falls (18), Thekkady male (18), Bangalore male (1920) dorsal and lateral view respectively; 2122, T. variabilis, male and female habitus, respectively." httpUri="https://zenodo.org/record/2609340/files/figure.png" pageId="4" pageNumber="505">Figs 2122</figureCitation>
,
<figureCitation id="132B8A9E8B2BEC64B45A44EEFEDE4395" box="[290,363,1922,1946]" captionStart="FIGURES 8090" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[243,1400,306,1780]" captionTargetId="figure@13.[151,1436,193,1906]" captionTargetPageId="13" captionText="FIGURES 8090. Tambila variabilis abdominal terminalia: 8081, male anal segments, dorsal and lateral views, respectively; 82, subgenital plate ventral view; 83, styles and connective, dorsal view; 84, style, dorsal view; 85, pygofer, lateral view; 86, connective, dorsal view; 8788, aedeagus, lateral view; 89, aedeagus, ventral view; 90, female sternite VII, ventral view. Topotypic male (8089) and female (90)." httpUri="https://zenodo.org/record/2609352/files/figure.png" pageId="4" pageNumber="505">8090</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30AC5908B2BEC65B5EF44A5FEC54550" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505" type="reference_group">
<paragraph id="8BAF961B8B2BEC65B5EF44A5FCE044A1" blockId="4.[151,534,1993,2015]" lastBlockId="5.[151,853,152,174]" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505">
<treatmentCitationGroup id="AB00B1358B2BEC65B5EF44A5FCE044A1" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505">
<taxonomicName id="4C10ED988B2BEC64B5EF44A5FDA743D0" ID-CoL="8JBTG" authority="Distant 1918: 18" authorityName="Distant" authorityPageNumber="18" authorityYear="1918" box="[151,530,1993,2015]" class="Insecta" family="Cicadellidae" genus="Penthimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B9644A098B2BEC64B5EF44A5FED343D0" box="[151,358,1993,2015]" italics="true" pageId="4" pageNumber="505">Penthimia variabilis</emphasis>
<treatmentCitation id="0AB1B00A8B2BEC64B41444A5FDA743D0" author="Distant, W. L." box="[364,530,1993,2015]" page="18" pageId="4" pageNumber="505" year="1918">
<bibRefCitation id="EF81EBEA8B2BEC64B41444A5FE5943D0" author="Distant, W. L." box="[364,492,1993,2015]" pageId="4" pageNumber="505" pagination="1 - 210" refId="ref4944" refString="Distant, W. L. (1918) Rhynchota Homoptera: Appendix. Heteroptera: Addenda. In: Shipley, A. E. &amp; Marshall, G. A. K. (Eds.), The Fauna of British India including Ceylon and Burma. Uol. 7. Taylor &amp; Francis, London, pp. i-viii + 1 - 210." type="book chapter" year="1918">Distant1918</bibRefCitation>
: 18
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="4C10ED988B2AEC65B5EF43F4FE0744A1" ID-CoL="8JZHY" authority="(Distant)" baseAuthorityName="Distant" box="[151,434,152,174]" class="Insecta" family="Cicadellidae" genus="Tambila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B9644A098B2AEC65B5EF43F4FEE544A1" box="[151,336,152,174]" italics="true" pageId="5" pageNumber="506">Tambila variabilis</emphasis>
(Distant)
</taxonomicName>
,
<bibRefCitation id="EF81EBEA8B2AEC65B4C543F4FCE444A1" author="Shobharani, M. &amp; Viraktamath, C. A. &amp; Webb, M. D." box="[445,849,152,174]" pageId="5" pageNumber="506" pagination="1 - 45" refId="ref5064" refString="Shobharani, M., Viraktamath, C. A. &amp; Webb, M. D. (2018) Review of the leafhopper genus Penthimia Germar (Hemiptera: Cicadellidae) from the Indian subcontinent with description of seven new species. Zootaxa, 4369 (1), 1 - 45. https: // doi. org / 10.11646 / zootaxa. 4369.1.1" type="journal article" year="2018">Shobharani, Viraktamath &amp; Webb 2018</bibRefCitation>
.
