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<mods:title id="D3E816519DB271586F656C329EFA9B38">Revision of the genus Epexochus Reitter, with description of three new species (Coleoptera: Curculionidae: Lixinae: Cleonini)</mods:title>
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<mods:date id="01546E3EA615E05644BCB888AE087A77">2009</mods:date>
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<paragraph id="8BB41A3A430C3364FF60D32F4D1AE2E2" blockId="3.[151,559,1912,1939]" box="[151,559,1912,1939]" pageId="3" pageNumber="50">
<heading id="D0FCAD56430C3364FF60D32F4D1AE2E2" bold="true" box="[151,559,1912,1939]" fontSize="11" level="1" pageId="3" pageNumber="50" reason="1">
<taxonomicName id="4C0B61B9430C3364FF60D32F4D1AE2E2" authority="Menetries" authorityName="Menetries" box="[151,559,1912,1939]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC628430C3364FF60D32F4D1AE2E2" bold="true" box="[151,559,1912,1939]" pageId="3" pageNumber="50">
<emphasis id="B97FC628430C3364FF60D32F4EA2E2E3" bold="true" box="[151,407,1913,1939]" italics="true" pageId="3" pageNumber="50">Epexochus lehmanni</emphasis>
(Ménétries)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31149B1430C3363FF60D3E94CD0E7F6" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50" type="reference_group">
<paragraph id="8BB41A3A430C3364FF60D3E94D64E2A5" blockId="3.[151,1150,1982,2038]" box="[151,593,1982,2006]" pageId="3" pageNumber="50">
<treatmentCitationGroup id="AB1B3D14430C3364FF60D3E94D64E2A5" box="[151,593,1982,2006]" pageId="3" pageNumber="50">
<taxonomicName id="4C0B61B9430C3364FF60D3E94D79E2A5" ID-CoL="8HGF6" authority="Lehmanni Menetries, 1849: 251" authorityName="Lehmanni Menetries" authorityPageNumber="251" authorityYear="1849" box="[151,588,1982,2006]" class="Insecta" family="Curculionidae" genus="Cleonus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC628430C3364FF60D3E94E55E2A6" box="[151,352,1983,2006]" italics="true" pageId="3" pageNumber="50">Cleonus Lehmanni</emphasis>
<treatmentCitation id="0AAA3C2B430C3364FE91D3E84D79E2A5" author="Menetries" box="[358,588,1982,2005]" page="251" pageId="3" pageNumber="50" year="1849">
<bibRefCitation id="EF9A67CB430C3364FE91D3E84D22E2A5" author="Menetries" box="[358,535,1982,2005]" pageId="3" pageNumber="67" refString="Menetries, M. (1849) Catalogue des insectes recueillis par feu M. Lehmann avec les descriptions des nouvelles especes. Seconde et derniere partie. Tetramere. Curculionites. Memoires de l'Academie Imperiale des Sciences de Saint- Petersbourg. Sixieme Serie. Sciences Naturelles, VI: 249 - 263." type="book chapter" year="1849">Ménétries, 1849</bibRefCitation>
: 251
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8BB41A3A430C3364FF60D3894B4BE285" blockId="3.[151,1150,1982,2038]" box="[151,1150,2014,2038]" pageId="3" pageNumber="50">
<treatmentCitationGroup id="AB1B3D14430C3364FF60D3894B4BE285" box="[151,1150,2014,2038]" pageId="3" pageNumber="50">
<taxonomicName id="4C0B61B9430C3364FF60D3894D1EE285" ID-CoL="8HLKB" authority="Lehmanni (Menetries)" authorityName="Lehmanni (Menetries" box="[151,555,2014,2038]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC628430C3364FF60D3894E96E286" box="[151,419,2015,2038]" italics="true" pageId="3" pageNumber="50">Leucochromus Lehmanni</emphasis>
(Ménétries)
</taxonomicName>
:
<treatmentCitation id="0AAA3C2B430C3364FDC2D3884DC0E285" author="Chevrolat" box="[565,757,2014,2037]" page="2" pageId="3" pageNumber="50" year="1873">
<bibRefCitation id="EF9A67CB430C3364FDC2D3884DEBE285" author="Chevrolat" box="[565,734,2014,2037]" pageId="3" pageNumber="67" refString="Chevrolat, A. (1873) Memoire sur les Cleonides. Memoires de la Societe Royale des Sciences de Liege, 2 (5): I - VIII + 1 - 118." type="journal article" year="1873">Chevrolat 1873</bibRefCitation>
: 2
</treatmentCitation>
[as
<emphasis id="B97FC628430C3364FCD5D3894CB8E286" box="[802,909,2015,2038]" italics="true" pageId="3" pageNumber="50">Lhemanni</emphasis>
], 99;
<treatmentCitation id="0AAA3C2B430C3364FC3BD3884B4FE285" author="Faust" box="[972,1146,2014,2037]" page="191" pageId="3" pageNumber="50" year="1904">
<bibRefCitation id="EF9A67CB430C3364FC3BD3884B70E285" author="Faust" box="[972,1093,2014,2037]" pageId="3" pageNumber="67" refString="Faust, J. (1904) Revision der Gruppe Cleonides vrais. Deutsche Entomologische Zeitschrift, 1904 (1): 177 - 284." type="journal article" year="1904">Faust 1904</bibRefCitation>
: 191
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.
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</paragraph>
<paragraph id="8BB41A3A430B3363FF60D4CE4D9AE5DF" blockId="4.[151,1083,152,495]" box="[151,687,152,175]" pageId="4" pageNumber="51">
<treatmentCitationGroup id="AB1B3D14430B3363FF60D4CE4D9AE5DF" box="[151,687,152,175]" pageId="4" pageNumber="51">
<taxonomicName id="4C0B61B9430B3363FF60D4CE4EC7E5DF" ID-CoL="8KWV7" authority="Lehmanni (Menetries)" authorityName="Lehmanni (Menetries" box="[151,498,152,175]" class="Insecta" family="Carabidae" genus="Epilectus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC628430B3363FF60D4CE4E5EE5DF" box="[151,363,152,175]" italics="true" pageId="4" pageNumber="51">Epilectus Lehmanni</emphasis>
(Ménétries)
</taxonomicName>
:
<treatmentCitation id="0AAA3C2B430B3363FE0AD4CE4D9EE5DF" author="Faust" box="[509,683,152,175]" page="208" pageId="4" pageNumber="51" year="1904">
<bibRefCitation id="EF9A67CB430B3363FE0AD4CE4D43E5DF" author="Faust" box="[509,630,152,175]" pageId="4" pageNumber="67" refString="Faust, J. (1904) Revision der Gruppe Cleonides vrais. Deutsche Entomologische Zeitschrift, 1904 (1): 177 - 284." type="journal article" year="1904">Faust 1904</bibRefCitation>
: 208
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BB41A3A430B3363FF60D4EE4C15E59F" blockId="4.[151,1083,152,495]" pageId="4" pageNumber="51">
<treatmentCitationGroup id="AB1B3D14430B3363FF60D4EE4C15E59F" pageId="4" pageNumber="51">
<taxonomicName id="4C0B61B9430B3363FF60D4EE4ECEE5BF" authority="Menetries" authorityName="Menetries" box="[151,507,184,207]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC628430B3363FF60D4EE4E41E5BF" box="[151,372,184,207]" italics="true" pageId="4" pageNumber="51">Epexochus lehmanni</emphasis>
(Ménétries)
</taxonomicName>
:
<treatmentCitation id="0AAA3C2B430B3363FDF2D4EE4D83E5BF" author="Reitter" box="[517,694,184,207]" page="40" pageId="4" pageNumber="51" year="1913">
<bibRefCitation id="EF9A67CB430B3363FDF2D4EE4DBBE5BF" author="Reitter" box="[517,654,184,207]" pageId="4" pageNumber="67" refString="Reitter, E. (1913) Bestimmungs-Schlussel der mir bekannten europaischen Gattungen der Curculionidae, mit Einschluss der mir bekannten Gattungen aus dem palaearctischen Gebiete. Verhandlungen des naturforschenden Vereines in Brunn, 52: 129 - 242." type="journal article" year="1913">Reitter 1913</bibRefCitation>
: 40
</treatmentCitation>
;
<treatmentCitation id="0AAA3C2B430B3363FD36D4EE4CFFE5BF" author="Ter-Minasyan" box="[705,970,184,207]" page="516" pageId="4" pageNumber="51" year="1968">
<bibRefCitation id="EF9A67CB430B3363FD36D4EE4CA0E5BF" author="Ter-Minasyan" box="[705,917,184,207]" pageId="4" pageNumber="68" refString="Ter-Minasyan, M. E .. (1968) Obzor zhukov-dolgonosikov triby Cleonini (Coleoptera, Curculionidae) fauny Srediey Azii i Kazakhstana. Entomologicheskoe Obozrennie 47 (3): 512 - 522." type="journal article" year="1968">Ter-Minasyan 1968</bibRefCitation>
: 516
</treatmentCitation>
; 1988: 46.
<taxonomicName id="4C0B61B9430B3363FF60D48E4D47E59F" authority="Lehmanni (Menetries)" authorityName="Lehmanni (Menetries" box="[151,626,216,239]" class="Insecta" family="Curculionidae" genus="Cleonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="subGenus" subGenus="Epexochus">
<emphasis id="B97FC628430B3363FF60D48E4ED9E59F" box="[151,492,216,239]" italics="true" pageId="4" pageNumber="51">Cleonus (Epexochus) Lehmanni</emphasis>
(Ménétries)
</taxonomicName>
:
<treatmentCitation id="0AAA3C2B430B3363FD8AD48E4C28E59F" author="Csiki" box="[637,797,216,239]" page="39" pageId="4" pageNumber="51" year="1934">
<bibRefCitation id="EF9A67CB430B3363FD8AD48E4DC1E59F" author="Csiki" box="[637,756,216,239]" pageId="4" pageNumber="67" refString="Csiki, E. (1934). Curculionidae. Subfam. Cleoninae. In: Schenkling S. (Ed.). Coleopterorum Catalogus auspiciis et auxilio W. Junk, 134: pp. 152." type="journal article" year="1934">Csiki 1934</bibRefCitation>
: 39
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB41A3A430B3363FF60D4AE4DBEE47F" blockId="4.[151,1083,152,495]" box="[151,651,247,271]" pageId="4" pageNumber="51">
<treatmentCitationGroup id="AB1B3D14430B3363FF60D4AE4DBEE47F" box="[151,651,247,271]" pageId="4" pageNumber="51">
<taxonomicName id="4C0B61B9430B3363FF60D4AE4ECEE47F" authority="Chevrolat, 1873: 3" authorityName="Chevrolat" authorityPageNumber="3" authorityYear="1873" box="[151,507,248,271]" class="Insecta" family="Curculionidae" genus="Exochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC628430B3363FF60D4AE4E19E47F" box="[151,300,248,271]" italics="true" pageId="4" pageNumber="51">Exochus latus</emphasis>
<treatmentCitation id="0AAA3C2B430B3363FEC5D4AE4ECEE47F" author="Chevrolat" box="[306,507,248,271]" page="3" pageId="4" pageNumber="51" year="1873">
<bibRefCitation id="EF9A67CB430B3363FEC5D4AE4ED7E47F" author="Chevrolat" box="[306,482,248,271]" pageId="4" pageNumber="67" refString="Chevrolat, A. (1873) Memoire sur les Cleonides. Memoires de la Societe Royale des Sciences de Liege, 2 (5): I - VIII + 1 - 118." type="journal article" year="1873">Chevrolat, 1873</bibRefCitation>
: 3
</treatmentCitation>
</taxonomicName>
; 99
<emphasis id="B97FC628430B3363FDDED4A14DB6E47F" bold="true" box="[553,643,247,271]" pageId="4" pageNumber="51">(syn. n.)</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB41A3A430B3363FF60D54E4DC1E45F" blockId="4.[151,1083,152,495]" box="[151,756,280,303]" pageId="4" pageNumber="51">
<treatmentCitationGroup id="AB1B3D14430B3363FF60D54E4DC1E45F" box="[151,756,280,303]" pageId="4" pageNumber="51">
<taxonomicName id="4C0B61B9430B3363FF60D54E4E5EE45F" ID-CoL="8KWV7" authority="Lehmanni" authorityName="Lehmanni" box="[151,363,280,303]" class="Insecta" family="Carabidae" genus="Epilectus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC628430B3363FF60D54E4E5EE45F" box="[151,363,280,303]" italics="true" pageId="4" pageNumber="51">Epilectus Lehmanni</emphasis>
</taxonomicName>
?
<taxonomicName id="4C0B61B9430B3363FE89D54E4D02E45F" authority="Chevrolat" authorityName="Chevrolat" box="[382,567,280,303]" class="Insecta" family="Curculionidae" genus="Exochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC628430B3363FE89D54E4E87E45F" box="[382,434,280,303]" italics="true" pageId="4" pageNumber="51">latus</emphasis>
(Chevrolat)
</taxonomicName>
:
<treatmentCitation id="0AAA3C2B430B3363FDB6D54E4DC5E45F" author="Faust" box="[577,752,280,303]" page="208" pageId="4" pageNumber="51" year="1904">
<bibRefCitation id="EF9A67CB430B3363FDB6D54E4D8EE45F" author="Faust" box="[577,699,280,303]" pageId="4" pageNumber="67" refString="Faust, J. (1904) Revision der Gruppe Cleonides vrais. Deutsche Entomologische Zeitschrift, 1904 (1): 177 - 284." type="journal article" year="1904">Faust 1904</bibRefCitation>
: 208
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB41A3A430B3363FF60D56E4DFCE49F" blockId="4.[151,1083,152,495]" pageId="4" pageNumber="51">
<treatmentCitationGroup id="AB1B3D14430B3363FF60D56E4DFCE49F" pageId="4" pageNumber="51">
<taxonomicName id="4C0B61B9430B3363FF60D56E4ED9E43F" authority="Lehmanni" authorityName="Lehmanni" box="[151,492,312,335]" class="Insecta" family="Curculionidae" genus="Cleonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="subGenus" subGenus="Epexochus">
<emphasis id="B97FC628430B3363FF60D56E4ED9E43F" box="[151,492,312,335]" italics="true" pageId="4" pageNumber="51">Cleonus (Epexochus) Lehmanni</emphasis>
</taxonomicName>
?
<taxonomicName id="4C0B61B9430B3363FE09D56E4D82E43F" authority="Chevrolat" authorityName="Chevrolat" box="[510,695,312,335]" class="Insecta" family="Curculionidae" genus="Exochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC628430B3363FE09D56E4D07E43F" box="[510,562,312,335]" italics="true" pageId="4" pageNumber="51">latus</emphasis>
(Chevrolat)
</taxonomicName>
:
<treatmentCitation id="0AAA3C2B430B3363FD36D56E4C57E43F" author="Csiki" box="[705,866,312,335]" page="39" pageId="4" pageNumber="51" year="1934">
<bibRefCitation id="EF9A67CB430B3363FD36D56E4C0CE43F" author="Csiki" box="[705,825,312,335]" pageId="4" pageNumber="67" refString="Csiki, E. (1934). Curculionidae. Subfam. Cleoninae. In: Schenkling S. (Ed.). Coleopterorum Catalogus auspiciis et auxilio W. Junk, 134: pp. 152." type="journal article" year="1934">Csiki 1934</bibRefCitation>
: 39
</treatmentCitation>
.
