643 lines
79 KiB
XML
643 lines
79 KiB
XML
<document id="D854C964348405F1CC02ACACBFDB0877" ID-CLB-Dataset="25959" ID-DOI="10.11646/zootaxa.4658.1.4" ID-GBIF-Dataset="7138d838-3b94-44f6-92ca-fb6c3b42832e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3373129" ID-ZooBank="11726879-264F-4643-B6A0-B240C1D8CE10" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566365215565" checkinUser="plazi" docAuthor="Shen, Huei-Ping, Chang, Chih-Han & Chih, Wen-Jay" docDate="2019" docId="039087F7FFBDFFE5FF14FB69FDF8FE94" docLanguage="en" docName="zootaxa.4658.1.4.pdf" docOrigin="Zootaxa 4658 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amynthas luridus Shen & Chang & Chih 2019, sp. nov." docType="treatment" docVersion="11" lastPageNumber="106" masterDocId="FFA9FF8FFFBEFFECFF83FFE0FF84FFD7" masterDocTitle="Two new earthworm species of the genus Amynthas (Oligochaeta: Megascolecidae) from central Taiwan, with comments on some recent species assignments in Amynthas and Metaphire" masterLastPageNumber="123" masterPageNumber="101" pageNumber="104" updateTime="1698751945313" updateUser="ExternalLinkService">
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<mods:mods id="14D3BE92B1C37A0E598F133BB83D0CF3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2F3A5509B970BF2DE9BD1BD5DD5CD34C">
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<mods:title id="BF4F4EBAD8CC2062CA2B57C424E9BC29">Two new earthworm species of the genus Amynthas (Oligochaeta: Megascolecidae) from central Taiwan, with comments on some recent species assignments in Amynthas and Metaphire</mods:title>
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</mods:titleInfo>
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<mods:name id="ACA2B9DA0205DBB83C9445926A1DB3DB" type="personal">
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<mods:roleTerm id="08348A09E9AB2A7A8D6F6867E2F96F1A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0C0DB5AB2722A538F1E4723DB0EDA51F">Shen, Huei-Ping</mods:namePart>
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</mods:name>
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<mods:roleTerm id="29544EA99FB4373C6D826C1885540670">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7E866C6634BEF5F44942D8AB26A9C347">Chang, Chih-Han</mods:namePart>
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</mods:name>
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<mods:name id="A4D97594EE09BF5B7F5A5860625A9425" type="personal">
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<mods:role id="B3217278E9B92CA27FC586A4605D7A8D">
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<mods:roleTerm id="DE185E5D98E0CDB0BC95BA1B1A1F21CE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6263A4C157A3D3D6573F9513DF6E1B8C">Chih, Wen-Jay</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E844B001C7631EC5B13EE0B7640F842C">text</mods:typeOfResource>
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<mods:titleInfo id="5BF91877D94947E4D36369740A532138">
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<mods:title id="02169760C64D41EFCC278A87BA2F89E5">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="50AE032C5CB22F5CEB2A1CAD6014D170">
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<mods:date id="84B6B5F98B0B0A94C5D6C3347F9A13ED">2019</mods:date>
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<mods:detail id="E1861A3A71765722E777FE300870D5B6" type="pubDate">
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<mods:number id="CF348E04A895F4CA958C929111024916">2019-08-21</mods:number>
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<mods:detail id="BA69CB69DEE45ADA6C2A5992A31F52DE" type="volume">
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<mods:number id="8851DD8231860AA7F4C9262FE872E8EA">4658</mods:number>
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<mods:detail id="6A7C49EE61034249A6EAD39033D003D9" type="issue">
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<mods:number id="BFD27EDDE5A03B0B77463BA63BAF8374">1</mods:number>
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<mods:start id="274EF4B3A79784848E019653C7035547">101</mods:start>
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<mods:end id="78E3D2F0164FFD35391A8741A8EE4F3B">123</mods:end>
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<mods:classification id="60D9B8DF2EC5DF12A966A89D1F3A6CA5">journal article</mods:classification>
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<mods:identifier id="9A4535956C4AD41BB8351E91C4A922B4" type="CLB-Dataset">25959</mods:identifier>
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<mods:identifier id="8076A7CED9DA65D6677D59BFBFD4A8CB" type="DOI">10.11646/zootaxa.4658.1.4</mods:identifier>
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<mods:identifier id="732492C30B25991E2465E58BFEAC0F00" type="GBIF-Dataset">7138d838-3b94-44f6-92ca-fb6c3b42832e</mods:identifier>
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<mods:identifier id="1D682110C77A819DFB3711194A6C6463" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="FFFFB8116F7BD4A888AEE888A1848CC8" type="Zenodo-Dep">3373129</mods:identifier>
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<mods:identifier id="9050B63399D8155739CF6FFCAC52DC15" type="ZooBank">11726879-264F-4643-B6A0-B240C1D8CE10</mods:identifier>
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</mods:mods>
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<treatment id="039087F7FFBDFFE5FF14FB69FDF8FE94" ID-GBIF-Taxon="159217275" LSID="urn:lsid:plazi:treatment:039087F7FFBDFFE5FF14FB69FDF8FE94" httpUri="http://treatment.plazi.org/id/039087F7FFBDFFE5FF14FB69FDF8FE94" lastPageId="9" lastPageNumber="106" pageId="3" pageNumber="104">
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<subSubSection id="C323656AFFBDFFEFFF14FB69FD1FFB73" box="[151,667,1161,1188]" pageId="3" pageNumber="104" type="nomenclature">
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<paragraph id="8B8636E1FFBDFFEFFF14FB69FD1FFB73" blockId="3.[151,667,1161,1223]" box="[151,667,1161,1188]" pageId="3" pageNumber="104">
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<heading id="D0CE818DFFBDFFEFFF14FB69FD1FFB73" bold="true" box="[151,667,1161,1188]" fontSize="11" level="1" pageId="3" pageNumber="104" reason="1">
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<emphasis id="B94DEAF3FFBDFFEFFF14FB69FD1FFB73" bold="true" box="[151,667,1161,1188]" pageId="3" pageNumber="104">
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<taxonomicName id="4C394D62FFBDFFEFFF14FB69FDB5FB74" authority="Shen & Chang" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[151,561,1161,1188]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="104" phylum="Annelida" rank="species" species="luridus" status="sp. nov.">
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<emphasis id="B94DEAF3FFBDFFEFFF14FB69FEEBFB73" bold="true" box="[151,367,1161,1188]" italics="true" pageId="3" pageNumber="104">Amynthas luridus</emphasis>
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Shen & Chang
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27E5788FFBDFFEFFDBEFB6AFD1FFB73" box="[573,667,1162,1188]" pageId="3" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C323656AFFBDFFEFFF14FB4DFE8CFB10" box="[151,264,1197,1223]" pageId="3" pageNumber="104" type="description">
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<paragraph id="8B8636E1FFBDFFEFFF14FB4DFE8CFB10" blockId="3.[151,667,1161,1223]" box="[151,264,1197,1223]" pageId="3" pageNumber="104">
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||
(
|
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<figureCitation id="13022A64FFBDFFEFFF1CFB4DFF79FB10" box="[159,253,1197,1223]" captionStart="FIGURE 1" captionStartId="4.[151,250,1543,1568]" captionTargetBox="[196,1417,211,1516]" captionTargetId="figure@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 1. Amynthas luridus sp. nov. A. Ventral view of spermathecal pore region of Holotype (TESRI-O-H-54) (sp, spermathecal pore; gp, genital papilla). B. Ventral view of male pore region of holotype (mp, male porophore). C. Ventral view of male pore region of paratype. D. Dorsal view of right spermathecae of holotype (amp, ampulla; dv, diverticulum; ag, accessory gland). E. Dorsal view of right prostate gland of holotype (pd, prostatic duct). Scale bars = 1 mm." httpUri="https://zenodo.org/record/3373131/files/figure.png" pageId="3" pageNumber="104">Figure 1</figureCitation>
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)
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C323656AFFBDFFE5FF14FB16FDF8FE94" lastPageId="9" lastPageNumber="110" pageId="3" pageNumber="104" type="materials_examined">
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||
<paragraph id="8B8636E1FFBDFFEFFF14FB16FEDBFA80" blockId="3.[151,1437,1269,1980]" pageId="3" pageNumber="104">
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||
<materialsCitation id="3B513CBCFFBDFFEFFF14FB16FEDBFA80" ID-GBIF-Occurrence="2350486285" collectingDate="2009-04-15" collectorName="H. - P. Chen" elevation="1526" latitude="24.11333" location="Ruiyen Creek Nature Reserve" longLatPrecision="1" longitude="121.1697" municipality="Clitellate" pageId="3" pageNumber="104" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
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||
<emphasis id="B94DEAF3FFBDFFEFFF14FB16FE89FAC7" bold="true" box="[151,269,1270,1296]" pageId="3" pageNumber="104">
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||
<typeStatus id="54828843FFBDFFEFFF14FB16FE8CFAC7" box="[151,264,1270,1296]" pageId="3" pageNumber="104" type="holotype">Holotype</typeStatus>
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||
:
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</emphasis>
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||
<collectingMunicipality id="6BE2AC9BFFBDFFEFFE96FB15FEF8FAD8" box="[277,380,1269,1295]" pageId="3" pageNumber="104">Clitellate</collectingMunicipality>
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(mature) specimen (
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||
<quantity id="4CC19B04FFBDFFEFFDE6FB15FD32FAC7" box="[613,694,1269,1296]" metricMagnitude="-2" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="104" unit="mm" value="97.0">97 mm</quantity>
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||
in total length, dissected), from roadside ditches in
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||
<location id="8EE6603AFFBDFFEFFA82FB15FECCFAE3" LSID="urn:lsid:plazi:treatment:039087F7FFBDFFE5FF14FB69FDF8FE94:8EE6603AFFBDFFEFFA82FB15FECCFAE3" latitude="24.11333" longLatPrecision="1" longitude="121.1697" municipality="Clitellate" name="Ruiyen Creek Nature Reserve" pageId="3" pageNumber="104" stateProvince="Nantou">Ruiyen Creek Nature Reserve</location>
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||
,
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||
<collectingRegion id="49FDF803FFBDFFEFFED7FAFAFD86FAE4" box="[340,514,1305,1332]" country="Taiwan" name="Nantou" pageId="3" pageNumber="104">Nantou County</collectingRegion>
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,
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||
<geoCoordinate id="EE0D5026FFBDFFEFFD8DFAF9FD33FAE4" box="[526,695,1305,1331]" degrees="24" direction="north" minutes="06" orientation="latitude" pageId="3" pageNumber="104" precision="1" seconds="47.99" value="24.11333">24°06'47.99''N</geoCoordinate>
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||
,
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<geoCoordinate id="EE0D5026FFBDFFEFFD41FAF9FCF1FAE4" box="[706,885,1305,1331]" degrees="121" direction="east" minutes="10" orientation="longitude" pageId="3" pageNumber="104" precision="1" seconds="10.93" value="121.1697">121°10'10.93''E</geoCoordinate>
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,
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||
<elevation id="0014D1D2FFBDFFEFFC02FAF9FB84FAE4" box="[897,1024,1305,1332]" metricMagnitude="3" metricUnit="m" metricValue="1.526" pageId="3" pageNumber="104" unit="m" value="1526.0">
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<quantity id="4CC19B04FFBDFFEFFC02FAF9FC53FAE3" box="[897,983,1305,1332]" metricMagnitude="3" metricUnit="m" metricValue="1.526" pageId="3" pageNumber="104" unit="m" value="1526.0">1526 m</quantity>
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||
asl
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||
</elevation>
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||
,
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||
<date id="FF871021FFBDFFEFFB8FFAF9FB2FFAE4" box="[1036,1195,1305,1332]" pageId="3" pageNumber="104" value="2009-04-15">
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||
<collectingDate id="EFC3E9C9FFBDFFEFFB8FFAF9FB2FFAE4" box="[1036,1195,1305,1332]" pageId="3" pageNumber="104" value="2009-04-15">15 April 2009</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="26CC5337FFBDFFEFFB34FAFAFAB0FAE4" box="[1207,1332,1305,1332]" pageId="3" pageNumber="104">H.-P. Chen</collectorName>
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||
(TESRI- O-H-54; RY021).
