343 lines
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XML
343 lines
51 KiB
XML
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<mods:title id="379A29706AAE5DE4083D266A577C70C4">Revision of the Mesobuthus caucasicus complex from Central Asia with descriptions of six new species (Scorpiones Buthidae)</mods:title>
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<taxonomicName id="4C0B3AC77A556A7DFED458DEFDA1418C" authority="Fet & Kovařík & Gantenbein & Kaiser & Stewart & Graham, 2018" authorityName="Fet & Kovařík & Gantenbein & Kaiser & Stewart & Graham" authorityYear="2018" box="[343,577,689,716]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="brutus" status="sp. nov.">Mesobuthus brutus</taxonomicName>
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<figureCitation id="13305DC17A556A7DFF4958BCFEDD41AA" box="[202,317,723,746]" captionStart-0="Figures 1–4" captionStart-1="Figures 5–14" captionStart-2="Figures 15–20" captionStartId-0="5.[192,278,1533,1557]" captionStartId-1="6.[192,278,1635,1659]" captionStartId-2="7.[192,278,1637,1661]" captionTargetBox-0="[205,1422,185,1495]" captionTargetBox-1="[191,1440,191,1606]" captionTargetBox-2="[202,1432,206,1597]" captionTargetId-0="figure-6@5.[192,1437,168,1504]" captionTargetId-1="figure-11@6.[192,1439,192,1605]" captionTargetId-2="figure-6@7.[192,1439,195,1608]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="Figures 1–4: Mesobuthus brutus sp. n. Figures 1–2. Holotype male, dorsal (1) and ventral (2) views. Figures 3–4. Paratype female, dorsal (3) and ventral (4) views." captionText-1="Figures 5–14: Mesobuthus brutus sp. n. Figures 5, 7, 9–11. Holotype male, chelicerae, carapace and tergites I–III (5), sternopectinal region and sternite III (7), pedipalp chela, dorsal (9) and external (10), pedipalp patella dorsal (11). The trichobothrial pattern is indicated in Figures 9–10. Figures 6, 8, 12–14. Paratype female, chelicerae, carapace and tergites I–III (6), sternopectinal region and sternite III (8), pedipalp chela, dorsal (12) and external (13), pedipalp patella dorsal (14)." captionText-2="Figures 15–20: Mesobuthus brutus sp. n. Figures 15–17. Holotype male, metasoma and telson, lateral (15), ventral (16), and dorsal (17) views. Figures 18–20. Paratype female, metasoma and telson, lateral (18), ventral (19), and dorsal (20) views." pageId="8" pageNumber="7">Figs. 1–20</figureCitation>
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,
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<figureCitation id="13305DC17A556A7DFECA58BCFE4941AA" box="[329,425,723,746]" captionStart="Figures 181–188" captionStartId="39.[192,278,1168,1192]" captionTargetBox="[196,1430,203,1134]" captionTargetId="figure-59@39.[192,1437,192,1139]" captionTargetPageId="39" captionText="Figures 181–188: Figures 181–182: Mesobuthus kreuzbergi sp. n., metasoma V and telson lateral of male (181) and female (182). Figures 183–188: Mesobuthus brutus sp. n., metasoma V and telson lateral of male holotype (183) and female paratype (184), and left legs I–IV, retrolateral aspect of male holotype (185–188)." pageId="8" pageNumber="7">183–188</figureCitation>
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<figureCitation id="13305DC17A556A7DFE3058BCFE3E41AA" box="[435,478,723,746]" captionStart="Figures 265–273" captionStartId="56.[192,278,1569,1593]" captionTargetBox="[208,1427,176,1539]" captionTargetId="figure-49@56.[192,1439,168,1540]" captionTargetPageId="56" captionText="Figures 265–273: Mesobuthus, metasoma and telson ventral of males. Figure 265. M. brutus sp. n., holotype. Figure 266. M. caucasicus, Turkey. Figure 267. M. gorelovi sp. n., holotype. Figure 268. M. fuscus, Tajikistan. Figure 269. M. kaznakovi, Tajikistan. Figure 270. M. kreuzbergi sp. n., holotype. Figure 271. M. mischi sp. n., holotype. Figure 272. M. nenilini sp. n., holotype. Figure 273. M. parthorum, Afghanistan." pageId="8" pageNumber="7">265</figureCitation>
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<figureCitation id="13305DC17A556A7DFE6A58BCFDF441AA" box="[489,532,723,746]" captionStart="Figures 274–284" captionStartId="57.[192,278,1468,1492]" captionTargetBox="[196,1428,206,1431]" captionTargetId="figure-6@57.[192,1440,192,1439]" captionTargetPageId="57" captionText="Figures 274–284: Mesobuthus, tarsomeres of third leg. Figure 274. M. brutus sp. n., male holotype. Figure 275. M. caucasicus, female from Turkey. Figure 276. M. elenae sp. n., female holotype. Figure 277. M. gorelovi sp. n., female paratype from type locality. Figure 278. M. fuscus, female from Tajikistan. Figure 279. M. kaznakovi, male from Tajikistan. Figure 280. M. kreuzbergi sp. n., female paratype. Figure 281. M. mischi sp. n., male holotype. Figure 282. M. nenilini sp. n., female paratype. Figure 283–284. M. parthorum, male from Ghazni, Afghanistan (283) and female from Kholm, Afghanistan (284)." pageId="8" pageNumber="7">274</figureCitation>