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</paragraph>
<paragraph id="8BAF961B8B2AEC65B5EF43B6FEC54550" blockId="5.[151,1437,218,1110]" pageId="5" pageNumber="506">Dark yellowish brown to pale yellowish brown, irregularly mottled, suffused with black. Pronotum brownish yellow with median pale yellow irregular band. Scutellum dull brownish yellow, with dark spot near each basal angle, two small dark spots on apical area. Face pale yellow, basal half mottled with black. Forewing pale brown, mottled with black.</paragraph>
</subSubSection>
<subSubSection id="C30AC5908B2AEC65B5BF4206FAA246AD" pageId="5" pageNumber="506" type="description">
<paragraph id="8BAF961B8B2AEC65B5BF4206FB58461C" blockId="5.[151,1437,218,1110]" pageId="5" pageNumber="506">
<emphasis id="B9644A098B2AEC65B5BF4206FECF458C" bold="true" box="[199,378,362,387]" pageId="5" pageNumber="506">Male genitalia.</emphasis>
Pygofer lobes rounded posteriorly with stout setae ventrally. Xth segment weakly produced anteriorly (
<figureCitation id="132B8A9E8B2AEC65B46B42E2FED645A8" box="[275,355,398,423]" captionStart="FIGURES 8090" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[243,1400,306,1780]" captionTargetId="figure@13.[151,1436,193,1906]" captionTargetPageId="13" captionText="FIGURES 8090. Tambila variabilis abdominal terminalia: 8081, male anal segments, dorsal and lateral views, respectively; 82, subgenital plate ventral view; 83, styles and connective, dorsal view; 84, style, dorsal view; 85, pygofer, lateral view; 86, connective, dorsal view; 8788, aedeagus, lateral view; 89, aedeagus, ventral view; 90, female sternite VII, ventral view. Topotypic male (8089) and female (90)." httpUri="https://zenodo.org/record/2609352/files/figure.png" pageId="5" pageNumber="506">Fig. 81</figureCitation>
) Subgenital plate obtusely produced on lateral margin near base with apex rounded, two rows of stout setae on lateral margin. Style with well developed preapical lobe, apophysis long curved laterally with median protuberance laterally. Connective short, with arms as long as stem. Aedeagus simple, dorsal apodeme well developed, preatrium reduced, shaft curved posterodorsally with large gonopore on ventral margin.
</paragraph>
<paragraph id="8BAF961B8B2AEC65B5EF4172FDAE4670" blockId="5.[151,1437,218,1110]" pageId="5" pageNumber="506">
<emphasis id="B9644A098B2AEC65B5EF4172FED64638" bold="true" box="[151,355,542,567]" pageId="5" pageNumber="506">Female genitalia.</emphasis>
Seventh sternite longer medially than sixth, lateral angles of posterior margin rounded, medially produced without median concavity, area adjacent to median projection shallowly concave, median projection longer than lateral rounded angles.
</paragraph>
<paragraph id="8BAF961B8B2AEC65B5BF41E6FAA246AD" blockId="5.[151,1437,218,1110]" box="[199,1303,650,675]" pageId="5" pageNumber="506">
<emphasis id="B9644A098B2AEC65B5BF41E6FEC846AC" bold="true" box="[199,381,650,675]" pageId="5" pageNumber="506">Measurements.</emphasis>
Male
<quantity id="4CE83BFE8B2AEC65B4BD41E6FDFE46AD" box="[453,587,650,675]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.7" metricValueMin="4.6" pageId="5" pageNumber="506" unit="mm" value="4.65" valueMax="4.7" valueMin="4.6">4.64.7 mm</quantity>
long,
<quantity id="4CE83BFE8B2AEC65B7EA41E7FD5D46AD" box="[658,744,651,675]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="506" unit="mm" value="1.7">1.7 mm</quantity>
wide across head,
<quantity id="4CE83BFE8B2AEC65B6C441E7FBA746AD" box="[956,1042,651,675]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="506" unit="mm" value="1.9">1.9 mm</quantity>
wide across pronotum.
</paragraph>
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<emphasis id="B9644A098B2AEC65B5EF41C2FE3046C8" bold="true" box="[151,389,686,711]" pageId="5" pageNumber="506">Material examined.</emphasis>
<collectingCountry id="F307D68B8B2AEC65B4E841C3FE5146C8" box="[400,484,687,711]" name="India" pageId="5" pageNumber="506">INDIA</collectingCountry>
: Kodaikanal:
<materialsCitation id="3B789C468B2AEC65B7F041C3FB8746E4" ID-GBIF-Occurrence="2028438437" collectingDate="1914" collectingDateMax="1918" collectingDateMin="1914" collectionCode="BMNH" collectorName="Campbell" country="India" pageId="5" pageNumber="506" specimenCount="23" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D165D928B2AEC65B7F041C3FC9146C8" box="[648,804,686,711]" pageId="5" pageNumber="506" type="generic">21 specimens</specimenCount>
(
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and
<specimenCount id="9D165D928B2AEC65B6F641C3FC1446C8" box="[910,929,687,711]" pageId="5" pageNumber="506" type="female"></specimenCount>
), various months and years
<collectingDate id="EFEA49338B2AEC65B18E41C3FAC146C8" box="[1270,1396,687,711]" pageId="5" pageNumber="506" value="1914" valueMax="1918">
<date id="FFAEB0DB8B2AEC65B18E41C3FAC146C8" box="[1270,1396,687,711]" pageId="5" pageNumber="506" value="1914" valueMax="1918">19141918</date>
</collectingDate>
on reverse of specimen mount (e.g., 5.17), T.V.