|
||
</materialsCitation>
|
||
</paragraph>
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||
<paragraph id="8B8636E1FFBDFFEFFF44FA82FB4CFA48" blockId="3.[151,1437,1269,1980]" pageId="3" pageNumber="104">
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||
<materialsCitation id="3B513CBCFFBDFFEFFF44FA82FB40FA48" ID-GBIF-Occurrence="2350486286" collectingDate="2009-03-26" collectorName="T. - J. Lin & D. - H. Chen" elevation="2198" latitude="24.109959" location="Ruiyen Creek Nature Reserve" longLatPrecision="1" longitude="121.18894" pageId="3" pageNumber="104" specimenCount="1" stateProvince="Nantou" typeStatus="paratype">
|
||
<emphasis id="B94DEAF3FFBDFFEFFF44FA82FEB9FAAB" bold="true" box="[199,317,1378,1404]" pageId="3" pageNumber="104">
|
||
<typeStatus id="54828843FFBDFFEFFF44FA82FEBCFAAB" box="[199,312,1378,1404]" pageId="3" pageNumber="104" type="paratype">Paratype</typeStatus>
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||
:
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</emphasis>
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1 aclitellate, from roadside slopes in
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||
<location id="8EE6603AFFBDFFEFFD5EFA81FBA9FAAB" LSID="urn:lsid:plazi:treatment:039087F7FFBDFFE5FF14FB69FDF8FE94:8EE6603AFFBDFFEFFD5EFA81FBA9FAAB" box="[733,1069,1377,1404]" latitude="24.109959" longLatPrecision="1" longitude="121.18894" name="Ruiyen Creek Nature Reserve" pageId="3" pageNumber="104" stateProvince="Nantou">Ruiyen Creek Nature Reserve</location>
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||
,
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||
<collectingRegion id="49FDF803FFBDFFEFFBB4FA82FB61FAAC" box="[1079,1253,1377,1404]" country="Taiwan" name="Nantou" pageId="3" pageNumber="104">Nantou County</collectingRegion>
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||
,
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||
<geoCoordinate id="EE0D5026FFBDFFEFFB73FA81FA1CFAAB" box="[1264,1432,1377,1404]" degrees="24" direction="north" minutes="06" orientation="latitude" pageId="3" pageNumber="104" precision="1" seconds="35.85" value="24.109959">24°06'35.85''N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE0D5026FFBDFFEFFF14FA65FECDFA48" box="[151,329,1413,1439]" degrees="121" direction="east" minutes="11" orientation="longitude" pageId="3" pageNumber="104" precision="1" seconds="20.19" value="121.18894">121°11'20.19''E</geoCoordinate>
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||
,
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||
<elevation id="0014D1D2FFBDFFEFFED6FA65FE56FA48" box="[341,466,1413,1440]" metricMagnitude="3" metricUnit="m" metricValue="2.198" pageId="3" pageNumber="104" unit="m" value="2198.0">
|
||
<quantity id="4CC19B04FFBDFFEFFED6FA65FE2EFA77" box="[341,426,1413,1440]" metricMagnitude="3" metricUnit="m" metricValue="2.198" pageId="3" pageNumber="104" unit="m" value="2198.0">2198 m</quantity>
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||
asl
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||
</elevation>
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||
,
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||
<date id="FF871021FFBDFFEFFE5EFA65FD08FA48" box="[477,652,1413,1439]" pageId="3" pageNumber="104" value="2009-03-26">
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||
<collectingDate id="EFC3E9C9FFBDFFEFFE5EFA65FD08FA48" box="[477,652,1413,1439]" pageId="3" pageNumber="104" value="2009-03-26">26 March 2009</collectingDate>
|
||
</date>
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||
,
|
||
<collectorName id="26CC5337FFBDFFEFFD14FA66FD7CFA48" box="[663,760,1413,1440]" pageId="3" pageNumber="104">T.-J. Lin</collectorName>
|
||
&
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||
<collectorName id="26CC5337FFBDFFEFFC9FFA66FC19FA48" box="[796,925,1413,1440]" pageId="3" pageNumber="104">D.-H. Chen</collectorName>
|
||
(TESRI-O-P-48; RY020)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8636E1FFBDFFEFFF44FA49FF73F968" blockId="3.[151,1437,1269,1980]" pageId="3" pageNumber="104">
|
||
<emphasis id="B94DEAF3FFBDFFEFFF44FA49FE05FA13" bold="true" box="[199,385,1449,1476]" pageId="3" pageNumber="104">Other material.</emphasis>
|
||
1 clitellate (dissected) and 5 aclitellates (one dissected) from Mt. Beidongyen,
|
||
<collectingRegion id="49FDF803FFBDFFEFFB68FA4AFA1CFA14" box="[1259,1432,1449,1476]" country="Taiwan" name="Nantou" pageId="3" pageNumber="104">Nantou County</collectingRegion>
|
||
,
|
||
<quantity id="4CC19B04FFBDFFEFFF14FA2DFF69FA3F" box="[151,237,1485,1512]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="104" unit="m" value="1800.0">1800 m</quantity>
|
||
asl,
|
||
<date id="FF871021FFBDFFEFFEA7FA2DFE74FA30" box="[292,496,1485,1511]" pageId="3" pageNumber="104" value="1999-12-08">8 December 1999</date>
|
||
,
|
||
<materialsCitation id="3B513CBCFFBDFFEFFE7FFA2DFA84F9DB" ID-GBIF-Occurrence="2350486442" collectingDate="1999-06-29" collectingDateMax="2005-08-19" collectingDateMin="1999-06-29" collectorName="C. - F. Tsai & S. - C. Tsai & P. - H. Ho & Shen" elevation="1817" latitude="24.08307" location="Mt. Beidongyen" longLatPrecision="1" longitude="121.13671" pageId="3" pageNumber="104" specimenCount="1" stateProvince="Nantou">
|
||
<collectorName id="26CC5337FFBDFFEFFE7FFA2DFDF5FA30" box="[508,625,1485,1511]" pageId="3" pageNumber="104">C.-F. Tsai</collectorName>
|
||
,
|
||
<collectorName id="26CC5337FFBDFFEFFDFEFA2DFD76FA30" box="[637,754,1485,1511]" pageId="3" pageNumber="104">S.-C. Tsai</collectorName>
|
||
,
|
||
<collectorName id="26CC5337FFBDFFEFFD7DFA2EFCE0FA3F" box="[766,868,1486,1512]" pageId="3" pageNumber="104">P.-H. Ho</collectorName>
|
||
& H.-P.
|
||
<collectorName id="26CC5337FFBDFFEFFC4EFA2DFB81FA30" box="[973,1029,1485,1511]" pageId="3" pageNumber="104">Shen</collectorName>
|
||
(TESRI-O-
|
||
<date id="FF871021FFBDFFEFFB10FA2DFA81FA3F" box="[1171,1285,1485,1512]" pageId="3" pageNumber="104" value="1999-06-29">
|
||
<collectingDate id="EFC3E9C9FFBDFFEFFB10FA2DFA81FA3F" box="[1171,1285,1485,1512]" pageId="3" pageNumber="104" value="1999-06-29">1999-29-6</collectingDate>
|
||
</date>
|
||
); 1 aclitellate from
|
||
<location id="8EE6603AFFBDFFEFFF57FA12FE09F9DC" LSID="urn:lsid:plazi:treatment:039087F7FFBDFFE5FF14FB69FDF8FE94:8EE6603AFFBDFFEFFF57FA12FE09F9DC" box="[212,397,1521,1548]" latitude="24.08307" longLatPrecision="1" longitude="121.13671" name="Mt. Beidongyen" pageId="3" pageNumber="104" stateProvince="Nantou">Mt. Beidongyen</location>
|
||
,
|
||
<collectingRegion id="49FDF803FFBDFFEFFE1BFA12FDC2F9DC" box="[408,582,1521,1548]" country="Taiwan" name="Nantou" pageId="3" pageNumber="104">Nantou County</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE0D5026FFBDFFEFFDD2FA11FD7DF9DB" box="[593,761,1521,1548]" degrees="24" direction="north" minutes="04" orientation="latitude" pageId="3" pageNumber="104" precision="1" seconds="59.05" value="24.08307">24°04'59.05''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0D5026FFBDFFEFFC87FA11FC33F9DC" box="[772,951,1521,1547]" degrees="121" direction="east" minutes="08" orientation="longitude" pageId="3" pageNumber="104" precision="1" seconds="12.18" value="121.13671">121°08'12.18''E</geoCoordinate>
|
||
,
|
||
<elevation id="0014D1D2FFBDFFEFFC41FA11FBC4F9DC" box="[962,1088,1521,1548]" metricMagnitude="3" metricUnit="m" metricValue="1.8169999999999997" pageId="3" pageNumber="104" unit="m" value="1817.0">
|
||
<quantity id="4CC19B04FFBDFFEFFC41FA11FB93F9DB" box="[962,1047,1521,1548]" metricMagnitude="3" metricUnit="m" metricValue="1.8169999999999997" pageId="3" pageNumber="104" unit="m" value="1817.0">1817 m</quantity>
|
||
asl
|
||
</elevation>
|
||
,
|
||
<date id="FF871021FFBDFFEFFBC9FA11FA84F9DB" box="[1098,1280,1521,1548]" pageId="3" pageNumber="104" value="2005-08-19">
|
||
<collectingDate id="EFC3E9C9FFBDFFEFFBC9FA11FA84F9DB" box="[1098,1280,1521,1548]" pageId="3" pageNumber="104" value="2005-08-19">19 August 2005</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B513CBCFFBDFFEFFA89FA12FA8BF984" ID-GBIF-Occurrence="2350486443" collectingDate="2005-03-09" collectingDateMax="2009-03-25" collectingDateMin="2005-03-09" collectorName="T. - L. Huang & I. - M. Hsiao & Shen" elevation="2195" latitude="24.10397" location="Ruiyen Creek Nature Reserve" longLatPrecision="1" longitude="121.17611" pageId="3" pageNumber="104" specimenCount="1" stateProvince="Nantou">
|
||
<collectorName id="26CC5337FFBDFFEFFA89FA12FA1CF9DB" box="[1290,1432,1522,1548]" pageId="3" pageNumber="104">T.-L. Huang</collectorName>
|
||
,
|
||
<collectorName id="26CC5337FFBDFFEFFF14F9F6FE9DF9F8" box="[151,281,1557,1584]" pageId="3" pageNumber="104">I.-M. Hsiao</collectorName>
|
||
& H.-P.
|
||
<collectorName id="26CC5337FFBDFFEFFE01F9F5FE3EF9F8" box="[386,442,1557,1583]" pageId="3" pageNumber="104">Shen</collectorName>
|
||
(TESRI-O-
|
||
<date id="FF871021FFBDFFEFFDC4F9F5FD2FF9E7" box="[583,683,1557,1584]" pageId="3" pageNumber="104" value="2005-03-09">
|
||
<collectingDate id="EFC3E9C9FFBDFFEFFDC4F9F5FD2FF9E7" box="[583,683,1557,1584]" pageId="3" pageNumber="104" value="2005-03-09">2005-9-3</collectingDate>
|
||
</date>
|
||
); 1 clitellate (RY019, PCR failed) from roadside ditches in
|
||
<location id="8EE6603AFFBDFFEFFAC8F9F5FE10F983" LSID="urn:lsid:plazi:treatment:039087F7FFBDFFE5FF14FB69FDF8FE94:8EE6603AFFBDFFEFFAC8F9F5FE10F983" latitude="24.10397" longLatPrecision="1" longitude="121.17611" name="Ruiyen Creek Nature Reserve" pageId="3" pageNumber="104" stateProvince="Nantou">Ruiyen Creek Nature Reserve</location>
|
||
,
|
||
<collectingRegion id="49FDF803FFBDFFEFFE23F9DAFDD4F984" box="[416,592,1593,1620]" country="Taiwan" name="Nantou" pageId="3" pageNumber="104">Nantou County</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE0D5026FFBDFFEFFDDEF9D9FC81F984" box="[605,773,1593,1619]" degrees="24" direction="north" minutes="06" orientation="latitude" pageId="3" pageNumber="104" precision="1" seconds="14.29" value="24.10397">24°06'14.29''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0D5026FFBDFFEFFC91F9D9FC41F984" box="[786,965,1593,1619]" degrees="121" direction="east" minutes="10" orientation="longitude" pageId="3" pageNumber="104" precision="1" seconds="34.01" value="121.17611">121°10'34.01''E</geoCoordinate>
|
||
,
|
||
<elevation id="0014D1D2FFBDFFEFFC51F9D9FBD6F984" box="[978,1106,1593,1620]" metricMagnitude="3" metricUnit="m" metricValue="2.195" pageId="3" pageNumber="104" unit="m" value="2195.0">
|
||
<quantity id="4CC19B04FFBDFFEFFC51F9D9FBACF983" box="[978,1064,1593,1620]" metricMagnitude="3" metricUnit="m" metricValue="2.195" pageId="3" pageNumber="104" unit="m" value="2195.0">2195 m</quantity>
|
||
asl
|
||
</elevation>
|
||
,
|
||
<date id="FF871021FFBDFFEFFBDDF9D9FA8BF984" box="[1118,1295,1593,1620]" pageId="3" pageNumber="104" value="2009-03-25">
|
||
<collectingDate id="EFC3E9C9FFBDFFEFFBDDF9D9FA8BF984" box="[1118,1295,1593,1620]" pageId="3" pageNumber="104" value="2009-03-25">25 March 2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B513CBCFFBDFFEFFA98F9DAFB04F94C" ID-GBIF-Occurrence="2350486384" collectingDate="2009-03-25" collectingDateMax="2009-05-06" collectingDateMin="2009-03-25" collectorName="T. - J. Lin & D. - H. Chen" elevation="2308" latitude="24.108267" location="Ruiyen Creek Nature Reserve" longLatPrecision="1" longitude="121.19002" pageId="3" pageNumber="104" specimenCount="1" stateProvince="Nantou">
|
||
<collectorName id="26CC5337FFBDFFEFFA98F9DAFAFAF984" box="[1307,1406,1593,1620]" pageId="3" pageNumber="104">T.-J. Lin</collectorName>
|
||
&
|
||
<collectorName id="26CC5337FFBDFFEFFF14F9BEFE9CF9A0" box="[151,280,1629,1656]" pageId="3" pageNumber="104">D.-H. Chen</collectorName>
|
||
(TESRI-O-
|
||
<date id="FF871021FFBDFFEFFE27F9BDFD8CF9AF" box="[420,520,1629,1656]" pageId="3" pageNumber="104" value="2009-05-06">
|
||
<collectingDate id="EFC3E9C9FFBDFFEFFE27F9BDFD8CF9AF" box="[420,520,1629,1656]" pageId="3" pageNumber="104" value="2009-05-06">2009-6-5</collectingDate>
|
||
</date>
|
||
); 1 clitellate (RY018) from ditches around forestry station in
|
||
<location id="8EE6603AFFBDFFEFFB33F9BDFF70F94B" LSID="urn:lsid:plazi:treatment:039087F7FFBDFFE5FF14FB69FDF8FE94:8EE6603AFFBDFFEFFB33F9BDFF70F94B" latitude="24.108267" longLatPrecision="1" longitude="121.19002" name="Ruiyen Creek Nature Reserve" pageId="3" pageNumber="104" stateProvince="Nantou">Ruiyen Creek Nature Reserve</location>
|
||
,
|
||
<collectingRegion id="49FDF803FFBDFFEFFE81F962FE30F94C" box="[258,436,1665,1692]" country="Taiwan" name="Nantou" pageId="3" pageNumber="104">Nantou County</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE0D5026FFBDFFEFFE41F961FDEFF94C" box="[450,619,1665,1691]" degrees="24" direction="north" minutes="06" orientation="latitude" pageId="3" pageNumber="104" precision="1" seconds="29.76" value="24.108267">24°06'29.76''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0D5026FFBDFFEFFDFAF961FCAFF94C" box="[633,811,1665,1691]" degrees="121" direction="east" minutes="11" orientation="longitude" pageId="3" pageNumber="104" precision="1" seconds="24.07" value="121.19002">121°11'24.07''E</geoCoordinate>
|
||
,
|
||
<elevation id="0014D1D2FFBDFFEFFCBAF961FC39F94C" box="[825,957,1665,1692]" metricMagnitude="3" metricUnit="m" metricValue="2.308" pageId="3" pageNumber="104" unit="m" value="2308.0">
|
||
<quantity id="4CC19B04FFBDFFEFFCBAF961FC16F94B" box="[825,914,1665,1692]" metricMagnitude="3" metricUnit="m" metricValue="2.308" pageId="3" pageNumber="104" unit="m" value="2308.0">2308 m</quantity>
|
||
asl
|
||
</elevation>
|
||
,
|
||
<date id="FF871021FFBDFFEFFC48F961FB04F94C" box="[971,1152,1665,1692]" pageId="3" pageNumber="104" value="2009-03-25">
|
||
<collectingDate id="EFC3E9C9FFBDFFEFFC48F961FB04F94C" box="[971,1152,1665,1692]" pageId="3" pageNumber="104" value="2009-03-25">25 March 2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
, M.-H. Shen (TESRI-O- 2009-7).
|
||
</paragraph>
|
||
<paragraph id="8B8636E1FFBDFFEFFF44F929FAFCF86C" blockId="3.[151,1437,1269,1980]" pageId="3" pageNumber="104">
|
||
<emphasis id="B94DEAF3FFBDFFEFFF44F929FEC6F933" bold="true" box="[199,322,1737,1764]" pageId="3" pageNumber="104">Diagnosis.</emphasis>
|
||
Length (clitellates)
|
||
<quantity id="4CC19B04FFBDFFEFFDA4F929FD37F933" box="[551,691,1737,1764]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" metricValueMax="10.7" metricValueMin="8.3" pageId="3" pageNumber="104" unit="mm" value="95.0" valueMax="107.0" valueMin="83.0">83–107 mm</quantity>
|
||
. Segments numbering 94–132. Setae
|
||
<quantity id="4CC19B04FFBDFFEFFBDDF929FB46F934" box="[1118,1218,1737,1764]" metricMagnitude="0" metricUnit="m" metricValue="1.3208" metricValueMax="1.4986" metricValueMin="1.143" pageId="3" pageNumber="104" unit="in" value="52.0" valueMax="59.0" valueMin="45.0">45–59 in</quantity>
|
||
VII,
|
||
<quantity id="4CC19B04FFBDFFEFFA83F929FAE1F934" box="[1280,1381,1737,1764]" metricMagnitude="0" metricUnit="m" metricValue="1.4478" metricValueMax="1.6509999999999998" metricValueMin="1.2446" pageId="3" pageNumber="104" unit="in" value="57.0" valueMax="65.0" valueMin="49.0">49–65 in</quantity>
|
||
XX, 9–13 between male pores. First dorsal pore in 12/13. Clitellum XIV–XVI. Spermathecal pores four pairs in 5/6–8/9, 0.28–0.32 body circumference ventrally apart. Male pores 0.26–0.27 body circumference ventrally apart in XVIII, each on a round porophore about
|
||
<quantity id="4CC19B04FFBDFFEFFD8AF8D5FDDAF887" box="[521,606,1845,1872]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="104" unit="mm" value="0.5">0.5 mm</quantity>
|
||
in diameter, with an anterior and/or a posterior genital papillae. Additional papillae often widely paired in postsetal XVII or occasionally paired in presetal XIX. Spermathecae four pairs in VI– IX. Seminal vesicles vestigial, two pairs in XI and XII. Prostate glands lobed, paired in
|
||
<collectionCode id="ED28AE24FFBDFFEFFBECF89EFB6AF84F" box="[1135,1262,1918,1944]" pageId="3" pageNumber="104">XVII–XIX</collectionCode>
|
||
. Prostatic ducts small, short, C-shaped in XVIII. Accessory glands sessile or stalked, corresponding to external genital papillae.
|
||
</paragraph>
|
||
<caption id="DF466669FFBAFFE8FF14F9E7FB5AF95C" httpUri="https://zenodo.org/record/3373131/files/figure.png" pageId="4" pageNumber="105" startId="4.[151,250,1543,1568]" targetBox="[196,1417,211,1516]" targetPageId="4">
|
||
<paragraph id="8B8636E1FFBAFFE8FF14F9E7FB5AF95C" blockId="4.[151,1436,1543,1676]" pageId="4" pageNumber="105">
|
||
<emphasis id="B94DEAF3FFBAFFE8FF14F9E7FE9CF9F7" bold="true" box="[151,280,1543,1568]" pageId="4" pageNumber="105">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C394D62FFBAFFE8FEA0F9E7FE5EF9C8" authority="Shen & Chang & Chih, 2019" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[291,474,1543,1567]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="105" phylum="Annelida" rank="species" species="luridus" status="sp. nov.">
|
||
<emphasis id="B94DEAF3FFBAFFE8FEA0F9E7FE5EF9C8" box="[291,474,1543,1567]" italics="true" pageId="4" pageNumber="105">Amynthas luridus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAF3FFBAFFE8FE66F9E8FDE4F9F7" bold="true" box="[485,608,1543,1568]" pageId="4" pageNumber="105">
|
||
<taxonomicNameLabel id="A27E5788FFBAFFE8FE66F9E8FDB9F9F7" box="[485,573,1544,1568]" pageId="4" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A.
|
||
</emphasis>
|
||
Ventral view of spermathecal pore region of Holotype (TESRI-O-H-54) (sp, spermathecal pore; gp, genital papilla).
|
||
<emphasis id="B94DEAF3FFBAFFE8FDB5F9CCFDC9F993" bold="true" box="[566,589,1580,1604]" pageId="4" pageNumber="105">B.</emphasis>
|
||
Ventral view of male pore region of holotype (mp, male porophore).
|
||
<emphasis id="B94DEAF3FFBAFFE8FAACF9CBFAC3F993" bold="true" box="[1327,1351,1579,1604]" pageId="4" pageNumber="105">C.</emphasis>
|
||
Ventral view of male pore region of paratype.
|
||
<emphasis id="B94DEAF3FFBAFFE8FDA1F9B0FDBFF9BF" bold="true" box="[546,571,1616,1640]" pageId="4" pageNumber="105">D.</emphasis>
|
||
Dorsal view of right spermathecae of holotype (amp, ampulla; dv, diverticulum; ag, accessory gland).
|
||
<emphasis id="B94DEAF3FFBAFFE8FECEF994FEE1F95B" bold="true" box="[333,357,1652,1676]" pageId="4" pageNumber="105">E.</emphasis>
|
||
Dorsal view of right prostate gland of holotype (pd, prostatic duct). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8636E1FFBAFFE9FF44F959FA06FE5C" blockId="4.[151,1437,1721,2036]" lastBlockId="5.[151,1437,153,2016]" lastPageId="5" lastPageNumber="106" pageId="4" pageNumber="105">
|
||
<emphasis id="B94DEAF3FFBAFFE8FF44F959FEDDF903" bold="true" box="[199,345,1721,1748]" pageId="4" pageNumber="105">Description.</emphasis>
|
||
External: Length (clitellates)
|
||
<quantity id="4CC19B04FFBAFFE8FD1EF95AFCA7F903" box="[669,803,1722,1748]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" metricValueMax="10.7" metricValueMin="8.3" pageId="4" pageNumber="105" unit="mm" value="95.0" valueMax="107.0" valueMin="83.0">83–107 mm</quantity>
|
||
, weight
|
||
<quantity id="4CC19B04FFBAFFE8FCFFF959FC7BF903" box="[892,1023,1721,1748]" metricMagnitude="-3" metricUnit="kg" metricValue="1.26" metricValueMax="1.54" metricValueMin="0.98" pageId="4" pageNumber="105" unit="g" value="1.26" valueMax="1.54" valueMin="0.98">0.98–1.54 g</quantity>
|
||
in 95%ethanol. Segments numbering 94–132. Clitellum XIV–XVI, setae and dorsal pores absent,
|
||
<quantity id="4CC19B04FFBAFFE8FCC7F93EFC6CF92F" box="[836,1000,1758,1784]" metricMagnitude="-3" metricUnit="m" metricValue="3.525" metricValueMax="4.22" metricValueMin="2.83" pageId="4" pageNumber="105" unit="mm" value="3.525" valueMax="4.22" valueMin="2.83">2.83–4.22 mm</quantity>
|
||
in length and
|
||
<quantity id="4CC19B04FFBAFFE8FB0FF93EFAB4F92F" box="[1164,1328,1758,1784]" metricMagnitude="-3" metricUnit="m" metricValue="3.925" metricValueMax="4.38" metricValueMin="3.47" pageId="4" pageNumber="105" unit="mm" value="3.925" valueMax="4.38" valueMin="3.47">3.47–4.38 mm</quantity>
|
||
in width. Prostomium epilobous. Three annuli (secondary segments) per segment in VI–XIII. Setal number
|
||
<quantity id="4CC19B04FFBAFFE8FA83F8E1FAE1F8CB" box="[1280,1381,1793,1820]" metricMagnitude="0" metricUnit="m" metricValue="1.3208" metricValueMax="1.4986" metricValueMin="1.143" pageId="4" pageNumber="105" unit="in" value="52.0" valueMax="59.0" valueMin="45.0">45–59 in</quantity>
|
||
VII,
|
||
<quantity id="4CC19B04FFBAFFE8FF14F8C5FF7FF897" box="[151,251,1829,1856]" metricMagnitude="0" metricUnit="m" metricValue="1.4478" metricValueMax="1.6509999999999998" metricValueMin="1.2446" pageId="4" pageNumber="105" unit="in" value="57.0" valueMax="65.0" valueMin="49.0">49–65 in</quantity>
|
||
XX, and 9–13 between male pores in XVIII. First dorsal pore in 12/13. Spermathecal pores four pairs in intersegmental furrows of 5/6–8/9, distance between paired pores 0.28–0.32 body circumference ventrally apart, each pore small, often with a small papilla posteromedial and another anteromedial to each pore, papilla
|
||
<quantity id="4CC19B04FFBAFFE8FAB9F88DFF47F87B" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="4" pageNumber="105" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.25–0.3 mm</quantity>
|
||
in diameter (
|
||
<figureCitation id="13022A64FFBAFFE8FEDEF872FE33F87B" box="[349,439,1938,1964]" captionStart="FIGURE 1" captionStartId="4.[151,250,1543,1568]" captionTargetBox="[196,1417,211,1516]" captionTargetId="figure@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 1. Amynthas luridus sp. nov. A. Ventral view of spermathecal pore region of Holotype (TESRI-O-H-54) (sp, spermathecal pore; gp, genital papilla). B. Ventral view of male pore region of holotype (mp, male porophore). C. Ventral view of male pore region of paratype. D. Dorsal view of right spermathecae of holotype (amp, ampulla; dv, diverticulum; ag, accessory gland). E. Dorsal view of right prostate gland of holotype (pd, prostatic duct). Scale bars = 1 mm." httpUri="https://zenodo.org/record/3373131/files/figure.png" pageId="4" pageNumber="105">Fig. 1A</figureCitation>
|
||
). No other genital papillae in the preclitellar region. Female pore single, mid-ventral in XIV. Male pores inconspicuous, paired in XVIII, 0.26–0.27 body circumference ventrally apart, each situated on a round porophore about
|
||
<quantity id="4CC19B04FFBAFFE8FE39F839FD96F823" box="[442,530,2009,2036]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="105" unit="mm" value="0.5">0.5 mm</quantity>
|
||
in diameter. Genital papillae anterior and/or posterior to male pores, paired, in line with and smaller than male porophore. Additional papillae often widely paired in line with male porophore in postsetal XVII or occasionally paired in presetal XIX. For all
|
||
<specimenCount id="9D3FFD68FFBBFFE9FCC5FF5DFC5AFF00" box="[838,990,189,215]" pageId="5" pageNumber="106" type="generic">11 specimens</specimenCount>
|
||
examined, male pores with anterior and posterior papillae for two specimens and one of them (RY021) with an additional pair in postsetal XVII (
|
||
<figureCitation id="13022A64FFBBFFE9FAB5FF01FA09FF2C" box="[1334,1421,225,251]" captionStart="FIGURE 1" captionStartId="4.[151,250,1543,1568]" captionTargetBox="[196,1417,211,1516]" captionTargetId="figure@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 1. Amynthas luridus sp. nov. A. Ventral view of spermathecal pore region of Holotype (TESRI-O-H-54) (sp, spermathecal pore; gp, genital papilla). B. Ventral view of male pore region of holotype (mp, male porophore). C. Ventral view of male pore region of paratype. D. Dorsal view of right spermathecae of holotype (amp, ampulla; dv, diverticulum; ag, accessory gland). E. Dorsal view of right prostate gland of holotype (pd, prostatic duct). Scale bars = 1 mm." httpUri="https://zenodo.org/record/3373131/files/figure.png" pageId="5" pageNumber="106">Fig. 1B</figureCitation>
|
||
); male pores with anterior papillae for two specimens and one of them (RY020) with an additional pair in presetal XIX (
|
||
<figureCitation id="13022A64FFBBFFE9FF5AFEC9FEB4FE94" box="[217,304,297,323]" captionStart="FIGURE 1" captionStartId="4.[151,250,1543,1568]" captionTargetBox="[196,1417,211,1516]" captionTargetId="figure@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 1. Amynthas luridus sp. nov. A. Ventral view of spermathecal pore region of Holotype (TESRI-O-H-54) (sp, spermathecal pore; gp, genital papilla). B. Ventral view of male pore region of holotype (mp, male porophore). C. Ventral view of male pore region of paratype. D. Dorsal view of right spermathecae of holotype (amp, ampulla; dv, diverticulum; ag, accessory gland). E. Dorsal view of right prostate gland of holotype (pd, prostatic duct). Scale bars = 1 mm." httpUri="https://zenodo.org/record/3373131/files/figure.png" pageId="5" pageNumber="106">Fig. 1C</figureCitation>
|
||
); male pores with posterior papillae for seven specimens and five of them with an additional pair in postsetal XVII. Each papilla round, with a slightly concave center, about
|
||
<quantity id="4CC19B04FFBBFFE9FC64FEACFBC9FEB0" box="[999,1101,332,359]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="106" unit="mm" value="0.35">0.35 mm</quantity>
|
||
in diameter. Genital papillae absent in the mid-ventral region. Live worms white. Preserved specimens white and slightly darker on clitellum.
|
||
</paragraph>
|
||
<paragraph id="8B8636E1FFBBFFE9FF44FE75FA95FD24" blockId="5.[151,1437,153,2016]" pageId="5" pageNumber="106">
|
||
Internal: Septa 5/6–7/8 thick,
|
||
<date id="FF871021FFBBFFE9FD9EFE75FDE1FE78" box="[541,613,405,431]" pageId="5" pageNumber="106" value="2010-09-08">8/9/10</date>
|
||
absent, 10/11 thick,
|
||
<date id="FF871021FFBBFFE9FCD4FE75FC61FE78" box="[855,997,405,431]" pageId="5" pageNumber="106" value="2014-11-12" valueMax="2014-11-13" valueMin="2014-11-12">11/12–13/14</date>
|
||
muscular. Nephridial tufts on anterior faces of 5/6/7. Gizzard large, round in
|
||
<collectionCode id="ED28AE24FFBBFFE9FDDCFE59FD31FE04" box="[607,693,441,467]" pageId="5" pageNumber="106">VIII–X</collectionCode>
|
||
. Intestine enlarged from XVI. Intestinal caeca paired in XXVII, simple, extending anteriorly to XXIV or XXIII. Esophageal hearts paired in XI–XIII. Spermathecae four pairs in VI–IX (
|
||
<figureCitation id="13022A64FFBBFFE9FF71FDE1FEC8FDCC" box="[242,332,513,539]" captionStart="FIGURE 1" captionStartId="4.[151,250,1543,1568]" captionTargetBox="[196,1417,211,1516]" captionTargetId="figure@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 1. Amynthas luridus sp. nov. A. Ventral view of spermathecal pore region of Holotype (TESRI-O-H-54) (sp, spermathecal pore; gp, genital papilla). B. Ventral view of male pore region of holotype (mp, male porophore). C. Ventral view of male pore region of paratype. D. Dorsal view of right spermathecae of holotype (amp, ampulla; dv, diverticulum; ag, accessory gland). E. Dorsal view of right prostate gland of holotype (pd, prostatic duct). Scale bars = 1 mm." httpUri="https://zenodo.org/record/3373131/files/figure.png" pageId="5" pageNumber="106">Fig. 1D</figureCitation>
|
||
), ampulla elongated oval-shaped, surface wrinkled,
|
||
<quantity id="4CC19B04FFBBFFE9FC27FDE0FBCDFDCC" box="[932,1097,512,539]" metricMagnitude="-3" metricUnit="m" metricValue="1.355" metricValueMax="1.45" metricValueMin="1.26" pageId="5" pageNumber="106" unit="mm" value="1.355" valueMax="1.45" valueMin="1.26">1.26–1.45 mm</quantity>
|
||
long and
|
||
<quantity id="4CC19B04FFBBFFE9FB3CFDE1FAD2FDCC" box="[1215,1366,513,539]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" metricValueMax="8.4" metricValueMin="6.0" pageId="5" pageNumber="106" unit="mm" value="0.72" valueMax="0.84" valueMin="0.6">0.6–0.84 mm</quantity>
|
||
wide, spermathecal duct stout,
|
||
<quantity id="4CC19B04FFBBFFE9FE2FFDC5FDB6FDE8" box="[428,562,549,575]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="5" pageNumber="106" unit="mm" value="0.44999999999999996" valueMax="0.6" valueMin="0.3">0.3–0.6 mm</quantity>
|
||
in length. Diverticulum stalk slender, seminal chamber rudimentary or absent, not iridescent,
|
||
<quantity id="4CC19B04FFBBFFE9FEC3FDA9FE52FDB4" box="[320,470,585,611]" metricMagnitude="-3" metricUnit="m" metricValue="1.06" metricValueMax="1.32" metricValueMin="0.8" pageId="5" pageNumber="106" unit="mm" value="1.06" valueMax="1.32" valueMin="0.8">0.8–1.32 mm</quantity>
|
||
in total length. Accessory glands sessile or stalked,
|
||
<quantity id="4CC19B04FFBBFFE9FBA8FDA8FB37FDB4" box="[1067,1203,584,611]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="106" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 mm</quantity>
|
||
in total length, each corresponding to external genital papilla. Holandric. Testes small, round, two pairs in ventrally joined sacs in X and XI. Seminal vesicles vestigial, two pairs in XI and XII, occupying less than half of the segmental compartment. Prostate glands paired in
|
||
<collectionCode id="ED28AE24FFBBFFE9FE32FD55FDA8FD18" box="[433,556,693,719]" pageId="5" pageNumber="106">XVII–XIX</collectionCode>
|
||
(
|
||
<figureCitation id="13022A64FFBBFFE9FDB8FD55FD0BFD18" box="[571,655,693,719]" captionStart="FIGURE 1" captionStartId="4.[151,250,1543,1568]" captionTargetBox="[196,1417,211,1516]" captionTargetId="figure@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 1. Amynthas luridus sp. nov. A. Ventral view of spermathecal pore region of Holotype (TESRI-O-H-54) (sp, spermathecal pore; gp, genital papilla). B. Ventral view of male pore region of holotype (mp, male porophore). C. Ventral view of male pore region of paratype. D. Dorsal view of right spermathecae of holotype (amp, ampulla; dv, diverticulum; ag, accessory gland). E. Dorsal view of right prostate gland of holotype (pd, prostatic duct). Scale bars = 1 mm." httpUri="https://zenodo.org/record/3373131/files/figure.png" pageId="5" pageNumber="106">Fig. 1E</figureCitation>
|
||
), wrinkled and lobed. Prostatic ducts short, small, C-shaped in XVIII. Accessory glands round, sessile, about
|
||
<quantity id="4CC19B04FFBBFFE9FDCCFD39FD21FD24" box="[591,677,729,755]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="106" unit="mm" value="0.3">0.3 mm</quantity>
|
||
in diameter, corresponding to external genital papillae.
|
||
</paragraph>
|
||
<paragraph id="8B8636E1FFBBFFE9FF44FD1CFDA3FCEC" blockId="5.[151,1437,153,2016]" pageId="5" pageNumber="106">
|
||
<emphasis id="B94DEAF3FFBBFFE9FF44FD1CFEFAFCC0" bold="true" box="[199,382,764,791]" pageId="5" pageNumber="106">DNA barcodes.</emphasis>
|
||
GenBank accession numbers
|
||
<accessionNumber id="946AAB02FFBBFFE9FD57FD1CFCD0FCC0" box="[724,852,764,791]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK251504" pageId="5" pageNumber="106">
|
||
<collectionCode id="ED28AE24FFBBFFE9FD57FD1CFC85FCC0" box="[724,769,764,791]" country="Kenya" name="National Museum of Kenya" pageId="5" pageNumber="106">MK</collectionCode>
|
||
251504
|
||
</accessionNumber>
|
||
(RY021,
|
||
<typeStatus id="54828843FFBBFFE9FC45FD1DFBA8FCCF" box="[966,1068,765,792]" pageId="5" pageNumber="106" type="holotype">holotype</typeStatus>
|
||
),
|
||
<accessionNumber id="946AAB02FFBBFFE9FBC3FD1CFB46FCC0" box="[1088,1218,764,791]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK251503" pageId="5" pageNumber="106">
|
||
<collectionCode id="ED28AE24FFBBFFE9FBC3FD1CFBE9FCC0" box="[1088,1133,764,791]" country="Kenya" name="National Museum of Kenya" pageId="5" pageNumber="106">MK</collectionCode>
|
||
251503
|
||
</accessionNumber>
|
||
(RY020,
|
||
<typeStatus id="54828843FFBBFFE9FAB1FD1DFA10FCC0" box="[1330,1428,765,791]" pageId="5" pageNumber="106" type="paratype">paratype</typeStatus>
|
||
) and
|
||
<accessionNumber id="946AAB02FFBBFFE9FF44FCC0FECCFCEC" box="[199,328,800,827]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK251502" pageId="5" pageNumber="106">
|
||
<collectionCode id="ED28AE24FFBBFFE9FF44FCC0FF70FCEC" box="[199,244,800,827]" country="Kenya" name="National Museum of Kenya" pageId="5" pageNumber="106">MK</collectionCode>
|
||
251502
|
||
</accessionNumber>
|
||
(RY018) (
|
||
<tableCitation id="C6BB035AFFBBFFE9FE42FCC1FD92FCEC" box="[449,534,801,827]" captionStart="TABLE 1" captionStartId="2.[151,239,298,322]" captionTargetBox="[161,1424,334,1961]" captionText="TABLE 1. GenBank accession numbers of COI sequences of specimens reported in this study." httpUri="http://table.plazi.org/id/DF466669FFBCFFEEFF14FECAFB0EFE95" pageId="5" pageNumber="106" tableUuid="DF466669FFBCFFEEFF14FECAFB0EFE95">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8636E1FFBBFFE9FF44FCA5FBD3FC88" blockId="5.[151,1437,153,2016]" box="[199,1111,837,863]" pageId="5" pageNumber="106">
|
||
<emphasis id="B94DEAF3FFBBFFE9FF44FCA5FECBFC88" bold="true" box="[199,335,837,863]" pageId="5" pageNumber="106">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="4C394D62FFBBFFE9FE4FFCA5FD9FFC88" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[460,539,837,863]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="luridus">
|
||
<emphasis id="B94DEAF3FFBBFFE9FE4FFCA5FD9FFC88" box="[460,539,837,863]" italics="true" pageId="5" pageNumber="106">luridus</emphasis>
|
||
</taxonomicName>
|
||
refers to the white, pale body color of this species.
|
||
</paragraph>
|
||
<paragraph id="8B8636E1FFBBFFE9FF44FC89FD6CF9DC" blockId="5.[151,1437,153,2016]" pageId="5" pageNumber="106">
|
||
<emphasis id="B94DEAF3FFBBFFE9FF44FC89FEBFFC54" bold="true" box="[199,315,873,899]" pageId="5" pageNumber="106">Remarks.</emphasis>
|
||
The postclitellar genital papilla arrangement of
|
||
<taxonomicName id="4C394D62FFBBFFE9FCDDFC89FBA0FC54" authority="Shen & Chang & Chih, 2019" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[862,1060,873,899]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="luridus" status="sp. nov.">
|
||
<emphasis id="B94DEAF3FFBBFFE9FCDDFC89FBA0FC54" box="[862,1060,873,899]" italics="true" pageId="5" pageNumber="106">Amynthas luridus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94DEAF3FFBBFFE9FBAFFC89FB0CFC54" bold="true" box="[1068,1160,873,899]" pageId="5" pageNumber="106">
|
||
<taxonomicNameLabel id="A27E5788FFBBFFE9FBAFFC89FB0CFC54" box="[1068,1160,873,899]" pageId="5" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
looks somewhat similar to that of
|
||
<taxonomicName id="4C394D62FFBBFFE9FE80FC6DFE1CFC70" box="[259,408,909,935]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="aquilonius">
|
||
<emphasis id="B94DEAF3FFBBFFE9FE80FC6DFE1CFC70" box="[259,408,909,935]" italics="true" pageId="5" pageNumber="106">A. aquilonius</emphasis>
|
||
</taxonomicName>
|
||
from central
|
||
<collectingCountry id="F32E7671FFBBFFE9FDACFC6DFD02FC70" box="[559,646,909,935]" name="Taiwan" pageId="5" pageNumber="106">Taiwan</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C394D62FFBBFFE9FD12FC6DFBF2FC70" authority="Tsai & Shen, 2010" authorityName="Tsai & Shen" authorityYear="2010" box="[657,1142,909,935]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="hongyehensis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FD12FC6DFC1AFC70" box="[657,926,909,935]" italics="true" pageId="5" pageNumber="106">Amynthas hongyehensis</emphasis>
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FC27FC6DFBF2FC70" author="Tsai, C. - F. & Shen, H. - P. & Tsai, S. - C." box="[932,1142,909,935]" pageId="5" pageNumber="106" pagination="1251 - 1267" refId="ref15771" refString="Tsai, C. - F., Shen, H. - P. & Tsai, S. - C. (2010) Four new species of Amynthas earthworms (Oligochaeta: Megascolecidae) from the Central Mountain Range of southern Taiwan. Journal of Natural History, 44 (21 - 24), 1251 - 1267. https: // doi. org / 10.1080 / 00222931003624788" type="journal article" year="2010">Tsai & Shen, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D62FFBBFFE9FB2FFC6DFF4BFC1C" authority="Shen, 2012" authorityName="Shen" authorityYear="2012" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="amis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FB2FFC6DFAD2FC70" box="[1196,1366,909,935]" italics="true" pageId="5" pageNumber="106">Amynthas amis</emphasis>
|
||
Shen, 2012
|
||
</taxonomicName>
|
||
from eastern
|
||
<collectingCountry id="F32E7671FFBBFFE9FEE4FC51FE3AFC1C" box="[359,446,945,971]" name="Taiwan" pageId="5" pageNumber="106">Taiwan</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C394D62FFBBFFE9FE75FC51FC4FFC1C" authority="Shen & Chih, 2016" authorityName="Shen & Chih" authorityYear="2016" box="[502,971,945,971]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="dinghuensis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FE75FC51FD74FC1C" box="[502,752,945,971]" italics="true" pageId="5" pageNumber="106">Amynthas dinghuensis</emphasis>
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FD75FC51FC4FFC1C" author="Shen, H. - P. & Chang, C. - H. & Chih, W. - J." box="[758,971,945,971]" pageId="5" pageNumber="106" pagination="1889 - 1910" refId="ref14786" refString="Shen, H. - P., Chang, C. - H. & Chih, W. - J. (2016) Four new earthworm species of the genus Amynthas (Megascolecidae: Oligochaeta) from southwestern Taiwan with re-description of Amynthas tungpuensis Tsai, Shen and Tsai, 1999. Journal of Natural History, 50 (29 - 30), 1889 - 1910. https: // doi. org / 10.1080 / 00222933.2016.1180721" type="journal article" year="2016">Shen & Chih, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
from southwestern
|
||
<collectingCountry id="F32E7671FFBBFFE9FB24FC51FB7BFC1C" box="[1191,1279,945,971]" name="Taiwan" pageId="5" pageNumber="106">Taiwan</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C394D62FFBBFFE9FA8BFC51FA18FC1C" box="[1288,1436,945,971]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="aquilonius">
|
||
<emphasis id="B94DEAF3FFBBFFE9FA8BFC51FA18FC1C" box="[1288,1436,945,971]" italics="true" pageId="5" pageNumber="106">A. aquilonius</emphasis>
|
||
</taxonomicName>
|
||
has postclitellar papillae widely paired in line with male porophore in presetal and postsetal
|
||
<collectionCode id="ED28AE24FFBBFFE9FB2AFC35FA9FFC38" box="[1193,1307,981,1007]" pageId="5" pageNumber="106">XVII–XX</collectionCode>
|
||
(
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FAA9FC35FF57FBC4" author="Tsai, C. - F. & Shen, H. - P. & Tsai, S. - C." pageId="5" pageNumber="106" pagination="276 - 288" refId="ref15584" refString="Tsai, C. - F., Shen, H. - P. & Tsai, S. - C. (2001) Some new earthworms of the genus Amynthas (Oligochaeta: Megascolecidae) from Mt. Hohuan of Taiwan. Zoological Studies, 40 (4), 276 - 288." type="journal article" year="2001">
|
||
Tsai
|
||
<emphasis id="B94DEAF3FFBBFFE9FAE7FC36FA11FC38" box="[1380,1429,981,1007]" italics="true" pageId="5" pageNumber="106">et al</emphasis>
|
||
. 2001
|
||
</bibRefCitation>
|
||
);
|
||
<taxonomicName id="4C394D62FFBBFFE9FF6BFC19FE1BFBC4" authorityName="Tsai & Shen" authorityYear="2010" box="[232,415,1017,1043]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="hongyehensis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FF6BFC19FE1BFBC4" box="[232,415,1017,1043]" italics="true" pageId="5" pageNumber="106">A. hongyehensis</emphasis>
|
||
</taxonomicName>
|
||
has postclitellar papillae widely paired in line with male porophore in presetal
|
||
<collectionCode id="ED28AE24FFBBFFE9FAA2FC19FA18FBC4" box="[1313,1436,1017,1043]" pageId="5" pageNumber="106">XVIII–XX</collectionCode>
|
||
(
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FF1CFBFDFECBFBE0" author="Tsai, C. - F. & Shen, H. - P. & Tsai, S. - C." box="[159,335,1053,1079]" pageId="5" pageNumber="106" pagination="1251 - 1267" refId="ref15771" refString="Tsai, C. - F., Shen, H. - P. & Tsai, S. - C. (2010) Four new species of Amynthas earthworms (Oligochaeta: Megascolecidae) from the Central Mountain Range of southern Taiwan. Journal of Natural History, 44 (21 - 24), 1251 - 1267. https: // doi. org / 10.1080 / 00222931003624788" type="journal article" year="2010">
|
||
Tsai
|
||
<emphasis id="B94DEAF3FFBBFFE9FF54FBFEFE83FBE0" box="[215,263,1053,1079]" italics="true" pageId="5" pageNumber="106">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
; Shen 2012);
|
||
<taxonomicName id="4C394D62FFBBFFE9FE65FBFDFDBDFBE0" authorityName="Shen" authorityYear="2012" box="[486,569,1053,1079]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="amis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FE65FBFDFDBDFBE0" box="[486,569,1053,1079]" italics="true" pageId="5" pageNumber="106">A. amis</emphasis>
|
||
</taxonomicName>
|
||
occasionally has a papilla antero-medial to each male porophore (Shen 2012);
|
||
<taxonomicName id="4C394D62FFBBFFE9FF14FBA1FEBAFB8C" authorityName="Shen & Chih" authorityYear="2016" box="[151,318,1089,1115]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="dinghuensis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FF14FBA1FEBAFB8C" box="[151,318,1089,1115]" italics="true" pageId="5" pageNumber="106">A. dinghuensis</emphasis>
|
||
</taxonomicName>
|
||
has postclitellar papillae widely paired in line with male porophore in postsetal XVII and presetal XIX (
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FF5AFB85FE1EFB57" author="Shen, H. - P. & Chang, C. - H. & Chih, W. - J." box="[217,410,1125,1152]" pageId="5" pageNumber="106" pagination="1889 - 1910" refId="ref14786" refString="Shen, H. - P., Chang, C. - H. & Chih, W. - J. (2016) Four new earthworm species of the genus Amynthas (Megascolecidae: Oligochaeta) from southwestern Taiwan with re-description of Amynthas tungpuensis Tsai, Shen and Tsai, 1999. Journal of Natural History, 50 (29 - 30), 1889 - 1910. https: // doi. org / 10.1080 / 00222933.2016.1180721" type="journal article" year="2016">
|
||
Shen
|
||
<emphasis id="B94DEAF3FFBBFFE9FE98FB86FECAFBA8" box="[283,334,1125,1151]" italics="true" pageId="5" pageNumber="106">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
). These species are octothecal with four pairs of spermathecae in VI–IX (for an exception in
|
||
<taxonomicName id="4C394D62FFBBFFE9FF34FB69FEF4FB74" authorityName="Tsai & Shen" authorityYear="2010" box="[183,368,1161,1187]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="hongyehensis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FF34FB69FEF4FB74" box="[183,368,1161,1187]" italics="true" pageId="5" pageNumber="106">A. hongyehensis</emphasis>
|
||
</taxonomicName>
|
||
see below). Their characters are compared in
|
||
<tableCitation id="C6BB035AFFBBFFE9FC0EFB69FC6CFB74" box="[909,1000,1161,1187]" captionStart="TABLE 2" captionStartId="7.[151,239,503,529]" captionTargetBox="[160,2010,577,1645]" captionText="TABLE 2. A comparison of characters among Amynthas luridus sp. nov., A. aquilonius Tsai et al., 2001, A. hongyehensis Tsai & Shen, 2010, A. amis Shen, 2012, A. dinghuensis Shen & Chih, 2016, A. oyamai (Ohfuchi, 1937), and A. morii (Kobayashi, 1938)." httpUri="http://table.plazi.org/id/DF466669FFB9FFEBFF14FE17FC73FDE2" pageId="5" pageNumber="106" tableUuid="DF466669FFB9FFEBFF14FE17FC73FDE2">Table 2</tableCitation>
|
||
. All four species are morphologically distinguishable from
|
||
<taxonomicName id="4C394D62FFBBFFE9FE0CFB4DFD86FB10" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[399,514,1197,1223]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="luridus">
|
||
<emphasis id="B94DEAF3FFBBFFE9FE0CFB4DFD86FB10" box="[399,514,1197,1223]" italics="true" pageId="5" pageNumber="106">A. luridus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C394D62FFBBFFE9FD95FB4DFD2BFB10" box="[534,687,1197,1223]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="aquilonius">
|
||
<emphasis id="B94DEAF3FFBBFFE9FD95FB4DFD2BFB10" box="[534,687,1197,1223]" italics="true" pageId="5" pageNumber="106">A. aquilonius</emphasis>
|
||
</taxonomicName>
|
||
is much smaller, and has fewer segments, lower setal number, anteriorly placed first dorsal pore in 6/7, large seminal vesicles and large prostate glands (
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FB33FB31FAE9FB3C" author="Tsai, C. - F. & Shen, H. - P. & Tsai, S. - C." box="[1200,1389,1233,1259]" pageId="5" pageNumber="106" pagination="276 - 288" refId="ref15584" refString="Tsai, C. - F., Shen, H. - P. & Tsai, S. - C. (2001) Some new earthworms of the genus Amynthas (Oligochaeta: Megascolecidae) from Mt. Hohuan of Taiwan. Zoological Studies, 40 (4), 276 - 288." type="journal article" year="2001">
|
||
Tsai
|
||
<emphasis id="B94DEAF3FFBBFFE9FB6FFB32FAA4FB3C" box="[1260,1312,1233,1259]" italics="true" pageId="5" pageNumber="106">et al</emphasis>
|
||
. 2001
|
||
</bibRefCitation>
|
||
);
|
||
<taxonomicName id="4C394D62FFBBFFE9FA07FB31FEAAFAD8" authorityName="Tsai & Shen" authorityYear="2010" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="hongyehensis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FA07FB31FEAAFAD8" italics="true" pageId="5" pageNumber="106">A. hongyehensis</emphasis>
|
||
</taxonomicName>
|
||
is much larger, and has variable numbers of spermathecae from three pairs in VII–IX to four pairs in VI–IX with normal, iridescent diverticula, and large testes, large seminal vesicles and large prostate glands (
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FAD3FAF9FF72FA80" author="Tsai, C. - F. & Shen, H. - P. & Tsai, S. - C." pageId="5" pageNumber="106" pagination="1251 - 1267" refId="ref15771" refString="Tsai, C. - F., Shen, H. - P. & Tsai, S. - C. (2010) Four new species of Amynthas earthworms (Oligochaeta: Megascolecidae) from the Central Mountain Range of southern Taiwan. Journal of Natural History, 44 (21 - 24), 1251 - 1267. https: // doi. org / 10.1080 / 00222931003624788" type="journal article" year="2010">
|
||
Tsai
|
||
<emphasis id="B94DEAF3FFBBFFE9FA0BFAFAFF29FA80" italics="true" pageId="5" pageNumber="106">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
; Shen 2012);
|
||
<taxonomicName id="4C394D62FFBBFFE9FE0FFADDFE5BFA80" authorityName="Shen" authorityYear="2012" box="[396,479,1341,1367]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="amis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FE0FFADDFE5BFA80" box="[396,479,1341,1367]" italics="true" pageId="5" pageNumber="106">A. amis</emphasis>
|
||
</taxonomicName>
|
||
has fewer setae, preclitellar genital papillae but no postclitellar papillae, spermathecae with normal, iridescent diverticula, and large testes, large seminal vesicles and large prostate glands (Shen 2012);
|
||
<taxonomicName id="4C394D62FFBBFFE9FF14FA65FEBAFA48" authorityName="Shen & Chih" authorityYear="2016" box="[151,318,1413,1439]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="dinghuensis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FF14FA65FEBAFA48" box="[151,318,1413,1439]" italics="true" pageId="5" pageNumber="106">A. dinghuensis</emphasis>
|
||
</taxonomicName>
|
||
is much smaller, and has fewer setae, spermathecae with normal, iridescent diverticula, and large testes, large seminal vesicles and large prostate glands (
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FC98FA49FC5BFA13" author="Shen, H. - P. & Chang, C. - H. & Chih, W. - J." box="[795,991,1449,1476]" pageId="5" pageNumber="106" pagination="1889 - 1910" refId="ref14786" refString="Shen, H. - P., Chang, C. - H. & Chih, W. - J. (2016) Four new earthworm species of the genus Amynthas (Megascolecidae: Oligochaeta) from southwestern Taiwan with re-description of Amynthas tungpuensis Tsai, Shen and Tsai, 1999. Journal of Natural History, 50 (29 - 30), 1889 - 1910. https: // doi. org / 10.1080 / 00222933.2016.1180721" type="journal article" year="2016">
|
||
Shen
|
||
<emphasis id="B94DEAF3FFBBFFE9FCDDFA4AFC16FA14" box="[862,914,1449,1475]" italics="true" pageId="5" pageNumber="106">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
). All these species are also genetically distinct (
|
||
<figureCitation id="13022A64FFBBFFE9FF79FA2DFEC4FA30" box="[250,320,1485,1511]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1823]" captionTargetBox="[273,1306,173,1771]" captionTargetId="figure@6.[268,1319,166,1779]" captionTargetPageId="6" captionText="FIGURE 2. Neighbor-joining tree of Amynthas luridus and representatives of Amynthas aquilonius Tsai et al., 2001, Amynthas catenus Tsai et al., 2001, Amynthas proasacceus Tsai et al., 2001, Amynthas hohuanmontis Tsai et al., 2002, Amynthas tessellatus Shen et al., 2002, Amynthas fenestrus Shen et al., 2003, Amynthas tantulus Shen et al., 2003, Amynthas uvaglandularis Shen et al., 2003, Amynthas hongyehensis Tsai & Shen, 2010, Amynthas amis Shen, 2012, and Amynthas dinghuensis Shen & Chih, 2016 from Taiwan. The tree is based on COI gene and Kimura’s two-parameter model, and rooted using Amynthas corticis. Specimens from this study and sequences retrieved from GenBank are shown by their voucher numbers and GenBank accession numbers, respectively. Type materials are indicated in parentheses. Numbers around nodes are bootstrap values above 50." httpUri="https://zenodo.org/record/3373133/files/figure.png" pageId="5" pageNumber="106">Fig. 2</figureCitation>
|
||
). Additionally, the GenBank accession number of the
|
||
<typeStatus id="54828843FFBBFFE9FC2BFA2DFB8EFA30" box="[936,1034,1485,1511]" pageId="5" pageNumber="106" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C394D62FFBBFFE9FBB1FA2DFB03FA30" authorityName="Shen" authorityYear="2012" box="[1074,1159,1485,1511]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="amis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FBB1FA2DFB03FA30" box="[1074,1159,1485,1511]" italics="true" pageId="5" pageNumber="106">A. amis</emphasis>
|
||
</taxonomicName>
|
||
reported in Shen (2012) was erroneous and should be corrected to
|
||
<accessionNumber id="946AAB02FFBBFFE9FDEDFA11FD61F9DC" box="[622,741,1521,1547]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JX290409" pageId="5" pageNumber="106">
|
||
JX
|
||
<date id="FF871021FFBBFFE9FD0DFA11FD61F9DC" box="[654,741,1521,1547]" pageId="5" pageNumber="106" value="09-04-29">290409</date>
|
||
</accessionNumber>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8636E1FFBBFFE5FF44F9F5FDF8FE94" blockId="5.[151,1437,153,2016]" lastBlockId="9.[151,1436,153,323]" lastPageId="9" lastPageNumber="110" pageId="5" pageNumber="106">
|
||
The genital papilla arrangement in the male pore region of
|
||
<taxonomicName id="4C394D62FFBBFFE9FCD5F9F5FC47F9F8" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[854,963,1557,1583]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="luridus">
|
||
<emphasis id="B94DEAF3FFBBFFE9FCD5F9F5FC47F9F8" box="[854,963,1557,1583]" italics="true" pageId="5" pageNumber="106">A. luridus</emphasis>
|
||
</taxonomicName>
|
||
is also similar to that of
|
||
<taxonomicName id="4C394D62FFBBFFE9FB56F9F5FECEF984" authority="(Ohfuchi, 1937)" authorityName="Ohfuchi" authorityYear="1937" baseAuthorityName="Ohfuchi" baseAuthorityYear="1937" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="oyamai">
|
||
<emphasis id="B94DEAF3FFBBFFE9FB56F9F5FA18F9F8" box="[1237,1436,1557,1583]" italics="true" pageId="5" pageNumber="106">Amynthas oyamai</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FF1CF9D9FEC6F984" author="Ohfuchi, S." box="[159,322,1593,1619]" pageId="5" pageNumber="106" pagination="31 - 136" refId="ref14453" refString="Ohfuchi, S. (1937) On the species possessing four pairs of spermathecae in the genus Pheretima, together with the variability of some external and internal characteristics. Saito Ho-on Kai Museum Research Bulletin, 12, 31 - 136." type="journal article" year="1937">Ohfuchi, 1937</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from northern
|
||
<collectingCountry id="F32E7671FFBBFFE9FE72F9D9FDB0F984" box="[497,564,1593,1619]" name="Japan" pageId="5" pageNumber="106">Japan</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C394D62FFBBFFE9FDBDF9D9FC40F984" authority="(Kobayashi, 1938)" authorityName="Kobayashi" authorityYear="1938" baseAuthorityName="Kobayashi" baseAuthorityYear="1938" box="[574,964,1593,1619]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="morii">
|
||
<emphasis id="B94DEAF3FFBBFFE9FDBDF9D9FD6BF984" box="[574,751,1593,1619]" italics="true" pageId="5" pageNumber="106">Amynthas morii</emphasis>
|
||
(Kobayashi, 1938)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F32E7671FFBBFFE9FB85F9D9FBC9F984" box="[1030,1101,1593,1619]" name="South Korea" pageId="5" pageNumber="106">Korea</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C394D62FFBBFFE9FBD4F9D9FA18F984" authority="(Thai, 1984)" authorityName="Thai" authorityYear="1984" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[1111,1436,1593,1619]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="dangi">
|
||
<emphasis id="B94DEAF3FFBBFFE9FBD4F9D9FA88F984" box="[1111,1292,1593,1619]" italics="true" pageId="5" pageNumber="106">Amynthas dangi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FA9AF9D9FA10F984" author="Thai, T. B." box="[1305,1428,1593,1619]" pageId="5" pageNumber="106" pagination="1317 - 1328" refId="ref15444" refString="Thai, T. B. (1984) New species of the genus Pheretima in Vietnam. Zoologicheskii Jurnal, 63 (9), 1317 - 1328. [in Vietnamese with English abstract]" type="journal article" year="1984">Thai, 1984</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F32E7671FFBBFFE9FF55F9BDFEBCF9A0" box="[214,312,1629,1655]" name="Vietnam" pageId="5" pageNumber="106">Vietnam</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C394D62FFBBFFE9FEF5F9BDFDEEF9A0" authorityName="Hong, Inkhavilay & James" authorityYear="2018" box="[374,618,1629,1655]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="nametensis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FEF5F9BDFDEEF9A0" box="[374,618,1629,1655]" italics="true" pageId="5" pageNumber="106">Amynthas nametensis</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FDF0F9BDFCB9F9A0" author="Hong, Y. & Inkhavilay, K. & James, S. W." box="[627,829,1629,1655]" pageId="5" pageNumber="106" pagination="238 - 250" refId="ref13618" refString="Hong, Y., Inkhavilay, K. & James, S. W. (2018) Nine new species of the " aeruginosus " - group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National Protected Area, Laos. Zootaxa, 4496 (1), 238 - 250. https: // doi. org / 10.11646 / zootaxa. 4496.1.19" type="journal article" year="2018">
|
||
Hong
|
||
<emphasis id="B94DEAF3FFBBFFE9FD38F9BEFD6AF9A0" box="[699,750,1629,1655]" italics="true" pageId="5" pageNumber="106">et al</emphasis>
|
||
., 2018
|
||
</bibRefCitation>
|
||
from
|
||
<collectingCountry id="F32E7671FFBBFFE9FC05F9BDFC44F9A0" box="[902,960,1629,1655]" name="Laos" pageId="5" pageNumber="106">Laos</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C394D62FFBBFFE9FC4FF9BDFBBBF9A0" baseAuthorityName="Ohfuchi" baseAuthorityYear="1937" box="[972,1087,1629,1655]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="oyamai">
|
||
<emphasis id="B94DEAF3FFBBFFE9FC4FF9BDFBBBF9A0" box="[972,1087,1629,1655]" italics="true" pageId="5" pageNumber="106">A. oyamai</emphasis>
|
||
</taxonomicName>
|
||
has a papilla antero-medial to each male porophore (
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FE0CF961FDABF94C" author="Ohfuchi, S." box="[399,559,1665,1691]" pageId="5" pageNumber="106" pagination="31 - 136" refId="ref14453" refString="Ohfuchi, S. (1937) On the species possessing four pairs of spermathecae in the genus Pheretima, together with the variability of some external and internal characteristics. Saito Ho-on Kai Museum Research Bulletin, 12, 31 - 136." type="journal article" year="1937">Ohfuchi 1937</bibRefCitation>
|
||
);
|
||
<taxonomicName id="4C394D62FFBBFFE9FDC1F961FD1AF94C" baseAuthorityName="Kobayashi" baseAuthorityYear="1938" box="[578,670,1665,1691]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="morii">
|
||
<emphasis id="B94DEAF3FFBBFFE9FDC1F961FD1AF94C" box="[578,670,1665,1691]" italics="true" pageId="5" pageNumber="106">A. morii</emphasis>
|
||
</taxonomicName>
|
||
has a papilla anterior to each male porophore (
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FB24F961FAF5F94B" author="Kobayashi, S." box="[1191,1393,1665,1692]" pageId="5" pageNumber="106" pagination="89 - 170" refId="ref13903" refString="Kobayashi, S. (1938 b) Earthworms of Korea. I. Science Reports of the Tohoku Imperial University, 13, 89 - 170." type="journal article" year="1938">Kobayashi 1938b</bibRefCitation>
|
||
);
|
||
<taxonomicName id="4C394D62FFBBFFE9FA07F961FF53F968" baseAuthorityName="Thai" baseAuthorityYear="1984" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="dangi">
|
||
<emphasis id="B94DEAF3FFBBFFE9FA07F961FF53F968" italics="true" pageId="5" pageNumber="106">A. dangi</emphasis>
|
||
</taxonomicName>
|
||
has papillae widely paired in line with male porophore in presetal
|
||
<collectionCode id="ED28AE24FFBBFFE9FC52F945FBD1F968" box="[977,1109,1701,1727]" pageId="5" pageNumber="106">XVIII–XIX</collectionCode>
|
||
and a papilla medial to each male pore (
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FE91F929FE0CF934" author="Thai, T. B." box="[274,392,1737,1763]" pageId="5" pageNumber="106" pagination="1317 - 1328" refId="ref15444" refString="Thai, T. B. (1984) New species of the genus Pheretima in Vietnam. Zoologicheskii Jurnal, 63 (9), 1317 - 1328. [in Vietnamese with English abstract]" type="journal article" year="1984">Thai 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FE11F929FDF3F933" author="Nguyen, T. T. & Lam, D. H. & Nguyen, A. D." box="[402,631,1737,1764]" pageId="5" pageNumber="106" pagination="52" refId="ref14174" refString="Nguyen, T. T., Lam, D. H. & Nguyen, A. D. (2016 a) On the giant pheretimoid earthworms from Vietnam (Clitellata: Megascolecidae), with descriptions of three new species. Zoological Studies, 55, 52. https: // doi. org / 10.6620 / ZS. 2016.55 - 52" type="journal article" year="2016">
|
||
Nguyen
|
||
<emphasis id="B94DEAF3FFBBFFE9FE72F92AFDA5F934" box="[497,545,1737,1763]" italics="true" pageId="5" pageNumber="106">et al</emphasis>
|
||
. 2016a
|
||
</bibRefCitation>
|
||
);
|
||
<taxonomicName id="4C394D62FFBBFFE9FD0BF929FCA6F934" authorityName="Hong, Inkhavilay & James" authorityYear="2018" box="[648,802,1737,1763]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="nametensis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FD0BF929FCA6F934" box="[648,802,1737,1763]" italics="true" pageId="5" pageNumber="106">A. nametensis</emphasis>
|
||
</taxonomicName>
|
||
has papillae widely paired in line with male porophore in presetal
|
||
<collectionCode id="ED28AE24FFBBFFE9FF76F90DFEFDF8D0" box="[245,377,1773,1799]" pageId="5" pageNumber="106">XVIII–XIX</collectionCode>
|
||
(
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FE0BF90DFDCEF8D0" author="Hong, Y. & Inkhavilay, K. & James, S. W." box="[392,586,1773,1799]" pageId="5" pageNumber="106" pagination="238 - 250" refId="ref13618" refString="Hong, Y., Inkhavilay, K. & James, S. W. (2018) Nine new species of the " aeruginosus " - group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National Protected Area, Laos. Zootaxa, 4496 (1), 238 - 250. https: // doi. org / 10.11646 / zootaxa. 4496.1.19" type="journal article" year="2018">
|
||
Hong
|
||
<emphasis id="B94DEAF3FFBBFFE9FE4CF90EFD84F8D0" box="[463,512,1773,1799]" italics="true" pageId="5" pageNumber="106">et al</emphasis>
|
||
. 2018
|
||
</bibRefCitation>
|
||
). Like
|
||
<taxonomicName id="4C394D62FFBBFFE9FD14F90DFC81F8D0" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[663,773,1773,1799]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="luridus">
|
||
<emphasis id="B94DEAF3FFBBFFE9FD14F90DFC81F8D0" box="[663,773,1773,1799]" italics="true" pageId="5" pageNumber="106">A. luridus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C394D62FFBBFFE9FC90F90DFC00F8D0" baseAuthorityName="Ohfuchi" baseAuthorityYear="1937" box="[787,900,1773,1799]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="oyamai">
|
||
<emphasis id="B94DEAF3FFBBFFE9FC90F90DFC00F8D0" box="[787,900,1773,1799]" italics="true" pageId="5" pageNumber="106">A. oyamai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394D62FFBBFFE9FC38F90DFB93F8D0" baseAuthorityName="Kobayashi" baseAuthorityYear="1938" box="[955,1047,1773,1799]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="morii">
|
||
<emphasis id="B94DEAF3FFBBFFE9FC38F90DFB93F8D0" box="[955,1047,1773,1799]" italics="true" pageId="5" pageNumber="106">A. morii</emphasis>
|
||
</taxonomicName>
|
||
are also octothecal and have small or little developed seminal vesicles and small, short prostatic duct (
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FC08F8F1FBA8F8FC" author="Ohfuchi, S." box="[907,1068,1809,1835]" pageId="5" pageNumber="106" pagination="31 - 136" refId="ref14453" refString="Ohfuchi, S. (1937) On the species possessing four pairs of spermathecae in the genus Pheretima, together with the variability of some external and internal characteristics. Saito Ho-on Kai Museum Research Bulletin, 12, 31 - 136." type="journal article" year="1937">Ohfuchi 1937</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FBBBF8F1FB7AF8FC" author="Kobayashi, S." box="[1080,1278,1809,1835]" pageId="5" pageNumber="106" pagination="405 - 417" refId="ref13879" refString="Kobayashi, S. (1938 a) Earthworms from Hakodate, Hokkaido. Annotationes Zoologicae Japonenses, 17, 405 - 417." type="journal article" year="1938">Kobayashi 1938a</bibRefCitation>
|
||
, b). However, their characters of larger body size, lower setal numbers, absence of postclitellar genital papillae in XVII and XIX, and absent or poorly developed prostate glands are different from those of
|
||
<taxonomicName id="4C394D62FFBBFFE9FC66F8B9FBD7F8A4" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[997,1107,1881,1907]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="luridus">
|
||
<emphasis id="B94DEAF3FFBBFFE9FC66F8B9FBD7F8A4" box="[997,1107,1881,1907]" italics="true" pageId="5" pageNumber="106">A. luridus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C6BB035AFFBBFFE9FBE0F8B9FB3FF8A4" box="[1123,1211,1881,1907]" captionStart="TABLE 2" captionStartId="7.[151,239,503,529]" captionTargetBox="[160,2010,577,1645]" captionText="TABLE 2. A comparison of characters among Amynthas luridus sp. nov., A. aquilonius Tsai et al., 2001, A. hongyehensis Tsai & Shen, 2010, A. amis Shen, 2012, A. dinghuensis Shen & Chih, 2016, A. oyamai (Ohfuchi, 1937), and A. morii (Kobayashi, 1938)." httpUri="http://table.plazi.org/id/DF466669FFB9FFEBFF14FE17FC73FDE2" pageId="5" pageNumber="106" tableUuid="DF466669FFB9FFEBFF14FE17FC73FDE2">Table 2</tableCitation>
|
||
). As to
|
||
<taxonomicName id="4C394D62FFBBFFE9FA90F8B9FAF7F8A4" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[1299,1395,1881,1907]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="dangi">
|
||
<emphasis id="B94DEAF3FFBBFFE9FA90F8B9FAF7F8A4" box="[1299,1395,1881,1907]" italics="true" pageId="5" pageNumber="106">A. dangi</emphasis>
|
||
</taxonomicName>
|
||
, its characters of large body size (length>
|
||
<quantity id="4CC19B04FFBBFFE9FDCFF89DFD29F840" box="[588,685,1917,1943]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="106" unit="mm" value="300.0">300 mm</quantity>
|
||
), much higher setal numbers (59 setae in segment II and more than 100 setae per segment after segment VIII) and two pairs of spermathecal pores in 7/8/9 (
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FBFCF841FB73F86C" author="Thai, T. B." box="[1151,1271,1953,1979]" pageId="5" pageNumber="106" pagination="1317 - 1328" refId="ref15444" refString="Thai, T. B. (1984) New species of the genus Pheretima in Vietnam. Zoologicheskii Jurnal, 63 (9), 1317 - 1328. [in Vietnamese with English abstract]" type="journal article" year="1984">Thai 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA84B10FFBBFFE9FA80F841FF59F837" author="Nguyen, T. T. & Lam, D. H. & Nguyen, A. D." pageId="5" pageNumber="106" pagination="52" refId="ref14174" refString="Nguyen, T. T., Lam, D. H. & Nguyen, A. D. (2016 a) On the giant pheretimoid earthworms from Vietnam (Clitellata: Megascolecidae), with descriptions of three new species. Zoological Studies, 55, 52. https: // doi. org / 10.6620 / ZS. 2016.55 - 52" type="journal article" year="2016">
|
||
Nguyen
|
||
<emphasis id="B94DEAF3FFBBFFE9FAE7F842FA11F86C" box="[1380,1429,1953,1979]" italics="true" pageId="5" pageNumber="106">et al</emphasis>
|
||
. 2016a
|
||
</bibRefCitation>
|
||
) clearly separate it from
|
||
<taxonomicName id="4C394D62FFBBFFE9FE78F825FDEEF808" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[507,618,1989,2015]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="luridus">
|
||
<emphasis id="B94DEAF3FFBBFFE9FE78F825FDEEF808" box="[507,618,1989,2015]" italics="true" pageId="5" pageNumber="106">A. luridus</emphasis>
|
||
</taxonomicName>
|
||
. The body size and setal numbers of
|
||
<taxonomicName id="4C394D62FFBBFFE9FB97F825FB34F808" authorityName="Hong, Inkhavilay & James" authorityYear="2018" box="[1044,1200,1989,2015]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="106" phylum="Annelida" rank="species" species="nametensis">
|
||
<emphasis id="B94DEAF3FFBBFFE9FB97F825FB34F808" box="[1044,1200,1989,2015]" italics="true" pageId="5" pageNumber="106">A. nametensis</emphasis>
|
||
</taxonomicName>
|
||
are close to those of
|
||
<taxonomicName id="4C394D62FFB7FFE5FF14FF79FE81FF64" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[151,261,153,179]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="110" phylum="Annelida" rank="species" species="luridus">
|
||
<emphasis id="B94DEAF3FFB7FFE5FF14FF79FE81FF64" box="[151,261,153,179]" italics="true" pageId="9" pageNumber="110">A. luridus</emphasis>
|
||
</taxonomicName>
|
||
, but
|
||
<taxonomicName id="4C394D62FFB7FFE5FEBCFF79FE5EFF64" authorityName="Hong, Inkhavilay & James" authorityYear="2018" box="[319,474,153,179]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="110" phylum="Annelida" rank="species" species="nametensis">
|
||
<emphasis id="B94DEAF3FFB7FFE5FEBCFF79FE5EFF64" box="[319,474,153,179]" italics="true" pageId="9" pageNumber="110">A. nametensis</emphasis>
|
||
</taxonomicName>
|
||
is quadrithecal with two pairs of spermathecal pores in 7/8/9. Furthermore,
|
||
<taxonomicName id="4C394D62FFB7FFE5FAACFF79FF4BFF00" authorityName="Hong, Inkhavilay & James" authorityYear="2018" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="110" phylum="Annelida" rank="species" species="nametensis">
|
||
<emphasis id="B94DEAF3FFB7FFE5FAACFF79FF4BFF00" italics="true" pageId="9" pageNumber="110">A. nametensis</emphasis>
|
||
</taxonomicName>
|
||
has globular ampulla, diverticulum coiled into irregular ovate block, and prostate glands extending from XVII to XXIII with long, stout, coiled ducts (
|
||
<bibRefCitation id="EFA84B10FFB7FFE5FDD4FF01FC9FFF2C" author="Hong, Y. & Inkhavilay, K. & James, S. W." box="[599,795,225,251]" pageId="9" pageNumber="110" pagination="238 - 250" refId="ref13618" refString="Hong, Y., Inkhavilay, K. & James, S. W. (2018) Nine new species of the " aeruginosus " - group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National Protected Area, Laos. Zootaxa, 4496 (1), 238 - 250. https: // doi. org / 10.11646 / zootaxa. 4496.1.19" type="journal article" year="2018">
|
||
Hong
|
||
<emphasis id="B94DEAF3FFB7FFE5FD1CFF02FD55FF2C" box="[671,721,225,251]" italics="true" pageId="9" pageNumber="110">et al</emphasis>
|
||
. 2018
|
||
</bibRefCitation>
|
||
), whereas
|
||
<taxonomicName id="4C394D62FFB7FFE5FC10FF01FB86FF2C" authorityName="Shen & Chang & Chih" authorityYear="2019" box="[915,1026,225,251]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="110" phylum="Annelida" rank="species" species="luridus">
|
||
<emphasis id="B94DEAF3FFB7FFE5FC10FF01FB86FF2C" box="[915,1026,225,251]" italics="true" pageId="9" pageNumber="110">A. luridus</emphasis>
|
||
</taxonomicName>
|
||
has elongated oval-shaped ampulla, diverticulum with a slender stalk and a seminal chamber rudimentary or absent, and prostate glands in
|
||
<collectionCode id="ED28AE24FFB7FFE5FAA2FEE5FA18FEC8" box="[1313,1436,261,287]" pageId="9" pageNumber="110">XVII–XIX</collectionCode>
|
||
with short, small, C-shaped prostatic ducts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |