439 lines
79 KiB
XML
439 lines
79 KiB
XML
<document id="748D8EA4C67283902C90FB0A859CDBE3" ID-CLB-Dataset="28353" ID-DOI="10.11646/zootaxa.4567.2.3" ID-GBIF-Dataset="7bd398c4-9441-4286-b150-0c421f91ec7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="2594934" ID-ZooBank="76FF8984-7ABC-4751-8FA9-943501F9D63A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552656074612" checkinUser="plazi" docAuthor="Rafael, José Albertino & Marques, Dayse Willkenia Almeida" docDate="2019" docId="039D09066519FFF4FF38FF26FB76F828" docLanguage="en" docName="zootaxa.4567.2.3.pdf" docOrigin="Zootaxa 4567 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrostomus montanus Rafael & Marques 2019, sp. nov." docType="treatment" docUuid="9EB41EDC-134E-4C71-802C-6A6E9ACDB185" docUuidSource="ZooBank" docVersion="10" lastPageNumber="261" masterDocId="FFA4717E6511FFFEFFAFFFB1FFCEFFE6" masterDocTitle="Five new species of Macrostomus Wiedemann and a checklist of Empididae (s. str.) (Diptera) from Bolivia" masterLastPageNumber="275" masterPageNumber="251" pageNumber="259" updateTime="1698701900396" updateUser="ExternalLinkService">
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<mods:title id="7E4C2B606B3CD70A8BF2B8B1A9DFCEAE">Five new species of Macrostomus Wiedemann and a checklist of Empididae (s. str.) (Diptera) from Bolivia</mods:title>
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<mods:name id="D260E348D6C736DC8D0947CF71089EC0" type="personal">
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<mods:namePart id="8C8DDC941AE18DD46B9E69FA7D857F05">Rafael, José Albertino</mods:namePart>
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<mods:namePart id="A374E3EF571F5ABC7431E495CE884262">Marques, Dayse Willkenia Almeida</mods:namePart>
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<mods:typeOfResource id="27FD1235551E407360D9571990E8B0AC">text</mods:typeOfResource>
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<mods:title id="F389114FFBEF21EC431A882B138D0E64">Zootaxa</mods:title>
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<mods:part id="957267E7063D5492097DAB22E27B4285">
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<mods:date id="2B5CCAD870857D8AC9B251D3769CF526">2019</mods:date>
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<mods:number id="C51C16A1C99872BD99943909DEFE11F7">2019-03-15</mods:number>
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<mods:number id="BBC6B7C7458A3B74EB9D7276B551ADE3">4567</mods:number>
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<treatment id="039D09066519FFF4FF38FF26FB76F828" ID-DOI="http://doi.org/10.5281/zenodo.5925350" ID-GBIF-Taxon="154402308" ID-Zenodo-Dep="5925350" LSID="urn:lsid:zoobank.org:act:9EB41EDC-134E-4C71-802C-6A6E9ACDB185" httpUri="http://treatment.plazi.org/id/039D09066519FFF4FF38FF26FB76F828" lastPageId="10" lastPageNumber="261" pageId="8" pageNumber="259">
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<subSubSection id="C32EEB9B6519FFF6FF38FF26FDEEFF54" box="[151,544,151,178]" pageId="8" pageNumber="259" type="nomenclature">
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<paragraph id="8B8BB8106519FFF6FF38FF26FDEEFF54" blockId="8.[151,982,151,245]" box="[151,544,151,178]" pageId="8" pageNumber="259">
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<heading id="D0C30F7C6519FFF6FF38FF26FDEEFF54" bold="true" box="[151,544,151,178]" fontSize="11" level="1" pageId="8" pageNumber="259" reason="1">
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<emphasis id="B94064026519FFF6FF38FF26FDEEFF54" bold="true" box="[151,544,151,178]" pageId="8" pageNumber="259">
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<taxonomicName id="4C34C3936519FFF6FF38FF26FE75FF57" authority="Rafael & Marques, 2019" authorityName="Rafael & Marques" authorityYear="2019" box="[151,443,151,177]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">
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<emphasis id="B94064026519FFF6FF38FF26FE75FF57" bold="true" box="[151,443,151,177]" italics="true" pageId="8" pageNumber="259">Macrostomus montanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A273D9796519FFF6FE6DFF29FDEEFF54" box="[450,544,152,178]" pageId="8" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EEB9B6519FFF6FF38FF0BFC18FF13" pageId="8" pageNumber="259" type="description">
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<paragraph id="8B8BB8106519FFF6FF38FF0BFEBBFF35" blockId="8.[151,982,151,245]" box="[151,373,186,211]" pageId="8" pageNumber="259">
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(
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<figureCitation id="130FA4956519FFF6FF30FF0BFF2AFF35" box="[159,228,186,211]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Figs 6</figureCitation>
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A–N, 7A–E)
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</paragraph>
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<paragraph id="8B8BB8106519FFF6FF38FF6DFC18FF13" blockId="8.[151,982,151,245]" box="[151,982,220,245]" pageId="8" pageNumber="259">
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<uri id="FFA5B4126519FFF6FF38FF6DFC18FF13" box="[151,982,220,245]" pageId="8" pageNumber="259">
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urn:lsid:zoobank.org:act:
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<uuid id="FF9282C56519FFF6FE05FF6DFC18FF13" box="[426,982,220,245]" pageId="8" pageNumber="259">9EB41EDC-134E-4C71-802C-6A6E9ACDB185</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EEB9B6519FFF6FF38FEADFA84FE5A" pageId="8" pageNumber="259" type="diagnosis">
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<paragraph id="8B8BB8106519FFF6FF38FEADFA84FE5A" blockId="8.[151,1437,284,1950]" pageId="8" pageNumber="259">
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<emphasis id="B94064026519FFF6FF38FEADFEDFFED3" bold="true" box="[151,273,284,309]" pageId="8" pageNumber="259">Diagnosis.</emphasis>
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Gena and postgena with setae somewhat stout; wing almost entirely hyaline, slightly brown infuscate at apex of costal cell, pterostigma and apex of cell r
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<subScript id="17B0BA556519FFF6FD4CFEFDFD35FEBC" attach="left" box="[739,763,332,346]" fontSize="6" pageId="8" pageNumber="259">2+3</subScript>
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; last section of vein M
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<subScript id="17B0BA556519FFF6FBBDFEFDFBD4FEBC" attach="left" box="[1042,1050,332,346]" fontSize="6" pageId="8" pageNumber="259">4</subScript>
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longer than vein dm-m; cell dm somewhat acute; tergite 8 short with distal sinus; epandrium with posterodorsal lobe longer than anterodorsal lobe, with longer setae more concentrated posteriorly on posterodorsal lobe; posterior cercus inwardly curved, trifid, apparently glabrous; hypandrium well sclerotized, slightly wider medially at level of posterior longer setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EEB9B6519FFF6FF68FE74FD56F953" pageId="8" pageNumber="259" type="description">
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<paragraph id="8B8BB8106519FFF6FF68FE74FB0FFA2A" blockId="8.[151,1437,284,1950]" pageId="8" pageNumber="259">
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<emphasis id="B94064026519FFF6FF68FE74FE97FE38" bold="true" box="[199,345,453,478]" pageId="8" pageNumber="259">Description.</emphasis>
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<typeStatus id="548F06B26519FFF6FECEFE74FE07FE38" box="[353,457,453,478]" pageId="8" pageNumber="259" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D3273996519FFF6FE7DFE77FE29FE3B" box="[466,487,454,477]" pageId="8" pageNumber="259" type="male">♂</specimenCount>
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(
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<figureCitation id="130FA4956519FFF6FE58FE74FD9DFE38" box="[503,595,453,478]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6A</figureCitation>
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). Body length
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<quantity id="4CCC15F56519FFF6FD52FE74FC99FE3B" box="[765,855,453,478]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="259" unit="mm" value="3.5">3.5 mm</quantity>
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; wing length
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<quantity id="4CCC15F56519FFF6FC5BFE77FB81FE3B" box="[1012,1103,454,478]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="259" unit="mm" value="3.2">3.2 mm</quantity>
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. Head dichoptic. Frons shiny black, slightly wider than anterior ocellus width. Two pairs of proclinate ocellar setae, posterior pair weaker. Face narrower than frons width, same as anterior ocellus width. Postcranium shiny black, entirely and sparsely gray pruinose. Inner vertical seta stouter than outer vertical seta, latter subequal in length to uniseriate postocular row of setae; 2–3 pairs of dorsal occipital setae. Gena with 3 setae and postgena with 2 somewhat stout setae. Antenna velvety black with postpedicel short pubescent. Proboscis shiny, brown basally, yellow apically. Palpus black. Thorax shiny black, gray pruinose. Thoracic chaetotaxy: 4 antepronotals; 1 postpronotal stout and 2 minutes; 4 dorsocentrals (right row with 5); 1 presutural supra-alar; 1 postsutural supra-alar; 1 postalar stout, 1 minute; 4 proepisternals weak; 1 notopleural stout and 1 weak; 2 pairs of scutellars, outer pair slightly weaker; 7 laterotergitals. Legs yellow with distal 3/4 of fore tibia, apex of mid femur, entire mid tibia, distal half of hind femur, entire hind tibia, all tarsi brown to black. Legs longer setae: fore tibia with 1 sub-basal and 1 median anterodorsal slightly longer; mid femur with 1 subapical anterior slightly longer; mid tibia with alternating anterodorsal and posterodorsal equally spaced; hind femur with 1 anterior subapical; hind tibia (
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<figureCitation id="130FA4956519FFF6FB7EFCE4FAE4FC88" box="[1233,1322,853,878]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6A</figureCitation>
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) with 4–5 slender dorsal and hind tarsomere 1 with 1 longer median dorsal seta. Wing (
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<figureCitation id="130FA4956519FFF6FB8FFCC7FBB5FC69" box="[1056,1147,886,911]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6A</figureCitation>
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) almost entirely hyaline, slightly brown infuscate at apex of costal cell, pterostigma and apex of cell r
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<subScript id="17B0BA556519FFF6FC5CFC16FBC5FC53" attach="left" box="[1011,1035,935,949]" fontSize="6" pageId="8" pageNumber="259">2+3</subScript>
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. Veins M
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<subScript id="17B0BA556519FFF6FBD7FC16FB4EFC53" attach="left" box="[1144,1152,935,949]" fontSize="6" pageId="8" pageNumber="259">1</subScript>
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and M
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<subScript id="17B0BA556519FFF6FB7FFC16FB16FC53" attach="left" box="[1232,1240,935,949]" fontSize="6" pageId="8" pageNumber="259">2</subScript>
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weaker and apex of CuA+CuP evanescent. Last section of vein M
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<subScript id="17B0BA556519FFF6FD69FC7AFD00FC3F" attach="left" box="[710,718,971,985]" fontSize="6" pageId="8" pageNumber="259">4</subScript>
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longer than vein dm-m. Cell dm somewhat acute. Halter light yellow. Abdomen (
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<figureCitation id="130FA4956519FFF6FEDFFC51FE09FC1F" box="[368,455,992,1017]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6B</figureCitation>
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) shiny black; tergites 1–3 with longer setae posteriorly. Tergite 8 (
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<figureCitation id="130FA4956519FFF6FB12FC51FADBFC1F" box="[1213,1301,992,1017]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6C</figureCitation>
|
||
) with distal sinus. Sternite 8 (
|
||
<figureCitation id="130FA4956519FFF6FECFFBB0FE77FBFC" box="[352,441,1025,1050]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6D</figureCitation>
|
||
) with distinct saddle-shaped projection posterodorsally and deep basal sinus. Tergite 8 articulated with sternite 8 somewhat horizontally (
|
||
<figureCitation id="130FA4956519FFF6FD75FB92FCFDFBDA" box="[730,819,1059,1084]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6E</figureCitation>
|
||
). Epandrium (
|
||
<figureCitation id="130FA4956519FFF6FC77FB92FBE3FBDA" box="[984,1069,1059,1084]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6F</figureCitation>
|
||
) with posterodorsal lobe longer than anterodorsal lobe, with longer setae more concentrated posteriorly on posterodorsal lobe; anterodorsal cleft minute, inconspicuous (smaller seta on
|
||
<figureCitation id="130FA4956519FFF6FDCFFBD4FD76FB98" box="[608,696,1125,1150]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6F</figureCitation>
|
||
); anteroventral cleft distinct (stronger seta on
|
||
<figureCitation id="130FA4956519FFF6FB60FBD4FAE8FB98" box="[1231,1318,1125,1150]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6F</figureCitation>
|
||
). Anterior cercus (
|
||
<figureCitation id="130FA4956519FFF6FF41FB37FEAFFB79" box="[238,353,1158,1183]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Figs 6G, J</figureCitation>
|
||
) small in lateral view, well sclerotized, with longer setae dorsally and posteriorly; when viewed anteriorly (
|
||
<figureCitation id="130FA4956519FFF6FEB8FB19FEBDFB27" box="[279,371,1192,1217]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6H</figureCitation>
|
||
) emitting mesially large and flat cercal bridge connected medially; cercal bridge expanded posteriorly and fused with upper hypoproctal lobe when viewed dorsally (
|
||
<figureCitation id="130FA4956519FFF6FBBFFB78FBABFB04" box="[1040,1125,1225,1250]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">
|
||
Fig.
|
||
<date id="FF8A9ED06519FFF6FBE4FB78FBABFB04" box="[1099,1125,1225,1250]" pageId="8" pageNumber="259">6I</date>
|
||
</figureCitation>
|
||
). Median cercus (
|
||
<figureCitation id="130FA4956519FFF6FA90FB78FA5AFB04" box="[1343,1428,1225,1250]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6J</figureCitation>
|
||
) somewhat cup-shaped with longer setae dorsally. Posterior cercus (
|
||
<figureCitation id="130FA4956519FFF6FC07FB5AFC35FAE2" box="[936,1019,1259,1284]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6J</figureCitation>
|
||
) inwardly curved, trifid, apparently glabrous. Ventral projection of cercus (
|
||
<figureCitation id="130FA4956519FFF6FDFCFABDFD62FAC3" box="[595,684,1292,1317]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6K</figureCitation>
|
||
) horizontally placed, distally expanded and setose. Dorsal lobe of hypoproct somewhat inconspicuous, fused with cercal bridge expansion of anterior cercus; ventral lobe of hypoproct distinctly setose, placed between setose lobes of ventral projection of cercus and cercal bridge of anterior cercus. Hypandrium (
|
||
<figureCitation id="130FA4956519FFF6FE48FAC1FDA9FA6F" box="[487,615,1392,1417]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Figs 6L, M</figureCitation>
|
||
) well sclerotized, almost same width throughout, slightly wider medially at level of posterior longer setae; distally with two pairs of curved hooks, distal pair stouter. Ejaculatory apodeme (
|
||
<figureCitation id="130FA4956519FFF6FF30FA02FF3DFA2A" box="[159,243,1459,1484]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6L</figureCitation>
|
||
) tetralamellar, all lamella subequal. Phallus (
|
||
<figureCitation id="130FA4956519FFF6FD42FA02FC8FFA2A" box="[749,833,1459,1484]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6L</figureCitation>
|
||
) slightly longer than hypandrium.
|
||
</paragraph>
|
||
<paragraph id="8B8BB8106519FFF6FF68FA65FA89F972" blockId="8.[151,1437,284,1950]" pageId="8" pageNumber="259">
|
||
<emphasis id="B94064026519FFF6FF68FA65FEEBFA0B" bold="true" box="[199,293,1492,1517]" pageId="8" pageNumber="259">Female.</emphasis>
|
||
Frons wider than width of anterior ocellus, wider than in male. Face slightly narrower than frons, approximately as wide as width of anterior ocellus. Wing distinctly more infuscated at costal and distal margins (
|
||
<figureCitation id="130FA4956519FFF6FF30F9A6FF35F9C9" box="[159,251,1559,1584]" captionStart="FIGURE 7" captionStartId="10.[151,250,1316,1338]" captionTargetBox="[181,1399,194,1294]" captionTargetId="figure@10.[181,1405,193,1294]" captionTargetPageId="10" captionText="FIGURE 7. Macrostomus montanus sp. nov. paratype ♀, A. Habitus, lateral view (specimen from Sarampioni, CBF collection); B–E. Specimen from Chulumani, NHMUK 010864430. B. Apex of abdomen, lateral view; C. Apex of abdomen, dorsal view; D. Sternite 8, genital fork and genital chamber, lateral view; E. Sternite 8 with genital fork and genital chamber placed laterally. Scale bars, figure A = 1 mm; figures B–E = 0.2 mm. Abbreviations: Ce, cercus; GC, genital chamber; GF, genital fork; St, sternite; Tg, tergite." httpUri="https://zenodo.org/record/2594949/files/figure.png" pageId="8" pageNumber="259">Fig. 7A</figureCitation>
|
||
). Tergite 8 (
|
||
<figureCitation id="130FA4956519FFF6FE27F9A6FDC9F9C9" box="[392,519,1559,1584]" captionStart="FIGURE 7" captionStartId="10.[151,250,1316,1338]" captionTargetBox="[181,1399,194,1294]" captionTargetId="figure@10.[181,1405,193,1294]" captionTargetPageId="10" captionText="FIGURE 7. Macrostomus montanus sp. nov. paratype ♀, A. Habitus, lateral view (specimen from Sarampioni, CBF collection); B–E. Specimen from Chulumani, NHMUK 010864430. B. Apex of abdomen, lateral view; C. Apex of abdomen, dorsal view; D. Sternite 8, genital fork and genital chamber, lateral view; E. Sternite 8 with genital fork and genital chamber placed laterally. Scale bars, figure A = 1 mm; figures B–E = 0.2 mm. Abbreviations: Ce, cercus; GC, genital chamber; GF, genital fork; St, sternite; Tg, tergite." httpUri="https://zenodo.org/record/2594949/files/figure.png" pageId="8" pageNumber="259">Figs 7B, C</figureCitation>
|
||
) wider basally in lateral view; with shallow basal sinus and deep distal sinus in dorsal view. Sternite 8 (
|
||
<figureCitation id="130FA4956519FFF6FE03F989FD9FF9B7" box="[428,593,1592,1617]" captionStart="FIGURE 7" captionStartId="10.[151,250,1316,1338]" captionTargetBox="[181,1399,194,1294]" captionTargetId="figure@10.[181,1405,193,1294]" captionTargetPageId="10" captionText="FIGURE 7. Macrostomus montanus sp. nov. paratype ♀, A. Habitus, lateral view (specimen from Sarampioni, CBF collection); B–E. Specimen from Chulumani, NHMUK 010864430. B. Apex of abdomen, lateral view; C. Apex of abdomen, dorsal view; D. Sternite 8, genital fork and genital chamber, lateral view; E. Sternite 8 with genital fork and genital chamber placed laterally. Scale bars, figure A = 1 mm; figures B–E = 0.2 mm. Abbreviations: Ce, cercus; GC, genital chamber; GF, genital fork; St, sternite; Tg, tergite." httpUri="https://zenodo.org/record/2594949/files/figure.png" pageId="8" pageNumber="259">Figs 7B, D, E</figureCitation>
|
||
) highly modified; in lateral and ventral views widened distally and with ventral subapical small translucid protuberance. Genital fork (
|
||
<figureCitation id="130FA4956519FFF6FCFAF9E8FC1CF994" box="[853,978,1625,1650]" captionStart="FIGURE 7" captionStartId="10.[151,250,1316,1338]" captionTargetBox="[181,1399,194,1294]" captionTargetId="figure@10.[181,1405,193,1294]" captionTargetPageId="10" captionText="FIGURE 7. Macrostomus montanus sp. nov. paratype ♀, A. Habitus, lateral view (specimen from Sarampioni, CBF collection); B–E. Specimen from Chulumani, NHMUK 010864430. B. Apex of abdomen, lateral view; C. Apex of abdomen, dorsal view; D. Sternite 8, genital fork and genital chamber, lateral view; E. Sternite 8 with genital fork and genital chamber placed laterally. Scale bars, figure A = 1 mm; figures B–E = 0.2 mm. Abbreviations: Ce, cercus; GC, genital chamber; GF, genital fork; St, sternite; Tg, tergite." httpUri="https://zenodo.org/record/2594949/files/figure.png" pageId="8" pageNumber="259">Figs 7D, E</figureCitation>
|
||
) T-shaped with arms upwardly directed. Genital chamber (
|
||
<figureCitation id="130FA4956519FFF6FECFF9CAFE19F975" box="[352,471,1659,1684]" captionStart="FIGURE 7" captionStartId="10.[151,250,1316,1338]" captionTargetBox="[181,1399,194,1294]" captionTargetId="figure@10.[181,1405,193,1294]" captionTargetPageId="10" captionText="FIGURE 7. Macrostomus montanus sp. nov. paratype ♀, A. Habitus, lateral view (specimen from Sarampioni, CBF collection); B–E. Specimen from Chulumani, NHMUK 010864430. B. Apex of abdomen, lateral view; C. Apex of abdomen, dorsal view; D. Sternite 8, genital fork and genital chamber, lateral view; E. Sternite 8 with genital fork and genital chamber placed laterally. Scale bars, figure A = 1 mm; figures B–E = 0.2 mm. Abbreviations: Ce, cercus; GC, genital chamber; GF, genital fork; St, sternite; Tg, tergite." httpUri="https://zenodo.org/record/2594949/files/figure.png" pageId="8" pageNumber="259">Fig. 7D, E</figureCitation>
|
||
) well sclerotized, wide, U-shaped, with basal protuberance upwardly directed.
|
||
</paragraph>
|
||
<paragraph id="8B8BB8106519FFF6FF68F92DFD56F953" blockId="8.[151,1437,284,1950]" box="[199,664,1692,1717]" pageId="8" pageNumber="259">
|
||
<emphasis id="B94064026519FFF6FF68F92DFE1EF953" bold="true" box="[199,464,1692,1717]" pageId="8" pageNumber="259">Geographical records.</emphasis>
|
||
<collectingCountry id="F323F8806519FFF6FE78F92DFDE7F953" box="[471,553,1692,1717]" name="Bolivia" pageId="8" pageNumber="259">Bolivia</collectingCountry>
|
||
(
|
||
<collectingRegion id="49F076F26519FFF6FD97F92CFD45F953" box="[568,651,1693,1717]" country="Bolivia" name="La Paz" pageId="8" pageNumber="259">La Paz</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32EEB9B6519FFF4FF68F90CFECBF9F8" lastPageId="10" lastPageNumber="261" pageId="8" pageNumber="259" type="materials_examined">
|
||
<paragraph id="8B8BB8106519FFF6FF68F90CFACCF878" blockId="8.[151,1437,284,1950]" pageId="8" pageNumber="259">
|
||
<emphasis id="B94064026519FFF6FF68F90CFD57F930" bold="true" box="[199,665,1725,1750]" pageId="8" pageNumber="259">
|
||
Type material examined.
|
||
<materialsCitation id="3B5CB24D6519FFF6FE55F90CFD57F930" ID-GBIF-Occurrence="2013156522" box="[506,665,1725,1750]" collectingDate="5. iv." collectionCode="NHMUK" country="Bolivia" county="Chulumani" elevation="2000" location="Chulumani" municipality="In" pageId="8" pageNumber="259" specimenCode="NHMUK 010664427" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="La Paz" typeStatus="holotype">
|
||
<typeStatus id="548F06B26519FFF6FE55F90CFD57F930" box="[506,665,1725,1750]" pageId="8" pageNumber="259" type="holotype">HOLOTYPE</typeStatus>
|
||
</materialsCitation>
|
||
</emphasis>
|
||
<specimenCount id="9D3273996519FFF6FD0DF90FFD77F933" box="[674,697,1726,1749]" pageId="8" pageNumber="259" type="male">♂</specimenCount>
|
||
,
|
||
<typeStatus id="548F06B26519FFF6FD69F90FFCE8F930" box="[710,806,1726,1750]" pageId="8" pageNumber="259" type="paratype">paratype</typeStatus>
|
||
<specimenCount id="9D3273996519FFF6FC80F90FFC8AF933" box="[815,836,1726,1749]" pageId="8" pageNumber="259" type="female">♀</specimenCount>
|
||
, on same pin: “
|
||
<collectingCountry id="F323F8806519FFF6FC55F90FFBA1F930" box="[1018,1135,1726,1750]" name="Bolivia" pageId="8" pageNumber="259">BOLIVIA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49F076F26519FFF6FBD4F90FFB04F933" box="[1147,1226,1726,1749]" country="Bolivia" name="La Paz" pageId="8" pageNumber="259">La Paz</collectingRegion>
|
||
,
|
||
<collectingCounty id="62EAC09C6519FFF6FB78F90CFA9DF930" box="[1239,1363,1725,1750]" pageId="8" pageNumber="259">Chulumani</collectingCounty>
|
||
, cloud forest, above
|
||
<quantity id="4CCC15F56519FFF6FE9FF96EFE47F911" box="[304,393,1759,1783]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="259" unit="m" value="2000.0">
|
||
<elevation id="00195F236519FFF6FE9FF96EFE47F911" box="[304,393,1759,1783]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="259" unit="m" value="2000.0">2000 m</elevation>
|
||
</quantity>
|
||
[eters]”; “
|
||
<collectingMunicipality id="6BEF226A6519FFF6FE5BF96EFDC3F911" box="[500,525,1759,1783]" pageId="8" pageNumber="259">In</collectingMunicipality>
|
||
copula,
|
||
<location id="8EEBEECB6519FFF6FDDCF96EFD3CF91E" LSID="urn:lsid:plazi:treatment:039D09066519FFF4FF38FF26FB76F828:8EEBEECB6519FFF6FDDCF96EFD3CF91E" box="[627,754,1759,1784]" country="Bolivia" county="Chulumani" municipality="In" name="Chulumani" pageId="8" pageNumber="259" stateProvince="La Paz">Chulumani</location>
|
||
,
|
||
<date id="FF8A9ED06519FFF6FCAEF96EFCFFF91E" box="[769,817,1759,1784]" pageId="8" pageNumber="259">
|
||
<collectingDate id="EFCE67386519FFF6FCAEF96EFCFFF91E" box="[769,817,1759,1784]" pageId="8" pageNumber="259">5.iv.</collectingDate>
|
||
</date>
|
||
[19]79”; “QR code:
|
||
<specimenCode id="DB92106B6519FFF6FB8FF951FAD9F91E" box="[1056,1303,1759,1784]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK 010664427</specimenCode>
|
||
”; “Holot
|
||
<specimenCount id="9D3273996519FFF6FA28F951FA52F911" box="[1415,1436,1760,1783]" pageId="8" pageNumber="259" type="male">♂</specimenCount>
|
||
Parat
|
||
<specimenCount id="9D3273996519FFF6FF76F8B0FF20F8FE" box="[217,238,1793,1816]" pageId="8" pageNumber="259" type="female">♀</specimenCount>
|
||
<taxonomicName id="4C34C3936519FFF6FF59F8B0FD2EF8FF" authority="Rafael & Marques" authorityName="Rafael & Marques" authorityYear="2019" box="[246,736,1792,1817]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B94064026519FFF6FF59F8B0FDCDF8FE" box="[246,515,1793,1816]" italics="true" pageId="8" pageNumber="259">Macrostomus montanus</emphasis>
|
||
Rafael & Marques
|
||
</taxonomicName>
|
||
” (
|
||
<figureCitation id="130FA4956519FFF6FD55F8B1FC9DF8FF" box="[762,851,1792,1817]" captionStart="FIGURE 6" captionStartId="9.[151,250,1736,1758]" captionTargetBox="[256,1311,194,1719]" captionTargetId="figure@9.[256,1329,193,1720]" captionTargetPageId="9" captionText="FIGURE 6. Macrostomus montanus sp. nov. paratype ♂. A. Habitus, before abdomen dissected, lateral view; B. Abdomen, dorsal view; C. Tergite 8, dorsal view; D. Sternite 8, ventral view; E. Tergite and sternite 8, lateral view; F. Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft); G. Epandrium and cerci, lateral view (pieces outlined); H. Cercal bridge of anterior cercus and subepandrial sclerite, anterior view; I. Cercal bridge of anterior cercus, dorsal view; J. Anterior, median and posterior lobes of cerci, lateral view; K. Dorsal and ventral hypoproctal lobes, posterior view; L. Hypandrium, ejaculatory apodeme and phallus, lateral view; M. Hypandrium, posterior view; N. Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe." httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="8" pageNumber="259">Fig. 6N</figureCitation>
|
||
) (NHMUK).
|
||
<materialsCitation id="3B5CB24D6519FFF6FC5FF8B1FD86F8DC" ID-GBIF-Occurrence="2013156533" collectingDate="2. iv." collectionCode="NHMUK" country="Bolivia" county="Chulumani" elevation="2000" location="Bolivia" municipality="In" pageId="8" pageNumber="259" specimenCount="1" stateProvince="La Paz" typeStatus="paratype">
|
||
<emphasis id="B94064026519FFF6FC5FF8B1FAC9F8FF" bold="true" box="[1008,1287,1792,1817]" pageId="8" pageNumber="259">
|
||
<typeStatus id="548F06B26519FFF6FC5FF8B1FB52F8FF" box="[1008,1180,1792,1817]" pageId="8" pageNumber="259" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<location id="8EEBEECB6519FFF6FB05F8B1FACCF8FF" LSID="urn:lsid:plazi:treatment:039D09066519FFF4FF38FF26FB76F828:8EEBEECB6519FFF6FB05F8B1FACCF8FF" box="[1194,1282,1792,1817]" name="Bolivia" pageId="8" pageNumber="259">Bolivia</location>
|
||
:
|
||
</emphasis>
|
||
same data as holotype, except
|
||
<date id="FF8A9ED06519FFF6FEF8F890FE48F8DC" box="[343,390,1825,1850]" pageId="8" pageNumber="259">
|
||
<collectingDate id="EFCE67386519FFF6FEF8F890FE48F8DC" box="[343,390,1825,1850]" pageId="8" pageNumber="259">2.iv.</collectingDate>
|
||
</date>
|
||
[19]79, QR code
|
||
</materialsCitation>
|
||
: NHMUK
|
||
<materialsCitation id="3B5CB24D6519FFF6FD66F890FBCBF8DC" ID-GBIF-Occurrence="2013156519" box="[713,1029,1825,1850]" collectionCode="NHMUK" country="Bolivia" pageId="8" pageNumber="259" specimenCount="1" specimenCount-male="1" stateProvince="La Paz" typeStatus="paratype">
|
||
0 10864440 (
|
||
<specimenCount id="9D3273996519FFF6FCF4F893FCB1F8DC" box="[859,895,1826,1850]" pageId="8" pageNumber="259" type="male">1♂</specimenCount>
|
||
,
|
||
<collectionCode id="ED2520D56519FFF6FC21F893FC33F8DC" box="[910,1021,1826,1850]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5CB24D6519FFF6FBBCF893FBB3F8DF" ID-GBIF-Occurrence="2013156540" box="[1043,1149,1826,1849]" collectionCode="NHMUK" country="Bolivia" isTailEnumeration="true" pageId="8" pageNumber="259" specimenCount="1" stateProvince="La Paz" typeStatus="paratype">
|
||
<collectionCode id="ED2520D56519FFF6FBBCF893FBB3F8DF" box="[1043,1149,1826,1849]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5CB24D6519FFF6FB28F890FA56F8DC" ID-GBIF-Occurrence="2013156520" box="[1159,1432,1825,1850]" collectionCode="INPA" country="Bolivia" pageId="8" pageNumber="259" specimenCount="1" specimenCount-female="1" stateProvince="La Paz" typeStatus="paratype">
|
||
0 10664353 (
|
||
<specimenCount id="9D3273996519FFF6FAB4F893FAF2F8DC" box="[1307,1340,1826,1850]" pageId="8" pageNumber="259" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED2520D56519FFF6FAE3F893FA5EF8DC" box="[1356,1424,1826,1850]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="8" pageNumber="259">INPA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5CB24D6519FFF6FF38F8F5FECFF8BD" ID-GBIF-Occurrence="2013156543" box="[151,257,1860,1883]" collectionCode="NHMUK" country="Bolivia" isTailEnumeration="true" pageId="8" pageNumber="259" specimenCount="1" stateProvince="La Paz" typeStatus="paratype">
|
||
<collectionCode id="ED2520D56519FFF6FF38F8F5FECFF8BD" box="[151,257,1860,1883]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5CB24D6519FFF6FEBBF8F2FDACF8BD" ID-GBIF-Occurrence="2013156546" box="[276,610,1859,1884]" collectionCode="NHMUK" country="Bolivia" pageId="8" pageNumber="259" specimenCount="1" specimenCount-female="1" stateProvince="La Paz" typeStatus="paratype">
|
||
0 10664429 (
|
||
<specimenCount id="9D3273996519FFF6FE1FF8F2FE1CF8BD" box="[432,466,1859,1883]" pageId="8" pageNumber="259" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED2520D56519FFF6FE44F8F2FD94F8BD" box="[491,602,1859,1883]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5CB24D6519FFF6FDD6F8F5FD2DF8BD" ID-GBIF-Occurrence="2013156534" box="[633,739,1860,1883]" collectionCode="NHMUK" country="Bolivia" isTailEnumeration="true" pageId="8" pageNumber="259" specimenCount="1" stateProvince="La Paz" typeStatus="paratype">
|
||
<collectionCode id="ED2520D56519FFF6FDD6F8F5FD2DF8BD" box="[633,739,1860,1883]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5CB24D6519FFF6FD59F8F2FB8AF8BD" ID-GBIF-Occurrence="2013156521" box="[758,1092,1859,1884]" collectionCode="NHMUK" country="Bolivia" pageId="8" pageNumber="259" specimenCount="1" specimenCount-female="1" stateProvince="La Paz" typeStatus="paratype">
|
||
0 10664436 (
|
||
<specimenCount id="9D3273996519FFF6FC3CF8F2FC7AF8BD" box="[915,948,1859,1883]" pageId="8" pageNumber="259" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED2520D56519FFF6FC62F8F2FBF2F8BD" box="[973,1084,1859,1883]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5CB24D6519FFF6FBF4F8F5FB0BF8BD" ID-GBIF-Occurrence="2013156524" box="[1115,1221,1860,1883]" collectionCode="NHMUK" country="Bolivia" isTailEnumeration="true" pageId="8" pageNumber="259" specimenCount="1" stateProvince="La Paz" typeStatus="paratype">
|
||
<collectionCode id="ED2520D56519FFF6FBF4F8F5FB0BF8BD" box="[1115,1221,1860,1883]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5CB24D6519FFF6FB77F8F2FEC0F89B" ID-GBIF-Occurrence="2013156526" collectionCode="NHMUK" country="Bolivia" pageId="8" pageNumber="259" specimenCount="1" specimenCount-female="1" stateProvince="La Paz" typeStatus="paratype">
|
||
0 10664441 (
|
||
<specimenCount id="9D3273996519FFF6FADAF8F2FA58F8BD" box="[1397,1430,1859,1883]" pageId="8" pageNumber="259" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED2520D56519FFF6FF38F8D4FEC8F89B" box="[151,262,1893,1917]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5CB24D6519FFF6FE8CF8D4FE43F89A" ID-GBIF-Occurrence="2013156553" box="[291,397,1893,1916]" collectionCode="NHMUK" country="Bolivia" isTailEnumeration="true" pageId="8" pageNumber="259" specimenCount="1" stateProvince="La Paz" typeStatus="paratype">
|
||
<collectionCode id="ED2520D56519FFF6FE8CF8D4FE43F89A" box="[291,397,1893,1916]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5CB24D6519FFF6FE32F8D5FB84F89B" ID-GBIF-Occurrence="2013156523" box="[413,1098,1892,1917]" collectingDate="25. iii." collectionCode="INPA" country="Bolivia" isTailEnumeration="true" pageId="8" pageNumber="259" specimenCount="1" specimenCount-female="1" stateProvince="La Paz" typeStatus="paratype">
|
||
0 10664442 (
|
||
<specimenCount id="9D3273996519FFF6FD97F8D4FD97F89B" box="[568,601,1893,1917]" pageId="8" pageNumber="259" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED2520D56519FFF6FDDFF8D4FD7AF89B" box="[624,692,1893,1917]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="8" pageNumber="259">INPA</collectionCode>
|
||
); same data, except
|
||
<date id="FF8A9ED06519FFF6FC13F8D5FC30F89B" box="[956,1022,1892,1917]" pageId="8" pageNumber="259">
|
||
<collectingDate id="EFCE67386519FFF6FC13F8D5FC30F89B" box="[956,1022,1892,1917]" pageId="8" pageNumber="259">25.iii.</collectingDate>
|
||
</date>
|
||
[19]79
|
||
</materialsCitation>
|
||
, NHMUK
|
||
<materialsCitation id="3B5CB24D6519FFF6FB75F8D5FE4DF878" ID-GBIF-Occurrence="2013156525" collectionCode="NHMUK" country="Bolivia" pageId="8" pageNumber="259" specimenCount="1" specimenCount-female="1" stateProvince="La Paz" typeStatus="paratype">
|
||
0 10664430 (
|
||
<specimenCount id="9D3273996519FFF6FADAF8D4FA58F89B" box="[1397,1430,1893,1917]" pageId="8" pageNumber="259" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED2520D56519FFF6FF38F837FECAF87B" box="[151,260,1926,1949]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="259" type="Museum">NHMUK</collectionCode>
|
||
, dissected)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5CB24D6519FFF6FE22F837FB30F878" ID-GBIF-Occurrence="2013156536" box="[397,1278,1925,1950]" collectingDate="2005-02-23" collectionCode="CBF" collectorName="Arm. Malaise, J & Rodriguez" country="Bolivia" location="Sarampionim" pageId="8" pageNumber="259" specimenCount="1" specimenCount-female="1" stateProvince="La Paz" typeStatus="paratype">
|
||
<collectingRegion id="49F076F26519FFF6FE22F837FE13F87B" box="[397,477,1926,1949]" country="Bolivia" name="La Paz" pageId="8" pageNumber="259">La Paz</collectingRegion>
|
||
,
|
||
<location id="8EEBEECB6519FFF6FE47F834FDB1F878" LSID="urn:lsid:plazi:treatment:039D09066519FFF4FF38FF26FB76F828:8EEBEECB6519FFF6FE47F834FDB1F878" box="[488,639,1925,1950]" country="Bolivia" name="Sarampionim" pageId="8" pageNumber="259" stateProvince="La Paz">Sarampionim</location>
|
||
[sic],
|
||
<date id="FF8A9ED06519FFF6FD69F834FCF7F878" box="[710,825,1925,1950]" pageId="8" pageNumber="259" value="2005-02-23">
|
||
<collectingDate id="EFCE67386519FFF6FD69F834FCF7F878" box="[710,825,1925,1950]" pageId="8" pageNumber="259" value="2005-02-23">23.ii.2005</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26C1DDC66519FFF6FCE9F837FC32F87B" box="[838,1020,1925,1950]" pageId="8" pageNumber="259">Arm. Malaise, J</collectorName>
|
||
.
|
||
<collectorName id="26C1DDC66519FFF6FBA9F834FBB5F878" box="[1030,1147,1925,1950]" pageId="8" pageNumber="259">Rodriguez</collectorName>
|
||
(
|
||
<specimenCount id="9D3273996519FFF6FB21F837FB61F878" box="[1166,1199,1926,1950]" pageId="8" pageNumber="259" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED2520D56519FFF6FB13F837FB38F878" box="[1212,1270,1926,1950]" country="Bolivia" httpUri="http://grbio.org/cool/ky5m-b42n" name="Coleccion Boliviana de Fauna" pageId="8" pageNumber="259">CBF</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF4BE8986518FFF7FF38F979FC7CF814" httpUri="https://zenodo.org/record/2594947/files/figure.png" pageId="9" pageNumber="260" startId="9.[151,250,1736,1758]" targetBox="[256,1311,194,1719]" targetPageId="9">
|
||
<paragraph id="8B8BB8106518FFF7FF38F979FC7CF814" blockId="9.[151,1436,1736,2034]" pageId="9" pageNumber="260">
|
||
<emphasis id="B94064026518FFF7FF38F979FEDAF93B" bold="true" box="[151,276,1736,1758]" pageId="9" pageNumber="260">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C34C3936518FFF7FEB3F978FDDEF938" authority="Rafael & Marques, 2019" authorityName="Rafael & Marques" authorityYear="2019" box="[284,528,1737,1758]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">
|
||
<emphasis id="B94064026518FFF7FEB3F978FDDEF938" box="[284,528,1737,1758]" italics="true" pageId="9" pageNumber="260">Macrostomus montanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94064026518FFF7FDB7F979FDA4F93B" bold="true" box="[536,618,1736,1757]" pageId="9" pageNumber="260">
|
||
<taxonomicNameLabel id="A273D9796518FFF7FDB7F979FDA4F93B" box="[536,618,1736,1757]" pageId="9" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
paratype ♂.
|
||
<emphasis id="B94064026518FFF7FD5BF979FCC3F93B" bold="true" box="[756,781,1736,1757]" pageId="9" pageNumber="260">A.</emphasis>
|
||
Habitus, before abdomen dissected, lateral view;
|
||
<emphasis id="B94064026518FFF7FABDF979FAE7F93B" bold="true" box="[1298,1321,1736,1757]" pageId="9" pageNumber="260">B.</emphasis>
|
||
Abdomen, dorsal view;
|
||
<emphasis id="B94064026518FFF7FE8DF956FEF4F91B" bold="true" box="[290,314,1767,1789]" pageId="9" pageNumber="260">C.</emphasis>
|
||
Tergite 8, dorsal view;
|
||
<emphasis id="B94064026518FFF7FD90F956FD96F91A" bold="true" box="[575,600,1767,1788]" pageId="9" pageNumber="260">D.</emphasis>
|
||
Sternite 8, ventral view;
|
||
<emphasis id="B94064026518FFF7FCC3F956FC4AF91A" bold="true" box="[876,900,1767,1788]" pageId="9" pageNumber="260">E.</emphasis>
|
||
Tergite and sternite 8, lateral view;
|
||
<emphasis id="B94064026518FFF7FAA2F956FAD1F91A" bold="true" box="[1293,1311,1767,1788]" pageId="9" pageNumber="260">F.</emphasis>
|
||
Epandrium detached, lateral view (superior arrow indicating anterodorsal cleft, inferior arrow indicating anteroventral cleft);
|
||
<emphasis id="B94064026518FFF7FAA1F8B4FAEDF8FD" bold="true" box="[1294,1315,1797,1819]" pageId="9" pageNumber="260">G.</emphasis>
|
||
Epandrium and cerci, lateral view (pieces outlined);
|
||
<emphasis id="B94064026518FFF7FD97F895FD9CF8DF" bold="true" box="[568,594,1828,1849]" pageId="9" pageNumber="260">H.</emphasis>
|
||
Cercal bridge of anterior cercus and subepandrial sclerite, anterior view;
|
||
<emphasis id="B94064026518FFF7FAECF895FA9DF8DF" bold="true" box="[1347,1363,1828,1849]" pageId="9" pageNumber="260">I.</emphasis>
|
||
Cercal bridge of anterior cercus, dorsal view;
|
||
<emphasis id="B94064026518FFF7FD81F8F2FD8FF8BE" bold="true" box="[558,577,1859,1880]" pageId="9" pageNumber="260">J.</emphasis>
|
||
Anterior, median and posterior lobes of cerci, lateral view;
|
||
<emphasis id="B94064026518FFF7FB16F8F2FB1DF8BE" bold="true" box="[1209,1235,1859,1880]" pageId="9" pageNumber="260">K.</emphasis>
|
||
Dorsal and ventral hypoproctal lobes, posterior view;
|
||
<emphasis id="B94064026518FFF7FE53F8D3FDDAF891" bold="true" box="[508,532,1890,1911]" pageId="9" pageNumber="260">L.</emphasis>
|
||
Hypandrium, ejaculatory apodeme and phallus, lateral view;
|
||
<emphasis id="B94064026518FFF7FB21F8D3FB63F891" bold="true" box="[1166,1197,1890,1911]" pageId="9" pageNumber="260">M.</emphasis>
|
||
Hypandrium, posterior view;
|
||
<emphasis id="B94064026518FFF7FF76F831FF3CF873" bold="true" box="[217,242,1920,1941]" pageId="9" pageNumber="260">N.</emphasis>
|
||
Holotype labels. Scale bars, figures A–B = 1 mm; C–M = 0.2 mm. Abbreviations: ACe, anterior cercus; ADL, anterodorsal lobe of epandrium; cbrd, cercal bridge; DHyL, dorsal hypoproctal lobe; Ej Apod, ejaculatory apodeme; Hypd, hypandrium; MCe, median cercus; PCe, posterior cercus; PDL, posterodorsal lobe of epandrium; Ph, phallus; sbep scl, subepandrial sclerite; St, sternite; Tg, tergite, VHyL, Ventral hypoproctal lobe.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4BE898651BFFF4FF38FA95FDCDFA52" httpUri="https://zenodo.org/record/2594949/files/figure.png" pageId="10" pageNumber="261" startId="10.[151,250,1316,1338]" targetBox="[181,1399,194,1294]" targetPageId="10">
|
||
<paragraph id="8B8BB810651BFFF4FF38FA95FDCDFA52" blockId="10.[151,1436,1316,1460]" pageId="10" pageNumber="261">
|
||
<emphasis id="B9406402651BFFF4FF38FA95FED7FADF" bold="true" box="[151,281,1316,1338]" pageId="10" pageNumber="261">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C34C393651BFFF4FE88FA94FDEEFADC" authority="Rafael & Marques, 2019" authorityName="Rafael & Marques" authorityYear="2019" box="[295,544,1317,1338]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">
|
||
<emphasis id="B9406402651BFFF4FE88FA94FDEEFADC" box="[295,544,1317,1338]" italics="true" pageId="10" pageNumber="261">Macrostomus montanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9406402651BFFF4FD83FA95FD4AFADF" bold="true" box="[556,644,1316,1337]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="A273D979651BFFF4FD83FA95FD4AFADF" box="[556,644,1316,1337]" pageId="10" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
paratype ♀,
|
||
<emphasis id="B9406402651BFFF4FCB2FA95FCF8FADF" bold="true" box="[797,822,1316,1337]" pageId="10" pageNumber="261">A.</emphasis>
|
||
Habitus, lateral view (specimen from Sarampioni, CBF collection);
|
||
<emphasis id="B9406402651BFFF4FEBEFAF2FE88FABE" bold="true" box="[273,326,1347,1369]" pageId="10" pageNumber="261">B–E.</emphasis>
|
||
Specimen from Chulumani, NHMUK 0 10864430.
|
||
<emphasis id="B9406402651BFFF4FCFAFAF2FCA2FABE" bold="true" box="[853,876,1347,1368]" pageId="10" pageNumber="261">B.</emphasis>
|
||
Apex of abdomen, lateral view;
|
||
<emphasis id="B9406402651BFFF4FB11FAF2FB19FABF" bold="true" box="[1214,1239,1347,1369]" pageId="10" pageNumber="261">C.</emphasis>
|
||
Apex of abdomen, dorsal view;
|
||
<emphasis id="B9406402651BFFF4FEB3FAD0FEFBFA90" bold="true" box="[284,309,1377,1398]" pageId="10" pageNumber="261">D.</emphasis>
|
||
Sternite 8, genital fork and genital chamber, lateral view;
|
||
<emphasis id="B9406402651BFFF4FC20FAD0FC68FA90" bold="true" box="[911,934,1377,1398]" pageId="10" pageNumber="261">E.</emphasis>
|
||
Sternite 8 with genital fork and genital chamber placed laterally. Scale bars, figure A = 1 mm; figures B–E = 0.2 mm. Abbreviations: Ce, cercus; GC, genital chamber; GF, genital fork; St, sternite; Tg, tergite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8BB810651BFFF4FF68FA50FECBF9F8" blockId="10.[151,1437,1505,1998]" pageId="10" pageNumber="261">
|
||
<emphasis id="B9406402651BFFF4FF68FA50FE78FA1C" bold="true" box="[199,438,1505,1530]" pageId="10" pageNumber="261">
|
||
<typeStatus id="548F06B2651BFFF4FF68FA50FEFAFA1C" box="[199,308,1505,1530]" pageId="10" pageNumber="261" type="holotype">Holotype</typeStatus>
|
||
condition.
|
||
</emphasis>
|
||
Good. Left hind leg lost; right hind leg glued on piece of white card; abdomen not dissected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32EEB9B651BFFF4FF68F998FECAF980" pageId="10" pageNumber="261" type="etymology">
|
||
<paragraph id="8B8BB810651BFFF4FF68F998FECAF980" blockId="10.[151,1437,1505,1998]" pageId="10" pageNumber="261">
|
||
<emphasis id="B9406402651BFFF4FF68F998FE80F9A4" bold="true" box="[199,334,1577,1602]" pageId="10" pageNumber="261">Etymology.</emphasis>
|
||
From the Latin
|
||
<taxonomicName id="4C34C393651BFFF4FDA7F99BFDBBF9A7" authorityName="Rafael & Marques" authorityYear="2019" box="[520,629,1578,1601]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B9406402651BFFF4FDA7F99BFDBBF9A7" box="[520,629,1578,1601]" italics="true" pageId="10" pageNumber="261">montanus</emphasis>
|
||
</taxonomicName>
|
||
= mountain and refers to the high elevation where the specimens were collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32EEB9B651BFFF4FF68F9C0FD6BF948" pageId="10" pageNumber="261" type="description">
|
||
<paragraph id="8B8BB810651BFFF4FF68F9C0FD6BF948" blockId="10.[151,1437,1505,1998]" pageId="10" pageNumber="261">
|
||
<emphasis id="B9406402651BFFF4FF68F9C0FEF2F96C" bold="true" box="[199,316,1649,1674]" pageId="10" pageNumber="261">Variation.</emphasis>
|
||
One specimen (QR code: NHMUK 010864440) with three pairs of ocellar setae; the additional anterior pair slightly longer than posterior pair.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32EEB9B651BFFF4FF68F908FB76F828" pageId="10" pageNumber="261" type="discussion">
|
||
<paragraph id="8B8BB810651BFFF4FF68F908FB76F828" blockId="10.[151,1437,1505,1998]" pageId="10" pageNumber="261">
|
||
<emphasis id="B9406402651BFFF4FF68F908FEF5F934" bold="true" box="[199,315,1721,1746]" pageId="10" pageNumber="261">Remarks.</emphasis>
|
||
<taxonomicName id="4C34C393651BFFF4FEE7F90BFD95F937" authority="Rafael & Marques, 2019" authorityName="Rafael & Marques" authorityYear="2019" box="[328,603,1722,1745]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">
|
||
<emphasis id="B9406402651BFFF4FEE7F90BFD95F937" box="[328,603,1722,1745]" italics="true" pageId="10" pageNumber="261">Macrostomus montanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9406402651BFFF4FDC7F908FD07F934" bold="true" box="[616,713,1721,1746]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="A273D979651BFFF4FDC7F908FD07F934" box="[616,713,1721,1746]" pageId="10" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is tentatively included in the
|
||
<taxonomicName id="4C34C393651BFFF4FB92F90BFB22F937" box="[1085,1260,1721,1745]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="limbipennis">
|
||
<emphasis id="B9406402651BFFF4FB92F90BFB22F937" box="[1085,1260,1721,1745]" italics="true" pageId="10" pageNumber="261">M. limbipennis</emphasis>
|
||
</taxonomicName>
|
||
species-group, however, it differs mainly by having epandrium with posterodorsal lobe longer than the anterodorsal lobe. In the key presented by
|
||
<bibRefCitation id="EFA5C5E1651BFFF4FE82F8B0FDAFF8FF" author="Rafael, J. A. & Cumming, J. M." box="[301,609,1793,1818]" pageId="10" pageNumber="261" pagination="45 - 55" refId="ref14826" refString="Rafael, J. A. & Cumming, J. M. (2012) Revision of the genus Macrostomus Wiedemann (Diptera, Empididae, Empidinae). III. The limbipennis species-group. Zootaxa, 3361, 45 - 55. https: // doi. org / 10.1590 / S 0044 - 59672010000300023" type="journal article" year="2012">Rafael & Cumming (2012)</bibRefCitation>
|
||
, this species runs to couplet 3 separating
|
||
<taxonomicName id="4C34C393651BFFF4FB9AF8B3FAF6F8FC" authority="(Bezzi)" baseAuthorityName="Bezzi" box="[1077,1336,1793,1818]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="limbipennis">
|
||
<emphasis id="B9406402651BFFF4FB9AF8B3FB10F8FF" box="[1077,1246,1793,1817]" italics="true" pageId="10" pageNumber="261">M. limbipennis</emphasis>
|
||
(Bezzi)
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C34C393651BFFF4FAD2F8B3FE0EF8D8" authority="Rafael & Cumming" authorityName="Rafael & Cumming" authorityYear="2006" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="penai">
|
||
<emphasis id="B9406402651BFFF4FAD2F8B3FF1BF8DB" italics="true" pageId="10" pageNumber="261">M. penai</emphasis>
|
||
Rafael & Cumming
|
||
</taxonomicName>
|
||
, but it differs from the first species by male tergite 8 not humped and from the second species by all cercal lobes shortened (versus elongate and upwardly directed in
|
||
<taxonomicName id="4C34C393651BFFF4FBFDF8FBFB75F887" authorityName="Rafael & Cumming" authorityYear="2006" box="[1106,1211,1866,1889]" class="Insecta" family="Empididae" genus="Macrostomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="penai">
|
||
<emphasis id="B9406402651BFFF4FBFDF8FBFB75F887" box="[1106,1211,1866,1889]" italics="true" pageId="10" pageNumber="261">M. penai</emphasis>
|
||
</taxonomicName>
|
||
), posterior lobe of epandrium longer than anterior lobe (versus same length) and by female sternite 8 with a ventral subapical small translucid protuberance (versus absent), genital fork T-shaped with arms upwardly directed (versus small rod, without lateral arms), genital chamber wide, U-shaped, well sclerotized (versus membranous).
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||
</paragraph>
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||
</subSubSection>
|
||
</treatment>
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||
</document> |