<collectorName id="26E5F3CD8B2AEC65B7DD41BEFCA046E4" box="[677,789,722,747]" pageId="5" pageNumber="506">Campbell</collectorName>
, BM 1928 478 (
<collectionCode id="ED010EDE8B2AEC65B6A841BFFB9F46E4" box="[976,1066,723,747]" pageId="5" pageNumber="506">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B789C468B2AEC65B14741BFFA974700" ID-GBIF-Occurrence="2028438412" collectionCode="TNAU" collectorName="Pulney Hills &amp; May &amp; T. V. Campbell" country="India" elevation="1372" pageId="5" pageNumber="506" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D165D928B2AEC65B14741BFFBD446E4" box="[1087,1121,723,747]" pageId="5" pageNumber="506" type="male">2♂</specimenCount>
<collectorName id="26E5F3CD8B2AEC65B11241BEFB4C46E4" box="[1130,1273,722,747]" pageId="5" pageNumber="506">Pulney Hills</collectorName>
,
<elevation id="003D71288B2AEC65B07E41BFFF714700" metricMagnitude="3" metricUnit="m" metricValue="1.3716" metricValueMax="1.8288" metricValueMin="0.9144" pageId="5" pageNumber="506" unit="ft" value="4500.0" valueMax="6000.0" valueMin="3000.0">
<quantity id="4CE83BFE8B2AEC65B07E41BFFF714700" metricMagnitude="3" metricUnit="m" metricValue="1.3716" metricValueMax="1.8288" metricValueMin="0.9144" pageId="5" pageNumber="506" unit="ft" value="4500.0" valueMax="6000.0" valueMin="3000.0">3000 to 6000 feet</quantity>
</elevation>
,
<collectorName id="26E5F3CD8B2AEC65B5A8419BFEB14700" box="[208,260,759,783]" pageId="5" pageNumber="506">May</collectorName>
, 1917,
<collectorName id="26E5F3CD8B2AEC65B421419BFDB24700" box="[345,519,758,783]" pageId="5" pageNumber="506">T. V. Campbell</collectorName>
,
<taxonomicName id="4C10ED988B2AEC65B76C419AFCD24700" authority="(Distant)" baseAuthorityName="Distant" box="[532,871,758,783]" class="Insecta" family="Cicadellidae" genus="Penthimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B9644A098B2AEC65B76C419AFD4E4700" box="[532,763,758,783]" italics="true" pageId="5" pageNumber="506">Penthimia variabilis</emphasis>
(Distant)
</taxonomicName>
[Distant determination label] (
<collectionCode id="ED010EDE8B2AEC65B1BE419BFAAF4700" box="[1222,1306,759,783]" country="India" httpUri="http://grbio.org/cool/xxr9-fdbj" name="Tamil Nadu Agricultural University" pageId="5" pageNumber="506">TNAU</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D165D928B2AEC65B056419BFAE94701" box="[1326,1372,759,783]" pageId="5" pageNumber="506" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D165D928B2AEC65B011419BFA204701" box="[1385,1429,759,783]" pageId="5" pageNumber="506" type="female">1 ♀</specimenCount>
, Shambaganur,
<quantity id="4CE83BFE8B2AEC65B4454071FE21473A" box="[317,404,797,822]" metricMagnitude="3" metricUnit="m" metricValue="1.6949999999999998" pageId="5" pageNumber="506" unit="m" value="1695.0">1695 m</quantity>
, 10
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1458.5 N, 77
<superScript id="7C653B538B2AEC65B7104076FDDA4727" attach="none" box="[616,623,794,808]" fontSize="6" pageId="5" pageNumber="506">o</superScript>
3113.9E,
<date id="FFAEB0DB8B2AEC65B7964072FCD64739" box="[750,867,798,822]" pageId="5" pageNumber="506" value="2012-05-22">22.V.2012</date>
, Prathapan, K.D.;
<materialsCitation id="3B789C468B2AEC65B1494072FCCC4755" ID-GBIF-Occurrence="2028438434" collectingDate="2012-05-24" collectionCode="UASB" collectorName="Prathapan, K. D." country="India" elevation="1833" longitude="77.36086" pageId="5" pageNumber="506" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D165D928B2AEC65B1494072FBE04739" box="[1073,1109,798,822]" pageId="5" pageNumber="506" type="male">1♂</specimenCount>
,
<specimenCount id="9D165D928B2AEC65B11A4072FB39473A" box="[1122,1164,798,822]" pageId="5" pageNumber="506" type="female">1 ♀</specimenCount>
, Kukkal Shola,
<elevation id="003D71288B2AEC65B0384072FA22473A" box="[1344,1431,798,822]" metricMagnitude="3" metricUnit="m" metricValue="1.833" pageId="5" pageNumber="506" unit="m" value="1833.0">
<quantity id="4CE83BFE8B2AEC65B0384072FA22473A" box="[1344,1431,798,822]" metricMagnitude="3" metricUnit="m" metricValue="1.833" pageId="5" pageNumber="506" unit="m" value="1833.0">1833 m</quantity>
</elevation>
, 10o179.1N,
<geoCoordinate id="EE24F0DC8B2AEC65B44C402EFE604755" box="[308,469,834,858]" direction="east" orientation="longitude" pageId="5" pageNumber="506" precision="1" value="77.36086">77o2139.1”E</geoCoordinate>
,
<collectingDate id="EFEA49338B2AEC65B498402EFDE74755" box="[480,594,834,858]" pageId="5" pageNumber="506" value="2012-05-24">
<date id="FFAEB0DB8B2AEC65B498402EFDE74755" box="[480,594,834,858]" pageId="5" pageNumber="506" value="2012-05-24">24.v.2012</date>
</collectingDate>
,
<collectorName id="26E5F3CD8B2AEC65B725402DFCA44756" box="[605,785,833,858]" pageId="5" pageNumber="506">Prathapan, K.D.</collectorName>
(
<collectionCode id="ED010EDE8B2AEC65B658402EFCC44755" box="[800,881,834,858]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="5" pageNumber="506">UASB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30AC5908B2AEC65B5BF4009FE4D4059" pageId="5" pageNumber="506" type="discussion">
<paragraph id="8BAF961B8B2AEC65B5BF4009FE4D4059" blockId="5.[151,1437,218,1110]" pageId="5" pageNumber="506">
<emphasis id="B9644A098B2AEC65B5BF4009FE8E4771" bold="true" box="[199,315,869,894]" pageId="5" pageNumber="506">Remarks.</emphasis>
This species was described from an unknown number of specimens of both sexes (syntypic) with the following data: South
<collectingCountry id="F307D68B8B2AEC65B4B840E5FE4847AD" box="[448,509,905,930]" name="India" pageId="5" pageNumber="506">India</collectingCountry>
; Kodaikanal (T.V. Campbell). No specimen of this species, bearing Distants usual
<typeStatus id="54AB28B98B2AEC65B5EF40C2FF7247C9" box="[151,199,942,966]" pageId="5" pageNumber="506">type</typeStatus>
label, has been found in the BMNH. Although there are several topotypic specimens collected by T.V. Campbell in the BMNH (see above) they have a modern label and are not considered
<typeStatus id="54AB28B98B2AEC65B1F540BEFB7D47E5" box="[1165,1224,978,1002]" pageId="5" pageNumber="506">types</typeStatus>
(see Material and methods). However, two specimens (labeled Pulney Hills, Kodaikanal) found in TNAU agree with the description and do have Distants determination label. The basal lobe of the aedeagal shaft, present in both
<taxonomicName id="4C10ED988B2AEC65B1AE4776FAFA403D" authorityName="Distant" authorityYear="1918" box="[1238,1359,1049,1074]" class="Insecta" family="Cicadellidae" genus="Tambila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="B9644A098B2AEC65B1AE4776FAFA403D" box="[1238,1359,1049,1074]" italics="true" pageId="5" pageNumber="506">T. gravelyi</emphasis>
</taxonomicName>
and
<emphasis id="B9644A098B2AEC65B0FF4776FF51405A" italics="true" pageId="5" pageNumber="506">
<taxonomicName id="4C10ED988B2AEC65B0FF4776FF54405A" authorityName="Distant" authorityYear="1908" class="Insecta" family="Cicadellidae" genus="Tambila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="greeni">T. greeni</taxonomicName>
,
</emphasis>
is absent in this species.
</paragraph>
</subSubSection>
</treatment>
</document>