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<figureCitation id="13305DC17A556A7DFD9D58BCFD9E41AA" box="[542,638,723,746]" captionStart="Figures 285–303" captionStartId="58.[192,278,1281,1305]" captionTargetBox="[198,1431,202,1246]" captionTargetId="figure-11@58.[192,1440,192,1252]" captionTargetPageId="58" captionText="Figures 285–303: Mesobuthus, pedipalp chela dorsal. Figure 285–286. M. brutus sp. n., male holotype (285) and female paratype (286). Figure 287–288. M. caucasicus, male (287) and female (288) from Turkey. Figure 289. M. elenae sp. n., female holotype. Figure 290–291. M. fuscus, differently colored male (290) and female (291) from Tajikistan. Figure 292–293. M. gorelovi sp. n., male holotype (292) and female paratype (293). Figure 294–295. M. kaznakovi, male (294) and female (295) from Tajikistan. Figure 296–297. M. kreuzbergi sp. n., male holotype (296) and female paratype (297). Figure 298–299. M. mischi sp. n., male holotype (298) and female paratype (299). Figure 300–301. M. nenilini sp. n., male holotype (300) and female paratype (301). Figure 302–303. M. parthorum, male (302) and female (303) from Afghanistan." pageId="8" pageNumber="7">285–286</figureCitation>
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<collectingCountry id="F31C01D47A556A7DFE1959A7FE2D40A0" box="[410,461,968,992]" name="Iran" pageId="8" pageNumber="7">Iran</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B97F9D567A556A7DFE6859A6FD4D40A0" box="[491,685,969,992]" italics="true" pageId="8" pageNumber="7">
|
||
<collectingRegion id="49CF8FA67A556A7DFE6859A6FD4D40A0" box="[491,685,969,992]" country="Iran" name="Qazvin" pageId="8" pageNumber="7">Qazvin Province</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<location id="8ED4179F7A556A7DFD4859A6FCFE40A0" LSID="urn:lsid:plazi:treatment:03A2F0527A556A7FFED458DEFD454077:8ED4179F7A556A7DFD4859A6FCFE40A0" box="[715,798,969,992]" country="Iran" latitude="36.44" longLatPrecision="712" longitude="50.59" name="Alamut" pageId="8" pageNumber="7" stateProvince="Qazvin">Alamut</location>
|
||
,
|
||
<geoCoordinate id="EE3F27837A556A7DFF435988FEFA40BE" box="[192,282,999,1022]" degrees="36.44" direction="north" orientation="latitude" pageId="8" pageNumber="7" precision="555" value="36.44">36.44°N</geoCoordinate>
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||
<geoCoordinate id="EE3F27837A556A7DFEA15988FE9B40BE" box="[290,379,999,1022]" degrees="50.59" direction="east" orientation="longitude" pageId="8" pageNumber="7" precision="555" value="50.59">50.59°E</geoCoordinate>
|
||
,
|
||
<date id="FFB567847A556A7DFE045988FDED40BE" box="[391,525,999,1022]" pageId="8" pageNumber="7" value="2012-06-08">
|
||
<collectingDate id="EFF19E6C7A556A7DFE045988FDED40BE" box="[391,525,999,1022]" pageId="8" pageNumber="7" value="2012-06-08">8 June 2012</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<specimenCount id="9D0D8ACD7A556A7DFD9A5988FDDA40BE" box="[537,570,999,1022]" pageId="8" pageNumber="7" type="male">3♂</specimenCount>
|
||
(
|
||
<typeStatus id="54B0FFE67A556A7DFDCA5988FD4B40BE" box="[585,683,999,1022]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFD345988FCFD40BE" box="[695,797,999,1022]" captionStart="Figures 1–4" captionStartId="5.[192,278,1533,1557]" captionTargetBox="[205,1422,185,1495]" captionTargetId="figure-6@5.[192,1437,168,1504]" captionTargetPageId="5" captionText="Figures 1–4: Mesobuthus brutus sp. n. Figures 1–2. Holotype male, dorsal (1) and ventral (2) views. Figures 3–4. Paratype female, dorsal (3) and ventral (4) views." pageId="8" pageNumber="7">Figs. 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFF435E69FED9475D" box="[192,313,1030,1053]" captionStart="Figures 5–14" captionStartId="6.[192,278,1635,1659]" captionTargetBox="[191,1440,191,1606]" captionTargetId="figure-11@6.[192,1439,192,1605]" captionTargetPageId="6" captionText="Figures 5–14: Mesobuthus brutus sp. n. Figures 5, 7, 9–11. Holotype male, chelicerae, carapace and tergites I–III (5), sternopectinal region and sternite III (7), pedipalp chela, dorsal (9) and external (10), pedipalp patella dorsal (11). The trichobothrial pattern is indicated in Figures 9–10. Figures 6, 8, 12–14. Paratype female, chelicerae, carapace and tergites I–III (6), sternopectinal region and sternite III (8), pedipalp chela, dorsal (12) and external (13), pedipalp patella dorsal (14)." pageId="8" pageNumber="7">5, 7, 9–11</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFEC85E69FE71475D" box="[331,401,1030,1053]" captionStart="Figures 15–20" captionStartId="7.[192,278,1637,1661]" captionTargetBox="[202,1432,206,1597]" captionTargetId="figure-6@7.[192,1439,195,1608]" captionTargetPageId="7" captionText="Figures 15–20: Mesobuthus brutus sp. n. Figures 15–17. Holotype male, metasoma and telson, lateral (15), ventral (16), and dorsal (17) views. Figures 18–20. Paratype female, metasoma and telson, lateral (18), ventral (19), and dorsal (20) views." pageId="8" pageNumber="7">15–17</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFE215E69FDDD475D" box="[418,573,1030,1053]" captionStart="Figures 181–188" captionStartId="39.[192,278,1168,1192]" captionTargetBox="[196,1430,203,1134]" captionTargetId="figure-59@39.[192,1437,192,1139]" captionTargetPageId="39" captionText="Figures 181–188: Figures 181–182: Mesobuthus kreuzbergi sp. n., metasoma V and telson lateral of male (181) and female (182). Figures 183–188: Mesobuthus brutus sp. n., metasoma V and telson lateral of male holotype (183) and female paratype (184), and left legs I–IV, retrolateral aspect of male holotype (185–188)." pageId="8" pageNumber="7">183, 185–188</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFDCD5E69FD99475D" box="[590,633,1030,1053]" captionStart="Figures 265–273" captionStartId="56.[192,278,1569,1593]" captionTargetBox="[208,1427,176,1539]" captionTargetId="figure-49@56.[192,1439,168,1540]" captionTargetPageId="56" captionText="Figures 265–273: Mesobuthus, metasoma and telson ventral of males. Figure 265. M. brutus sp. n., holotype. Figure 266. M. caucasicus, Turkey. Figure 267. M. gorelovi sp. n., holotype. Figure 268. M. fuscus, Tajikistan. Figure 269. M. kaznakovi, Tajikistan. Figure 270. M. kreuzbergi sp. n., holotype. Figure 271. M. mischi sp. n., holotype. Figure 272. M. nenilini sp. n., holotype. Figure 273. M. parthorum, Afghanistan." pageId="8" pageNumber="7">265</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFD095E69FD55475D" box="[650,693,1030,1053]" captionStart="Figures 274–284" captionStartId="57.[192,278,1468,1492]" captionTargetBox="[196,1428,206,1431]" captionTargetId="figure-6@57.[192,1440,192,1439]" captionTargetPageId="57" captionText="Figures 274–284: Mesobuthus, tarsomeres of third leg. Figure 274. M. brutus sp. n., male holotype. Figure 275. M. caucasicus, female from Turkey. Figure 276. M. elenae sp. n., female holotype. Figure 277. M. gorelovi sp. n., female paratype from type locality. Figure 278. M. fuscus, female from Tajikistan. Figure 279. M. kaznakovi, male from Tajikistan. Figure 280. M. kreuzbergi sp. n., female paratype. Figure 281. M. mischi sp. n., male holotype. Figure 282. M. nenilini sp. n., female paratype. Figure 283–284. M. parthorum, male from Ghazni, Afghanistan (283) and female from Kholm, Afghanistan (284)." pageId="8" pageNumber="7">274</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFD455E69FD0E475D" box="[710,750,1030,1053]" captionStart="Figures 285–303" captionStartId="58.[192,278,1281,1305]" captionTargetBox="[198,1431,202,1246]" captionTargetId="figure-11@58.[192,1440,192,1252]" captionTargetPageId="58" captionText="Figures 285–303: Mesobuthus, pedipalp chela dorsal. Figure 285–286. M. brutus sp. n., male holotype (285) and female paratype (286). Figure 287–288. M. caucasicus, male (287) and female (288) from Turkey. Figure 289. M. elenae sp. n., female holotype. Figure 290–291. M. fuscus, differently colored male (290) and female (291) from Tajikistan. Figure 292–293. M. gorelovi sp. n., male holotype (292) and female paratype (293). Figure 294–295. M. kaznakovi, male (294) and female (295) from Tajikistan. Figure 296–297. M. kreuzbergi sp. n., male holotype (296) and female paratype (297). Figure 298–299. M. mischi sp. n., male holotype (298) and female paratype (299). Figure 300–301. M. nenilini sp. n., male holotype (300) and female paratype (301). Figure 302–303. M. parthorum, male (302) and female (303) from Afghanistan." pageId="8" pageNumber="7">285</figureCitation>
|
||
and
|
||
<typeStatus id="54B0FFE67A556A7DFF435E4AFEC9477C" box="[192,297,1061,1084]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFEBB5E4AFE76477C" box="[312,406,1061,1084]" captionStart="Figures 304–315" captionStartId="59.[192,278,953,977]" captionTargetBox="[208,1427,193,918]" captionTargetId="figure-6@59.[192,1437,192,924]" captionTargetPageId="59" captionText="Figures 304–315: Mesobuthus, pedipalp movable (304–313) and fixed (314–315) fingers. Figure 304. M. brutus sp. n., male paratype. Figure 305. M. caucasicus, male from Turkey. Figure 306. M. elenae sp. n., female holotype. Figure 307. M. gorelovi sp. n., male holotype. Figure 308. M. fuscus, female from Tajikistan. Figure 309. M. kaznakovi, male from Tajikistan. Figure 310, 314. M. kreuzbergi sp. n., male holotype. Figure 311. M. mischi sp. n., male holotype. Figure 312, 315. M. nenilini sp. n., male holotype. Figure 313. M. parthorum, male from Afghanistan." pageId="8" pageNumber="7">Fig. 304</figureCitation>
|
||
)
|
||
<materialsCitation id="3B634B197A556A7DFE2B5E4AFD15471A" collectingDate="2012-06-10" collectionCode="FKCP" country="Iran" latitude="36.95" location="Tarom" longLatPrecision="4474" longitude="48.9" pageId="8" pageNumber="7" specimenCount="2" specimenCount-male="2" stateProvince="Zanjan" typeStatus="paratype">
|
||
<specimenCount id="9D0D8ACD7A556A7DFE2B5E4AFE29477C" box="[424,457,1061,1084]" pageId="8" pageNumber="7" type="male">2♂</specimenCount>
|
||
juvs. (
|
||
<typeStatus id="54B0FFE67A556A7DFD8D5E4AFD97477C" box="[526,631,1061,1084]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode id="ED1AD9817A556A7DFD135E4AFD39477C" box="[656,729,1061,1084]" pageId="8" pageNumber="7">FKCP</collectionCode>
|
||
);
|
||
<emphasis id="B97F9D567A556A7DFD6C5E4AFEA9471A" italics="true" pageId="8" pageNumber="7">
|
||
<collectingRegion id="49CF8FA67A556A7DFD6C5E4AFEA9471A" country="Iran" name="Zanjan" pageId="8" pageNumber="7">Zanjan Province</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<location id="8ED4179F7A556A7DFED45E2CFE41471A" LSID="urn:lsid:plazi:treatment:03A2F0527A556A7FFED458DEFD454077:8ED4179F7A556A7DFED45E2CFE41471A" box="[343,417,1091,1114]" country="Iran" latitude="36.95" longLatPrecision="4474" longitude="48.9" name="Tarom" pageId="8" pageNumber="7" stateProvince="Zanjan">Tarom</location>
|
||
,
|
||
<geoCoordinate id="EE3F27837A556A7DFE2F5E2CFDE6471A" box="[428,518,1091,1114]" degrees="36.95" direction="north" orientation="latitude" pageId="8" pageNumber="7" precision="555" value="36.95">36.95°N</geoCoordinate>
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||
<geoCoordinate id="EE3F27837A556A7DFD8F5E2CFDB8471A" box="[524,600,1091,1114]" degrees="48.9" direction="east" orientation="longitude" pageId="8" pageNumber="7" precision="5555" value="48.9">48.9°E</geoCoordinate>
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||
,
|
||
<date id="FFB567847A556A7DFDE75E2CFD15471A" box="[612,757,1091,1114]" pageId="8" pageNumber="7" value="2012-06-10">
|
||
<collectingDate id="EFF19E6C7A556A7DFDE75E2CFD15471A" box="[612,757,1091,1114]" pageId="8" pageNumber="7" value="2012-06-10">10 June 2012</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B634B197A556A7DFC835E2CFEF847D8" collectionCode="FKCP" country="Iran" pageId="8" pageNumber="7" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||
<specimenCount id="9D0D8ACD7A556A7DFC835E2CFCC2471A" box="[768,802,1091,1114]" pageId="8" pageNumber="7" type="female">2♀</specimenCount>
|
||
(
|
||
<typeStatus id="54B0FFE67A556A7DFF445E0DFED24739" box="[199,306,1122,1145]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFEC75E0DFE504739" box="[324,432,1122,1145]" captionStart="Figures 1–4" captionStartId="5.[192,278,1533,1557]" captionTargetBox="[205,1422,185,1495]" captionTargetId="figure-6@5.[192,1437,168,1504]" captionTargetPageId="5" captionText="Figures 1–4: Mesobuthus brutus sp. n. Figures 1–2. Holotype male, dorsal (1) and ventral (2) views. Figures 3–4. Paratype female, dorsal (3) and ventral (4) views." pageId="8" pageNumber="7">Figs. 3–4</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFE405E0DFDAB4739" box="[451,587,1122,1145]" captionStart="Figures 5–14" captionStartId="6.[192,278,1635,1659]" captionTargetBox="[191,1440,191,1606]" captionTargetId="figure-11@6.[192,1439,192,1605]" captionTargetPageId="6" captionText="Figures 5–14: Mesobuthus brutus sp. n. Figures 5, 7, 9–11. Holotype male, chelicerae, carapace and tergites I–III (5), sternopectinal region and sternite III (7), pedipalp chela, dorsal (9) and external (10), pedipalp patella dorsal (11). The trichobothrial pattern is indicated in Figures 9–10. Figures 6, 8, 12–14. Paratype female, chelicerae, carapace and tergites I–III (6), sternopectinal region and sternite III (8), pedipalp chela, dorsal (12) and external (13), pedipalp patella dorsal (14)." pageId="8" pageNumber="7">6, 8, 12–14</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFDDE5E0DFD434739" box="[605,675,1122,1145]" captionStart="Figures 15–20" captionStartId="7.[192,278,1637,1661]" captionTargetBox="[202,1432,206,1597]" captionTargetId="figure-6@7.[192,1439,195,1608]" captionTargetPageId="7" captionText="Figures 15–20: Mesobuthus brutus sp. n. Figures 15–17. Holotype male, metasoma and telson, lateral (15), ventral (16), and dorsal (17) views. Figures 18–20. Paratype female, metasoma and telson, lateral (18), ventral (19), and dorsal (20) views." pageId="8" pageNumber="7">18–20</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFD375E0DFD3E4739" box="[692,734,1122,1145]" captionStart="Figures 181–188" captionStartId="39.[192,278,1168,1192]" captionTargetBox="[196,1430,203,1134]" captionTargetId="figure-59@39.[192,1437,192,1139]" captionTargetPageId="39" captionText="Figures 181–188: Figures 181–182: Mesobuthus kreuzbergi sp. n., metasoma V and telson lateral of male (181) and female (182). Figures 183–188: Mesobuthus brutus sp. n., metasoma V and telson lateral of male holotype (183) and female paratype (184), and left legs I–IV, retrolateral aspect of male holotype (185–188)." pageId="8" pageNumber="7">184</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFD725E0DFCFA4739" box="[753,794,1122,1145]" captionStart="Figures 285–303" captionStartId="58.[192,278,1281,1305]" captionTargetBox="[198,1431,202,1246]" captionTargetId="figure-11@58.[192,1440,192,1252]" captionTargetPageId="58" captionText="Figures 285–303: Mesobuthus, pedipalp chela dorsal. Figure 285–286. M. brutus sp. n., male holotype (285) and female paratype (286). Figure 287–288. M. caucasicus, male (287) and female (288) from Turkey. Figure 289. M. elenae sp. n., female holotype. Figure 290–291. M. fuscus, differently colored male (290) and female (291) from Tajikistan. Figure 292–293. M. gorelovi sp. n., male holotype (292) and female paratype (293). Figure 294–295. M. kaznakovi, male (294) and female (295) from Tajikistan. Figure 296–297. M. kreuzbergi sp. n., male holotype (296) and female paratype (297). Figure 298–299. M. mischi sp. n., male holotype (298) and female paratype (299). Figure 300–301. M. nenilini sp. n., male holotype (300) and female paratype (301). Figure 302–303. M. parthorum, male (302) and female (303) from Afghanistan." pageId="8" pageNumber="7">286</figureCitation>
|
||
) (
|
||
<collectionCode id="ED1AD9817A556A7DFF4B5EEEFEF147D8" box="[200,273,1153,1176]" pageId="8" pageNumber="7">FKCP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31112CF7A556A7DFF435ED4FDBB47B1" pageId="8" pageNumber="7" type="etymology">
|
||
<paragraph id="8BB441447A556A7DFF435ED4FDBB47B1" blockId="8.[192,802,1211,1265]" pageId="8" pageNumber="7">ETYMOLOGY. The species is named in honor of Brutus, the famous Czech rock musical group.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31112CF7A556A7DFF435F78FDA2466E" box="[192,578,1303,1326]" pageId="8" pageNumber="7" type="distribution">
|
||
<paragraph id="8BB441447A556A7DFF435F78FDA2466E" blockId="8.[192,578,1303,1326]" box="[192,578,1303,1326]" pageId="8" pageNumber="7">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="F31C01D47A556A7DFEE45F78FE72466E" box="[359,402,1303,1326]" name="Iran" pageId="8" pageNumber="7">Iran</collectingCountry>
|
||
(
|
||
<figureCitation id="13305DC17A556A7DFE235F78FDE6466E" box="[416,518,1303,1326]" captionStart="Figure 328" captionStartId="63.[192,268,1348,1372]" captionTargetBox="[187,1924,187,1319]" captionTargetId="figure-24@63.[192,1920,192,1314]" captionTargetPageId="63" captionText="Figure 328: Distribution map showing all species examined in this study. See Appendix A for distribution maps of each species." pageId="8" pageNumber="7">Figs. 328</figureCitation>
|
||
, A
|
||
<figureCitation id="13305DC17A556A7DFDA95F78FDD4466E" box="[554,564,1303,1326]" captionStart="Figures 1–4" captionStartId="5.[192,278,1533,1557]" captionTargetBox="[205,1422,185,1495]" captionTargetId="figure-6@5.[192,1437,168,1504]" captionTargetPageId="5" captionText="Figures 1–4: Mesobuthus brutus sp. n. Figures 1–2. Holotype male, dorsal (1) and ventral (2) views. Figures 3–4. Paratype female, dorsal (3) and ventral (4) views." pageId="8" pageNumber="7">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31112CF7A556A7DFF435F3BFBE842D6" pageId="8" pageNumber="7" type="diagnosis">
|
||
<paragraph id="8BB441447A556A7DFF435F3BFBE842D6" blockId="8.[192,802,1364,1909]" lastBlockId="8.[830,1441,199,406]" pageId="8" pageNumber="7">
|
||
DIAGNOSIS. Total length of adult males
|
||
<quantity id="4CF3ECA17A556A7DFDF55F3BFD0D462B" box="[630,749,1364,1387]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" metricValueMax="6.1" metricValueMin="5.0" pageId="8" pageNumber="7" unit="mm" value="55.5" valueMax="61.0" valueMin="50.0">50–61 mm</quantity>
|
||
, 60–
|
||
<specimenCount id="9D0D8ACD7A556A7DFF435F1CFEDB46CA" box="[192,315,1395,1418]" pageId="8" pageNumber="7" type="female">62 females</specimenCount>
|
||
. Trichobothrium
|
||
<emphasis id="B97F9D567A556A7DFE7F5F1CFDF746CA" box="[508,535,1395,1418]" italics="true" pageId="8" pageNumber="7">db</emphasis>
|
||
on fixed finger of pedipalp situated between trichobothria
|
||
<emphasis id="B97F9D567A556A7DFDCD5FFDFD8C46E9" box="[590,620,1426,1449]" italics="true" pageId="8" pageNumber="7">est</emphasis>
|
||
and
|
||
<emphasis id="B97F9D567A556A7DFD275FFDFD2846E9" box="[676,712,1426,1449]" italics="true" pageId="8" pageNumber="7">esb</emphasis>
|
||
, near to
|
||
<emphasis id="B97F9D567A556A7DFF435FDFFF3E4687" box="[192,222,1456,1479]" italics="true" pageId="8" pageNumber="7">est</emphasis>
|
||
. Male with fingers proximally more twisted than female. Pedipalp chela length/width ratio
|
||
<quantity id="4CF3ECA17A556A7DFD155FA0FCC246A6" box="[662,802,1487,1510]" metricMagnitude="-2" metricUnit="m" metricValue="7.975599999999999" metricValueMax="8.001" metricValueMin="7.9502" pageId="8" pageNumber="7" unit="in" value="3.1399999999999997" valueMax="3.15" valueMin="3.13">3.13–3.15 in</quantity>
|
||
males and
|
||
<quantity id="4CF3ECA17A556A7DFEB95F81FE244545" box="[314,452,1518,1541]" metricMagnitude="-2" metricUnit="m" metricValue="9.1821" metricValueMax="9.270999999999999" metricValueMin="9.0932" pageId="8" pageNumber="7" unit="in" value="3.615" valueMax="3.65" valueMin="3.58">3.58–3.65 in</quantity>
|
||
females. Pectinal teeth number
|
||
<quantity id="4CF3ECA17A556A7DFF435C63FEC44563" box="[192,292,1548,1571]" metricMagnitude="-1" metricUnit="m" metricValue="6.731" metricValueMax="7.112" metricValueMin="6.35" pageId="8" pageNumber="7" unit="in" value="26.5" valueMax="28.0" valueMin="25.0">25–28 in</quantity>
|
||
males,
|
||
<quantity id="4CF3ECA17A556A7DFE035C63FE044563" box="[384,484,1548,1571]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" metricValueMax="5.588" metricValueMin="5.08" pageId="8" pageNumber="7" unit="in" value="21.0" valueMax="22.0" valueMin="20.0">20–22 in</quantity>
|
||
females. Chelicerae yellow, without reticulation. Pedipalps and metasoma very sparsely hirsute. Carapace and tergites yellowish brown, black pigmented; metasoma, telson, pedipalps and legs yellowish brown, only anterior part of metasomal segment V black. Femur of pedipalp with 4–5 granulate carinae. Patella with 8 granulate or smooth carinae. Chela lacks carinae. Movable fingers of pedipalps with 13 cutting rows of denticles and 5 terminal denticles. Seventh sternite bears 4 well marked granulate carinae. First metasomal segment with 10 carinae; second to fourth with 8 carinae, other two carinae are indicated by incomplete row of denticles on metasomal segments II and III; fourth with 8 carinae; fifth with 5 carinae. All carinae granulated. Length to width ratio of fourth metasomal segment
|
||
<quantity id="4CF3ECA17A556A7DFC765B4CFB63427A" box="[1013,1155,291,314]" metricMagnitude="-2" metricUnit="m" metricValue="3.937" metricValueMax="3.9878" metricValueMin="3.8862" pageId="8" pageNumber="7" unit="in" value="1.55" valueMax="1.57" valueMin="1.53">1.53–1.57 in</quantity>
|
||
males,
|
||
<quantity id="4CF3ECA17A556A7DFB655B4CFA94427A" box="[1254,1396,291,314]" metricMagnitude="-2" metricUnit="m" metricValue="3.937" metricValueMax="4.1148" metricValueMin="3.7592" pageId="8" pageNumber="7" unit="in" value="1.55" valueMax="1.62" valueMin="1.48">1.48–1.62 in</quantity>
|
||
females. Telotarsus III ventral setation represented by short and strong spiniform setae. Pedal spur of legs with solitary setae only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31112CF7A556A7FFCBD5BD3FD454077" lastPageId="10" lastPageNumber="9" pageId="8" pageNumber="7" type="description">
|
||
<paragraph id="8BB441447A556A7DFCBD5BD3FACE41EA" blockId="8.[830,1442,444,1909]" pageId="8" pageNumber="7">
|
||
DESCRIPTION. The total length of adult males
|
||
<quantity id="4CF3ECA17A556A7DFADE5BD3FC8A42B2" metricMagnitude="-2" metricUnit="m" metricValue="5.55" metricValueMax="6.1" metricValueMin="5.0" pageId="8" pageNumber="7" unit="mm" value="55.5" valueMax="61.0" valueMin="50.0">50–61 mm</quantity>
|
||
, 60–
|
||
<specimenCount id="9D0D8ACD7A556A7DFC235BB4FBFA42B2" box="[928,1050,475,498]" pageId="8" pageNumber="7" type="female">62 females</specimenCount>
|
||
. Trichobothrium
|
||
<emphasis id="B97F9D567A556A7DFB5B5BB4FB1342B2" box="[1240,1267,475,498]" italics="true" pageId="8" pageNumber="7">db</emphasis>
|
||
on fixed finger of pedipalp is situated between trichobothria
|
||
<emphasis id="B97F9D567A556A7DFAA05B95FAA14151" box="[1315,1345,506,529]" italics="true" pageId="8" pageNumber="7">est</emphasis>
|
||
and
|
||
<emphasis id="B97F9D567A556A7DFAF55B95FA7A4151" box="[1398,1434,506,529]" italics="true" pageId="8" pageNumber="7">esb</emphasis>
|
||
, near to
|
||
<emphasis id="B97F9D567A556A7DFC125877FC4F416F" box="[913,943,536,559]" italics="true" pageId="8" pageNumber="7">est</emphasis>
|
||
. Male has the fingers proximally a little more twisted than female. Female has longer and slightly narrower chela of pedipalps. Chelicerae yellow, without reticulation, the tips of teeth on cheliceral fingers are black. For measurements see tables 1 and 4–5.
|
||
</paragraph>
|
||
<paragraph id="8BB441447A556A7DFCBD58DDFC754003" blockId="8.[830,1442,444,1909]" pageId="8" pageNumber="7">
|
||
COLORATION (
|
||
<figureCitation id="13305DC17A556A7DFC5E58DDFBA74189" box="[989,1095,690,713]" captionStart="Figures 1–4" captionStartId="5.[192,278,1533,1557]" captionTargetBox="[205,1422,185,1495]" captionTargetId="figure-6@5.[192,1437,168,1504]" captionTargetPageId="5" captionText="Figures 1–4: Mesobuthus brutus sp. n. Figures 1–2. Holotype male, dorsal (1) and ventral (2) views. Figures 3–4. Paratype female, dorsal (3) and ventral (4) views." pageId="8" pageNumber="7">Figs. 1–4</figureCitation>
|
||
). The carapace and tergites are yellowish brown, strongly black pigmented. The metasoma, telson, pedipalps and legs are yellowish brown. The metasoma ventral, mainly segment V black pigmented.
|
||
</paragraph>
|
||
<paragraph id="8BB441447A556A7DFCBD5924FC79466E" blockId="8.[830,1442,444,1909]" pageId="8" pageNumber="7">
|
||
MESOSOMA AND CARAPACE (
|
||
<figureCitation id="13305DC17A556A7DFBF85924FB044022" box="[1147,1252,843,866]" captionStart="Figures 5–14" captionStartId="6.[192,278,1635,1659]" captionTargetBox="[191,1440,191,1606]" captionTargetId="figure-11@6.[192,1439,192,1605]" captionTargetPageId="6" captionText="Figures 5–14: Mesobuthus brutus sp. n. Figures 5, 7, 9–11. Holotype male, chelicerae, carapace and tergites I–III (5), sternopectinal region and sternite III (7), pedipalp chela, dorsal (9) and external (10), pedipalp patella dorsal (11). The trichobothrial pattern is indicated in Figures 9–10. Figures 6, 8, 12–14. Paratype female, chelicerae, carapace and tergites I–III (6), sternopectinal region and sternite III (8), pedipalp chela, dorsal (12) and external (13), pedipalp patella dorsal (14)." pageId="8" pageNumber="7">Figs. 5–8</figureCitation>
|
||
). The carapace is carinate and unevenly covered by granules of varying size; much of the granulation is fine, but some granules are larger and distinctly rounded. Tergites I–VI bear three carinae and are granulated, with some intercarinal granules small and others larger and rounded. Tergite VII is pentacarinate. The pectinal tooth count is
|
||
<quantity id="4CF3ECA17A556A7DFAC15E6CFA41475A" box="[1346,1441,1027,1050]" metricMagnitude="-1" metricUnit="m" metricValue="6.731" metricValueMax="7.112" metricValueMin="6.35" pageId="8" pageNumber="7" unit="in" value="26.5" valueMax="28.0" valueMin="25.0">25–28 in</quantity>
|
||
males,
|
||
<quantity id="4CF3ECA17A556A7DFC115E4DFC194779" box="[914,1017,1058,1081]" metricMagnitude="-1" metricUnit="m" metricValue="5.715" metricValueMax="6.35" metricValueMin="5.08" pageId="8" pageNumber="7" unit="in" value="22.5" valueMax="25.0" valueMin="20.0">20–25 in</quantity>
|
||
females. The pectinal marginal tips extend to about end of the sixth sternite in males and third of the sixth sternite in females. The pectines have three marginal lamellae and sixth to eight middle lamellae. The lamellae bear numerous long setae, each fulcrum with three to five setae. All sternites are smooth and sparsely hirsute. The seventh sternite bears four well marked granulate carinae. The other sternites bear two furrows.
|
||
</paragraph>
|
||
<paragraph id="8BB441447A556A7DFCBD5F59FA9A4502" blockId="8.[830,1442,444,1909]" pageId="8" pageNumber="7">
|
||
PEDIPALPS (
|
||
<figureCitation id="13305DC17A556A7DFC435F59FBD9460D" box="[960,1081,1334,1357]" captionStart="Figures 5–14" captionStartId="6.[192,278,1635,1659]" captionTargetBox="[191,1440,191,1606]" captionTargetId="figure-11@6.[192,1439,192,1605]" captionTargetPageId="6" captionText="Figures 5–14: Mesobuthus brutus sp. n. Figures 5, 7, 9–11. Holotype male, chelicerae, carapace and tergites I–III (5), sternopectinal region and sternite III (7), pedipalp chela, dorsal (9) and external (10), pedipalp patella dorsal (11). The trichobothrial pattern is indicated in Figures 9–10. Figures 6, 8, 12–14. Paratype female, chelicerae, carapace and tergites I–III (6), sternopectinal region and sternite III (8), pedipalp chela, dorsal (12) and external (13), pedipalp patella dorsal (14)." pageId="8" pageNumber="7">Figs. 9–14</figureCitation>
|
||
). The pedipalps are sparsely hirsute and smooth. The femur bears four to five granulated carinae, the middle carina on internal surface could be incomplete indicated by several strong denticles. The patella bears eight carinae from which internal in both sexes and dorsal in females are granulated. The chela is without carinae. The movable fingers of pedipalps bear 13 cutting rows of denticles, every with external and internal denticles present, and five terminal denticles.
|
||
</paragraph>
|
||
<paragraph id="8BB441447A556A7DFCBD5C25FB034459" blockId="8.[830,1442,444,1909]" pageId="8" pageNumber="7">
|
||
LEGS (
|
||
<figureCitation id="13305DC17A556A7DFC055C25FBC64521" box="[902,1062,1610,1633]" captionStart="Figures 181–188" captionStartId="39.[192,278,1168,1192]" captionTargetBox="[196,1430,203,1134]" captionTargetId="figure-59@39.[192,1437,192,1139]" captionTargetPageId="39" captionText="Figures 181–188: Figures 181–182: Mesobuthus kreuzbergi sp. n., metasoma V and telson lateral of male (181) and female (182). Figures 183–188: Mesobuthus brutus sp. n., metasoma V and telson lateral of male holotype (183) and female paratype (184), and left legs I–IV, retrolateral aspect of male holotype (185–188)." pageId="8" pageNumber="7">Figs. 185–188</figureCitation>
|
||
). The tarsomeres bear two rows of short and strong spiniform setae on the ventral surface and numerous macrosetae on the other surfaces. Pedal spur of legs with solitary setae only. Femur bears only several macrosetae. Femur and patella with carinae well developed. Tibial spurs present and long on third and fourth legs and absent in the other legs.
|
||
</paragraph>
|
||
<paragraph id="8BB441447A556A7FFCBD5D4FFD1542B2" blockId="8.[830,1442,444,1909]" lastBlockId="10.[192,802,199,498]" lastPageId="10" lastPageNumber="9" pageId="8" pageNumber="7">
|
||
METASOMA AND TELSON (
|
||
<figureCitation id="13305DC17A556A7DFBE95D4FFB124477" box="[1130,1266,1824,1847]" captionStart="Figures 15–20" captionStartId="7.[192,278,1637,1661]" captionTargetBox="[202,1432,206,1597]" captionTargetId="figure-6@7.[192,1439,195,1608]" captionTargetPageId="7" captionText="Figures 15–20: Mesobuthus brutus sp. n. Figures 15–17. Holotype male, metasoma and telson, lateral (15), ventral (16), and dorsal (17) views. Figures 18–20. Paratype female, metasoma and telson, lateral (18), ventral (19), and dorsal (20) views." pageId="8" pageNumber="7">Figs. 15–20</figureCitation>
|
||
,
|
||
<figureCitation id="13305DC17A556A7DFA875D4FFA864477" box="[1284,1382,1824,1847]" captionStart="Figures 181–188" captionStartId="39.[192,278,1168,1192]" captionTargetBox="[196,1430,203,1134]" captionTargetId="figure-59@39.[192,1437,192,1139]" captionTargetPageId="39" captionText="Figures 181–188: Figures 181–182: Mesobuthus kreuzbergi sp. n., metasoma V and telson lateral of male (181) and female (182). Figures 183–188: Mesobuthus brutus sp. n., metasoma V and telson lateral of male holotype (183) and female paratype (184), and left legs I–IV, retrolateral aspect of male holotype (185–188)." pageId="8" pageNumber="7">183–184</figureCitation>
|
||
). All metasomal segments are only very sparsely hirsute. The metasomal segment I with 10 carinae, II–III with 8 carinae but other two latero median carinae are indicated by incomplete row of denticles, IV with 8 carinae, and V with 5 carinae. All carinae with consistent denticles, larger denticles are present in ventral carinae on segment I–III. The dorsal surface of all segments is smooth in the middle and bumpy on margins. Other surfaces are bumpy or smooth with several solitary granules. Ventrolateral carinae of metasomal segment V posteriorly with several large lobated denticles. The telson is only sparsely hirsute, rather elongate, bumpy and smooth.
|
||
</paragraph>
|
||
<caption id="DF7411CC7A546A7CFF435C9AFEDF447E" pageId="9" pageNumber="8" startId="9.[192,278,1781,1805]" targetBox="[198,1439,224,1743]" targetPageId="9">
|
||
<paragraph id="8BB441447A546A7CFF435C9AFEDF447E" blockId="9.[192,1440,1781,1854]" pageId="9" pageNumber="8">
|
||
<emphasis id="B97F9D567A546A7CFF435C9AFE8A444D" bold="true" box="[192,362,1781,1805]" pageId="9" pageNumber="8">Figures 21–25:</emphasis>
|
||
<taxonomicName id="4C0B3AC77A546A7CFEF75C98FDB8444C" baseAuthorityName="Nordmann" baseAuthorityYear="1840" box="[372,600,1783,1804]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="caucasicus">
|
||
<emphasis id="B97F9D567A546A7CFEF75C98FDB8444C" box="[372,600,1783,1804]" italics="true" pageId="9" pageNumber="8">Mesobuthus caucasicus</emphasis>
|
||
</taxonomicName>
|
||
, holotype female. Dorsal (21) and ventral (22) views. Metasoma V and telson lateral (23), and metasoma and telson, lateral (24) and dorsal (25) views. The original labels are also included in the plate. Scale bar: 10 mm (21–22).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB441447A576A7FFF43587DFEEF4169" blockId="10.[192,271,530,553]" box="[192,271,530,553]" pageId="10" pageNumber="9">NOTES.</paragraph>
|
||
<paragraph id="8BB441447A576A7FFF5E5826FD454077" blockId="10.[221,802,585,823]" pageId="10" pageNumber="9">
|
||
<taxonomicName id="4C0B3AC77A576A7FFF5E5826FE4C4120" authority="Fet & Kovařík & Gantenbein & Kaiser & Stewart & Graham, 2018" authorityName="Fet & Kovařík & Gantenbein & Kaiser & Stewart & Graham" authorityYear="2018" box="[221,428,585,608]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="brutus" status="sp. nov.">
|
||
<emphasis id="B97F9D567A576A7FFF5E5826FE4C4120" box="[221,428,585,608]" italics="true" pageId="10" pageNumber="9">Mesobuthus brutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24C202D7A576A7FFE355826FE134120" box="[438,499,585,608]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
|
||
is the only species studied by us for which no DNA data are yet available, therefore we cannot place it into our phylogeny. The species is currently known only from the
|
||
<collectingRegion id="49CF8FA67A576A7FFD5858CAFCC241FC" box="[731,802,677,700]" country="Iran" name="Alborz" pageId="10" pageNumber="9">Elburz</collectingRegion>
|
||
Mountains of the northern
|
||
<collectingCountry id="F31C01D47A576A7FFD8B58ABFDD6419B" box="[520,566,708,731]" name="Iran" pageId="10" pageNumber="9">Iran</collectingCountry>
|
||
, south from the Caspian Sea, a biogeographic transition zone between the Caucasus to the west (where
|
||
<taxonomicName id="4C0B3AC77A576A7FFDAD596EFD2C4058" baseAuthorityName="Nordmann" baseAuthorityYear="1840" box="[558,716,769,792]" class="Arachnida" family="Buthidae" genus="Mesobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" sensu="stricto" species="caucasicus">
|
||
<emphasis id="B97F9D567A576A7FFDAD596EFD2C4058" box="[558,716,769,792]" italics="true" pageId="10" pageNumber="9">M. caucasicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24C202D7A576A7FFD5B596EFCE44058" box="[728,772,769,792]" pageId="10" pageNumber="9" sensu="stricto">s.str</taxonomicNameLabel>
|
||
is found) and the Kopetdagh Mts. to the east.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |