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<document id="2FF4598B44AF940CFE5C953A76A8C099" ID-DOI="10.5281/zenodo.214889" ID-GBIF-Dataset="621e7e0f-2512-4cde-8b3a-863b0656e9e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="214889" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460530037874" checkinUser="plazi" docAuthor="Nadolny, Anton A., Ponomarev, Alexandr V. &amp; Dvadnenko, Konstantin V." docDate="2012" docId="03A278408C7BFF956BD7F9E2FD62F871" docLanguage="en" docName="zt03484p088.pdf" docOrigin="Zootaxa 3484" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Alopecosa kovblyuki Nadolny &amp; Ponomarev, sp. nov." docType="treatment" docVersion="9" lastPageNumber="86" masterDocId="FF9B00388C7BFF966B40FFD9FFB0FFA1" masterDocTitle="A new wolf spider species in the genus Alopecosa Simon, 1885 (Araneae: Lycosidae) from Eastern Europe" masterLastPageNumber="88" masterPageNumber="83" pageNumber="83" updateTime="1698315659748" updateUser="plazi">
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<mods:title id="B1605FCAB7D53A2CCE180DB72F1312DD">A new wolf spider species in the genus Alopecosa Simon, 1885 (Araneae: Lycosidae) from Eastern Europe</mods:title>
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<mods:namePart id="F37F99A0365EC1878D028C72FDB03CC6">Nadolny, Anton A.</mods:namePart>
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<mods:namePart id="5D6069BC57ED7870DE14DCF691C2B008">Ponomarev, Alexandr V.</mods:namePart>
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<mods:namePart id="092845F97FC798CDBA9F99EF1217BEE0">Dvadnenko, Konstantin V.</mods:namePart>
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<mods:date id="FDBCE00BD9BB9385878F81AC7CE98D10">2012</mods:date>
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<treatment id="03A278408C7BFF956BD7F9E2FD62F871" ID-DOI="http://doi.org/10.5281/zenodo.6166791" ID-GBIF-Taxon="119526848" ID-Zenodo-Dep="6166791" LSID="urn:lsid:plazi:treatment:03A278408C7BFF956BD7F9E2FD62F871" httpUri="http://treatment.plazi.org/id/03A278408C7BFF956BD7F9E2FD62F871" lastPageId="3" lastPageNumber="86" pageId="0" pageNumber="83">
<subSubSection id="C3119ADD8C7BFF966BD7F9E2FE9AF9D4" pageId="0" pageNumber="83" type="nomenclature">
<paragraph id="8BB4C9568C7BFF966BD7F9E2FC90F9F4" blockId="0.[151,800,1595,1653]" box="[151,800,1595,1621]" pageId="0" pageNumber="83">
<heading id="D0FC7E3A8C7BFF966BD7F9E2FC90F9F4" bold="true" box="[151,800,1595,1621]" fontSize="11" level="1" pageId="0" pageNumber="83" reason="1">
<emphasis id="B97F15448C7BFF966BD7F9E2FC90F9F4" bold="true" box="[151,800,1595,1621]" pageId="0" pageNumber="83">
<taxonomicName id="4C0BB2D58C7BFF966BD7F9E2FD06F9F4" ID-CoL="C4W6" authority="Nadolny &amp; Ponomarev" authorityName="Nadolny &amp; Ponomarev" box="[151,694,1595,1621]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="kovblyuki" status="sp. nov.">
<emphasis id="B97F15448C7BFF966BD7F9E2FE3FF9F4" bold="true" box="[151,399,1595,1621]" italics="true" pageId="0" pageNumber="83">Alopecosa kovblyuki</emphasis>
Nadolny &amp; Ponomarev
</taxonomicName>
,
<taxonomicNameLabel id="A24CA83F8C7BFF966983F9E2FC90F9F4" box="[707,800,1595,1621]" pageId="0" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4C9568C7BFF966BD7F987FE9AF9D4" blockId="0.[151,800,1595,1653]" box="[151,298,1630,1653]" pageId="0" pageNumber="83">
<figureCitation id="1330D5D38C7BFF966BD7F987FE95F9D4" box="[151,293,1630,1653]" captionStart-0="FIGURES 1 12" captionStart-1="FIGURES 13 15" captionStartId-0="1.[151,258,1820,1841]" captionStartId-1="3.[151,258,731,752]" captionTargetBox-0="[151,1434,196,1796]" captionTargetBox-1="[151,1436,191,709]" captionTargetId-0="figure@1.[151,1436,194,1797]" captionTargetId-1="figure@3.[151,1436,191,709]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="FIGURES 1 12. Habitus and copulatory organs of Alopecosa kovblyuki sp. nov. from Ukraine (2 5, 7 8 — TNU 2819 / 1; 5, 9, 12 — TNU 2814 / 1; 10 — TNU SO 70; 11 — TNU SO 49 / 2). 1 — male in natural habitat (Crimea, Lebyazh'i Islands); 2 — male pedipalp, ventral; 3 — male pedipalp, prolatero-ventral; 4 — bulbus, retrolateral; 5 6 — tegular (= median) apophysis, ventral (variation); 7 — embolic division, ventral; 8 — embolic division, prolatero-caudal; 9 11 — epigyne, ventral (variation); 12 — epigyne, dorsal. Abbreviations: AEP — anterior epigynal pocket; E — embolus; ED — embolic division; S — epigynal septum; Se — synembolus; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." captionText-1="FIGURES 13 15. Male copulatory organs of Alopecosa kovblyuki sp. nov. and A. taeniopus from Russia (13 — ZMMU; 14 — CP 25.11.29 / 1) and A. taeniopus (15 — CP 25.11.5 / 34) from Russia. 13 — bulbus, ventral; 14 15 — bulbus, ventral (a little on front). Abbreviations: ED — embolic division; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri-0="https://zenodo.org/record/214890/files/figure.png" httpUri-1="https://zenodo.org/record/214891/files/figure.png" pageId="0" pageNumber="83">Figures 114</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3119ADD8C7BFF946BD7F97AFB72FDEE" lastPageId="2" lastPageNumber="85" pageId="0" pageNumber="83" type="materials_examined">
<paragraph id="8BB4C9568C7BFF966BD7F97AFBBBF97C" blockId="0.[151,1436,1699,2035]" pageId="0" pageNumber="83">
<emphasis id="B97F15448C7BFF966BD7F97AFEF3F91A" bold="true" box="[151,323,1699,1723]" pageId="0" pageNumber="83">
<typeStatus id="54B077F48C7BFF966BD7F97AFF7FF91A" box="[151,207,1699,1723]" pageId="0" pageNumber="83">Type</typeStatus>
material
</emphasis>
:
<typeStatus id="54B077F48C7BFF966A1EF97AFE71F91B" box="[350,449,1699,1722]" pageId="0" pageNumber="83" type="holotype">Holotype</typeStatus>
1 3 (
<collectionCode id="ED1A51938C7BFF966967F97AFDCDF91B" box="[551,637,1699,1722]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="0" pageNumber="83">ZMMU</collectionCode>
)
<collectingCountry id="F31C89C68C7BFF9669D9F97AFD4BF91B" box="[665,763,1699,1722]" name="Russia" pageId="0" pageNumber="83">RUSSIA</collectingCountry>
, Rostov area, Ust-Donetsk Distr., Razdorskaya Village, Pukhlyakovskie Sklony,
<geoCoordinate id="EE3FAF918C7BFF966ADFF91FFDABF97C" box="[415,539,1734,1757]" direction="north" orientation="latitude" pageId="0" pageNumber="83" precision="15" value="47.521667">47°31'18''N</geoCoordinate>
<geoCoordinate id="EE3FAF918C7BFF966962F91FFD2DF97C" box="[546,669,1734,1757]" direction="east" orientation="longitude" pageId="0" pageNumber="83" precision="15" value="40.610275">40°36'37''E</geoCoordinate>
, 7
<date id="FFB5EF968C7BFF966982F91FFC8BF97C" box="[706,827,1734,1757]" pageId="0" pageNumber="83" value="2004-04-14">14.04.2004</date>
(A.V. Ponomarev).
</paragraph>
<paragraph id="8BB4C9568C7BFF966B87F931FB58F853" blockId="0.[151,1436,1699,2035]" pageId="0" pageNumber="83">
<emphasis id="B97F15448C7BFF966B87F931FE89F8A1" bold="true" box="[199,313,1768,1792]" pageId="0" pageNumber="83">
<typeStatus id="54B077F48C7BFF966B87F931FE89F8A1" box="[199,313,1768,1792]" pageId="0" pageNumber="83" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="F31C89C68C7BFF966A06F931FE18F95E" box="[326,424,1768,1791]" name="Russia" pageId="0" pageNumber="83">RUSSIA</collectingCountry>
.
<emphasis id="B97F15448C7BFF966AF2F931FD83F8A1" bold="true" box="[434,563,1768,1792]" italics="true" pageId="0" pageNumber="83">Rostov area</emphasis>
: 1 3 (CP-
<date id="FFB5EF968C7BFF9669E0F931FCB1F95E" box="[672,769,1768,1791]" pageId="0" pageNumber="83" value="29-11-25">25.11.29</date>
/2), Rostov-on-Don, Shchepkinskiy Forestry,
<geoCoordinate id="EE3FAF918C7BFF966FA2F931FA8AF95E" box="[1250,1338,1768,1791]" direction="north" orientation="latitude" pageId="0" pageNumber="83" precision="925" value="47.333332">47°20'N</geoCoordinate>
<geoCoordinate id="EE3FAF918C7BFF966E00F931FA28F95E" box="[1344,1432,1768,1791]" direction="east" orientation="longitude" pageId="0" pageNumber="83" precision="925" value="39.75">39°45'E</geoCoordinate>
,
<date id="FFB5EF968C7BFF966BD7F8D2FEB5F883" box="[151,261,1803,1826]" pageId="0" pageNumber="83" value="2010-05">May 2010</date>
(V.V. Alexandrov). Ust-Donetsk Distr., Razdorskaya Village,
<geoCoordinate id="EE3FAF918C7BFF9668E8F8D2FB94F883" box="[936,1060,1803,1826]" direction="north" orientation="latitude" pageId="0" pageNumber="83" precision="15" value="47.541668">47°32'30''N</geoCoordinate>
<geoCoordinate id="EE3FAF918C7BFF966F6CF8D2FB19F883" box="[1068,1193,1803,1826]" direction="east" orientation="longitude" pageId="0" pageNumber="83" precision="15" value="40.647224">40°38'50''E</geoCoordinate>
: 1 3 (
<collectionCode id="ED1A51938C7BFF966FAEF8D2FAF7F883" box="[1262,1351,1803,1826]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="0" pageNumber="83">ZMMU</collectionCode>
), valley edge,
<taxonomicName id="4C0BB2D58C7BFF966B9BF8F7FE20F8E4" box="[219,400,1838,1861]" class="Magnoliopsida" family="Fabaceae" genus="Caragana" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="83" phylum="Tracheophyta" rank="species" species="frutex">
<emphasis id="B97F15448C7BFF966B9BF8F7FE20F8E4" box="[219,400,1838,1861]" italics="true" pageId="0" pageNumber="83">Caragana frutex</emphasis>
</taxonomicName>
, 17
<date id="FFB5EF968C7BFF966A87F8F7FDF2F8E4" box="[455,578,1838,1861]" pageId="0" pageNumber="83" value="2010-04-26">26.04.2010</date>
(A.V. Ponomarev); 1 3 (CP-
<date id="FFB5EF968C7BFF9668D1F8F7FC45F8E4" box="[913,1013,1838,1861]" pageId="0" pageNumber="83" value="29-11-25">25.11.29</date>
/1), valley edge,
<taxonomicName id="4C0BB2D58C7BFF966FECF8F7FAD2F8E4" box="[1196,1378,1838,1861]" class="Magnoliopsida" family="Fabaceae" genus="Caragana" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="83" phylum="Tracheophyta" rank="species" species="frutex">
<emphasis id="B97F15448C7BFF966FECF8F7FAD2F8E4" box="[1196,1378,1838,1861]" italics="true" pageId="0" pageNumber="83">Caragana frutex</emphasis>
</taxonomicName>
, 17
<date id="FFB5EF968C7BFF966BD7F889FF65F8C6" box="[151,213,1872,1895]" pageId="0" pageNumber="83" value="2010-04-26">26.04</date>
.2010 (A.V. Ponomarev).
<collectingCountry id="F31C89C68C7BFF966AAAF889FDD6F8C6" box="[490,614,1872,1895]" name="Ukraine" pageId="0" pageNumber="83">UKRAINE</collectingCountry>
.
<emphasis id="B97F15448C7BFF966931F889FD72F8C9" bold="true" box="[625,706,1872,1896]" italics="true" pageId="0" pageNumber="83">Crimea</emphasis>
, Dzhankoy Distr.:
<date id="FFB5EF968C7BFF9668D2F889FC42F8C6" box="[914,1010,1872,1896]" pageId="0" pageNumber="83" value="1914-07-03">14 3 7</date>
ƤƤ (TNU-SO1/1, SO5, SO6/1, SO11, SO14/1, SO26/3, SO27/3, SO28/2, SO45/2, SO46/2, SO49/2, SO56/5, SO58/3, SO66/3, SO70), environs of Solenoe Ozero Village,
<geoCoordinate id="EE3FAF918C7BFF966A78F84FFE20F80C" box="[312,400,1942,1965]" direction="north" orientation="latitude" pageId="0" pageNumber="83" precision="925" value="45.883335">45°53'N</geoCoordinate>
<geoCoordinate id="EE3FAF918C7BFF966AD6F84FFE5DF80C" box="[406,493,1942,1965]" direction="east" orientation="longitude" pageId="0" pageNumber="83" precision="925" value="34.45">34°27'E</geoCoordinate>
, hand collected and pitfall traps,
<date id="FFB5EF968C7BFF96680CF84FFC77F80C" box="[844,967,1942,1965]" pageId="0" pageNumber="83" value="2008-10-21">21.10.2008</date>
, 12.03
<date id="FFB5EF968C7BFF966F58F84FFB22F80C" box="[1048,1170,1942,1965]" pageId="0" pageNumber="83" value="2009-10-22">22.10.2009</date>
(A.A. Nadolny); 8 3 3 ƤƤ (
<collectionCode id="ED1A51938C7BFF966B90F861FE99F86E" box="[208,297,1976,1999]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="0" pageNumber="83">ZMMU</collectionCode>
),
<quantity id="4CF364B38C7BFF966A7EF861FE3BF86E" box="[318,395,1976,1999]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="0" pageNumber="83" unit="km" value="6.5">6.5 km</quantity>
N Solenoe Ozero Village,
<geoCoordinate id="EE3FAF918C7BFF9669F8F861FC85F86E" box="[696,821,1976,1999]" direction="north" orientation="latitude" pageId="0" pageNumber="83" precision="15" value="45.943058">45°56'35''N</geoCoordinate>
<geoCoordinate id="EE3FAF918C7BFF96687EF861FC7EF86E" box="[830,974,1976,1999]" direction="east" orientation="longitude" pageId="0" pageNumber="83" precision="1" value="34.451862">34°27'06.7''E</geoCoordinate>
,
<taxonomicName id="4C0BB2D58C7BFF96689CF860FB0DF871" box="[988,1213,1977,2000]" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" pageId="0" pageNumber="83" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B97F15448C7BFF96689CF860FB0DF871" box="[988,1213,1977,2000]" italics="true" pageId="0" pageNumber="83">Phragmites australis</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C0BB2D58C7BFF966FA5F860FA95F871" box="[1253,1317,1977,2000]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="0" pageNumber="83" phylum="Tracheophyta" rank="genus">
<emphasis id="B97F15448C7BFF966FA5F860FA95F871" box="[1253,1317,1977,2000]" italics="true" pageId="0" pageNumber="83">Carex</emphasis>
</taxonomicName>
sp. on the mollusks' shells bar, pitfall traps, 26.03-9.04.2009 (A.A. Nadolny). Razdol'noe Distr.: 1 3 (TNU),
<quantity id="4CF364B38C7BFF966FF2F802FB58F853" box="[1202,1256,2011,2034]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="0" pageNumber="83" unit="km" value="9.0">9 km</quantity>
</paragraph>
<caption id="DF7499DE8C7AFF976BD7F8C5FC2EF865" httpUri="https://zenodo.org/record/214890/files/figure.png" pageId="1" pageNumber="84" targetBox="[151,1434,196,1796]" targetPageId="1">
<paragraph id="8BB4C9568C7AFF976BD7F8C5FC2EF865" blockId="1.[151,1437,1820,1988]" pageId="1" pageNumber="84">
<emphasis id="B97F15448C7AFF976BD7F8C5FE8FF890" bold="true" box="[151,319,1820,1841]" pageId="1" pageNumber="84">FIGURES 112.</emphasis>
Habitus and copulatory organs of
<taxonomicName id="4C0BB2D58C7AFF9769D0F8C4FCE7F893" box="[656,855,1821,1842]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="kovblyuki" status="sp. nov.">
<emphasis id="B97F15448C7AFF9769D0F8C4FCE7F893" box="[656,855,1821,1842]" italics="true" pageId="1" pageNumber="84">Alopecosa kovblyuki</emphasis>
</taxonomicName>
<emphasis id="B97F15448C7AFF97681DF8C5FC1BF890" bold="true" box="[861,939,1820,1841]" pageId="1" pageNumber="84">
<taxonomicNameLabel id="A24CA83F8C7AFF97681DF8C5FC1BF890" box="[861,939,1820,1841]" pageId="1" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Ukraine (25, 78—TNU 2819/1; 5, 9, 12— TNU 2814/1; 10—TNU SO70; 11—TNU SO49/2). 1—male in natural habitat (Crimea, Lebyazh'i Islands); 2—male pedipalp, ventral; 3—male pedipalp, prolatero-ventral; 4—bulbus, retrolateral; 56—tegular (=median) apophysis, ventral (variation); 7—embolic division, ventral; 8—embolic division, prolatero-caudal; 911—epigyne, ventral (variation); 12—epigyne, dorsal. Abbreviations:
<emphasis id="B97F15448C7AFF976BD7F84BFF73F806" box="[151,195,1938,1959]" italics="true" pageId="1" pageNumber="84">AEP</emphasis>
—anterior epigynal pocket;
<emphasis id="B97F15448C7AFF976AB2F84BFDB0F806" box="[498,512,1938,1959]" italics="true" pageId="1" pageNumber="84">E</emphasis>
—embolus;
<emphasis id="B97F15448C7AFF9769C1F84BFD11F806" box="[641,673,1938,1959]" italics="true" pageId="1" pageNumber="84">ED</emphasis>
—embolic division;
<emphasis id="B97F15448C7AFF97683CF84BFC38F806" box="[892,904,1938,1959]" italics="true" pageId="1" pageNumber="84">S</emphasis>
—epigynal septum;
<emphasis id="B97F15448C7AFF976F21F84BFBC7F806" box="[1121,1143,1938,1959]" italics="true" pageId="1" pageNumber="84">Se</emphasis>
—synembolus;
<emphasis id="B97F15448C7AFF976E59F84BFAF1F806" box="[1305,1345,1938,1959]" italics="true" pageId="1" pageNumber="84">TgA</emphasis>
—tegular (=median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BB4C9568C79FF946BD7FF41FEA6FEDE" blockId="2.[151,1437,152,1284]" pageId="2" pageNumber="85">
N Razdol'noe, environs of Portovoe Village, Lebyazh'i Islands branch of Crimean State Nature Reserve,
<geoCoordinate id="EE3FAF918C79FF946E60FF41FA2CFF0E" box="[1312,1436,152,175]" direction="north" orientation="latitude" pageId="2" pageNumber="85" precision="15" value="45.85472">45°51'17''N</geoCoordinate>
<geoCoordinate id="EE3FAF918C79FF946BD7FF63FEA2FF70" box="[151,274,186,209]" direction="east" orientation="longitude" pageId="2" pageNumber="85" precision="15" value="33.494446">33°29'40''E</geoCoordinate>
,
<date id="FFB5EF968C79FF946A5FFF63FE27FF70" box="[287,407,186,209]" pageId="2" pageNumber="85" value="2012-04-19">19.04.2012</date>
(A.A. Nadolny).
<emphasis id="B97F15448C79FF946919FF63FD40FF73" bold="true" box="[601,752,186,210]" italics="true" pageId="2" pageNumber="85">Kherson area</emphasis>
, Genichesk Distr.: 5 3 5 ƤƤ (TNU-2804/3, 2812/2, 2814/1, 2819/1, 2887/9, 2894/2, 2903/4), environs of Genichesk,
<geoCoordinate id="EE3FAF918C79FF946856FF04FCDEFF55" box="[790,878,221,244]" direction="north" orientation="latitude" pageId="2" pageNumber="85" precision="925" value="46.166668">46°10'N</geoCoordinate>
<geoCoordinate id="EE3FAF918C79FF946839FF04FC60FF55" box="[889,976,221,244]" direction="east" orientation="longitude" pageId="2" pageNumber="85" precision="925" value="34.8">34°48'E</geoCoordinate>
, Arabatskaya Strelka, hand collected and pitfall traps, 31.05
<date id="FFB5EF968C79FF946A31FED9FE59FEB6" box="[369,489,256,279]" pageId="2" pageNumber="85" value="2010-11-26">26.11.2010</date>
(N.A. Stasyuk).
<emphasis id="B97F15448C79FF9469EDFED9FCE1FEB9" bold="true" box="[685,849,256,280]" italics="true" pageId="2" pageNumber="85">Mykolaiv area</emphasis>
, Pervomaysk Distr.: 3 ƤƤ (TNU), Myhiya Village,
<geoCoordinate id="EE3FAF918C79FF946BD7FEFBFF5FFE98" box="[151,239,290,313]" direction="north" orientation="latitude" pageId="2" pageNumber="85" precision="925" value="48.033333">48°02'N</geoCoordinate>
<geoCoordinate id="EE3FAF918C79FF946BB8FEFBFEFFFE98" box="[248,335,290,313]" direction="east" orientation="longitude" pageId="2" pageNumber="85" precision="925" value="30.95">30°57'E</geoCoordinate>
, dry meadow, 7.058.06.2007 (N.Yu. Polchaninova).
<emphasis id="B97F15448C79FF9468DCFEFBFB99FE9B" bold="true" box="[924,1065,290,314]" italics="true" pageId="2" pageNumber="85">Poltava area</emphasis>
, Novosanzharskiy Distr.: 2 3 3 ƤƤ (TNU), Sokolova Balka Village,
<geoCoordinate id="EE3FAF918C79FF946971FE9CFD39FEFD" box="[561,649,325,348]" direction="north" orientation="latitude" pageId="2" pageNumber="85" precision="925" value="49.2">49°12'N</geoCoordinate>
<geoCoordinate id="EE3FAF918C79FF9469D2FE9CFD59FEFD" box="[658,745,325,348]" direction="east" orientation="longitude" pageId="2" pageNumber="85" precision="925" value="34.616665">34°37'E</geoCoordinate>
, southern slope with ruderal vegetation, 27.04
<date id="FFB5EF968C79FF946FB2FE9CFAD8FEFD" box="[1266,1384,325,348]" pageId="2" pageNumber="85" value="2011-06-14">14.06.2011</date>
(I.P. Lezhenina).
</paragraph>
<paragraph id="8BB4C9568C79FF946B87FE53FB72FDEE" blockId="2.[151,1437,152,1284]" pageId="2" pageNumber="85">
<emphasis id="B97F15448C79FF946B87FE53FE7AFE03" bold="true" box="[199,458,394,418]" pageId="2" pageNumber="85">Comparative material</emphasis>
.
<taxonomicName id="4C0BB2D58C79FF946A9EFE53FCD4FE00" authority="Thorell, 1875" authorityName="Thorell" authorityYear="1875" box="[478,868,394,417]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="B97F15448C79FF946A9EFE53FD1AFE00" box="[478,682,394,417]" italics="true" pageId="2" pageNumber="85">Alopecosa beckeri</emphasis>
(Thorell, 1875)
</taxonomicName>
:
<collectingCountry id="F31C89C68C79FF946835FE53FC40FE00" box="[885,1008,394,417]" name="Ukraine" pageId="2" pageNumber="85">UKRAINE</collectingCountry>
.
<emphasis id="B97F15448C79FF946F41FE53FBE3FE03" bold="true" box="[1025,1107,394,418]" italics="true" pageId="2" pageNumber="85">Crimea</emphasis>
: see material in
<bibRefCitation id="EF9AB4A78C79FF946E5EFE53FEE8FE65" author="Nadolny" pageId="2" pageNumber="85" refString="Nadolny, A. A. &amp; Kovblyuk M. M. (2010) On two closely related wolf spider species Alopecosa beckeri (Thorell, 1875) and A. taeniopus (Kulczynski, 1895) (Aranei: Lycosidae). Arthropoda Selecta, 19, 237 - 247." type="journal article" year="2010">Nadolny &amp; Kovblyuk (2010)</bibRefCitation>
.
<taxonomicName id="4C0BB2D58C79FF946A28FE74FC9EFE65" authority="Kulczynski, 1895" authorityName="Kulczynski" authorityYear="1895" box="[360,814,429,452]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="taeniopus">
<emphasis id="B97F15448C79FF946A28FE74FDFBFE65" box="[360,587,429,452]" italics="true" pageId="2" pageNumber="85">Alopecosa taeniopus</emphasis>
(Kulczynski, 1895)
</taxonomicName>
:
<collectingCountry id="F31C89C68C79FF94687EFE74FC10FE65" box="[830,928,429,452]" name="Russia" pageId="2" pageNumber="85">RUSSIA</collectingCountry>
.
<emphasis id="B97F15448C79FF9468F0FE74FBD6FE64" bold="true" box="[944,1126,429,453]" italics="true" pageId="2" pageNumber="85">Astrachan' area</emphasis>
: 7 ƤƤ (CP),
<quantity id="4CF364B38C79FF946E4DFE74FAD3FE65" box="[1293,1379,429,452]" metricMagnitude="5" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="85" unit="km" value="230.0">230 km</quantity>
NW Astrachan', Bogdinsko-Baskunchakskiy Reserve, 13
<date id="FFB5EF968C79FF9469A9FE09FCD6FE46" box="[745,870,464,487]" pageId="2" pageNumber="85" value="2002-07-17">17.07.2002</date>
, 7
<date id="FFB5EF968C79FF9468D3FE09FBBCFE46" box="[915,1036,464,487]" pageId="2" pageNumber="85" value="2005-10-06">10.06.2005</date>
(E.A. Belosludtsev &amp; A.S. Tilli).
<emphasis id="B97F15448C79FF946BD7FE2BFE89FDAB" bold="true" box="[151,313,498,522]" italics="true" pageId="2" pageNumber="85">Belgorod area</emphasis>
: 2 3 (CP), Kustovoe Village,
<date id="FFB5EF968C79FF9469F4FE2BFC9BFDA8" box="[692,811,498,521]" pageId="2" pageNumber="85" value="1998-10-17">17.10.1998</date>
(A.V. Ponomarev).
<emphasis id="B97F15448C79FF946F53FE2BFB2BFDAB" bold="true" box="[1043,1179,498,522]" italics="true" pageId="2" pageNumber="85">Rostov area</emphasis>
: 1 3 (CP-25.11.5/34), Razdorskaya Village, Pukhlyakovskie Sklony,
<date id="FFB5EF968C79FF9469C9FDCCFCB1FD8D" box="[649,769,533,556]" pageId="2" pageNumber="85" value="2004-03-23">23.03.2004</date>
(A.V. Ponomarev); 4 3 1 Ƥ (CP), Don River delta,
<quantity id="4CF364B38C79FF946E7FFDCCFA32FD8D" box="[1343,1410,533,556]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="85" unit="km" value="12.0">12 km</quantity>
N Azov, 9
<date id="FFB5EF968C79FF946BB6FDE1FEDEFDEE" box="[246,366,568,591]" pageId="2" pageNumber="85" value="2008-04-16">16.04.2008</date>
(P.P. Ivliev).
<collectingCountry id="F31C89C68C79FF946ABEFDE1FDC9FDEE" box="[510,633,568,591]" name="Ukraine" pageId="2" pageNumber="85">UKRAINE</collectingCountry>
.
<emphasis id="B97F15448C79FF9469C4FDE1FD66FDF1" bold="true" box="[644,726,568,592]" italics="true" pageId="2" pageNumber="85">Crimea</emphasis>
: see material in
<bibRefCitation id="EF9AB4A78C79FF9468C5FDE1FB0FFDEE" author="Nadolny" box="[901,1215,568,591]" pageId="2" pageNumber="85" refString="Nadolny, A. A. &amp; Kovblyuk M. M. (2010) On two closely related wolf spider species Alopecosa beckeri (Thorell, 1875) and A. taeniopus (Kulczynski, 1895) (Aranei: Lycosidae). Arthropoda Selecta, 19, 237 - 247." type="journal article" year="2010">Nadolny &amp; Kovblyuk (2010)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3119ADD8C79FF946B87FD83FE3BFD35" pageId="2" pageNumber="85" type="etymology">
<paragraph id="8BB4C9568C79FF946B87FD83FE3BFD35" blockId="2.[151,1437,152,1284]" pageId="2" pageNumber="85">
<emphasis id="B97F15448C79FF946B87FD83FEF0FDD3" bold="true" box="[199,320,602,626]" pageId="2" pageNumber="85">Etymology</emphasis>
. The species name is a patronym in honour of our friend and colleague, Mykola M. Kovblyuk (Simferopol,
<collectingCountry id="F31C89C68C79FF946A65FDA4FE30FD35" box="[293,384,637,660]" name="Ukraine" pageId="2" pageNumber="85">Ukraine</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3119ADD8C79FF946B87FD46FD13FC08" pageId="2" pageNumber="85" type="diagnosis">
<paragraph id="8BB4C9568C79FF946B87FD46FD13FC08" blockId="2.[151,1437,152,1284]" pageId="2" pageNumber="85">
<emphasis id="B97F15448C79FF946B87FD46FE85FD16" bold="true" box="[199,309,671,695]" pageId="2" pageNumber="85">Diagnosis</emphasis>
. The new species is similar to
<taxonomicName id="4C0BB2D58C79FF9469E0FD79FCA1FD16" box="[672,785,672,695]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="B97F15448C79FF9469E0FD79FCA1FD16" box="[672,785,672,695]" italics="true" pageId="2" pageNumber="85">A. beckeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0BB2D58C79FF946864FD79FB9BFD16" authority="Dahl, 1908" authorityName="Dahl" authorityYear="1908" box="[804,1067,672,695]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="B97F15448C79FF946864FD79FC22FD16" box="[804,914,672,695]" italics="true" pageId="2" pageNumber="85">A. mariae</emphasis>
(Dahl, 1908)
</taxonomicName>
and
<taxonomicName id="4C0BB2D58C79FF946F29FD79FB43FD16" box="[1129,1267,672,695]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="taeniopus">
<emphasis id="B97F15448C79FF946F29FD79FB43FD16" box="[1129,1267,672,695]" italics="true" pageId="2" pageNumber="85">A. taeniopus</emphasis>
</taxonomicName>
. Males can be distinguished from these species by a rounded projection on the tegulum (
<figureCitation id="1330D5D38C79FF9468EBFD1BFBBBFD78" box="[939,1035,706,729]" captionStart="FIGURES 1 12" captionStartId="1.[151,258,1820,1841]" captionTargetBox="[151,1434,196,1796]" captionTargetId="figure@1.[151,1436,194,1797]" captionTargetPageId="1" captionText="FIGURES 1 12. Habitus and copulatory organs of Alopecosa kovblyuki sp. nov. from Ukraine (2 5, 7 8 — TNU 2819 / 1; 5, 9, 12 — TNU 2814 / 1; 10 — TNU SO 70; 11 — TNU SO 49 / 2). 1 — male in natural habitat (Crimea, Lebyazh'i Islands); 2 — male pedipalp, ventral; 3 — male pedipalp, prolatero-ventral; 4 — bulbus, retrolateral; 5 6 — tegular (= median) apophysis, ventral (variation); 7 — embolic division, ventral; 8 — embolic division, prolatero-caudal; 9 11 — epigyne, ventral (variation); 12 — epigyne, dorsal. Abbreviations: AEP — anterior epigynal pocket; E — embolus; ED — embolic division; S — epigynal septum; Se — synembolus; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214890/files/figure.png" pageId="2" pageNumber="85">Figs 23</figureCitation>
,
<figureCitation id="1330D5D38C79FF946F57FD1BFB84FD78" box="[1047,1076,706,729]" captionStart="FIGURES 13 15" captionStartId="3.[151,258,731,752]" captionTargetBox="[151,1436,191,709]" captionTargetId="figure@3.[151,1436,191,709]" captionTargetPageId="3" captionText="FIGURES 13 15. Male copulatory organs of Alopecosa kovblyuki sp. nov. and A. taeniopus from Russia (13 — ZMMU; 14 — CP 25.11.29 / 1) and A. taeniopus (15 — CP 25.11.5 / 34) from Russia. 13 — bulbus, ventral; 14 15 — bulbus, ventral (a little on front). Abbreviations: ED — embolic division; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214891/files/figure.png" pageId="2" pageNumber="85">13</figureCitation>
). In addition, the shape of tegular (=median) apophysis (
<figureCitation id="1330D5D38C79FF946AC9FD3CFE5AFD5D" box="[393,490,741,764]" captionStart="FIGURES 1 12" captionStartId="1.[151,258,1820,1841]" captionTargetBox="[151,1434,196,1796]" captionTargetId="figure@1.[151,1436,194,1797]" captionTargetPageId="1" captionText="FIGURES 1 12. Habitus and copulatory organs of Alopecosa kovblyuki sp. nov. from Ukraine (2 5, 7 8 — TNU 2819 / 1; 5, 9, 12 — TNU 2814 / 1; 10 — TNU SO 70; 11 — TNU SO 49 / 2). 1 — male in natural habitat (Crimea, Lebyazh'i Islands); 2 — male pedipalp, ventral; 3 — male pedipalp, prolatero-ventral; 4 — bulbus, retrolateral; 5 6 — tegular (= median) apophysis, ventral (variation); 7 — embolic division, ventral; 8 — embolic division, prolatero-caudal; 9 11 — epigyne, ventral (variation); 12 — epigyne, dorsal. Abbreviations: AEP — anterior epigynal pocket; E — embolus; ED — embolic division; S — epigynal septum; Se — synembolus; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214890/files/figure.png" pageId="2" pageNumber="85">Figs 26</figureCitation>
,
<figureCitation id="1330D5D38C79FF946AB7FD3CFD8BFD5D" box="[503,571,741,764]" captionStart="FIGURES 13 15" captionStartId="3.[151,258,731,752]" captionTargetBox="[151,1436,191,709]" captionTargetId="figure@3.[151,1436,191,709]" captionTargetPageId="3" captionText="FIGURES 13 15. Male copulatory organs of Alopecosa kovblyuki sp. nov. and A. taeniopus from Russia (13 — ZMMU; 14 — CP 25.11.29 / 1) and A. taeniopus (15 — CP 25.11.5 / 34) from Russia. 13 — bulbus, ventral; 14 15 — bulbus, ventral (a little on front). Abbreviations: ED — embolic division; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214891/files/figure.png" pageId="2" pageNumber="85">1314</figureCitation>
) and synembolus (
<figureCitation id="1330D5D38C79FF946849FD3CFCD9FD5D" box="[777,873,741,764]" captionStart="FIGURES 1 12" captionStartId="1.[151,258,1820,1841]" captionTargetBox="[151,1434,196,1796]" captionTargetId="figure@1.[151,1436,194,1797]" captionTargetPageId="1" captionText="FIGURES 1 12. Habitus and copulatory organs of Alopecosa kovblyuki sp. nov. from Ukraine (2 5, 7 8 — TNU 2819 / 1; 5, 9, 12 — TNU 2814 / 1; 10 — TNU SO 70; 11 — TNU SO 49 / 2). 1 — male in natural habitat (Crimea, Lebyazh'i Islands); 2 — male pedipalp, ventral; 3 — male pedipalp, prolatero-ventral; 4 — bulbus, retrolateral; 5 6 — tegular (= median) apophysis, ventral (variation); 7 — embolic division, ventral; 8 — embolic division, prolatero-caudal; 9 11 — epigyne, ventral (variation); 12 — epigyne, dorsal. Abbreviations: AEP — anterior epigynal pocket; E — embolus; ED — embolic division; S — epigynal septum; Se — synembolus; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214890/files/figure.png" pageId="2" pageNumber="85">Figs 78</figureCitation>
) is also distinct. Females of the new species and
<taxonomicName id="4C0BB2D58C79FF946EC5FD3CFF52FCBE" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="B97F15448C79FF946EC5FD3CFF52FCBE" italics="true" pageId="2" pageNumber="85">A. mariae</emphasis>
</taxonomicName>
clearly differ from
<taxonomicName id="4C0BB2D58C79FF946AF7FCD1FD93FCBE" box="[439,547,776,799]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="B97F15448C79FF946AF7FCD1FD93FCBE" box="[439,547,776,799]" italics="true" pageId="2" pageNumber="85">A. beckeri</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9AB4A78C79FF946973FCD1FC6EFCBE" author="Nadolny" box="[563,990,776,799]" pageId="2" pageNumber="85" refString="Nadolny, A. A. &amp; Kovblyuk M. M. (2010) On two closely related wolf spider species Alopecosa beckeri (Thorell, 1875) and A. taeniopus (Kulczynski, 1895) (Aranei: Lycosidae). Arthropoda Selecta, 19, 237 - 247." type="journal article" year="2010">Nadolny &amp; Kovblyuk 2010: figs 23, 27</bibRefCitation>
) and
<taxonomicName id="4C0BB2D58C79FF946F5BFCD1FB11FCBE" box="[1051,1185,776,799]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="taeniopus">
<emphasis id="B97F15448C79FF946F5BFCD1FB11FCBE" box="[1051,1185,776,799]" italics="true" pageId="2" pageNumber="85">A. taeniopus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9AB4A78C79FF946FF0FCD1FEE8FCE0" author="Nadolny" pageId="2" pageNumber="85" refString="Nadolny, A. A. &amp; Kovblyuk M. M. (2010) On two closely related wolf spider species Alopecosa beckeri (Thorell, 1875) and A. taeniopus (Kulczynski, 1895) (Aranei: Lycosidae). Arthropoda Selecta, 19, 237 - 247." type="journal article" year="2010">Nadolny &amp; Kovblyuk 2010: figs 25, 29</bibRefCitation>
) by the presence of two anterior epigynal pockets (
<figureCitation id="1330D5D38C79FF9468DEFCF3FBBFFCE0" box="[926,1039,810,833]" captionStart="FIGURES 1 12" captionStartId="1.[151,258,1820,1841]" captionTargetBox="[151,1434,196,1796]" captionTargetId="figure@1.[151,1436,194,1797]" captionTargetPageId="1" captionText="FIGURES 1 12. Habitus and copulatory organs of Alopecosa kovblyuki sp. nov. from Ukraine (2 5, 7 8 — TNU 2819 / 1; 5, 9, 12 — TNU 2814 / 1; 10 — TNU SO 70; 11 — TNU SO 49 / 2). 1 — male in natural habitat (Crimea, Lebyazh'i Islands); 2 — male pedipalp, ventral; 3 — male pedipalp, prolatero-ventral; 4 — bulbus, retrolateral; 5 6 — tegular (= median) apophysis, ventral (variation); 7 — embolic division, ventral; 8 — embolic division, prolatero-caudal; 9 11 — epigyne, ventral (variation); 12 — epigyne, dorsal. Abbreviations: AEP — anterior epigynal pocket; E — embolus; ED — embolic division; S — epigynal septum; Se — synembolus; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214890/files/figure.png" pageId="2" pageNumber="85">Figs 911</figureCitation>
;
<bibRefCitation id="EF9AB4A78C79FF946F5EFCF3FA94FCE0" author="Buchar" box="[1054,1316,810,833]" pageId="2" pageNumber="85" refString="Buchar, J. &amp; Thaler, K. (2004) Ein Artproblem bei Wolfspinnen: Zur Differenzierung und vikarianten Verbreitung von Alopecosa striatipes (C. L. Koch) und A. mariae (Dahl) (Araneae, Lycosidae). Denisia, 12, zugleich Kataloge der Ober t sterreichischen Landesmuseen, Neure Serie 14, 271 - 280." type="journal article" year="2004">Buchar &amp; Thaler, 2004</bibRefCitation>
: figs 56). Females of
<taxonomicName id="4C0BB2D58C79FF946A5DFC94FE16FCC5" box="[285,422,845,868]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="kovblyuki" status="sp. nov.">
<emphasis id="B97F15448C79FF946A5DFC94FE16FCC5" box="[285,422,845,868]" italics="true" pageId="2" pageNumber="85">A. kovblyuki</emphasis>
</taxonomicName>
<emphasis id="B97F15448C79FF946AF2FC95FDBDFCC5" bold="true" box="[434,525,844,868]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="A24CA83F8C79FF946AF2FC95FDBDFCC5" box="[434,525,844,868]" pageId="2" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C0BB2D58C79FF94690BFC94FD0AFCC5" box="[587,698,845,868]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="B97F15448C79FF94690BFC94FD0AFCC5" box="[587,698,845,868]" italics="true" pageId="2" pageNumber="85">A. mariae</emphasis>
</taxonomicName>
differ in the shape of the epigynal septum and the spermathecal reservoirs. The epigynal septum of
<taxonomicName id="4C0BB2D58C79FF94695DFCA9FD13FC26" box="[541,675,880,903]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="kovblyuki" status="sp. nov.">
<emphasis id="B97F15448C79FF94695DFCA9FD13FC26" box="[541,675,880,903]" italics="true" pageId="2" pageNumber="85">A. kovblyuki</emphasis>
</taxonomicName>
<emphasis id="B97F15448C79FF9469ECFCB6FCB4FC26" bold="true" box="[684,772,879,903]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="A24CA83F8C79FF9469ECFCB6FCB4FC26" box="[684,772,879,903]" pageId="2" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is highly variable (
<figureCitation id="1330D5D38C79FF94689BFCA9FBFBFC26" box="[987,1099,880,903]" captionStart="FIGURES 1 12" captionStartId="1.[151,258,1820,1841]" captionTargetBox="[151,1434,196,1796]" captionTargetId="figure@1.[151,1436,194,1797]" captionTargetPageId="1" captionText="FIGURES 1 12. Habitus and copulatory organs of Alopecosa kovblyuki sp. nov. from Ukraine (2 5, 7 8 — TNU 2819 / 1; 5, 9, 12 — TNU 2814 / 1; 10 — TNU SO 70; 11 — TNU SO 49 / 2). 1 — male in natural habitat (Crimea, Lebyazh'i Islands); 2 — male pedipalp, ventral; 3 — male pedipalp, prolatero-ventral; 4 — bulbus, retrolateral; 5 6 — tegular (= median) apophysis, ventral (variation); 7 — embolic division, ventral; 8 — embolic division, prolatero-caudal; 9 11 — epigyne, ventral (variation); 12 — epigyne, dorsal. Abbreviations: AEP — anterior epigynal pocket; E — embolus; ED — embolic division; S — epigynal septum; Se — synembolus; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214890/files/figure.png" pageId="2" pageNumber="85">Figs 911</figureCitation>
), therefore females of the new species and
<taxonomicName id="4C0BB2D58C79FF946A58FC4BFE31FC08" box="[280,385,914,937]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="B97F15448C79FF946A58FC4BFE31FC08" box="[280,385,914,937]" italics="true" pageId="2" pageNumber="85">A. mariae</emphasis>
</taxonomicName>
are difficult to distinguish.
</paragraph>
</subSubSection>
<subSubSection id="C3119ADD8C79FF946B87FC6DFE28F87F" pageId="2" pageNumber="85" type="description">
<paragraph id="8BB4C9568C79FF946B87FC6DFC66FB40" blockId="2.[151,1437,152,1284]" pageId="2" pageNumber="85">
<emphasis id="B97F15448C79FF946B87FC6DFEFBFC6D" bold="true" box="[199,331,948,972]" pageId="2" pageNumber="85">Description</emphasis>
.
<typeStatus id="54B077F48C79FF946A1EFC6CFE76FC6D" box="[350,454,949,972]" pageId="2" pageNumber="85" type="holotype">Holotype</typeStatus>
: total length 9.0; carapace 4.4 long, 3.2 wide. Median light brown band on carapace widening in anterior part, and reaches the border of the carapace at the level of posterior lateral eyes. Dark brown areas on each side of median band. Carapace with broad white marginal bands. Eye area black. Chelicerae dark brown. Cheliceral claw with hillock on proximal part. Sternum light brown, covered with black setae. Legs and pedipalps light brown. Pedipalpal tibia more than twice as long as wide. Median band with grey lanceolate spot; series of spots on the dorsal abdomen; median band bordered with brown band. Sides of abdomen yellow. Ventral abdomen with grey spot. Tibiae spination: I, II pl 1-1, rl 1-1,
<date id="FFB5EF968C79FF946973FB5CFDC5FB3D" box="[563,629,1157,1180]" pageId="2" pageNumber="85">v 2-2</date>
-2(a); III, IV d 1-1, pl 1-1, rl 1-1,
<date id="FFB5EF968C79FF946F4AFB5CFBF7FB3D" box="[1034,1095,1157,1180]" pageId="2" pageNumber="85">v 2-2</date>
-2(a).
<typeStatus id="54B077F48C79FF946FC9FB5CFB41FB3D" box="[1161,1265,1157,1180]" pageId="2" pageNumber="85" type="paratype">Paratypes</typeStatus>
(TNU-2814/1), measurements (male/female): total length 8.0 / 9.4; carapace 3.8 / 4.2 long, 2.7 / 2.9 wide. Length of pedipalp segments (male/female): femur 1.4 / 1.4, patella 0.8 / 0.8, tibia 0.7 / 0.7, tarsus 1.3 / 1.2.
</paragraph>
<paragraph id="8BB4C9568C79FF946B87FB35FDC6FAA5" blockId="2.[151,1437,152,1284]" box="[199,630,1260,1284]" pageId="2" pageNumber="85">
<emphasis id="B97F15448C79FF946B87FB35FDC6FAA5" bold="true" box="[199,630,1260,1284]" pageId="2" pageNumber="85">Length of leg segments (male/female):</emphasis>
</paragraph>
<paragraph id="8BB4C9568C79FF946BE5FAFCFAAEFA45" pageId="2" pageNumber="85">
<table id="F90B3BF68C7900696BE5FAFCFAAEFA45" box="[165,1310,1317,1508]" gridcols="5" gridrows="5" pageId="2" pageNumber="85">
<tr id="353BCB148C7900696BE5FAFCFAAEFA9B" box="[165,1310,1317,1338]" gridrow="0" pageId="2" pageNumber="85">
<th id="76EAA2688C7900696BE5FAFCFECEFA9B" box="[165,382,1317,1338]" gridcol="0" gridrow="0" pageId="2" pageNumber="85">Leg Femur</th>
<th id="76EAA2688C790069694AFAFCFDE9FA9B" box="[522,601,1317,1338]" gridcol="1" gridrow="0" pageId="2" pageNumber="85">Patella</th>
<th id="76EAA2688C79006969A5FAFCFC84FA9B" box="[741,820,1317,1338]" gridcol="2" gridrow="0" pageId="2" pageNumber="85">Tibia</th>
<th id="76EAA2688C7900696880FAFCFB99FA9B" box="[960,1065,1317,1338]" gridcol="3" gridrow="0" pageId="2" pageNumber="85">Metatarsus</th>
<th id="76EAA2688C7900696F8FFAFCFAAEFA9B" box="[1231,1310,1317,1338]" gridcol="4" gridrow="0" pageId="2" pageNumber="85">Tarsus</th>
</tr>
<tr id="353BCB148C7900696BE5FA96FAAEFAC5" box="[165,1310,1359,1380]" gridrow="1" pageId="2" pageNumber="85">
<th id="76EAA2688C7900696BE5FA96FECEFAC5" box="[165,382,1359,1380]" gridcol="0" gridrow="1" pageId="2" pageNumber="85">I 2.6 / 2.8</th>
<td id="76EAA2688C790069694AFA96FDE9FAC5" box="[522,601,1359,1380]" gridcol="1" gridrow="1" pageId="2" pageNumber="85">1.4 / 1.5</td>
<td id="76EAA2688C79006969A5FA96FC84FAC5" box="[741,820,1359,1380]" gridcol="2" gridrow="1" pageId="2" pageNumber="85">2.0 / 2.0</td>
<td id="76EAA2688C7900696880FA96FB99FAC5" box="[960,1065,1359,1380]" gridcol="3" gridrow="1" pageId="2" pageNumber="85">2.0 / 2.0</td>
<td id="76EAA2688C7900696F8FFA96FAAEFAC5" box="[1231,1310,1359,1380]" gridcol="4" gridrow="1" pageId="2" pageNumber="85">1.6 / 1.4</td>
</tr>
<tr id="353BCB148C7900696BE5FAA3FAAEFA2E" box="[165,1310,1402,1423]" gridrow="2" pageId="2" pageNumber="85">
<th id="76EAA2688C7900696BE5FAA3FECEFA2E" box="[165,382,1402,1423]" gridcol="0" gridrow="2" pageId="2" pageNumber="85">II 2.5 / 2.7</th>
<td id="76EAA2688C790069694AFAA3FDE9FA2E" box="[522,601,1402,1423]" gridcol="1" gridrow="2" pageId="2" pageNumber="85">1.3 / 1.4</td>
<td id="76EAA2688C79006969A5FAA3FC84FA2E" box="[741,820,1402,1423]" gridcol="2" gridrow="2" pageId="2" pageNumber="85">1.8 / 1.8</td>
<td id="76EAA2688C7900696880FAA3FB99FA2E" box="[960,1065,1402,1423]" gridcol="3" gridrow="2" pageId="2" pageNumber="85">2.0 / 2.0</td>
<td id="76EAA2688C7900696F8FFAA3FAAEFA2E" box="[1231,1310,1402,1423]" gridcol="4" gridrow="2" pageId="2" pageNumber="85">1.5 / 1.4</td>
</tr>
<tr id="353BCB148C7900696BE5FA7CFAAEFA1B" box="[165,1310,1445,1466]" gridrow="3" pageId="2" pageNumber="85">
<th id="76EAA2688C7900696BE5FA7CFECEFA1B" box="[165,382,1445,1466]" gridcol="0" gridrow="3" pageId="2" pageNumber="85">III 2.4 / 2.5</th>
<td id="76EAA2688C790069694AFA7CFDE9FA1B" box="[522,601,1445,1466]" gridcol="1" gridrow="3" pageId="2" pageNumber="85">1.2 / 1.2</td>
<td id="76EAA2688C79006969A5FA7CFC84FA1B" box="[741,820,1445,1466]" gridcol="2" gridrow="3" pageId="2" pageNumber="85">1.6 / 1.6</td>
<td id="76EAA2688C7900696880FA7CFB99FA1B" box="[960,1065,1445,1466]" gridcol="3" gridrow="3" pageId="2" pageNumber="85">2.1 / 2.2</td>
<td id="76EAA2688C7900696F8FFA7CFAAEFA1B" box="[1231,1310,1445,1466]" gridcol="4" gridrow="3" pageId="2" pageNumber="85">1.3 / 1.3</td>
</tr>
<tr id="353BCB148C7900696BE5FA16FAAEFA45" box="[165,1310,1487,1508]" gridrow="4" pageId="2" pageNumber="85">
<th id="76EAA2688C7900696BE5FA16FECEFA45" box="[165,382,1487,1508]" gridcol="0" gridrow="4" pageId="2" pageNumber="85">IV 3.1 / 3.4</th>
<td id="76EAA2688C790069694AFA16FDE9FA45" box="[522,601,1487,1508]" gridcol="1" gridrow="4" pageId="2" pageNumber="85">1.3 / 1.5</td>
<td id="76EAA2688C79006969A5FA16FC84FA45" box="[741,820,1487,1508]" gridcol="2" gridrow="4" pageId="2" pageNumber="85">2.3 / 2.5</td>
<td id="76EAA2688C7900696880FA16FB99FA45" box="[960,1065,1487,1508]" gridcol="3" gridrow="4" pageId="2" pageNumber="85">3.3 / 3.5</td>
<td id="76EAA2688C7900696F8FFA16FAAEFA45" box="[1231,1310,1487,1508]" gridcol="4" gridrow="4" pageId="2" pageNumber="85">1.8 / 1.7</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB4C9568C79FF946B87F9FEFDECF907" blockId="2.[151,1436,1575,2014]" pageId="2" pageNumber="85">
Male pedipalp: tegulum with rounded projection on ventral side, retrolateral side of tegular apophysis curved ventrally, embolus flat and wide, synembolus claw-like, distal part of embolus and synembolus hiden behind tegular apophysis (
<figureCitation id="1330D5D38C79FF946A53F9B5FEC7F922" box="[275,375,1644,1667]" captionStart="FIGURES 1 12" captionStartId="1.[151,258,1820,1841]" captionTargetBox="[151,1434,196,1796]" captionTargetId="figure@1.[151,1436,194,1797]" captionTargetPageId="1" captionText="FIGURES 1 12. Habitus and copulatory organs of Alopecosa kovblyuki sp. nov. from Ukraine (2 5, 7 8 — TNU 2819 / 1; 5, 9, 12 — TNU 2814 / 1; 10 — TNU SO 70; 11 — TNU SO 49 / 2). 1 — male in natural habitat (Crimea, Lebyazh'i Islands); 2 — male pedipalp, ventral; 3 — male pedipalp, prolatero-ventral; 4 — bulbus, retrolateral; 5 6 — tegular (= median) apophysis, ventral (variation); 7 — embolic division, ventral; 8 — embolic division, prolatero-caudal; 9 11 — epigyne, ventral (variation); 12 — epigyne, dorsal. Abbreviations: AEP — anterior epigynal pocket; E — embolus; ED — embolic division; S — epigynal septum; Se — synembolus; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214890/files/figure.png" pageId="2" pageNumber="85">Figs 28</figureCitation>
,
<figureCitation id="1330D5D38C79FF946AC6F9B5FE7DF922" box="[390,461,1644,1667]" captionStart="FIGURES 13 15" captionStartId="3.[151,258,731,752]" captionTargetBox="[151,1436,191,709]" captionTargetId="figure@3.[151,1436,191,709]" captionTargetPageId="3" captionText="FIGURES 13 15. Male copulatory organs of Alopecosa kovblyuki sp. nov. and A. taeniopus from Russia (13 — ZMMU; 14 — CP 25.11.29 / 1) and A. taeniopus (15 — CP 25.11.5 / 34) from Russia. 13 — bulbus, ventral; 14 15 — bulbus, ventral (a little on front). Abbreviations: ED — embolic division; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214891/files/figure.png" pageId="2" pageNumber="85">1314</figureCitation>
); epigyne with two anterior pockets, septum flat, width of septum more than 1.5 times length, spermatheca spherical (
<figureCitation id="1330D5D38C79FF946AA1F956FDE0F907" box="[481,592,1679,1702]" captionStart="FIGURES 1 12" captionStartId="1.[151,258,1820,1841]" captionTargetBox="[151,1434,196,1796]" captionTargetId="figure@1.[151,1436,194,1797]" captionTargetPageId="1" captionText="FIGURES 1 12. Habitus and copulatory organs of Alopecosa kovblyuki sp. nov. from Ukraine (2 5, 7 8 — TNU 2819 / 1; 5, 9, 12 — TNU 2814 / 1; 10 — TNU SO 70; 11 — TNU SO 49 / 2). 1 — male in natural habitat (Crimea, Lebyazh'i Islands); 2 — male pedipalp, ventral; 3 — male pedipalp, prolatero-ventral; 4 — bulbus, retrolateral; 5 6 — tegular (= median) apophysis, ventral (variation); 7 — embolic division, ventral; 8 — embolic division, prolatero-caudal; 9 11 — epigyne, ventral (variation); 12 — epigyne, dorsal. Abbreviations: AEP — anterior epigynal pocket; E — embolus; ED — embolic division; S — epigynal septum; Se — synembolus; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214890/files/figure.png" pageId="2" pageNumber="85">Figs 912</figureCitation>
).
</paragraph>
<paragraph id="8BB4C9568C79FF946B87F968FE55F8AF" blockId="2.[151,1436,1575,2014]" pageId="2" pageNumber="85">
<emphasis id="B97F15448C79FF946B87F968FE80F968" bold="true" box="[199,304,1713,1737]" pageId="2" pageNumber="85">Variation</emphasis>
. Males (n = 11): carapace 3.64.4 long and 2.63.2 wide. Tegular apophysis shape variable (
<figureCitation id="1330D5D38C79FF946E6DF96BFA20F968" box="[1325,1424,1714,1737]" captionStart="FIGURES 1 12" captionStartId="1.[151,258,1820,1841]" captionTargetBox="[151,1434,196,1796]" captionTargetId="figure@1.[151,1436,194,1797]" captionTargetPageId="1" captionText="FIGURES 1 12. Habitus and copulatory organs of Alopecosa kovblyuki sp. nov. from Ukraine (2 5, 7 8 — TNU 2819 / 1; 5, 9, 12 — TNU 2814 / 1; 10 — TNU SO 70; 11 — TNU SO 49 / 2). 1 — male in natural habitat (Crimea, Lebyazh'i Islands); 2 — male pedipalp, ventral; 3 — male pedipalp, prolatero-ventral; 4 — bulbus, retrolateral; 5 6 — tegular (= median) apophysis, ventral (variation); 7 — embolic division, ventral; 8 — embolic division, prolatero-caudal; 9 11 — epigyne, ventral (variation); 12 — epigyne, dorsal. Abbreviations: AEP — anterior epigynal pocket; E — embolus; ED — embolic division; S — epigynal septum; Se — synembolus; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214890/files/figure.png" pageId="2" pageNumber="85">Figs 56</figureCitation>
). Females (n = 10): carapace 4.14.6 long, 2.93.2 wide. Epigyne shape variable (
<figureCitation id="1330D5D38C79FF9468B0F90DFBEEF94A" box="[1008,1118,1748,1771]" captionStart="FIGURES 1 12" captionStartId="1.[151,258,1820,1841]" captionTargetBox="[151,1434,196,1796]" captionTargetId="figure@1.[151,1436,194,1797]" captionTargetPageId="1" captionText="FIGURES 1 12. Habitus and copulatory organs of Alopecosa kovblyuki sp. nov. from Ukraine (2 5, 7 8 — TNU 2819 / 1; 5, 9, 12 — TNU 2814 / 1; 10 — TNU SO 70; 11 — TNU SO 49 / 2). 1 — male in natural habitat (Crimea, Lebyazh'i Islands); 2 — male pedipalp, ventral; 3 — male pedipalp, prolatero-ventral; 4 — bulbus, retrolateral; 5 6 — tegular (= median) apophysis, ventral (variation); 7 — embolic division, ventral; 8 — embolic division, prolatero-caudal; 9 11 — epigyne, ventral (variation); 12 — epigyne, dorsal. Abbreviations: AEP — anterior epigynal pocket; E — embolus; ED — embolic division; S — epigynal septum; Se — synembolus; TgA — tegular (= median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/214890/files/figure.png" pageId="2" pageNumber="85">Figs 911</figureCitation>
). The grey spot on the ventral side of abdomen can be absent.
</paragraph>
<paragraph id="8BB4C9568C79FF946B87F8C0FEC2F8D7" blockId="2.[151,1436,1575,2014]" pageId="2" pageNumber="85">
<emphasis id="B97F15448C79FF946B87F8C0FEF3F890" bold="true" box="[199,323,1817,1841]" pageId="2" pageNumber="85">Phenology.</emphasis>
Males and females have been found from March to November, with peak activity in April. Females with egg sacs were found in May.
<taxonomicName id="4C0BB2D58C79FF946AA1F8E5FD70F8F2" box="[481,704,1852,1875]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="kovblyuki" status="sp. nov.">
<emphasis id="B97F15448C79FF946AA1F8E5FD70F8F2" box="[481,704,1852,1875]" italics="true" pageId="2" pageNumber="85">Alopecosa kovblyuki</emphasis>
</taxonomicName>
<emphasis id="B97F15448C79FF94698AF8E5FC93F8F5" bold="true" box="[714,803,1852,1876]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="A24CA83F8C79FF94698AF8E5FC93F8F5" box="[714,803,1852,1876]" pageId="2" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C0BB2D58C79FF94681DF8E5FC56F8F2" box="[861,998,1852,1875]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="taeniopus">
<emphasis id="B97F15448C79FF94681DF8E5FC56F8F2" box="[861,998,1852,1875]" italics="true" pageId="2" pageNumber="85">A. taeniopus</emphasis>
</taxonomicName>
occur sympatrically and have a similar phenology (
<figureCitation id="1330D5D38C79FF946A56F886FED6F8D7" box="[278,358,1887,1910]" captionStart="FIGURE 16" captionStartId="3.[151,244,1710,1731]" captionTargetBox="[173,1414,895,1687]" captionTargetId="figure@3.[172,1414,894,1689]" captionTargetPageId="3" captionText="FIGURE 16. Comparative phenology of Alopecosa kovblyuki sp. nov. and A. taeniopus based on specimens collected by pitfall traps in the environs of Solenoe Ozero Village, Crimea during 2009." httpUri="https://zenodo.org/record/214892/files/figure.png" pageId="2" pageNumber="85">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BB4C9568C79FF946B87F858FE28F87F" blockId="2.[151,1436,1575,2014]" pageId="2" pageNumber="85">
<emphasis id="B97F15448C79FF946B87F858FE99F838" bold="true" box="[199,297,1921,1945]" pageId="2" pageNumber="85">Habitats</emphasis>
. Most of specimens were collected in grasslands (steppe; meadows with
<taxonomicName id="4C0BB2D58C79FF946F74F85BFB2CF838" box="[1076,1180,1922,1945]" class="Magnoliopsida" family="Asteraceae" genus="Artemisia" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="B97F15448C79FF946F74F85BFB2CF838" box="[1076,1180,1922,1945]" italics="true" pageId="2" pageNumber="85">Artemisia</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C0BB2D58C79FF946FFDF85BFA99F838" box="[1213,1321,1922,1945]" class="Magnoliopsida" family="Plumbaginaceae" genus="Limonium" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="B97F15448C79FF946FFDF85BFA99F838" box="[1213,1321,1922,1945]" italics="true" pageId="2" pageNumber="85">Limonium</emphasis>
</taxonomicName>
; meadows with
<taxonomicName id="4C0BB2D58C79FF946B90F87DFEFBF81A" box="[208,331,1956,1979]" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" pageId="2" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="B97F15448C79FF946B90F87DFEFBF81A" box="[208,331,1956,1979]" italics="true" pageId="2" pageNumber="85">Phragmites</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C0BB2D58C79FF946A33F87DFE03F81A" box="[371,435,1956,1979]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="2" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="B97F15448C79FF946A33F87DFE03F81A" box="[371,435,1956,1979]" italics="true" pageId="2" pageNumber="85">Carex</emphasis>
</taxonomicName>
;
<taxonomicName id="4C0BB2D58C79FF946A84F87DFD83F81A" box="[452,563,1956,1979]" class="Magnoliopsida" family="Amaranthaceae" genus="Salicornia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="B97F15448C79FF946A84F87DFD83F81A" box="[452,563,1956,1979]" italics="true" pageId="2" pageNumber="85">Salicornia</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C0BB2D58C79FF94691BF87DFD59F81A" box="[603,745,1956,1979]" class="Magnoliopsida" family="Amaranthaceae" genus="Halocnemum" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="B97F15448C79FF94691BF87DFD59F81A" box="[603,745,1956,1979]" italics="true" pageId="2" pageNumber="85">Halocnemum</emphasis>
</taxonomicName>
on the silt). A few specimens were collected in plantations of various deciduous trees.
</paragraph>
</subSubSection>
<caption id="DF7499DE8C78FF956BD7FD02FA27FC8A" httpUri="https://zenodo.org/record/214891/files/figure.png" pageId="3" pageNumber="86" targetBox="[151,1436,191,709]" targetPageId="3">
<paragraph id="8BB4C9568C78FF956BD7FD02FA27FC8A" blockId="3.[151,1437,731,811]" pageId="3" pageNumber="86">
<emphasis id="B97F15448C78FF956BD7FD02FEFBFD51" bold="true" box="[151,331,731,752]" pageId="3" pageNumber="86">FIGURES 1315.</emphasis>
Male copulatory organs of
<taxonomicName id="4C0BB2D58C78FF956920FD05FC99FD50" box="[608,809,732,753]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="kovblyuki" status="sp. nov.">
<emphasis id="B97F15448C78FF956920FD05FC99FD50" box="[608,809,732,753]" italics="true" pageId="3" pageNumber="86">Alopecosa kovblyuki</emphasis>
</taxonomicName>
<emphasis id="B97F15448C78FF956871FD02FCCFFD51" bold="true" box="[817,895,731,752]" pageId="3" pageNumber="86">
<taxonomicNameLabel id="A24CA83F8C78FF956871FD02FCCFFD51" box="[817,895,731,752]" pageId="3" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C0BB2D58C78FF9568F3FD05FB9DFD50" box="[947,1069,732,753]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="taeniopus">
<emphasis id="B97F15448C78FF9568F3FD05FB9DFD50" box="[947,1069,732,753]" italics="true" pageId="3" pageNumber="86">A. taeniopus</emphasis>
</taxonomicName>
from Russia (13—ZMMU; 14—CP 25.11.29/1) and
<taxonomicName id="4C0BB2D58C78FF956A7CFD20FE08FCAF" box="[316,440,761,782]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="taeniopus">
<emphasis id="B97F15448C78FF956A7CFD20FE08FCAF" box="[316,440,761,782]" italics="true" pageId="3" pageNumber="86">A. taeniopus</emphasis>
</taxonomicName>
(15—CP 25.11.5/34) from Russia. 13—bulbus, ventral; 1415—bulbus, ventral (a little on front). Abbreviations:
<emphasis id="B97F15448C78FF956A6BFCCFFEFBFC8A" box="[299,331,790,811]" italics="true" pageId="3" pageNumber="86">ED</emphasis>
—embolic division;
<emphasis id="B97F15448C78FF95694EFCCFFD86FC8A" box="[526,566,790,811]" italics="true" pageId="3" pageNumber="86">TgA</emphasis>
—tegular (=median) apophysis. Arrow indicates projection of tegulum. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="DF7499DE8C78FF956BD7F977FD42F941" httpUri="https://zenodo.org/record/214892/files/figure.png" pageId="3" pageNumber="86" targetBox="[173,1414,895,1687]" targetPageId="3">
<paragraph id="8BB4C9568C78FF956BD7F977FD42F941" blockId="3.[151,1436,1710,1760]" pageId="3" pageNumber="86">
<emphasis id="B97F15448C78FF956BD7F977FEA9F962" bold="true" box="[151,281,1710,1731]" pageId="3" pageNumber="86">FIGURE 16.</emphasis>
Comparative phenology of
<taxonomicName id="4C0BB2D58C78FF956966F977FD5DF962" box="[550,749,1710,1731]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="kovblyuki" status="sp. nov.">
<emphasis id="B97F15448C78FF956966F977FD5DF962" box="[550,749,1710,1731]" italics="true" pageId="3" pageNumber="86">Alopecosa kovblyuki</emphasis>
</taxonomicName>
<emphasis id="B97F15448C78FF9569B3F977FC8FF962" bold="true" box="[755,831,1710,1731]" pageId="3" pageNumber="86">
<taxonomicNameLabel id="A24CA83F8C78FF9569B3F977FC8FF962" box="[755,831,1710,1731]" pageId="3" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C0BB2D58C78FF95682FF977FC57F962" box="[879,999,1710,1731]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="taeniopus">
<emphasis id="B97F15448C78FF95682FF977FC57F962" box="[879,999,1710,1731]" italics="true" pageId="3" pageNumber="86">A. taeniopus</emphasis>
</taxonomicName>
based on specimens collected by pitfall traps in the environs of Solenoe Ozero Village, Crimea during 2009.
</paragraph>
</caption>
<subSubSection id="C3119ADD8C78FF956B87F8F7FD66F8C9" pageId="3" pageNumber="86" type="distribution">
<paragraph id="8BB4C9568C78FF956B87F8F7FD66F8C9" blockId="3.[151,1436,1838,2001]" pageId="3" pageNumber="86">
<emphasis id="B97F15448C78FF956B87F8F7FEE2F8E7" bold="true" box="[199,338,1838,1862]" pageId="3" pageNumber="86">Distribution</emphasis>
.
<collectingCountry id="F31C89C68C78FF956A25F8F7FE0BF8E4" box="[357,443,1838,1861]" name="Ukraine" pageId="3" pageNumber="86">Ukraine</collectingCountry>
(Mykolaiv area, Crimea, Kherson area, Poltava area),
<collectingCountry id="F31C89C68C78FF956F70F8F7FBC7F8E4" box="[1072,1143,1838,1861]" name="Russia" pageId="3" pageNumber="86">Russia</collectingCountry>
(Rostov area). Collection records of
<taxonomicName id="4C0BB2D58C78FF956A49F888FE3CF8C9" box="[265,396,1873,1896]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="kovblyuki" status="sp. nov.">
<emphasis id="B97F15448C78FF956A49F888FE3CF8C9" box="[265,396,1873,1896]" italics="true" pageId="3" pageNumber="86">A. kovblyuki</emphasis>
</taxonomicName>
<emphasis id="B97F15448C78FF956AD3F888FE58F8C8" bold="true" box="[403,488,1873,1897]" pageId="3" pageNumber="86">
<taxonomicNameLabel id="A24CA83F8C78FF956AD3F888FE58F8C8" box="[403,488,1873,1897]" pageId="3" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are shown on
<figureCitation id="1330D5D38C78FF9569C4F888FD62F8C9" box="[644,722,1873,1896]" captionStart="FIGURE 17" captionStartId="4.[151,244,1137,1158]" captionTargetBox="[152,1435,194,1116]" captionTargetId="figure@4.[152,1436,194,1116]" captionTargetPageId="4" captionText="FIGURE 17. Localities of Alopecosa kovblyuki sp. nov. (squares) in Ukraine and Russia." httpUri="https://zenodo.org/record/214893/files/figure.png" pageId="3" pageNumber="86">Fig. 17</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3119ADD8C78FF956B87F8AAFD62F871" pageId="3" pageNumber="86" type="discussion">
<paragraph id="8BB4C9568C78FF956B87F8AAFD62F871" blockId="3.[151,1436,1838,2001]" pageId="3" pageNumber="86">
<emphasis id="B97F15448C78FF956B87F8AAFE85F82A" bold="true" box="[199,309,1907,1931]" pageId="3" pageNumber="86">Remarks.</emphasis>
<taxonomicName id="4C0BB2D58C78FF956A7CF8ADFDA8F82A" box="[316,536,1908,1931]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="kovblyuki" status="sp. nov.">
<emphasis id="B97F15448C78FF956A7CF8ADFDA8F82A" box="[316,536,1908,1931]" italics="true" pageId="3" pageNumber="86">Alopecosa kovblyuki</emphasis>
</taxonomicName>
<emphasis id="B97F15448C78FF95695EF8AAFDC4F82A" bold="true" box="[542,628,1907,1931]" pageId="3" pageNumber="86">
<taxonomicNameLabel id="A24CA83F8C78FF95695EF8AAFDC4F82A" box="[542,628,1907,1931]" pageId="3" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has similarities with other
<taxonomicName id="4C0BB2D58C78FF9568D5F8ADFBB4F82A" box="[917,1028,1908,1931]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B97F15448C78FF9568D5F8ADFBB4F82A" box="[917,1028,1908,1931]" italics="true" pageId="3" pageNumber="86">Alopecosa</emphasis>
</taxonomicName>
species. However, the structure of the pedipalp and epigyne differs from the
<typeStatus id="54B077F48C78FF956979F84FFDD7F80C" box="[569,615,1942,1965]" pageId="3" pageNumber="86">type</typeStatus>
species of the genus,
<taxonomicName id="4C0BB2D58C78FF95681AF84FFC76F80C" box="[858,966,1942,1965]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="fabrilis">
<emphasis id="B97F15448C78FF95681AF84FFC76F80C" box="[858,966,1942,1965]" italics="true" pageId="3" pageNumber="86">A. fabrilis</emphasis>
</taxonomicName>
. The correct placement of
<taxonomicName id="4C0BB2D58C78FF956FAEF84FFAC3F80C" box="[1262,1395,1942,1965]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="kovblyuki" status="sp. nov.">
<emphasis id="B97F15448C78FF956FAEF84FFAC3F80C" box="[1262,1395,1942,1965]" italics="true" pageId="3" pageNumber="86">A. kovblyuki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24CA83F8C78FF956E3CF84FFF76F870" pageId="3" pageNumber="86" rank="species">
<emphasis id="B97F15448C78FF956E3CF84FFF76F870" bold="true" pageId="3" pageNumber="86">sp. nov.</emphasis>
</taxonomicNameLabel>
can only be resolved when
<taxonomicName id="4C0BB2D58C78FF956AAFF860FDEEF871" box="[495,606,1977,2000]" class="Arachnida" family="Lycosidae" genus="Alopecosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B97F15448C78FF956AAFF860FDEEF871" box="[495,606,1977,2000]" italics="true" pageId="3" pageNumber="86">Alopecosa</emphasis>
</taxonomicName>
is revised.
</paragraph>
</subSubSection>
</treatment>
</document>