<taxonomicName id="4C0B61B9430B3363FF60D50E4E96E41F" ID-CoL="8HLKB" authority="Lehmanni" authorityName="Lehmanni" box="[151,419,344,367]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC628430B3363FF60D50E4E96E41F" box="[151,419,344,367]" italics="true" pageId="4" pageNumber="51">Leucochromus Lehmanni</emphasis>
</taxonomicName>
<taxonomicName id="4C0B61B9430B3363FE5DD50E4C23E41F" authority="Faust, 1904: 191" authorityName="Faust" authorityPageNumber="191" authorityYear="1904" box="[426,790,344,367]" pageId="4" pageNumber="51" rank="variety" variety="consobrinus">
var.
<emphasis id="B97FC628430B3363FE2ED50E4D6EE41F" box="[473,603,344,367]" italics="true" pageId="4" pageNumber="51">consobrinus</emphasis>
<treatmentCitation id="0AAA3C2B430B3363FD95D50E4C23E41F" author="Faust" box="[610,790,344,367]" page="191" pageId="4" pageNumber="51" year="1904">
<bibRefCitation id="EF9A67CB430B3363FD95D50E4DD7E41F" author="Faust" box="[610,738,344,367]" pageId="4" pageNumber="67" refString="Faust, J. (1904) Revision der Gruppe Cleonides vrais. Deutsche Entomologische Zeitschrift, 1904 (1): 177 - 284." type="journal article" year="1904">Faust, 1904</bibRefCitation>
: 191
</treatmentCitation>
</taxonomicName>
, 192.
<taxonomicName id="4C0B61B9430B3363FF60D52E4E59E4FF" ID-CoL="8KWV7" authority="Lehmanni" authorityName="Lehmanni" box="[151,364,376,399]" class="Insecta" family="Carabidae" genus="Epilectus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC628430B3363FF60D52E4E59E4FF" box="[151,364,376,399]" italics="true" pageId="4" pageNumber="51">Epilectus Lehmanni</emphasis>
</taxonomicName>
<taxonomicName id="4C0B61B9430B3363FE85D52E4D4DE4FF" authority="Faust" authorityName="Faust" box="[370,632,376,399]" pageId="4" pageNumber="51" rank="variety" variety="consobrinus">
var.
<emphasis id="B97FC628430B3363FE57D52E4D16E4FF" box="[416,547,376,399]" italics="true" pageId="4" pageNumber="51">consobrinus</emphasis>
(
<treatmentCitation id="0AAA3C2B430B3363FDC7D52E4D4DE4FF" author="Faust" box="[560,632,376,399]" page="208" pageId="4" pageNumber="51" year="1904">
<bibRefCitation id="EF9A67CB430B3363FDC7D52E4D4DE4FF" author="Faust" box="[560,632,376,399]" pageId="4" pageNumber="67" refString="Faust, J. (1904) Revision der Gruppe Cleonides vrais. Deutsche Entomologische Zeitschrift, 1904 (1): 177 - 284." type="journal article" year="1904">Faust)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
: Faust 1904: 208.
<taxonomicName id="4C0B61B9430B3363FF60D5CE4ED9E4DF" authority="Lehmanni" authorityName="Lehmanni" box="[151,492,408,431]" class="Insecta" family="Curculionidae" genus="Cleonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="subGenus" subGenus="Epexochus">
<emphasis id="B97FC628430B3363FF60D5CE4ED9E4DF" box="[151,492,408,431]" italics="true" pageId="4" pageNumber="51">Cleonus (Epexochus) Lehmanni</emphasis>
</taxonomicName>
ab.
<taxonomicName id="4C0B61B9430B3363FDEFD5CE4DC4E4DF" ID-CoL="6Q5M5" authority="Faust" authorityName="Faust" box="[536,753,408,431]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC628430B3363FDEFD5CE4DAEE4DF" box="[536,667,408,431]" italics="true" pageId="4" pageNumber="51">consobrinus</emphasis>
(Faust)
</taxonomicName>
:
<treatmentCitation id="0AAA3C2B430B3363FD0DD5CE4CAEE4DF" author="Csiki" box="[762,923,408,431]" page="39" pageId="4" pageNumber="51" year="1934">
<bibRefCitation id="EF9A67CB430B3363FD0DD5CE4C47E4DF" author="Csiki" box="[762,882,408,431]" pageId="4" pageNumber="67" refString="Csiki, E. (1934). Curculionidae. Subfam. Cleoninae. In: Schenkling S. (Ed.). Coleopterorum Catalogus auspiciis et auxilio W. Junk, 134: pp. 152." type="journal article" year="1934">Csiki 1934</bibRefCitation>
: 39
</treatmentCitation>
.
<taxonomicName id="4C0B61B9430B3363FF60D5EE4D27E4BF" ID-CoL="6Q5M5" authority="Faust" authorityName="Faust" box="[151,530,440,463]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC628430B3363FF60D5EE4E89E4BF" box="[151,444,440,463]" italics="true" pageId="4" pageNumber="51">Leucochromus consobrinus</emphasis>
(Faust)
</taxonomicName>
:
<treatmentCitation id="0AAA3C2B430B3363FDEBD5EE4C10E4BF" author="Ter-Minasyan" box="[540,805,440,463]" page="543" pageId="4" pageNumber="51" year="1972">
<bibRefCitation id="EF9A67CB430B3363FDEBD5EE4DC5E4BF" author="Ter-Minasyan" box="[540,752,440,463]" pageId="4" pageNumber="68" refString="Ter-Minasyan, M. E. (1972) Zhuki-dolgonosiki podsemeystva Cleoninae (Curculionidae, Coleoptera) sobrannye sovetsko - mongolskimi zoologichelskimi ekspeditsiyami v 1967 - 1969 gg. Nasekomye Mongolii. 1. L.: Nauka: 539 - 556." type="journal article" year="1972">Ter-Minasyan 1972</bibRefCitation>
: 543
</treatmentCitation>
; 1988: 43.
<taxonomicName id="4C0B61B9430B3363FF60D58E4EDFE49F" ID-CoL="6FNCD" authority="Faust" authorityName="Faust" box="[151,490,472,495]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC628430B3363FF60D58E4EA6E49F" box="[151,403,472,495]" italics="true" pageId="4" pageNumber="51">Epexochus consobrinus</emphasis>
(Faust)
</taxonomicName>
:
<treatmentCitation id="0AAA3C2B430B3363FE03D58E4DF1E49F" author="Arzanov" box="[500,708,472,495]" page="150" pageId="4" pageNumber="51" year="2005">
<bibRefCitation id="EF9A67CB430B3363FE03D58E4DBAE49F" author="Arzanov" box="[500,655,472,495]" pageId="4" pageNumber="67" refString="Arzanov, Ju. G. (2005) Taxonomic notes on weevils of the tribe Cleonini (Coleoptera, Curculionidae, Lixinae). Caucasian Entomological Bulletin 1 (2): 150." type="journal article" year="2005">Arzanov 2005</bibRefCitation>
: 150
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB41A3A430B3363FF60D6484CD0E7F6" blockId="4.[151,1437,542,1960]" pageId="4" pageNumber="51">
<emphasis id="B97FC628430B3363FF60D6484E7FE748" bold="true" box="[151,330,542,568]" pageId="4" pageNumber="51">Redescription</emphasis>
. Dimensions: Body length excluding rostrum:
<quantity id="4CF3B7DF430B3363FC72D6484B36E748" box="[901,1027,542,568]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="4" pageNumber="51" unit="mm" value="18.1">18.10 mm</quantity>
. Rostrum: length
<quantity id="4CF3B7DF430B3363FB2BD6484A7EE748" box="[1244,1355,542,568]" metricMagnitude="-3" metricUnit="m" metricValue="3.89" pageId="4" pageNumber="51" unit="mm" value="3.89">3.89 mm</quantity>
, width
<quantity id="4CF3B7DF430B3363FF60D6134FCAE72F" box="[151,255,581,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.99" pageId="4" pageNumber="51" unit="mm" value="1.99">1.99 mm</quantity>
(ratio 1.95). Pronotum: length
<quantity id="4CF3B7DF430B3363FD9CD6134DE2E72F" box="[619,727,581,607]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="4" pageNumber="51" unit="mm" value="5.1">5.10 mm</quantity>
, width
<quantity id="4CF3B7DF430B3363FCDBD6134CA1E72F" box="[812,916,581,607]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="4" pageNumber="51" unit="mm" value="5.9">5.90 mm</quantity>
(ratio 0.86). Elytra: length
<quantity id="4CF3B7DF430B3363FB23D6134A7BE72F" box="[1236,1358,581,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.306" pageId="4" pageNumber="51" unit="mm" value="13.06">13.06 mm</quantity>
, width
<quantity id="4CF3B7DF430B3363FF60D63A4FCAE7F6" box="[151,255,620,646]" metricMagnitude="-3" metricUnit="m" metricValue="8.19" pageId="4" pageNumber="51" unit="mm" value="8.19">8.19 mm</quantity>
(ratio 1.59). Ratio of elytral to pronotal length 2.56 (
<typeStatus id="54B0A498430B3363FC9BD63A4CECE7F6" box="[876,985,620,646]" pageId="4" pageNumber="51" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C31149B1430B3360FF32D6C44E30E3CC" lastPageId="7" lastPageNumber="54" pageId="4" pageNumber="51" type="description">
<paragraph id="8BB41A3A430B3363FF32D6C44FD0E78A" blockId="4.[151,1437,542,1960]" pageId="4" pageNumber="51">
Habitus. Body oval, broad, moderately convex (
<figureCitation id="133006BF430B3363FCF8D6C44C56E7DC" box="[783,867,658,684]" captionStart="FIGURES 1 9" captionStartId="5.[151,270,1918,1942]" captionTargetBox="[236,1367,197,1896]" captionTargetId="figure@5.[236,1370,197,1898]" captionTargetPageId="5" captionText="FIGURES 1 9. Epexochus lehmanni, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5, Exochus latus, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 8 9, specimen from Borokhoro. Scale bars: 10 mm." httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="4" pageNumber="51">Figs. 1</figureCitation>
, 10). Integument dark ferruginous to blackish, densely covered on dorsum with small, simple, acuminate scales and on underside with bifid to multifid scales.
</paragraph>
<paragraph id="8BB41A3A430B3363FF32D7504A47E08F" blockId="4.[151,1437,542,1960]" pageId="4" pageNumber="51">Rostrum straight, more than twice as long as wide, dorso-lateral margins rounded, not keeled, nearly straight from base to apex; epistoma prominent anteriorly, apex straight; dorsum with narrow glossy median keel extending to apex before epistoma, flanked by a broad triangular convex plate, whose sides are broadened anteriad, extending from base, where sides of convexity closely approach median keel, to antennal insertion, where sides nearly reach dorso-lateral margins; dorsal convexity delimiting narrow, relatively shallow dorsal furrows; upper margin of scrobes glossy, evenly curved, sinuate posteriorly, directed towards, and interrupted before, lower margin of eye; lower margin of scrobes parallel to upper margin, reaching underside near base, interrupted on underside at lateral margins of rostrum; underside with keels linearly converging, strongly so towards base and there confluent; surface weakly concave between keels; in lateral view rostrum straight, of equal thickness from base to apex, dorso-lateral margins slightly sinuate basally, median convexity distinctly higher than sides, median keel very weakly curved, sinuate basally, angle between forehead and rostrum strongly obtuse. Surface of rostrum glossy, minutely punctulate and with sparse, moderately larger, isolated punctures, more distinct basally and on sides before eyes. Vestiture consisting of dense, white, glossy, acuminate, simple scales, directed towards middle near base and progressively forward towards apex, progressively more elongate towards apical part, often with two minute and acute lateral basal projections, which towards sides are longer and transform part of the scales into a digitate form with three teeth; sides in front of eyes with sparse, brown trifid scales not hiding integument; pregenae and underside with elliptical, acuminate, glossy white scales; setae sparse, relatively thick, on dorsum short, appressed to integument and limited to basal part, on pregenae and underside longer and thinner, semi-erect, clearly distinct; epistoma punctulate, with microscopic golden setae (Figs. 1213, 16).</paragraph>
<paragraph id="8BB41A3A430B3363FF32D25A4CBCE397" blockId="4.[151,1437,542,1960]" pageId="4" pageNumber="51">Antenna slender. Scape narrow, very weakly thickened at apex, slightly curved anteriad. Funicle with segment 1 subquadrate, 2 cylindrical, as wide as and 1.5 times as long as 1; 36 similar to each other, transverse; 7 subquadrate, slightly larger and twice as long as 6, fused to club. Club elliptical, as long as funicle segments 47 together, club segment 3 as long as previous two segments together, annulus clearly distinct; scape with sparse, ovate or digitate scales and narrow setae, funicle with dense white scales, club with very dense, golden, hair-like scales, lacking setae (Fig. 17).</paragraph>
<paragraph id="8BB41A3A430B3363FF32D2A24C6CE2D8" blockId="4.[151,1437,542,1960]" pageId="4" pageNumber="51">Head broad, nearly as long as wide, not sharply differentiated from base of rostrum, interocular distance wider than rostrum at base, forehead nearly flat, with small interocular pit and narrow median glossy line extending to vertex; this moderately convex in middle. Vestiture consisting of dense, white scales and short, barely distinct, orange setae reclined on integument, inserted in small, spaced punctures. Eyes flat, large, moderately narrowed below, lower margin broadly rounded.</paragraph>
<caption id="DF744AB2430A3362FF60D3284DF5E2AB" httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="5" pageNumber="52" targetBox="[236,1367,197,1896]" targetPageId="5">
<paragraph id="8BB41A3A430A3362FF60D3284DF5E2AB" blockId="5.[151,1436,1918,2011]" pageId="5" pageNumber="52">
<emphasis id="B97FC628430A3362FF60D3284E08E2E6" bold="true" box="[151,317,1918,1942]" pageId="5" pageNumber="52">FIGURES 19</emphasis>
.
<taxonomicName id="4C0B61B9430A3362FEBCD3294D1EE2E6" box="[331,555,1919,1942]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC628430A3362FEBCD3294D1EE2E6" box="[331,555,1919,1942]" italics="true" pageId="5" pageNumber="52">Epexochus lehmanni</emphasis>
</taxonomicName>
, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5,
<taxonomicName id="4C0B61B9430A3362FDFFD3F74D94E2C8" box="[520,673,1953,1976]" class="Insecta" family="Curculionidae" genus="Exochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC628430A3362FDFFD3F74D94E2C8" box="[520,673,1953,1976]" italics="true" pageId="5" pageNumber="52">Exochus latus</emphasis>
</taxonomicName>
, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 89, specimen from Borokhoro. Scale bars: 10 mm.
</paragraph>
</caption>
<paragraph id="8BB41A3A43093361FF60D31D4B25E2BA" blockId="6.[151,1436,1867,1995]" pageId="6" pageNumber="53">
<emphasis id="B97FC62843093361FF60D31D4E69E213" bold="true" box="[151,348,1867,1891]" pageId="6" pageNumber="53">FIGURES 1020</emphasis>
.
<taxonomicName id="4C0B61B943093361FE9DD31A4D7BE213" box="[362,590,1868,1891]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC62843093361FE9DD31A4D7BE213" box="[362,590,1868,1891]" italics="true" pageId="6" pageNumber="53">Epexochus lehmanni</emphasis>
</taxonomicName>
, structural details (10, 1213, 1618:
<typeStatus id="54B0A49843093361FC01D31A4B6FE213" box="[1014,1114,1868,1891]" pageId="6" pageNumber="53" type="holotype">holotype</typeStatus>
; 11, 1415, 1920: specimen from Borokhoro). 10, habitus, lateral view; 11, habitus, ventral view; 1213, rostrum, dorsal and ventral views; 1415, tarsus, ventral and dorsal views; 16, rostrum, lateral view; 17, antenna; 18, pronotum; 1920, maxilla, dorsal and ventral views. Scale bars: 1011:
<quantity id="4CF3B7DF43093361FE5AD3E24ECEE2BB" box="[429,507,1972,1995]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="53" unit="mm" value="10.0">10 mm</quantity>
; 1215, 18:
<quantity id="4CF3B7DF43093361FD88D3E24DF5E2BB" box="[639,704,1972,1995]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="53" unit="mm" value="2.0">2 mm</quantity>
; 1617:
<quantity id="4CF3B7DF43093361FCECD3E24C69E2BB" box="[795,860,1972,1995]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="53" unit="mm" value="1.0">1 mm</quantity>
; 1920: 500 μm.
</paragraph>
<paragraph id="8BB41A3A43083360FF32D4CE4DAEE744" blockId="7.[151,1438,152,2034]" pageId="7" pageNumber="54">Pronotum broad, nearly flat, base subrectilinear, moderately and evenly arched, posteriorly not medially protruding towards elytra, sides nearly parallel or feebly diverging towards apical quarter, then moderately, evenly curved to apex; anterior margin very weakly sinuate; surface smooth, minutely punctulate and with sparse, slightly larger, round punctures, often with a small, glossy, round bare patch near lateral margin; median line glossy, very narrow, barely distinct at the extremes, usually slightly more evident and broadened on disc; pronotum in lateral view faintly convex, maximum height near base. Vestiture consisting of oval, acutely pointed scales inserted in minute punctures, usually white-yellowish on dorso-lateral part and on sides, dark brown on disc forming an irregular pattern, narrowed anteriorly and widened posteriorly, with some whitish spots inside, and also brown on part of sides, mainly forming an anterior patch; scales on sides progressively more slender, setae orange or light brown, short and completely appressed against integument, inserted in larger sparse punctures (Fig. 18).</paragraph>
<paragraph id="8BB41A3A43083360FF32D6174DAEE72B" blockId="7.[151,1438,152,2034]" box="[197,667,577,603]" pageId="7" pageNumber="54">Scutellum very small, triangular, setose.</paragraph>
<paragraph id="8BB41A3A43083360FF32D63E4B26E6AE" blockId="7.[151,1438,152,2034]" pageId="7" pageNumber="54">
Elytra ovalelliptical, in lateral view moderately convex, maximum height behind middle of length, declivity oblique; base weakly arched, humeri not prominent, sides weakly curvilinear, with maximum width behind middle of length, evenly rounded at apex. Intervals wide, even ones narrower than odd ones, 3 and 5 clearly higher than others, particularly in basal half, 6 and 8 not reaching base, 7 and 9 fused basally into a broad, weakly convex hump, separated by a narrow furrow from base of interval 5. Striae very narrow and shallow, with barely distinct punctures. Vestiture consisting of very dense, oval, acutely pointed, whitish alternating with brown scales in very variable patterns; the latter frequent on dorsum in a more or less continuous layer, where white scales form irregularly scattered patches; intervals 711 usually with whitish scales and seldom with brown patches; interval 11 with digitate scales; setae golden coloured, semi-erect, shorter than an interval width, more distinct on declivity and sides (
<figureCitation id="133006BF43083360FC59D7924B33E6AE" box="[942,1030,964,990]" captionStart="FIGURES 21 39" captionStartId="12.[151,269,1819,1843]" captionTargetBox="[260,1326,194,1795]" captionTargetId="figure@12.[260,1328,194,1795]" captionTargetPageId="12" captionText="FIGURES 21 39. Epexochus, genitalia, claws and scales. Female genitalia (21 23: specimen from Kzyl-Dzhar; 24 25: specimen from Aulie-ata). 21, 24, sternum VIII; 22, 25, spermatheca; 23, hemisternites. Claws. 26, specimen from Borokhoro. Aedeagus, dorsal and lateral views and detail of apex. 27, 30, 35, specimen from Koibun; 28 29, 32, specimen from Elton lake; 31, specimen from Dzharkentsk; 33, specimen from Betpak; 34, specimen from Borokhoro. Elytral scales. 36, holotype of E. lehmanni; 37, holotype of E. korotyaevi; 38, holotype of E. voriseki; 39, paratype of E. mongolicus. Scale bars: 21, 23 24, 27 31: 1 mm; 26, 32 35: 500 μm; 36 39: 100 μm." httpUri="https://zenodo.org/record/274689/files/figure.png" pageId="7" pageNumber="54">Fig. 36</figureCitation>
).
</paragraph>
<paragraph id="8BB41A3A43083360FF32D7BC4A07E063" blockId="7.[151,1438,152,2034]" pageId="7" pageNumber="54">
Legs slender. Femora slightly thickened medially. Tibiae narrow, fore tibia weakly sinuate, middle and hind tibiae straight, apex not expanded; vestiture consisting of dense, whitish scales, often replaced with brown scales near apex of femora, and dense long, semi-erect, white setae inserted in distinct, small punctures. Tarsi with segments 1 and 2 of fore and middle tarsi triangular, 2 broader than 1, 3 lobed, lobes very shortly expanded; segment 1 of hind tarsus longer, nearly three times as long as wide, 2 slightly longer than wide, 3 very short; all segments with white scales and setae expanded side- and downwards (Fig. 15); onychium elongated, slightly thickened in apical half; underside lacking adhesive pad, replaced by downward-directed, thick, orange setae (Fig. 14); claws slender, divergent, connate at base (
<figureCitation id="133006BF43083360FB3AD0AF4A10E063" box="[1229,1317,1273,1299]" captionStart="FIGURES 21 39" captionStartId="12.[151,269,1819,1843]" captionTargetBox="[260,1326,194,1795]" captionTargetId="figure@12.[260,1328,194,1795]" captionTargetPageId="12" captionText="FIGURES 21 39. Epexochus, genitalia, claws and scales. Female genitalia (21 23: specimen from Kzyl-Dzhar; 24 25: specimen from Aulie-ata). 21, 24, sternum VIII; 22, 25, spermatheca; 23, hemisternites. Claws. 26, specimen from Borokhoro. Aedeagus, dorsal and lateral views and detail of apex. 27, 30, 35, specimen from Koibun; 28 29, 32, specimen from Elton lake; 31, specimen from Dzharkentsk; 33, specimen from Betpak; 34, specimen from Borokhoro. Elytral scales. 36, holotype of E. lehmanni; 37, holotype of E. korotyaevi; 38, holotype of E. voriseki; 39, paratype of E. mongolicus. Scale bars: 21, 23 24, 27 31: 1 mm; 26, 32 35: 500 μm; 36 39: 100 μm." httpUri="https://zenodo.org/record/274689/files/figure.png" pageId="7" pageNumber="54">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="8BB41A3A43083360FF32D1764E49E352" blockId="7.[151,1438,152,2034]" pageId="7" pageNumber="54">Venter. Prosternum with low hump before fore coxae. Ventrite I relatively narrow, width between hind coxae and margin narrower than space between coxae; II slightly narrower than I; III and IV moderately narrower than II; V short, clearly impressed before apex; vestiture consisting of very dense coating of elongate, slender, bifid or multifid scales, white with a narrow brown basal patch, slightly expanded towards sides, on ventrites IIIV; ventrite V with a broader brown triangular patch, narrowed apically and nearly reaching apex; setae whitish, long, very narrow and relatively dense, inserted in barely visible, small punctures (Fig. 11).</paragraph>
<paragraph id="8BB41A3A43083360FF32D2784E30E3CC" blockId="7.[151,1438,152,2034]" pageId="7" pageNumber="54">
Genitalia. Aedeagus slender, tubular, curved, lamella shortly sub-triangular, apex subacute or narrowly rounded (
<figureCitation id="133006BF43083360FEFFD2034EACE31F" box="[264,409,1621,1647]" captionStart="FIGURES 21 39" captionStartId="12.[151,269,1819,1843]" captionTargetBox="[260,1326,194,1795]" captionTargetId="figure@12.[260,1328,194,1795]" captionTargetPageId="12" captionText="FIGURES 21 39. Epexochus, genitalia, claws and scales. Female genitalia (21 23: specimen from Kzyl-Dzhar; 24 25: specimen from Aulie-ata). 21, 24, sternum VIII; 22, 25, spermatheca; 23, hemisternites. Claws. 26, specimen from Borokhoro. Aedeagus, dorsal and lateral views and detail of apex. 27, 30, 35, specimen from Koibun; 28 29, 32, specimen from Elton lake; 31, specimen from Dzharkentsk; 33, specimen from Betpak; 34, specimen from Borokhoro. Elytral scales. 36, holotype of E. lehmanni; 37, holotype of E. korotyaevi; 38, holotype of E. voriseki; 39, paratype of E. mongolicus. Scale bars: 21, 23 24, 27 31: 1 mm; 26, 32 35: 500 μm; 36 39: 100 μm." httpUri="https://zenodo.org/record/274689/files/figure.png" pageId="7" pageNumber="54">Figs. 2735</figureCitation>
). Sternite VIII of Ψ with narrow arms, connate at base, lamina sclerotised only at apex of arms (
<figureCitation id="133006BF43083360FEF7D22A4E6DE3E6" box="[256,344,1660,1686]" captionStart="FIGURES 21 39" captionStartId="12.[151,269,1819,1843]" captionTargetBox="[260,1326,194,1795]" captionTargetId="figure@12.[260,1328,194,1795]" captionTargetPageId="12" captionText="FIGURES 21 39. Epexochus, genitalia, claws and scales. Female genitalia (21 23: specimen from Kzyl-Dzhar; 24 25: specimen from Aulie-ata). 21, 24, sternum VIII; 22, 25, spermatheca; 23, hemisternites. Claws. 26, specimen from Borokhoro. Aedeagus, dorsal and lateral views and detail of apex. 27, 30, 35, specimen from Koibun; 28 29, 32, specimen from Elton lake; 31, specimen from Dzharkentsk; 33, specimen from Betpak; 34, specimen from Borokhoro. Elytral scales. 36, holotype of E. lehmanni; 37, holotype of E. korotyaevi; 38, holotype of E. voriseki; 39, paratype of E. mongolicus. Scale bars: 21, 23 24, 27 31: 1 mm; 26, 32 35: 500 μm; 36 39: 100 μm." httpUri="https://zenodo.org/record/274689/files/figure.png" pageId="7" pageNumber="54">Fig. 21</figureCitation>
). Spermatheca with ramus usually sub-basal (
<figureCitation id="133006BF43083360FC85D22A4B34E3E6" box="[882,1025,1660,1686]" captionStart="FIGURES 21 39" captionStartId="12.[151,269,1819,1843]" captionTargetBox="[260,1326,194,1795]" captionTargetId="figure@12.[260,1328,194,1795]" captionTargetPageId="12" captionText="FIGURES 21 39. Epexochus, genitalia, claws and scales. Female genitalia (21 23: specimen from Kzyl-Dzhar; 24 25: specimen from Aulie-ata). 21, 24, sternum VIII; 22, 25, spermatheca; 23, hemisternites. Claws. 26, specimen from Borokhoro. Aedeagus, dorsal and lateral views and detail of apex. 27, 30, 35, specimen from Koibun; 28 29, 32, specimen from Elton lake; 31, specimen from Dzharkentsk; 33, specimen from Betpak; 34, specimen from Borokhoro. Elytral scales. 36, holotype of E. lehmanni; 37, holotype of E. korotyaevi; 38, holotype of E. voriseki; 39, paratype of E. mongolicus. Scale bars: 21, 23 24, 27 31: 1 mm; 26, 32 35: 500 μm; 36 39: 100 μm." httpUri="https://zenodo.org/record/274689/files/figure.png" pageId="7" pageNumber="54">Figs. 22, 25</figureCitation>
). Hemisternites as typical of genus (
<figureCitation id="133006BF43083360FF68D2F44FC2E3CC" box="[159,247,1698,1724]" captionStart="FIGURES 21 39" captionStartId="12.[151,269,1819,1843]" captionTargetBox="[260,1326,194,1795]" captionTargetId="figure@12.[260,1328,194,1795]" captionTargetPageId="12" captionText="FIGURES 21 39. Epexochus, genitalia, claws and scales. Female genitalia (21 23: specimen from Kzyl-Dzhar; 24 25: specimen from Aulie-ata). 21, 24, sternum VIII; 22, 25, spermatheca; 23, hemisternites. Claws. 26, specimen from Borokhoro. Aedeagus, dorsal and lateral views and detail of apex. 27, 30, 35, specimen from Koibun; 28 29, 32, specimen from Elton lake; 31, specimen from Dzharkentsk; 33, specimen from Betpak; 34, specimen from Borokhoro. Elytral scales. 36, holotype of E. lehmanni; 37, holotype of E. korotyaevi; 38, holotype of E. voriseki; 39, paratype of E. mongolicus. Scale bars: 21, 23 24, 27 31: 1 mm; 26, 32 35: 500 μm; 36 39: 100 μm." httpUri="https://zenodo.org/record/274689/files/figure.png" pageId="7" pageNumber="54">Fig. 23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31149B14308336FFF32D29E4BB7E174" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54" type="materials_examined">
<paragraph id="8BB41A3A43083360FF32D29E4D49E2D4" blockId="7.[151,1438,152,2034]" pageId="7" pageNumber="54">
<emphasis id="B97FC62843083360FF32D29E4EF0E392" bold="true" box="[197,453,1736,1762]" pageId="7" pageNumber="54">Material examined.</emphasis>
<emphasis id="B97FC62843083360FE39D29F4C7CE392" box="[462,841,1737,1762]" italics="true" pageId="7" pageNumber="54">
<typeStatus id="54B0A49843083360FE39D29F4D32E392" box="[462,519,1737,1762]" pageId="7" pageNumber="54">Type</typeStatus>
series.
<taxonomicName id="4C0B61B943083360FD9DD29F4C7CE392" authority="Menetries" authorityName="Menetries" box="[618,841,1737,1762]" class="Insecta" family="Curculionidae" genus="Cleonus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="lehmanni">Cleonus lehmanni</taxonomicName>
</emphasis>
Ménétries,
<typeStatus id="54B0A49843083360FC28D29F4B7EE393" box="[991,1099,1737,1763]" pageId="7" pageNumber="54" type="holotype">holotype</typeStatus>
Ψ:
<collectingCountry id="F31C5AAA43083360FB8ED29F4A11E393" box="[1145,1316,1737,1763]" name="Kazakhstan" pageId="7" pageNumber="54">KAZAKHSTAN</collectingCountry>
, AKTOBE
<collectionCode id="ED1A82FF43083360FF60D2A54FE6E278" box="[151,211,1779,1800]" pageId="7" pageNumber="54">PROV</collectionCode>
.: &quot;Emba-Steppe [approximately 48°N 57°E], … [illegible], 1840 /
<taxonomicName id="4C0B61B943083360FC13D2A64B8CE279" box="[996,1209,1776,1801]" class="Insecta" family="Curculionidae" genus="Cleonus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC62843083360FC13D2A64B8CE279" box="[996,1209,1776,1801]" italics="true" pageId="7" pageNumber="54">Cleonus lehmanni</emphasis>
</taxonomicName>
m. / Ménétries det. [pr.] /
<taxonomicName id="4C0B61B943083360FF1ED3414E53E240" box="[233,358,1815,1840]" class="Insecta" family="Curculionidae" genus="Cleonus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC62843083360FF1ED3414E53E240" box="[233,358,1815,1840]" italics="true" pageId="7" pageNumber="54">Lehmanni</emphasis>
</taxonomicName>
Ménétr. /
<taxonomicName id="4C0B61B943083360FE1BD3414C98E240" authority="Menetries 1849" authorityName="Menetries" authorityYear="1849" box="[492,941,1814,1840]" class="Insecta" family="Curculionidae" genus="Cleonus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC62843083360FE1BD3414DE7E240" box="[492,722,1815,1840]" italics="true" pageId="7" pageNumber="54">Cleonus lehmanni</emphasis>
,
<bibRefCitation id="EF9A67CB43083360FD13D3404C98E240" author="Menetries" box="[740,941,1814,1840]" pageId="7" pageNumber="67" refString="Menetries, M. (1849) Catalogue des insectes recueillis par feu M. Lehmann avec les descriptions des nouvelles especes. Seconde et derniere partie. Tetramere. Curculionites. Memoires de l'Academie Imperiale des Sciences de Saint- Petersbourg. Sixieme Serie. Sciences Naturelles, VI: 249 - 263." type="book chapter" year="1849">Ménétries 1849</bibRefCitation>
</taxonomicName>
,
<typeStatus id="54B0A49843083360FC4BD3404B76E240" box="[956,1091,1814,1840]" pageId="7" pageNumber="54" type="holotype">Holotypus</typeStatus>
, 2008 Meregalli vid. (red).
<taxonomicName id="4C0B61B943083360FF60D36B4EF4E226" box="[151,449,1853,1878]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC62843083360FF60D36B4EF4E226" box="[151,449,1853,1878]" italics="true" pageId="7" pageNumber="54">Leucochromus lehmanni</emphasis>
</taxonomicName>
<taxonomicName id="4C0B61B943083360FE3ED36B4DD5E227" authority="Faust" authorityName="Faust" box="[457,736,1853,1879]" pageId="7" pageNumber="54" rank="variety" variety="consobrinus">
var.
<emphasis id="B97FC62843083360FE09D36B4DA7E226" box="[510,658,1853,1878]" italics="true" pageId="7" pageNumber="54">consobrinus</emphasis>
Faust
</taxonomicName>
, 3
<typeStatus id="54B0A49843083360FCF3D36B4C46E227" box="[772,883,1853,1879]" pageId="7" pageNumber="54" type="syntype">syntypes</typeStatus>
:
<collectingCountry id="F31C5AAA43083360FC88D36B4B1DE227" box="[895,1064,1853,1879]" name="Kazakhstan" pageId="7" pageNumber="54">KAZAKHSTAN</collectingCountry>
, ZHAMBYL
<collectionCode id="ED1A82FF43083360FB4CD3174BCCE226" box="[1211,1273,1857,1878]" pageId="7" pageNumber="54">PROV</collectionCode>
.: &quot;Syr Daria, Aulie Ata&quot; [= Taraz,
<geoCoordinate id="EE3F7CFD43083360FE50D3324D25E20E" box="[423,528,1892,1918]" direction="north" orientation="latitude" pageId="7" pageNumber="54" precision="925" value="42.883335">42°53'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD43083360FDEED3324DB4E20E" box="[537,641,1892,1918]" direction="east" orientation="longitude" pageId="7" pageNumber="54" precision="925" value="71.36667">71°22'E</geoCoordinate>
] (
<collectionCode id="ED1A82FF43083360FD6CD3324DCCE20E" box="[667,761,1892,1918]" pageId="7" pageNumber="54">SMTD</collectionCode>
).
<taxonomicName id="4C0B61B943083360FCE7D3324B78E20E" authority="Chevrolat" authorityName="Chevrolat" box="[784,1101,1892,1918]" class="Insecta" family="Curculionidae" genus="Exochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC62843083360FCE7D3324CF7E20D" box="[784,962,1892,1917]" italics="true" pageId="7" pageNumber="54">Exochus latus</emphasis>
Chevrolat
</taxonomicName>
,
<typeStatus id="54B0A49843083360FBACD3324BFFE20E" box="[1115,1226,1892,1918]" pageId="7" pageNumber="54" type="holotype">holotype</typeStatus>
Ψ (see
<bibRefCitation id="EF9A67CB43083360FAD9D3324E62E2D4" author="Perrin" pageId="7" pageNumber="67" refString="Perrin, H. &amp; Meregalli, M. (2008) Designation de lectotypes de Cleonini decrits par Gebler et Chevrolat, dans les collections Entomologiques du Museum national d'Histoire naturelle (Coleoptera: Curculionidae: Lixinae). Nouvelle revue d'Entomologie, 29: 129 - 148." type="journal article" year="2008">Perrin &amp; Meregalli, 2008</bibRefCitation>
): &quot;Kirghis or.&quot; (
<collectionCode id="ED1A82FF43083360FDF8D3DC4D5AE2D4" LSID="urn:lsid:biocol.org:col:34988" box="[527,623,1930,1956]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="54">MNHN</collectionCode>
).
</paragraph>
<paragraph id="8BB41A3A4308336FFF32D3E74BB7E174" blockId="7.[151,1438,152,2034]" lastBlockId="8.[151,1438,151,2034]" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54">
<emphasis id="B97FC62843083360FF32D3E74EFEE2BA" box="[197,459,1969,1994]" italics="true" pageId="7" pageNumber="54">Non-type specimens.</emphasis>
<collectingCountry id="F31C5AAA43083360FE24D3E74D05E2BB" box="[467,560,1969,1995]" name="Russia" pageId="7" pageNumber="54">RUSSIA</collectingCountry>
, VOLGOGRAD
<collectionCode id="ED1A82FF43083360FD1FD3E34C1DE2BA" box="[744,808,1973,1994]" pageId="7" pageNumber="54">PROV</collectionCode>
.: &quot;Elton ok., ozera na doroge [
<geoCoordinate id="EE3F7CFD43083360FB45D3E74A29E2BB" box="[1202,1308,1969,1995]" direction="north" orientation="latitude" pageId="7" pageNumber="54" precision="925" value="49.116665">49°07'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD43083360FAD2D3E74AA5E2BB" box="[1317,1424,1969,1995]" direction="east" orientation="longitude" pageId="7" pageNumber="54" precision="925" value="46.566666">46°34'E</geoCoordinate>
],
<date id="FFB53CFA43083360FF60D38E4FEDE282" box="[151,216,2008,2034]" pageId="7" pageNumber="54">4.VI.</date>
[19]52, Buriascheva&quot;, 1 ɗ (
<collectionCode id="ED1A82FF43083360FDC0D38E4D47E282" box="[567,626,2008,2034]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="7" pageNumber="54">ZIN</collectionCode>
).
<collectingCountry id="F31C5AAA43083360FD7ED38E4C02E282" box="[649,823,2008,2034]" name="Kazakhstan" pageId="7" pageNumber="54">KAZAKHSTAN</collectingCountry>
: &quot;
<quantity id="4CF3B7DF43083360FCA4D38E4C83E282" box="[851,950,2008,2034]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="54" unit="km" value="200.0">200 km</quantity>
ot [from] Atbasara [Atbasar:
<geoCoordinate id="EE3F7CFD43083360FAC2D38E4AA8E282" box="[1333,1437,2008,2034]" direction="north" orientation="latitude" pageId="7" pageNumber="54" precision="925" value="51.283333">51°17'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD4307336FFF60D4CE4FC9E5C2" box="[151,252,152,178]" direction="east" orientation="longitude" pageId="8" pageNumber="55" precision="925" value="68.316666">68°19'E</geoCoordinate>
], Akmol. obl., A. Emeljanov,
<date id="FFB53CFA4307336FFD9FD4CE4D93E5C2" box="[616,678,152,178]" pageId="8" pageNumber="55">3.VI.</date>
[19]57&quot;, 1 ɗ (
<collectionCode id="ED1A82FF4307336FFCB8D4CE4CB2E5C2" box="[847,903,152,178]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
). &quot;Turg. obl., Myn-sai [approximately 49°N 65°E],
<date id="FFB53CFA4307336FFF1BD4E84E29E5A8" box="[236,284,190,216]" pageId="8" pageNumber="55">3.V.</date>
[19]14&quot;, 1 ɗ (
<collectionCode id="ED1A82FF4307336FFE36D4E84ECDE5A8" box="[449,504,190,216]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Betpak-Dala, Sary-Kamys [
<geoCoordinate id="EE3F7CFD4307336FFC96D4E84CF0E5A8" box="[865,965,190,216]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="46.033333">46°02'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD4307336FFC39D4E84B04E5A8" box="[974,1073,190,216]" direction="east" orientation="longitude" pageId="8" pageNumber="55" precision="925" value="70.2">70°12'E</geoCoordinate>
],
<date id="FFB53CFA4307336FFBB1D4E84B43E5A8" box="[1094,1142,190,216]" pageId="8" pageNumber="55">7.V.</date>
[19]55, L. Serkova&quot;, 1 ɗ (
<collectionCode id="ED1A82FF4307336FFF68D4B34FE3E58F" box="[159,214,229,255]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;
<quantity id="4CF3B7DF4307336FFF04D4B34E75E58F" box="[243,320,229,255]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="55" unit="km" value="40.0">40 km</quantity>
<collectingCountry id="F31C5AAA4307336FFEBFD4B34E68E58F" box="[328,349,229,255]" name="Iceland" pageId="8" pageNumber="55">O</collectingCountry>
st. Kzyl-Dzhar [
<geoCoordinate id="EE3F7CFD4307336FFDD7D4B34DB6E58F" box="[544,643,229,255]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="48.3">48°18'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD4307336FFD7CD4B34DD8E58F" box="[651,749,229,255]" direction="east" orientation="longitude" pageId="8" pageNumber="55" precision="925" value="69.63333">69°38'E</geoCoordinate>
], Karagand. obl., L. Arnoldi,
<date id="FFB53CFA4307336FFBB0D4B34BBCE58F" box="[1095,1161,229,255]" pageId="8" pageNumber="55">23.V.</date>
[1]962&quot;, 1 ɗ (
<collectionCode id="ED1A82FF4307336FFAD0D4B34A6BE58F" box="[1319,1374,229,255]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;
<quantity id="4CF3B7DF4307336FFA8BD4B34F88E456" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="55" unit="km" value="10.0">10 km</quantity>
SV [N-E] g. Balkhasha, Z. Matis,
<date id="FFB53CFA4307336FFD8DD55A4DC8E456" box="[634,765,268,294]" pageId="8" pageNumber="55" value="1966-05-23">23.V.1966</date>
&quot;, 1 ɗ (
<collectionCode id="ED1A82FF4307336FFCA6D55A4CBFE456" box="[849,906,268,294]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Syr Daria, Aulie Ata&quot; [= Taraz,
<geoCoordinate id="EE3F7CFD4307336FFACFD55A4AA8E456" box="[1336,1437,268,294]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="42.883335">42°53'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD4307336FFF60D5644FCCE43C" box="[151,249,306,332]" direction="east" orientation="longitude" pageId="8" pageNumber="55" precision="925" value="71.36667">71°22'E</geoCoordinate>
], 102 ex (100 exx
<collectionCode id="ED1A82FF4307336FFE38D5644D16E43C" box="[463,547,306,332]" pageId="8" pageNumber="55">SMTD</collectionCode>
, 2 ex MER); id, under
<taxonomicName id="4C0B61B94307336FFCD9D5644C89E43B" box="[814,956,306,331]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284307336FFCD9D5644C89E43B" box="[814,956,306,331]" italics="true" pageId="8" pageNumber="55">C. lehmanni</emphasis>
</taxonomicName>
<taxonomicName id="4C0B61B94307336FFC33D5644BFBE43C" authority="Faust" authorityName="Faust" box="[964,1230,306,332]" pageId="8" pageNumber="55" rank="variety" variety="consobrinus">
var.
<emphasis id="B97FC6284307336FFC02D5644BB1E43B" box="[1013,1156,306,331]" italics="true" pageId="8" pageNumber="55">consobrinus</emphasis>
Faust
</taxonomicName>
, 55 exx (
<collectionCode id="ED1A82FF4307336FFAC0D5644ABAE43C" box="[1335,1423,306,332]" pageId="8" pageNumber="55">SMTD</collectionCode>
); &quot;Turkest.: Aulie-ata, E. Wilberg&quot;, 2 Ψ (MER); &quot;Aulie Ata&quot;, 2 exx (MER); &quot;Semirechye [approximately 4346°N 7479°E], step k Balkhashy, Makhonin, 1909&quot; 1 ɗ (
<collectionCode id="ED1A82FF4307336FFC9ED5D64C95E4EA" box="[873,928,384,410]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Okr. oz. Balkhasha,
<date id="FFB53CFA4307336FFB5AD5D64A36E4EA" box="[1197,1283,384,410]" pageId="8" pageNumber="55" value="1907-05">V.1907</date>
, Mamisson&quot;, 1 ɗ (
<collectionCode id="ED1A82FF4307336FFF26D5F04E3DE4B0" box="[209,264,422,448]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Koibun, Semirjetschensk, coll. Winkler, 3 ɗ 1 Ψ. (MER); &quot;Balchasch, Matthiessen&quot;, 1 ɗ (POD); &quot;Yu-V [S-E]
<collectingCountry id="F31C5AAA4307336FFEC0D59B4EFDE497" box="[311,456,461,487]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
, Taldy - Kurgan. obl., pustynya Ak-Tau,
<collectingCountry id="F31C5AAA4307336FFC37D59B4B35E497" box="[960,1024,461,487]" name="Vietnam" pageId="8" pageNumber="55">V. N.</collectingCountry>
Plasolov,
<date id="FFB53CFA4307336FFB77D59B4A37E497" box="[1152,1282,461,487]" pageId="8" pageNumber="55" value="1994-05-10">10.V.1994</date>
&quot;, 1 Ψ (
<collectionCode id="ED1A82FF4307336FFAA1D59B4ABAE497" box="[1366,1423,461,487]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Okr. Kopala [
<geoCoordinate id="EE3F7CFD4307336FFEB5D5A24E90E77E" box="[322,421,500,526]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="45.133335">45°08'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD4307336FFE5AD5A24D25E77E" box="[429,528,500,526]" direction="east" orientation="longitude" pageId="8" pageNumber="55" precision="925" value="79.01667">79°01'E</geoCoordinate>
],
<date id="FFB53CFA4307336FFDD3D5A24D9DE77E" box="[548,680,500,526]" pageId="8" pageNumber="55" value="1910-04-18">18.IV.1910</date>
, Lukjanovitsch&quot;, 1 ɗ (
<collectionCode id="ED1A82FF4307336FFC45D5A24CDCE77E" box="[946,1001,500,526]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Kazakhst. m.-or., Saryesik-Atyrau des., Enbek [
<geoCoordinate id="EE3F7CFD4307336FFED9D64C4EA4E744" box="[302,401,538,564]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="45.25">45°15'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD4307336FFE6ED64C4EC9E744" box="[409,508,538,564]" direction="east" orientation="longitude" pageId="8" pageNumber="55" precision="925" value="75.48333">75°29'E</geoCoordinate>
],
<date id="FFB53CFA4307336FFDF8D64C4D4DE744" box="[527,632,538,564]" pageId="8" pageNumber="55" value="1990-05-01">1.5.1990</date>
, Beneš + Voříšek leg.&quot;, 2 Ψ (1 Ψ MER, 1 Ψ VOR); &quot;
<collectingCountry id="F31C5AAA4307336FFB29D64C4A5FE744" box="[1246,1386,538,564]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
SE, Bakanas vill. [
<geoCoordinate id="EE3F7CFD4307336FFECAD6174E95E72B" box="[317,416,577,603]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="44.8">44°48'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD4307336FFE50D6174D3CE72B" box="[423,521,577,603]" direction="east" orientation="longitude" pageId="8" pageNumber="55" precision="925" value="76.26667">76°16'E</geoCoordinate>
], 04/1993, leg. Saldaitiz&quot; 1 Ψ (
<collectionCode id="ED1A82FF4307336FFC9AD6174CF7E72B" box="[877,962,577,603]" pageId="8" pageNumber="55">CSNV</collectionCode>
); &quot;S.E.
<collectingCountry id="F31C5AAA4307336FFBEFD6174B91E72B" box="[1048,1188,577,603]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
, Ili riv., Bakanas vill,
<date id="FFB53CFA4307336FFF60D63E4E31E7F2" box="[151,260,616,642]" pageId="8" pageNumber="55" value="1996-05-02">2.5.1996</date>
&quot;, 1 ɗ (
<collectionCode id="ED1A82FF4307336FFEA6D63E4E90E7F2" box="[337,421,616,642]" pageId="8" pageNumber="55">CSNV</collectionCode>
); &quot;
<collectingCountry id="F31C5AAA4307336FFE35D63E4D64E7F2" box="[450,593,616,642]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
, Ily desert, S fr. Bakanas, 2/
<date id="FFB53CFA4307336FFC6BD63E4B33E7F2" box="[924,1030,616,642]" pageId="8" pageNumber="55" value="1997-05-04">4.V.1997</date>
, Klimenko A., 2 ɗ (1 ɗ MER, 1 ɗ
<collectionCode id="ED1A82FF4307336FFF60D6D84FDBE7D8" box="[151,238,654,680]" pageId="8" pageNumber="55">CSNV</collectionCode>
); &quot;
<collectingCountry id="F31C5AAA4307336FFEFBD6D84E97E7D8" box="[268,418,654,680]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
, Prov. Almaty,
<quantity id="4CF3B7DF4307336FFD94D6D84D97E7D8" box="[611,674,654,680]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="55" unit="km" value="5.0">5 km</quantity>
NE Burundisu,
<quantity id="4CF3B7DF4307336FFC9AD6D84C8BE7D8" box="[877,958,654,680]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="55" unit="m" value="550.0">550 m</quantity>
,
<geoCoordinate id="EE3F7CFD4307336FFC3CD6D84B1BE7D8" box="[971,1070,654,680]" direction="east" orientation="longitude" pageId="8" pageNumber="55" precision="925" value="78.63333">78°38'E</geoCoordinate>
<geoCoordinate id="EE3F7CFD4307336FFBC1D6D84BABE7D8" box="[1078,1182,654,680]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="43.716667">43°43'N</geoCoordinate>
,
<date id="FFB53CFA4307336FFB5BD6D84A1BE7D8" box="[1196,1326,654,680]" pageId="8" pageNumber="55" value="1994-05-17">17.V.1994</date>
, leg. Gy. Fábián &amp; I. Retezár&quot;, 2 ɗ 2 Ψ (2 ɗ 1 Ψ POD, 1 Ψ MER); &quot;Mzhd. Tshilikom i Tsharynom,
<date id="FFB53CFA4307336FFB2ED6E34A2CE7BF" box="[1241,1305,693,719]" pageId="8" pageNumber="55">13.V.</date>
[18]89, Gr. Grzhimailo&quot;, 1 ɗ (
<collectionCode id="ED1A82FF4307336FFE81D68A4E98E786" box="[374,429,732,758]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;
<collectingCountry id="F31C5AAA4307336FFE3BD68A4D68E786" box="[460,605,732,758]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
, 1965, r. Tsharyn, ur. Sartogai [
<geoCoordinate id="EE3F7CFD4307336FFC21D68A4B0EE786" box="[982,1083,732,758]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="43.516666">43°31'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD4307336FFBB4D68A4B90E786" box="[1091,1189,732,758]" direction="east" orientation="longitude" pageId="8" pageNumber="55" precision="925" value="79.23333">79°14'E</geoCoordinate>
],
<date id="FFB53CFA4307336FFB33D68A4A0AE786" box="[1220,1343,732,758]" pageId="8" pageNumber="55" value="1964-06-03">3.VI.1964</date>
, Kostin Badenko&quot;, 1 ɗ 1 Ψ (
<collectionCode id="ED1A82FF4307336FFE72D7544E89E66C" box="[389,444,770,796]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Dzharkentsk. utz., r. Ili [
<geoCoordinate id="EE3F7CFD4307336FFD0DD7544C6BE66C" box="[762,862,770,796]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="43.833332">43°50'N</geoCoordinate>
<geoCoordinate id="EE3F7CFD4307336FFC92D7544CF2E66C" box="[869,967,770,796]" direction="east" orientation="longitude" pageId="8" pageNumber="55" precision="925" value="80.0">80°00'E</geoCoordinate>
], 1/
<date id="FFB53CFA4307336FFC18D7544B55E66C" box="[1007,1120,770,796]" pageId="8" pageNumber="55" value="1909-05-02">2.V.1909</date>
&quot;, 1 Ψ (
<collectionCode id="ED1A82FF4307336FFB46D7544BDDE66C" box="[1201,1256,770,796]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Zailiskii Ala
<collectingCountry id="F31C5AAA4307336FFF60D77F4FF2E633" box="[151,199,809,835]" name="China" pageId="8" pageNumber="55">Tau</collectingCountry>
, sklon Moyun-saz (?), 20.06.[19]23, Pashina&quot; 1 ɗ (
<collectionCode id="ED1A82FF4307336FFCCBD77F4C40E633" box="[828,885,809,835]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
).
<collectingCountry id="F31C5AAA4307336FFC7DD77F4CEFE633" box="[906,986,809,835]" name="China" pageId="8" pageNumber="55">CHINA</collectingCountry>
, XINJIANG: &quot;Dzhungaria, Borokhoro Geb. [
<geoCoordinate id="EE3F7CFD4307336FFF15D7064E7FE61A" box="[226,330,848,874]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="43.916668">43°55'N</geoCoordinate>
82°06'], Coll. Hauser, 6.05&quot;, 59 ex (18 exx: MER; 2 ex: POD; 4 ex:
<collectionCode id="ED1A82FF4307336FFB5BD7064A3DE61A" box="[1196,1288,848,874]" pageId="8" pageNumber="55">ZMHB</collectionCode>
; 35
<collectionCode id="ED1A82FF4307336FFACBD7064ABAE61A" box="[1340,1423,848,874]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="55">NMW</collectionCode>
); &quot;Kuldsha [= Yining,
<geoCoordinate id="EE3F7CFD4307336FFE67D7204EC6E6E0" box="[400,499,886,912]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="43.883335">43°53'N</geoCoordinate>
81°16'], A. Regel&quot;, 1 ɗ (
<collectionCode id="ED1A82FF4307336FFCD5D7204C6CE6E0" box="[802,857,886,912]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Mzhd. Suidunom i Khorgossom,
<date id="FFB53CFA4307336FFAFFD7204A7CE6E0" box="[1288,1353,886,912]" pageId="8" pageNumber="55">28.V.</date>
[18]89, Gr. Grzhimailo&quot;, 1 Ψ (
<collectionCode id="ED1A82FF4307336FFE50D7CB4EEAE6C7" box="[423,479,925,951]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Dzhinkho [= Jinghe,
<geoCoordinate id="EE3F7CFD4307336FFCF4D7CB4C53E6C7" box="[771,870,925,951]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="44.6">44°36'N</geoCoordinate>
82°53'],
<date id="FFB53CFA4307336FFC20D7CB4B10E6C7" box="[983,1061,925,951]" pageId="8" pageNumber="55">13.VI.</date>
[18]89, Gr. Grzhimailo&quot;, 3 exx (
<collectionCode id="ED1A82FF4307336FFF68D7924FE3E6AE" box="[159,214,964,990]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); &quot;Dzhungar, Urumtschi&quot;[
<geoCoordinate id="EE3F7CFD4307336FFDF9D7924D5FE6AE" box="[526,618,964,990]" direction="north" orientation="latitude" pageId="8" pageNumber="55" precision="925" value="43.916668">43°55'N</geoCoordinate>
87°34'], 1 Ψ (MER). UNDETERMINED COUNTRY: &quot;Kirghiz, Eversm.&quot;, 1 ɗ 1 Ψ. (
<collectionCode id="ED1A82FF4307336FFEFFD7BC4E51E174" box="[264,356,1002,1028]" pageId="8" pageNumber="55">SMTD</collectionCode>
);&quot;R.m., Kirg. stepp.&quot; (red, pr.), 1 Ψ (
<collectionCode id="ED1A82FF4307336FFCFDD7BC4C74E174" box="[778,833,1002,1028]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="55">ZIN</collectionCode>
); no data, &quot;93&quot; 1 ɗ (MER).
</paragraph>
</subSubSection>
<subSubSection id="C31149B14307336FFF32D0464EDCE1EF" pageId="8" pageNumber="55" type="distribution">
<paragraph id="8BB41A3A4307336FFF32D0464EDCE1EF" blockId="8.[151,1438,151,2034]" pageId="8" pageNumber="55">
<emphasis id="B97FC6284307336FFF32D0464E53E15A" bold="true" box="[197,358,1040,1066]" pageId="8" pageNumber="55">Distribution</emphasis>
(
<figureCitation id="133006BF4307336FFE8FD0474EE3E15B" box="[376,470,1041,1067]" captionStart="FIGURE 82" captionStartId="19.[151,255,1833,1857]" captionTargetBox="[151,1435,1006,1808]" captionTargetId="figure@19.[151,1436,1006,1809]" captionTargetPageId="19" captionText="FIGURE 82. Distribution of the genus Epexochus [map taken from Encarta World Atlas 2000 (Microsoft Corporation), elaborated with Photoshop 7.0 (Adobe Systems Incorporated)]." httpUri="https://zenodo.org/record/274690/files/figure.png" pageId="8" pageNumber="55">Fig. 82</figureCitation>
).
<taxonomicName id="4C0B61B94307336FFE1BD0474DD8E15A" box="[492,749,1041,1066]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284307336FFE1BD0474DD8E15A" box="[492,749,1041,1066]" italics="true" pageId="8" pageNumber="55">Epexochus lehmanni</emphasis>
</taxonomicName>
has a broad range, which stretches more than
<quantity id="4CF3B7DF4307336FFAD8D0474AA8E15B" box="[1327,1437,1041,1067]" metricMagnitude="6" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="55" unit="km" value="3000.0">3000 km</quantity>
between its extremes. The
<typeStatus id="54B0A4984307336FFE2ED06E4D77E122" box="[473,578,1080,1106]" pageId="8" pageNumber="55" type="holotype">holotype</typeStatus>
is from the northernmost limit, but most of the known specimens were found in south-eastern
<collectingCountry id="F31C5AAA4307336FFE52D0084D1BE108" box="[421,558,1118,1144]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
and in Chinese
<collectingCountry id="F31C5AAA4307336FFD1BD0084C4DE108" box="[748,888,1118,1144]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
, in present-day north-western Xinjiang, on the Dzhungar Alatau mountains.
</paragraph>
</subSubSection>
<subSubSection id="C31149B14307336FFF32D0FD4CEEE063" pageId="8" pageNumber="55" type="biology_ecology">
<paragraph id="8BB41A3A4307336FFF32D0FD4CEEE063" blockId="8.[151,1438,151,2034]" pageId="8" pageNumber="55">
<emphasis id="B97FC6284307336FFF32D0FD4E11E1B5" bold="true" box="[197,292,1195,1221]" pageId="8" pageNumber="55">Biology</emphasis>
. Nothing is known about the host plants of the species, a single record existing of a specimen found &quot;on
<taxonomicName id="4C0B61B94307336FFEEED0854EBAE19C" box="[281,399,1235,1260]" class="Magnoliopsida" family="Asteraceae" genus="Artemisia" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="55" phylum="Tracheophyta" rank="genus">
<emphasis id="B97FC6284307336FFEEED0854EBAE19C" box="[281,399,1235,1260]" italics="true" pageId="8" pageNumber="55">Artemisia</emphasis>
</taxonomicName>
&quot;. This plant is associated with steppe habitats, growing at altitudes from sea level in northern and central
<collectingCountry id="F31C5AAA4307336FFE7DD0AF4D26E063" box="[394,531,1273,1299]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
to at least
<quantity id="4CF3B7DF4307336FFD66D0AF4DDFE063" box="[657,746,1273,1299]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="55" unit="m" value="1500.0">1500 m</quantity>
on the Alatau ridge.
</paragraph>
</subSubSection>
<subSubSection id="C31149B14307336DFF32D1494EA8E281" lastPageId="10" lastPageNumber="57" pageId="8" pageNumber="55" type="discussion">
<paragraph id="8BB41A3A4307336FFF32D1494B56E0F7" blockId="8.[151,1438,151,2034]" pageId="8" pageNumber="55">
<emphasis id="B97FC6284307336FFF32D1494E0BE049" bold="true" box="[197,318,1311,1337]" pageId="8" pageNumber="55">Remarks.</emphasis>
The description of the
<taxonomicName id="4C0B61B94307336FFDBAD1764DEFE049" box="[589,730,1312,1337]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284307336FFDBAD1764DEFE049" box="[589,730,1312,1337]" italics="true" pageId="8" pageNumber="55">E. lehmanni</emphasis>
</taxonomicName>
was based on a single specimen, found &quot;in springtime in the steppe near the Emba [river]&quot; (&quot;D'apres un seul individu pris au primtemps dans les steppes près de l'Emba&quot;,
<bibRefCitation id="EF9A67CB4307336FFF60D13B4E67E0F7" author="Menetries" box="[151,338,1389,1415]" pageId="8" pageNumber="67" refString="Menetries, M. (1849) Catalogue des insectes recueillis par feu M. Lehmann avec les descriptions des nouvelles especes. Seconde et derniere partie. Tetramere. Curculionites. Memoires de l'Academie Imperiale des Sciences de Saint- Petersbourg. Sixieme Serie. Sciences Naturelles, VI: 249 - 263." type="book chapter" year="1849">Ménétries 1849</bibRefCitation>
: 252). This
<typeStatus id="54B0A4984307336FFE2AD13B4D25E0F7" box="[477,528,1389,1415]" pageId="8" pageNumber="55">type</typeStatus>
specimen, housed in the ZIN, is thus the
<typeStatus id="54B0A4984307336FFC02D13B4B6AE0F7" box="[1013,1119,1389,1415]" pageId="8" pageNumber="55" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BB41A3A4307336FFF32D1C24DF3E20E" blockId="8.[151,1438,151,2034]" pageId="8" pageNumber="55">
Two more names are applicable to this species.
<bibRefCitation id="EF9A67CB4307336FFD00D1C24CF4E0DE" author="Chevrolat" box="[759,961,1428,1454]" pageId="8" pageNumber="67" refString="Chevrolat, A. (1873) Memoire sur les Cleonides. Memoires de la Societe Royale des Sciences de Liege, 2 (5): I - VIII + 1 - 118." type="journal article" year="1873">Chevrolat (1873)</bibRefCitation>
described
<taxonomicName id="4C0B61B94307336FFBB7D1C24BAFE0DD" box="[1088,1178,1428,1453]" class="Insecta" family="Curculionidae" genus="Exochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC6284307336FFBB7D1C24BAFE0DD" box="[1088,1178,1428,1453]" italics="true" pageId="8" pageNumber="55">E. latus</emphasis>
</taxonomicName>
, based on a specimen ex Gebler's collection, but since this
<typeStatus id="54B0A4984307336FFD98D1EC4D90E0A4" box="[623,677,1466,1492]" pageId="8" pageNumber="55">type</typeStatus>
specimen was not located again, the name has always been considered as of doubtful status.
<bibRefCitation id="EF9A67CB4307336FFDEED1B74D98E08B" author="Faust" box="[537,685,1505,1531]" pageId="8" pageNumber="67" refString="Faust, J. (1904) Revision der Gruppe Cleonides vrais. Deutsche Entomologische Zeitschrift, 1904 (1): 177 - 284." type="journal article" year="1904">Faust (1904)</bibRefCitation>
included it with a question mark under
<emphasis id="B97FC6284307336FFB76D1B74A5FE08A" box="[1153,1386,1505,1530]" italics="true" pageId="8" pageNumber="55">
<taxonomicName id="4C0B61B94307336FFB76D1B74A53E08A" box="[1153,1382,1505,1530]" class="Insecta" family="Carabidae" genus="Epilectus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="lehmanni">Epilectus lehmanni</taxonomicName>
,
</emphasis>
and since then it was never cited again in taxonomic works; in the
<emphasis id="B97FC6284307336FFC80D25E4B9CE351" box="[887,1193,1544,1569]" italics="true" pageId="8" pageNumber="55">Coleopterorum Catalogus</emphasis>
<bibRefCitation id="EF9A67CB4307336FFB47D25E4A71E352" author="Csiki" box="[1200,1348,1544,1570]" pageId="8" pageNumber="67" refString="Csiki, E. (1934). Curculionidae. Subfam. Cleoninae. In: Schenkling S. (Ed.). Coleopterorum Catalogus auspiciis et auxilio W. Junk, 134: pp. 152." type="journal article" year="1934">Csiki (1934)</bibRefCitation>
simply followed Faust in doubtfully referring it to
<taxonomicName id="4C0B61B94307336FFD39D2794C5EE338" box="[718,875,1583,1608]" class="Insecta" family="Carabidae" genus="Epilectus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284307336FFD39D2794C5EE338" box="[718,875,1583,1608]" italics="true" pageId="8" pageNumber="55">E. lehmanni</emphasis>
</taxonomicName>
. A part of Gebler's collection was recently rediscovered in the MNHN (
<bibRefCitation id="EF9A67CB4307336FFE00D2034C1AE31F" author="Perrin" box="[503,815,1621,1647]" pageId="8" pageNumber="67" refString="Perrin, H. &amp; Meregalli, M. (2008) Designation de lectotypes de Cleonini decrits par Gebler et Chevrolat, dans les collections Entomologiques du Museum national d'Histoire naturelle (Coleoptera: Curculionidae: Lixinae). Nouvelle revue d'Entomologie, 29: 129 - 148." type="journal article" year="2008">Perrin &amp; Meregalli 2008</bibRefCitation>
); it includes the
<typeStatus id="54B0A4984307336FFC0AD2034B07E31F" box="[1021,1074,1621,1647]" pageId="8" pageNumber="55">type</typeStatus>
of
<taxonomicName id="4C0B61B94307336FFBAAD2034BACE31E" box="[1117,1177,1621,1646]" class="Insecta" family="Curculionidae" genus="Exochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC6284307336FFBAAD2034BACE31E" box="[1117,1177,1621,1646]" italics="true" pageId="8" pageNumber="55">latus</emphasis>
</taxonomicName>
and its pertinence to
<taxonomicName id="4C0B61B94307336FFF60D22A4E17E3E5" box="[151,290,1660,1685]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC6284307336FFF60D22A4E17E3E5" box="[151,290,1660,1685]" italics="true" pageId="8" pageNumber="55">Epexochus</emphasis>
</taxonomicName>
was confirmed. The
<typeStatus id="54B0A4984307336FFDCFD22A4D45E3E6" box="[568,624,1660,1686]" pageId="8" pageNumber="55">type</typeStatus>
locality of
<taxonomicName id="4C0B61B94307336FFCFED22A4C7CE3E5" box="[777,841,1660,1685]" class="Insecta" family="Curculionidae" genus="Exochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC6284307336FFCFED22A4C7CE3E5" box="[777,841,1660,1685]" italics="true" pageId="8" pageNumber="55">latus</emphasis>
</taxonomicName>
was cited as &quot;
<emphasis id="B97FC6284307336FFBFFD22A4A2EE3E5" box="[1032,1307,1660,1685]" italics="true" pageId="8" pageNumber="55">Deserto Kirghisorum</emphasis>
&quot;, but this unfortunately is a very vague indication since the Kirghiz desert embraces a very large area of central Asia, including most of central and southern
<collectingCountry id="F31C5AAA4307336FFDA9D29F4DDEE393" box="[606,747,1737,1763]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
. Two specimens housed in the SMTD, extremely similar to the
<typeStatus id="54B0A4984307336FFF35D2A64FC0E27A" box="[194,245,1776,1802]" pageId="8" pageNumber="55">type</typeStatus>
of
<taxonomicName id="4C0B61B94307336FFEECD2A64E61E279" box="[283,340,1776,1801]" class="Insecta" family="Curculionidae" genus="Exochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC6284307336FFEECD2A64E61E279" box="[283,340,1776,1801]" italics="true" pageId="8" pageNumber="55">latus</emphasis>
</taxonomicName>
, are labelled, in Fausts' handwriting, &quot;Kirghiz, Eversmann&quot; and &quot;
<taxonomicName id="4C0B61B94307336FFBA7D2A64B88E279" box="[1104,1213,1776,1801]" class="Insecta" family="Carabidae" genus="Epilectus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284307336FFBA7D2A64B88E279" box="[1104,1213,1776,1801]" italics="true" pageId="8" pageNumber="55">lehmanni</emphasis>
</taxonomicName>
, cum typo comp.&quot;. They look somewhat intermediate between the central and southern
<collectingCountry id="F31C5AAA4307336FFC35D3404B79E240" box="[962,1100,1814,1840]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
populations of
<taxonomicName id="4C0B61B94307336FFAF2D3414AA1E240" box="[1285,1428,1815,1840]" class="Insecta" family="Carabidae" genus="Epilectus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284307336FFAF2D3414AA1E240" box="[1285,1428,1815,1840]" italics="true" pageId="8" pageNumber="55">E. lehmanni</emphasis>
</taxonomicName>
, being quite similar to the forms from Aulie-ata except for having shorter elytral setae. The status to be afforded to
<taxonomicName id="4C0B61B94307336FFEE8D3324EEBE20D" box="[287,478,1892,1917]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC6284307336FFEE8D3324EEBE20D" box="[287,478,1892,1917]" italics="true" pageId="8" pageNumber="55">Epexochus latus</emphasis>
</taxonomicName>
is discussed below.
</paragraph>
<paragraph id="8BB41A3A4307336EFF32D3DD4C25E633" blockId="8.[151,1438,151,2034]" lastBlockId="9.[151,1437,152,2010]" lastPageId="9" lastPageNumber="56" pageId="8" pageNumber="55">
The second species name associated with
<taxonomicName id="4C0B61B94307336FFD00D3DD4CA2E2D4" box="[759,919,1931,1956]" class="Insecta" family="Carabidae" genus="Epilectus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284307336FFD00D3DD4CA2E2D4" box="[759,919,1931,1956]" italics="true" pageId="8" pageNumber="55">E. lehmanni</emphasis>
</taxonomicName>
was proposed by
<bibRefCitation id="EF9A67CB4307336FFB7DD3DD4A4DE2D5" author="Faust" box="[1162,1400,1931,1957]" pageId="8" pageNumber="67" refString="Faust, J. (1904) Revision der Gruppe Cleonides vrais. Deutsche Entomologische Zeitschrift, 1904 (1): 177 - 284." type="journal article" year="1904">Faust (1904: 192)</bibRefCitation>
as
<taxonomicName id="4C0B61B94307336FFF60D3E74E8DE2BA" box="[151,440,1969,1994]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284307336FFF60D3E74E8DE2BA" box="[151,440,1969,1994]" italics="true" pageId="8" pageNumber="55">Leucochromus lehmanni</emphasis>
</taxonomicName>
<taxonomicName id="4C0B61B94307336FFE37D3E74DB7E2BA" box="[448,642,1969,1995]" pageId="8" pageNumber="55" rank="variety" variety="consobrinus">
var.
<emphasis id="B97FC6284307336FFE04D3E74DB7E2BA" box="[499,642,1969,1994]" italics="true" pageId="8" pageNumber="55">consobrinus</emphasis>
</taxonomicName>
, for specimens from Aulie-ata [= Djambul, now Taraz, in southern
<collectingCountry id="F31C5AAA4307336FFF60D38E4E1FE282" box="[151,298,2008,2034]" name="Kazakhstan" pageId="8" pageNumber="55">Kazakhstan</collectingCountry>
] characterised by the elytral vestiture consisting of predominantly greyish-white scales and occurring sympatrically with the typically brownish-coloured specimens. Later in the same work,
<bibRefCitation id="EF9A67CB4306336EFAADD4CE4E15E5A8" author="Faust" pageId="9" pageNumber="67" refString="Faust, J. (1904) Revision der Gruppe Cleonides vrais. Deutsche Entomologische Zeitschrift, 1904 (1): 177 - 284." type="journal article" year="1904">Faust (1904: 208)</bibRefCitation>
referred
<taxonomicName id="4C0B61B94306336EFE7DD4E94EC2E5A8" box="[394,503,191,216]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284306336EFE7DD4E94EC2E5A8" box="[394,503,191,216]" italics="true" pageId="9" pageNumber="56">lehmanni</emphasis>
</taxonomicName>
and its variety
<taxonomicName id="4C0B61B94306336EFD5DD4E94C0CE5A8" box="[682,825,191,216]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC6284306336EFD5DD4E94C0CE5A8" box="[682,825,191,216]" italics="true" pageId="9" pageNumber="56">consobrinus</emphasis>
</taxonomicName>
to the new genus
<taxonomicName id="4C0B61B94306336EFBFBD4E94A7CE5A8" authority="Faust. According" authorityName="Faust. According" box="[1036,1353,190,216]" class="Insecta" family="Carabidae" genus="Epilectus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC6284306336EFBFBD4E94B4CE5A8" box="[1036,1145,191,216]" italics="true" pageId="9" pageNumber="56">Epilectus</emphasis>
Faust. According
</taxonomicName>
to Art. 45.6.4 of the ICZN (1999),
<taxonomicName id="4C0B61B94306336EFE12D4B34D61E58E" box="[485,596,229,254]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284306336EFE12D4B34D61E58E" box="[485,596,229,254]" italics="true" pageId="9" pageNumber="56">lehmanni</emphasis>
</taxonomicName>
<taxonomicName id="4C0B61B94306336EFDABD4B34C59E58F" authority="Faust" authorityName="Faust" box="[604,876,229,255]" pageId="9" pageNumber="56" rank="variety" variety="consobrinus">
var.
<emphasis id="B97FC6284306336EFD66D4B34C16E58E" box="[657,803,229,254]" italics="true" pageId="9" pageNumber="56">consobrinus</emphasis>
Faust
</taxonomicName>
has infrasubspecific rank, since the content of Fausts work reveals that he used the name for a sympatric colour variety, i.e. the consequence of intrapopulational variability defined as an infrasubspecific entity in the glossary of the ICZN. Faust's name was not again considered until
<bibRefCitation id="EF9A67CB4306336EFDF3D50F4DC9E403" author="Ter-Minasyan" box="[516,764,345,371]" pageId="9" pageNumber="68" refString="Ter-Minasyan, M. E. (1972) Zhuki-dolgonosiki podsemeystva Cleoninae (Curculionidae, Coleoptera) sobrannye sovetsko - mongolskimi zoologichelskimi ekspeditsiyami v 1967 - 1969 gg. Nasekomye Mongolii. 1. L.: Nauka: 539 - 556." type="journal article" year="1972">Ter-Minasyan (1972)</bibRefCitation>
raised it to species level (as &quot;
<taxonomicName id="4C0B61B94306336EFBADD50F4FEEE4EA" authority="Faust" authorityName="Faust" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="consobrinus" status="stat. nov.">
<emphasis id="B97FC6284306336EFBADD50F4AA9E402" box="[1114,1436,345,370]" italics="true" pageId="9" pageNumber="56">Leucochromus consobrinus</emphasis>
Faust
</taxonomicName>
,
<emphasis id="B97FC6284306336EFF1FD5294E6FE4E9" bold="true" box="[232,346,383,409]" pageId="9" pageNumber="56">
<taxonomicNameLabel id="A24C7B534306336EFF1FD5294E6FE4E9" box="[232,346,383,409]" pageId="9" pageNumber="56">stat. nov.</taxonomicNameLabel>
</emphasis>
&quot;), applying the name to a specimen from
<collectingCountry id="F31C5AAA4306336EFCAED5D64CE4E4EA" box="[857,977,384,410]" name="Mongolia" pageId="9" pageNumber="56">Mongolia</collectingCountry>
. This generic attribution is surprising, because a) apart from its ambiguous original description,
<taxonomicName id="4C0B61B94306336EFCC1D5F14CF0E4B0" box="[822,965,423,448]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC6284306336EFCC1D5F14CF0E4B0" box="[822,965,423,448]" italics="true" pageId="9" pageNumber="56">consobrinus</emphasis>
</taxonomicName>
had always been referred to
<taxonomicName id="4C0B61B94306336EFAE2D5F14AA1E4B0" box="[1301,1428,423,448]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC6284306336EFAE2D5F14AA1E4B0" box="[1301,1428,423,448]" italics="true" pageId="9" pageNumber="56">Epexochus</emphasis>
</taxonomicName>
, which is clearly distinct from
<taxonomicName id="4C0B61B94306336EFDE7D59B4DF1E496" box="[528,708,461,486]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC6284306336EFDE7D59B4DF1E496" box="[528,708,461,486]" italics="true" pageId="9" pageNumber="56">Leucochromus</emphasis>
</taxonomicName>
, b) the Mongolian entity belongs to
<taxonomicName id="4C0B61B94306336EFB67D59B4A20E496" box="[1168,1301,461,486]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC6284306336EFB67D59B4A20E496" box="[1168,1301,461,486]" italics="true" pageId="9" pageNumber="56">Epexochus</emphasis>
</taxonomicName>
and not to
<taxonomicName id="4C0B61B94306336EFF60D5A24E77E77D" box="[151,322,500,525]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC6284306336EFF60D5A24E77E77D" box="[151,322,500,525]" italics="true" pageId="9" pageNumber="56">Leucochromus</emphasis>
</taxonomicName>
, and c) it is also a species different from Faust's
<taxonomicName id="4C0B61B94306336EFC8AD5A24B39E77D" box="[893,1036,500,525]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC6284306336EFC8AD5A24B39E77D" box="[893,1036,500,525]" italics="true" pageId="9" pageNumber="56">consobrinus</emphasis>
</taxonomicName>
from Aulie-ata. However, despite this misapplication to another taxon, according to Art. 45.6.4.1 Ter-Minasyan's adoption of Faust's name for a species makes
<taxonomicName id="4C0B61B94306336EFEB0D6174EE2E72A" box="[327,471,577,602]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC6284306336EFEB0D6174EE2E72A" box="[327,471,577,602]" italics="true" pageId="9" pageNumber="56">consobrinus</emphasis>
</taxonomicName>
an available name at subspecific rank, with its original authorship. Ter-Minasyan equipped the Mongolian specimen with a red label reading &quot;
<taxonomicName id="4C0B61B94306336EFC87D63E4A33E7F2" authority="Faust" authorityName="Faust" box="[880,1286,616,642]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC6284306336EFC87D63E4B8CE7F1" box="[880,1209,616,641]" italics="true" pageId="9" pageNumber="56">Leucochromus consobrinus</emphasis>
Faust
</taxonomicName>
,
<typeStatus id="54B0A4984306336EFAE5D63E4ABAE7F2" box="[1298,1423,616,642]" pageId="9" pageNumber="56" type="holotype">holotypus</typeStatus>
&quot;, but that is an erroneous designation since the
<typeStatus id="54B0A4984306336EFD50D6D84DEFE7D8" box="[679,730,654,680]" pageId="9" pageNumber="56">type</typeStatus>
series is from Aulie-ata, as in Faust's collection. The species erroneously attributed by Ter-Minasyan to
<taxonomicName id="4C0B61B94306336EFD7AD6E34B21E7BF" authority="Faust" authorityName="Faust" box="[653,1044,693,719]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC6284306336EFD7AD6E34CFBE7BE" box="[653,974,693,718]" italics="true" pageId="9" pageNumber="56">Leucochromus consobrinus</emphasis>
Faust
</taxonomicName>
is described below as
<taxonomicName id="4C0B61B94306336EFAEAD6E34E2BE785" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="mongolicus" status="sp. nov.">
<emphasis id="B97FC6284306336EFAEAD6E34E2BE785" italics="true" pageId="9" pageNumber="56">Epexochus mongolicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C7B534306336EFED2D68A4E51E786" box="[293,356,732,758]" pageId="9" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
Finally,
<bibRefCitation id="EF9A67CB4306336EFE3BD68A4DBFE786" author="Arzanov" box="[460,650,732,758]" pageId="9" pageNumber="67" refString="Arzanov, Ju. G. (2005) Taxonomic notes on weevils of the tribe Cleonini (Coleoptera, Curculionidae, Lixinae). Caucasian Entomological Bulletin 1 (2): 150." type="journal article" year="2005">Arzanov (2005)</bibRefCitation>
, based on some differences between
<taxonomicName id="4C0B61B94306336EFBCAD68A4BF9E785" box="[1085,1228,732,757]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC6284306336EFBCAD68A4BF9E785" box="[1085,1228,732,757]" italics="true" pageId="9" pageNumber="56">consobrinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B61B94306336EFAF1D68A4AA9E785" box="[1286,1436,732,757]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="imperialis">
<emphasis id="B97FC6284306336EFAF1D68A4AA9E785" box="[1286,1436,732,757]" italics="true" pageId="9" pageNumber="56">L. imperialis</emphasis>
</taxonomicName>
in the structure of the internal sac of the aedeagus, considered the former as a valid species in the genus
<taxonomicName id="4C0B61B94306336EFF60D77F4E23E632" box="[151,278,809,834]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC6284306336EFF60D77F4E23E632" box="[151,278,809,834]" italics="true" pageId="9" pageNumber="56">Epexochus</emphasis>
</taxonomicName>
and formally established this combination.
</paragraph>
<paragraph id="8BB41A3A4306336DFF32D7064C85E375" blockId="9.[151,1437,152,2010]" lastBlockId="10.[151,1437,152,2033]" lastPageId="10" lastPageNumber="57" pageId="9" pageNumber="56">
The evaluation of the status to be given to
<taxonomicName id="4C0B61B94306336EFD31D7064C62E619" box="[710,855,848,873]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC6284306336EFD31D7064C62E619" box="[710,855,848,873]" italics="true" pageId="9" pageNumber="56">consobrinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B61B94306336EFC64D7064CF8E619" box="[915,973,848,873]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC6284306336EFC64D7064CF8E619" box="[915,973,848,873]" italics="true" pageId="9" pageNumber="56">latus</emphasis>
</taxonomicName>
(species, subspecies or synonym of
<taxonomicName id="4C0B61B94306336EFA74D7064E31E6FF" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284306336EFA74D7064E31E6FF" italics="true" pageId="9" pageNumber="56">E. lehmanni</emphasis>
</taxonomicName>
) requires an analysis of the intra- and interpopulation variation of
<taxonomicName id="4C0B61B94306336EFBE7D7204BABE6FF" box="[1040,1182,886,911]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284306336EFBE7D7204BABE6FF" box="[1040,1182,886,911]" italics="true" pageId="9" pageNumber="56">E. lehmanni</emphasis>
</taxonomicName>
. Only few specimens could be examined from central and northern
<collectingCountry id="F31C5AAA4306336EFD45D7CD4C0BE6C5" box="[690,830,923,949]" name="Kazakhstan" pageId="9" pageNumber="56">Kazakhstan</collectingCountry>
, where the
<typeStatus id="54B0A4984306336EFC33D7CD4CC3E6C5" box="[964,1014,923,949]" pageId="9" pageNumber="56">type</typeStatus>
locality of
<taxonomicName id="4C0B61B94306336EFB89D7CA4BDEE6C5" box="[1150,1259,924,949]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284306336EFB89D7CA4BDEE6C5" box="[1150,1259,924,949]" italics="true" pageId="9" pageNumber="56">lehmanni</emphasis>
</taxonomicName>
lies. These are characterised by the dorsum being mainly covered with dark brown scales, with the white scales forming a few roundish spots on the disc and the top of the declivity. The declivity itself and the lateral parts of the elytra are generally whitish. The elytral setae vary from whitish, short and sparse (
<typeStatus id="54B0A4984306336EFBEED05B4BB1E157" box="[1049,1156,1037,1063]" pageId="9" pageNumber="56" type="holotype">holotype</typeStatus>
,
<figureCitation id="133006BF4306336EFB66D05B4BE3E157" box="[1169,1238,1037,1063]" captionStart="FIGURES 1 9" captionStartId="5.[151,270,1918,1942]" captionTargetBox="[236,1367,197,1896]" captionTargetId="figure@5.[236,1370,197,1898]" captionTargetPageId="5" captionText="FIGURES 1 9. Epexochus lehmanni, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5, Exochus latus, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 8 9, specimen from Borokhoro. Scale bars: 10 mm." httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="9" pageNumber="56">Fig. 1</figureCitation>
; Elton lake,
<figureCitation id="133006BF4306336EFA99D05B4F93E13D" captionStart="FIGURES 1 9" captionStartId="5.[151,270,1918,1942]" captionTargetBox="[236,1367,197,1896]" captionTargetId="figure@5.[236,1370,197,1898]" captionTargetPageId="5" captionText="FIGURES 1 9. Epexochus lehmanni, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5, Exochus latus, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 8 9, specimen from Borokhoro. Scale bars: 10 mm." httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="9" pageNumber="56">Fig. 2</figureCitation>
) to dense, golden, erect (particularly on the declivity), more than half as long as an interval width (Myn-Sai,
<figureCitation id="133006BF4306336EFF60D00F4FEAE103" box="[151,223,1113,1139]" captionStart="FIGURES 1 9" captionStartId="5.[151,270,1918,1942]" captionTargetBox="[236,1367,197,1896]" captionTargetId="figure@5.[236,1370,197,1898]" captionTargetPageId="5" captionText="FIGURES 1 9. Epexochus lehmanni, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5, Exochus latus, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 8 9, specimen from Borokhoro. Scale bars: 10 mm." httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="9" pageNumber="56">Fig. 3</figureCitation>
; Atbasa, Kzyl-Dzhar). The rostrum is usually short, narrower at the base and has a prominent median keel, with the anterior triangular plate usually clearly distinct, the dorso-lateral margins low and in lateral view more concave in their basal part. The base of the pronotum is usually medially slightly lobed, and the elytra are widest behind the middle and broader towards the apex. The aedeagus of the specimen from Elton lake has a short, acute lamella and is strongly curved in lateral view (
<figureCitation id="133006BF4306336EFBFFD0A64BE7E07A" box="[1032,1234,1264,1290]" captionStart="FIGURES 21 39" captionStartId="12.[151,269,1819,1843]" captionTargetBox="[260,1326,194,1795]" captionTargetId="figure@12.[260,1328,194,1795]" captionTargetPageId="12" captionText="FIGURES 21 39. Epexochus, genitalia, claws and scales. Female genitalia (21 23: specimen from Kzyl-Dzhar; 24 25: specimen from Aulie-ata). 21, 24, sternum VIII; 22, 25, spermatheca; 23, hemisternites. Claws. 26, specimen from Borokhoro. Aedeagus, dorsal and lateral views and detail of apex. 27, 30, 35, specimen from Koibun; 28 29, 32, specimen from Elton lake; 31, specimen from Dzharkentsk; 33, specimen from Betpak; 34, specimen from Borokhoro. Elytral scales. 36, holotype of E. lehmanni; 37, holotype of E. korotyaevi; 38, holotype of E. voriseki; 39, paratype of E. mongolicus. Scale bars: 21, 23 24, 27 31: 1 mm; 26, 32 35: 500 μm; 36 39: 100 μm." httpUri="https://zenodo.org/record/274689/files/figure.png" pageId="9" pageNumber="56">Figs. 2829, 32</figureCitation>
), but that of the specimens from Kzyl-Dzhar and Betpak-Dala has a better developed lamella and is less strongly curved (
<figureCitation id="133006BF4306336EFA9AD1404F8DE026" captionStart="FIGURES 21 39" captionStartId="12.[151,269,1819,1843]" captionTargetBox="[260,1326,194,1795]" captionTargetId="figure@12.[260,1328,194,1795]" captionTargetPageId="12" captionText="FIGURES 21 39. Epexochus, genitalia, claws and scales. Female genitalia (21 23: specimen from Kzyl-Dzhar; 24 25: specimen from Aulie-ata). 21, 24, sternum VIII; 22, 25, spermatheca; 23, hemisternites. Claws. 26, specimen from Borokhoro. Aedeagus, dorsal and lateral views and detail of apex. 27, 30, 35, specimen from Koibun; 28 29, 32, specimen from Elton lake; 31, specimen from Dzharkentsk; 33, specimen from Betpak; 34, specimen from Borokhoro. Elytral scales. 36, holotype of E. lehmanni; 37, holotype of E. korotyaevi; 38, holotype of E. voriseki; 39, paratype of E. mongolicus. Scale bars: 21, 23 24, 27 31: 1 mm; 26, 32 35: 500 μm; 36 39: 100 μm." httpUri="https://zenodo.org/record/274689/files/figure.png" pageId="9" pageNumber="56">Fig. 33</figureCitation>
). A specimen from the steppes north of the Balkhash lake and a few others generically labelled &quot;Balkhash lake&quot; have quite a similar pattern of elytral scales; these specimens are smaller and have shorter setae, although not as short as in the specimens from Yenbek and the medium to lower course of the Ili river, described below. Other specimens also generically labelled Balkhash, but from different collectors, have predominantly greyish scales (
<figureCitation id="133006BF4306336EFDE1D1854D51E09D" box="[534,612,1491,1517]" captionStart="FIGURES 1 9" captionStartId="5.[151,270,1918,1942]" captionTargetBox="[236,1367,197,1896]" captionTargetId="figure@5.[236,1370,197,1898]" captionTargetPageId="5" captionText="FIGURES 1 9. Epexochus lehmanni, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5, Exochus latus, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 8 9, specimen from Borokhoro. Scale bars: 10 mm." httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="9" pageNumber="56">Fig. 4</figureCitation>
), but the inaccuracy of their collection data precludes any further deliberations of the significance of this variation in colour pattern in the demes from the steppes around the Balkhash lake. The few specimens simply labelled &quot;Kirgiz., Eversmann&quot; and the
<typeStatus id="54B0A4984306336EFB9AD2494BE3E349" box="[1133,1238,1567,1593]" pageId="9" pageNumber="56" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C0B61B94306336EFB08D2764A0CE349" box="[1279,1337,1568,1593]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC6284306336EFB08D2764A0CE349" box="[1279,1337,1568,1593]" italics="true" pageId="9" pageNumber="56">latus</emphasis>
</taxonomicName>
(
<figureCitation id="133006BF4306336EFABDD2494AA7E349" box="[1354,1426,1567,1593]" captionStart="FIGURES 1 9" captionStartId="5.[151,270,1918,1942]" captionTargetBox="[236,1367,197,1896]" captionTargetId="figure@5.[236,1370,197,1898]" captionTargetPageId="5" captionText="FIGURES 1 9. Epexochus lehmanni, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5, Exochus latus, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 8 9, specimen from Borokhoro. Scale bars: 10 mm." httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="9" pageNumber="56">Fig. 5</figureCitation>
) have a more varied pattern, with the round pale spots less evident, and the whitish scales scattered in irregular patches and small stripes.
<collectingCountry id="F31C5AAA4306336EFE3DD23D4EDFE3F5" box="[458,490,1643,1669]" name="American Samoa" pageId="9" pageNumber="56">As</collectingCountry>
in the specimen from Myn-Sai, the elytra are broadly rounded towards the apex, the median keel of the rostrum has the triangular plate more raised and distinct, the base of the pronotum is completely linear, not lobed towards the scutellum, the elytra are parallel-sided from the base to at least half their length and the sides have denser, more erect setae. These specimens are somewhat intermediate between those from northern
<collectingCountry id="F31C5AAA4306336EFE7FD3544D24E26C" box="[392,529,1794,1820]" name="Kazakhstan" pageId="9" pageNumber="56">Kazakhstan</collectingCountry>
and those found at the end of the 19th century at the upper course of the Syr Darya, in the surroundings of Aulie-ata (
<figureCitation id="133006BF4306336EFD73D37E4DFBE232" box="[644,718,1832,1858]" captionStart="FIGURES 1 9" captionStartId="5.[151,270,1918,1942]" captionTargetBox="[236,1367,197,1896]" captionTargetId="figure@5.[236,1370,197,1898]" captionTargetPageId="5" captionText="FIGURES 1 9. Epexochus lehmanni, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5, Exochus latus, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 8 9, specimen from Borokhoro. Scale bars: 10 mm." httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="9" pageNumber="56">Fig. 6</figureCitation>
), the
<typeStatus id="54B0A4984306336EFCE4D37E4C72E232" box="[787,839,1832,1858]" pageId="9" pageNumber="56">type</typeStatus>
locality of
<taxonomicName id="4C0B61B94306336EFC25D37F4BE6E232" authority="Faust. A" authorityName="Faust. A" box="[978,1235,1832,1858]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC6284306336EFC25D37F4B51E232" box="[978,1124,1833,1858]" italics="true" pageId="9" pageNumber="56">consobrinus</emphasis>
Faust. A
</taxonomicName>
large number of specimens of this population could be examined. They have more globose elytra with rounded sides, very distinct hair-like, golden, semi-erect setae nearly as long as an interval width rather densely distributed over the entire body, and the colour pattern of the elytra consists of roughly equal areas of brown and white scales, with pale round patches absent but usually a sharp contrast between the dark and pale parts, with several alternate longitudinal stripes and spots. The rostrum is usually shorter, narrower at its base and has a prominent median keel; the anterior triangular anterior triangular plate is usually raised and the dorso-lateral margins are low, in lateral view slightly concave in their basal part. Several
<taxonomicName id="4C0B61B94305336DFBE7D4B24BBAE58D" box="[1040,1167,228,253]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC6284305336DFBE7D4B24BBAE58D" box="[1040,1167,228,253]" italics="true" pageId="10" pageNumber="57">Epexochus</emphasis>
</taxonomicName>
specimens were found in the Semirechye region in south-eastern
<collectingCountry id="F31C5AAA4305336DFD46D55F4C72E453" box="[689,839,265,291]" name="Kazakhstan" pageId="10" pageNumber="57">Kazakhstan</collectingCountry>
, south of the Balkhash lake (Koibun, Yenbeck, Bakanas, etc.). These are characterised by very short, barely visible setae, the rostrum longer and broader at the base, appearing slightly conical and with the dorso-lateral margins slightly convergent towards the antennal insertions and better developed and relatively sharply raised, the median keel sharper, the triangular plate less evident and in lateral view the rostrum often hardly curved. The elytra have predominantly pale scales, with brown scales mainly placed along the striae and denser before the declivity to reach the intervals and form a vague, darker transversal patch. Their sides are broadened, sometimes not very evenly curved. The length of the setae is not constant in the specimens from this area: of the two males collected together along the Ili river, south of Bakanas, one has dense and relatively long, erect setae on the elytra and the legs and the other almost indistinct setae on the elytra and very short and sparse on the legs; this specimen also has predominantly dark scales, with white scales limited to a few broadly rounded patches, and its pronotum is not completely straight at the base. There are no other differences in phenotypic traits, including the shape of the aedeagus. More populations are recorded from along the middle and upper course of the river Ili, in southeastern
<collectingCountry id="F31C5AAA4305336DFF05D6A34E49E67F" box="[242,380,757,783]" name="Kazakhstan" pageId="10" pageNumber="57">Kazakhstan</collectingCountry>
and north-western Xinjiang. The specimens from the steppe around the Tsharyn river, in south-eastern
<collectingCountry id="F31C5AAA4305336DFECDD74D4EF6E645" box="[314,451,795,821]" name="Kazakhstan" pageId="10" pageNumber="57">Kazakhstan</collectingCountry>
(
<figureCitation id="133006BF4305336DFE24D74D4D2FE645" box="[467,538,795,821]" captionStart="FIGURES 1 9" captionStartId="5.[151,270,1918,1942]" captionTargetBox="[236,1367,197,1896]" captionTargetId="figure@5.[236,1370,197,1898]" captionTargetPageId="5" captionText="FIGURES 1 9. Epexochus lehmanni, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5, Exochus latus, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 8 9, specimen from Borokhoro. Scale bars: 10 mm." httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="10" pageNumber="57">Fig. 7</figureCitation>
), are rather similar to those from the Semirechye region and have very short setae. Many
<taxonomicName id="4C0B61B94305336DFEDDD7174E9CE62A" box="[298,425,833,858]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B97FC6284305336DFEDDD7174E9CE62A" box="[298,425,833,858]" italics="true" pageId="10" pageNumber="57">Epexochus</emphasis>
</taxonomicName>
specimens were collected by Hauser in
<date id="FFB53CFA4305336DFC8BD7174CC6E62B" box="[892,1011,833,859]" pageId="10" pageNumber="57" value="1906-05">May 1906</date>
on the Borokhoro ridge, more to the east and at a higher altitude, along the upper course of the Kash river, a tributary of the Ili river, in present day Xinjiang. Most of these specimens are characterised by the whitish scales covering most of the elytra, with pale brownish scales confined to two broad transverse patches before the declivity, not sharply delimited, and to some small spots along the striae. The pronotum is usually dark, with white scales limited to the sides and along the median line (
<figureCitation id="133006BF4305336DFE50D7A94EC5E169" box="[423,496,1023,1049]" captionStart="FIGURES 1 9" captionStartId="5.[151,270,1918,1942]" captionTargetBox="[236,1367,197,1896]" captionTargetId="figure@5.[236,1370,197,1898]" captionTargetPageId="5" captionText="FIGURES 1 9. Epexochus lehmanni, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5, Exochus latus, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 8 9, specimen from Borokhoro. Scale bars: 10 mm." httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="10" pageNumber="57">Fig. 9</figureCitation>
). Two specimens have most of the elytral scales brownish (
<figureCitation id="133006BF4305336DFB45D7A94BCCE169" box="[1202,1273,1023,1049]" captionStart="FIGURES 1 9" captionStartId="5.[151,270,1918,1942]" captionTargetBox="[236,1367,197,1896]" captionTargetId="figure@5.[236,1370,197,1898]" captionTargetPageId="5" captionText="FIGURES 1 9. Epexochus lehmanni, habitus. 1, holotype; 2, specimen from Elton lake; 3, specimen from Myn-sai; 4, specimen from Balkhash lake; 5, Exochus latus, holotype; 6, specimen from Aulie-ata; 7, specimen from Burundisu; 8 9, specimen from Borokhoro. Scale bars: 10 mm." httpUri="https://zenodo.org/record/274688/files/figure.png" pageId="10" pageNumber="57">Fig. 8</figureCitation>
), whereas six are nearly completely white, including on the pronotum and before the declivity. In contrast to the other demes, the scales in this population are barely glossy. The elytral setae on the Borokhoro specimens are generally whitish, moderately erect and distinctly shorter than an interval width; their density and length vary, but they are always clearly distinct in profile, longer than the setae of the forms from the steppe north and east of Almaty. The rostrum is slightly variable in length and curvature, and the median triangular longitudinal keel is usually well differentiated, only flatter in few specimens. The base of the pronotum is generally truncate, without any curvature towards the scutellum. The shape of the elytra and the size of the body are relatively uniform in all these specimens, the size ranging from 16.81 to 20.00 mm (mean of the specimens examined
<quantity id="4CF3B7DF4305336DFEE3D1054EA5E01D" box="[276,400,1363,1389]" metricMagnitude="-2" metricUnit="m" metricValue="1.835" pageId="10" pageNumber="57" unit="mm" value="18.35">18.35 mm</quantity>
) and the elytra have subparallel sides, with the maximum width approximately in the middle and from there evenly converging towards the apex. The specimen labelled Urumtschi [= Ürümqi, Xinjiang] is also referable to this form, as is that from Kuldja [= Yining, Xinjiang], at the base of the Borokhoro ridge. In the specimens of south-eastern
<collectingCountry id="F31C5AAA4305336DFD08D1934CBDE0AF" box="[767,904,1477,1503]" name="Kazakhstan" pageId="10" pageNumber="57">Kazakhstan</collectingCountry>
and Xinjiang the aedeagus always is weakly curved and has a slightly rounded lamella (
<figureCitation id="133006BF4305336DFD78D1BD4C96E375" box="[655,931,1515,1541]" captionStart="FIGURES 21 39" captionStartId="12.[151,269,1819,1843]" captionTargetBox="[260,1326,194,1795]" captionTargetId="figure@12.[260,1328,194,1795]" captionTargetPageId="12" captionText="FIGURES 21 39. Epexochus, genitalia, claws and scales. Female genitalia (21 23: specimen from Kzyl-Dzhar; 24 25: specimen from Aulie-ata). 21, 24, sternum VIII; 22, 25, spermatheca; 23, hemisternites. Claws. 26, specimen from Borokhoro. Aedeagus, dorsal and lateral views and detail of apex. 27, 30, 35, specimen from Koibun; 28 29, 32, specimen from Elton lake; 31, specimen from Dzharkentsk; 33, specimen from Betpak; 34, specimen from Borokhoro. Elytral scales. 36, holotype of E. lehmanni; 37, holotype of E. korotyaevi; 38, holotype of E. voriseki; 39, paratype of E. mongolicus. Scale bars: 21, 23 24, 27 31: 1 mm; 26, 32 35: 500 μm; 36 39: 100 μm." httpUri="https://zenodo.org/record/274689/files/figure.png" pageId="10" pageNumber="57">Figs. 27, 3031, 3435</figureCitation>
).
</paragraph>
<paragraph id="8BB41A3A4305336DFF32D2474EA8E281" blockId="10.[151,1437,152,2033]" pageId="10" pageNumber="57">
In conclusion, it seems that each population shows peculiar characters, usually associated with the form of the elytra, the colour pattern of the scales and the length of the setae. The specimens from the
<collectingCountry id="F31C5AAA4305336DFAE7D2614AA9E321" box="[1296,1436,1591,1617]" name="Kazakhstan" pageId="10" pageNumber="57">Kazakhstan</collectingCountry>
steppe north and west of the Balkhash lake share the brown colour, usually with white scales in round spots, but vary in the length of the setae and in the shape of the rostrum; in southern
<collectingCountry id="F31C5AAA4305336DFBC0D2D44BF5E3EC" box="[1079,1216,1666,1692]" name="Kazakhstan" pageId="10" pageNumber="57">Kazakhstan</collectingCountry>
and north western Xinjiang the demes have the elytra mainly whitish with often relatively dense and erect setae and a sharply truncate pronotal base; in the Semirechye region most of the specimens have very short or nearly indistinct setae, the elytra with pale scales, the pronotal base often weakly curved and a longer rostrum. The shape of the aedeagus varies hardly at all, and the female genitalia are also not particularly differentiated among the various populations. It is hence impossible to recognise a clear and univocal differentiation of some demes, and the different morphological traits can only be regarded as variations at population level, not indicative of distinct taxa at species rank.
<taxonomicName id="4C0B61B94305336DFE0BD3DA4C3BE2D5" authority="Chevrolat, 1873" authorityName="Chevrolat" authorityYear="1873" box="[508,782,1931,1957]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC6284305336DFE0BD3DA4D0AE2D5" box="[508,575,1932,1957]" italics="true" pageId="10" pageNumber="57">Latus</emphasis>
<bibRefCitation id="EF9A67CB4305336DFDBFD3DD4C3BE2D5" author="Chevrolat" box="[584,782,1931,1957]" pageId="10" pageNumber="67" refString="Chevrolat, A. (1873) Memoire sur les Cleonides. Memoires de la Societe Royale des Sciences de Liege, 2 (5): I - VIII + 1 - 118." type="journal article" year="1873">Chevrolat, 1873</bibRefCitation>
</taxonomicName>
is therefore treated as a junior synonym of
<taxonomicName id="4C0B61B94305336DFADCD3DA4E54E2BB" authority="Menetries, 1849" authorityName="Menetries" authorityYear="1849" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284305336DFADCD3DA4AA9E2D5" box="[1323,1436,1932,1957]" italics="true" pageId="10" pageNumber="57">lehmanni</emphasis>
<bibRefCitation id="EF9A67CB4305336DFF60D3E74E54E2BB" author="Menetries" box="[151,353,1969,1995]" pageId="10" pageNumber="67" refString="Menetries, M. (1849) Catalogue des insectes recueillis par feu M. Lehmann avec les descriptions des nouvelles especes. Seconde et derniere partie. Tetramere. Curculionites. Memoires de l'Academie Imperiale des Sciences de Saint- Petersbourg. Sixieme Serie. Sciences Naturelles, VI: 249 - 263." type="book chapter" year="1849">Ménétries, 1849</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C0B61B94305336DFE55D3E74DE7E2BB" authority="Faust, 1904" authorityName="Faust" authorityYear="1904" box="[418,722,1969,1995]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B97FC6284305336DFE55D3E74D02E2BA" box="[418,567,1969,1994]" italics="true" pageId="10" pageNumber="57">consobrinus</emphasis>
<bibRefCitation id="EF9A67CB4305336DFDB7D3E74DE7E2BB" author="Faust" box="[576,722,1969,1995]" pageId="10" pageNumber="67" refString="Faust, J. (1904) Revision der Gruppe Cleonides vrais. Deutsche Entomologische Zeitschrift, 1904 (1): 177 - 284." type="journal article" year="1904">Faust, 1904</bibRefCitation>
</taxonomicName>
, which is itself referable to
<taxonomicName id="4C0B61B94305336DFBDCD3E74B52E2BA" box="[1067,1127,1969,1994]" class="Insecta" family="Curculionidae" genus="Epexochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B97FC6284305336DFBDCD3E74B52E2BA" box="[1067,1127,1969,1994]" italics="true" pageId="10" pageNumber="57">latus</emphasis>
</taxonomicName>
, is thus likewise a junior synonym of
<taxonomicName id="4C0B61B94305336DFEDED38E4EA3E281" box="[297,406,2008,2033]" class="Insecta" family="Curculionidae" genus="Leucochromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B97FC6284305336DFEDED38E4EA3E281" box="[297,406,2008,2033]" italics="true" pageId="10" pageNumber="57">lehmanni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>