treatments-xml/data/03/90/87/039087DEFFA76D5DBC069983662EA8B5.xml

<document ID-DOI="10.5733/afin.049.0210" ID-ISSN="2305-2562" ID-Zenodo-Dep="7917381" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683578209636" checkinUser="felipe" docAuthor="Londt, Jason G. H." docDate="2008" docId="039087DEFFA76D5DBC069983662EA8B5" docLanguage="en" docName="AfrInvertebr.49.2.171-226.pdf" docOrigin="African Invertebrates 49 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.049.0210" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Trichardis effrena Londt 2008, sp. n." docType="treatment" docVersion="3" lastPageNumber="183" masterDocId="FFA9FFA6FFAC6D51BE01987D6467AB6A" masterDocTitle="A review of Afrotropical Trichardis Hermann, 1906, and the description of the first Oriental representative of the genus (Diptera: Asilidae: Laphriinae)" masterLastPageNumber="171" masterPageNumber="171" pageNumber="182" updateTime="1683695595129" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>A review of Afrotropical Trichardis Hermann, 1906, and the description of the first Oriental representative of the genus (Diptera: Asilidae: Laphriinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Londt, Jason G. H.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
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<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-12-31</mods:number>
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<mods:detail type="volume">
<mods:number>49</mods:number>
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<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>171</mods:start>
<mods:end>171</mods:end>
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<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.049.0210</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.049.0210</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7917381</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:039087DEFFA76D5DBC069983662EA8B5" httpUri="http://treatment.plazi.org/id/039087DEFFA76D5DBC069983662EA8B5" lastPageId="12" lastPageNumber="183" pageId="11" pageNumber="182">
<subSubSection box="[519,804,510,535]" pageId="11" pageNumber="182" type="nomenclature">
<paragraph blockId="11.[181,1142,510,1673]" box="[519,804,510,535]" pageId="11" pageNumber="182">
<heading box="[519,804,510,535]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="182" reason="2">
<taxonomicName authority="Londt, 2008" authorityName="Londt" authorityYear="2008" box="[519,737,510,535]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="182" phylum="Arthropoda" rank="species" species="effrena" status="sp. n.">
<emphasis bold="true" box="[519,737,510,535]" pageId="11" pageNumber="182">Trichardis effrena</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[744,804,510,535]" pageId="11" pageNumber="182" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="183" pageId="11" pageNumber="182" type="description">
<paragraph blockId="11.[181,1142,510,1673]" box="[564,759,550,575]" pageId="11" pageNumber="182">
<figureCitation box="[564,635,550,575]" captionStart="Figs 1– 4" captionStartId="3.[182,221,1623,1643]" captionTargetBox="[181,1141,985,1604]" captionTargetId="figure-247@3.[181,1141,985,1604]" captionTargetPageId="3" captionText="Figs 1– 4. Trichardis species: (1–3) T. testacea (Macquart, 1838): (1) dorsal view of entire male, (2) lateral view of entire male, (3) wing; (4) T. effrena sp. n., wing. See text for measurements." figureDoi="http://doi.org/10.5281/zenodo.7917383" httpUri="https://zenodo.org/record/7917383/files/figure.png" pageId="11" pageNumber="182">Figs 4</figureCitation>
, 
<figureCitation box="[647,719,550,575]" captionStart="Figs 11–16" captionStartId="46.[144,184,1475,1495]" captionTargetBox="[167,1104,307,1448]" captionTargetId="figure-9@46.[167,1104,307,1448]" captionTargetPageId="46" captionText="Figs 11–16. Trichardis species, male genitalia: (11, 12) lateral and ventral of T. cribrata (Loew, 1858), Mhlopeni Nat. Res.; (13, 14) lateral and ventral of T. eburacta sp. n., paratype, Comoé National Park; (15, 16) lateral and ventral of T. effrena sp. n., paratype, Witsand Nat. Res. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7917387" httpUri="https://zenodo.org/record/7917387/files/figure.png" pageId="11" pageNumber="182">15, 16</figureCitation>
, 
<figureCitation box="[731,759,550,575]" captionStart="Fig 59" captionStartId="55.[183,213,872,892]" captionTargetBox="[287,1039,203,852]" captionTargetId="figure-10@55.[285,1041,203,852]" captionTargetPageId="55" captionText="Fig 59. Distributions of southern African Trichardis species: – T. cribrata (Loew, 1858), 9 – T. effrena sp. n., 1974, • – T. picta Hermann, 1906." figureDoi="http://doi.org/10.5281/zenodo.7917407" httpUri="https://zenodo.org/record/7917407/files/figure.png" pageId="11" pageNumber="182">59</figureCitation>
</paragraph>
<paragraph blockId="11.[181,1142,510,1673]" pageId="11" pageNumber="182">
Etymology: From Latin 
<taxonomicName authorityName="Londt" authorityYear="2008" box="[453,531,590,614]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="182" phylum="Arthropoda" rank="species" species="effrena">
<emphasis box="[453,531,590,614]" italics="true" pageId="11" pageNumber="182">effrena</emphasis>
</taxonomicName>
(unrestrained). Refers to the absence of the costal vein along the posterior margin of the wing.
</paragraph>
<paragraph blockId="11.[181,1142,510,1673]" box="[181,800,662,687]" pageId="11" pageNumber="182">
Description (based on 
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in excellent condition):
</paragraph>
<paragraph blockId="11.[181,1142,510,1673]" pageId="11" pageNumber="182">
<emphasis box="[181,242,702,726]" italics="true" pageId="11" pageNumber="182">Head</emphasis>
: Brown-orange, entirely fine silver pruinose. Antenna brown-orange except for distal part of postpedicel and style which are red-brown, setae orange; postpedicel clavate (L:D=2.4:1). Mystax uniformly shiny orange. Ocellar tubercle with 4 macrosetae. Proboscis proximally orange-brown distally dark red-brown, palpi orange-brown.
</paragraph>
<paragraph blockId="11.[181,1142,510,1673]" pageId="11" pageNumber="182">
<emphasis box="[181,262,838,862]" italics="true" pageId="11" pageNumber="182">Thorax</emphasis>
: Brown-orange with some red-brown areas, extensively fine silver pruinose. Postpronotum medially pruinose, laterally apruinose; mesonotum brown-orange with red-brown dorsal stripe and laterally situated broad bands, apruinose except for narrow lateral and posterior margins. Scutellum entirely apruinose.Anepisternum with orange posterior macroseta, extensively pruinose except for small anteroventral area. Proepimeron pruinose; katepisternum red-brown pruinose posteriorly, apruinose anteriorly; anepisternum entirely pruinose. Legs: Brown-orange (femora, tarsomere 5 and hind tibiae darker), pulvilli and empodium of similar length. Hind femur orange-brown, length:height ratio 3.4:1, ventral tubercles poorly developed. Hind tibia lacking ventrodistal spur. Wing: 4.6× 
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. Costal vein strongly developed only as far as wing tip, very weak or absent along entire posterior margin of wing. Membrane not extensively microtrichose—discal and r 
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cell almost entirely lacking microtrichiae.
</paragraph>
<paragraph blockId="11.[181,1142,510,1673]" pageId="11" pageNumber="182">
<emphasis box="[181,285,1230,1254]" italics="true" pageId="11" pageNumber="182">Abdomen</emphasis>
: Brown-orange, macrosetae orange, fine setulae pale yellow.T2 brown-orange, apruinose except for strong silver pruinose spot posterolaterally.
</paragraph>
<paragraph blockId="11.[181,1142,510,1673]" pageId="11" pageNumber="182">
<emphasis box="[208,225,1295,1319]" italics="true" pageId="11" pageNumber="182">ơ</emphasis>
genitalia (
<figureCitation box="[336,459,1294,1319]" captionStart="Figs 11–16" captionStartId="46.[144,184,1475,1495]" captionTargetBox="[167,1104,307,1448]" captionTargetId="figure-9@46.[167,1104,307,1448]" captionTargetPageId="46" captionText="Figs 11–16. Trichardis species, male genitalia: (11, 12) lateral and ventral of T. cribrata (Loew, 1858), Mhlopeni Nat. Res.; (13, 14) lateral and ventral of T. eburacta sp. n., paratype, Comoé National Park; (15, 16) lateral and ventral of T. effrena sp. n., paratype, Witsand Nat. Res. Scale lines = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7917387" httpUri="https://zenodo.org/record/7917387/files/figure.png" pageId="11" pageNumber="182">Figs 15, 16</figureCitation>
): Epandrium in lateral view longer than basal part of gonocoxite (i.e. excluding distal projection of gonocoxite and gonostylus). Proctiger moderately dorsoventrally compressed. Hypandrium greatly reduced and simple. Gonocoxite in ventral view without median projections distally and with a distal row of about 7 macrosetae; mediodistal projection stout, with strongly upturned and scerotised distal end. Gonostylus fairly stout, with broadly rounded, fairly straight distal end. Aedeagal prongs small, more or less straight and with a small terminal end.
</paragraph>
<paragraph blockId="11.[181,1142,510,1673]" pageId="11" pageNumber="182">
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: 
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: 
<emphasis box="[446,460,1525,1544]" italics="true" pageId="11" pageNumber="182">ơ</emphasis>
‘ 
<collectingMunicipality box="[467,587,1524,1544]" pageId="11" pageNumber="182">South Africa</collectingMunicipality>
: N 
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/ 
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: 
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/ 
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&amp; 
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/ 
<date box="[462,548,1548,1568]" pageId="11" pageNumber="182" value="2004-01-31">
<collectingDate box="[462,548,1548,1568]" pageId="11" pageNumber="182" value="2004-01-31">31.i.2004</collectingDate>
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<emphasis box="[553,616,1549,1569]" italics="true" pageId="11" pageNumber="182">Acacia</emphasis>
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mixed / woodland. 
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area’ (
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).
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: 
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: 
<emphasis box="[392,430,1580,1600]" italics="true" pageId="11" pageNumber="182">1^‘</emphasis>
Brit. S. W. – Africa / Kalahari / L. Schultze S.’, ‘ 
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/ 
<taxonomicName authorityName=": Engel" authorityYear="1924" box="[1009,1083,1581,1601]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="182" phylum="Arthropoda" rank="species" species="cribrata">
<emphasis box="[1009,1083,1581,1601]" italics="true" pageId="11" pageNumber="182">cribrata</emphasis>
</taxonomicName>
/ Lw / Kalahari / ı 968. a. / Det Dr. F. Hermann [sideways]’ (ZMHB). 
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: 2 
<emphasis box="[952,1011,1604,1624]" italics="true" pageId="11" pageNumber="182">ơ 2^‘</emphasis>
S:Africa: NW Province / Molopo Game Reserve / Phiri Camp area / 
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: 
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/ 
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<date box="[958,1056,1628,1648]" pageId="11" pageNumber="182" value="2003-03-14">14.iii.2003</date>
J Londt / 
<taxonomicName authority="Erogrostis" authorityName="Erogrostis" box="[181,345,1653,1673]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="11" pageNumber="182" phylum="Tracheophyta" rank="genus">
<emphasis box="[181,345,1653,1673]" italics="true" pageId="11" pageNumber="182">Acacia Erogrostis</emphasis>
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savannah’; 
<emphasis box="[455,494,1652,1672]" italics="true" pageId="11" pageNumber="182">1^‘</emphasis>
S Africa: N 
<collectingRegion box="[607,654,1652,1672]" country="South Africa" name="Western Cape" pageId="11" pageNumber="182">Cape</collectingRegion>
#15 / 
<quantity box="[713,771,1652,1672]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="182" unit="km" value="14.0">14 km</quantity>
S of Hotazel / 
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: 
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<quantity box="[1060,1129,1652,1672]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="11" pageNumber="182" unit="m" value="1050.0">1050 m</quantity>
/ Date: 
<date box="[201,299,204,224]" pageId="12" pageNumber="183" value="1991-03-14">14.iii.1991</date>
/ Londt &amp; Whittington / Ga-Mogara River bed’; 
<emphasis box="[766,804,204,224]" italics="true" pageId="12" pageNumber="183">1^‘</emphasis>
South Africa: N 
<collectingRegion box="[963,1010,204,224]" country="South Africa" name="Western Cape" pageId="12" pageNumber="183">Cape</collectingRegion>
/ Vaalbos National Park / Riverside Picnic site 
<quantity box="[479,548,228,248]" metricMagnitude="3" metricUnit="m" metricValue="1.055" pageId="12" pageNumber="183" unit="m" value="1055.0">1055 m</quantity>
/ 
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<geoCoordinate box="[681,805,226,249]" degrees="024" direction="east" minutes="19.994" orientation="longitude" pageId="12" pageNumber="183" precision="1" value="24.333233">024°19.994'E</geoCoordinate>
/ 
<date box="[823,942,228,248]" pageId="12" pageNumber="183" value="2004-01-28" valueMax="2004-01-29" valueMin="2004-01-28">28–29.i.2004</date>
JGH Londt &amp; / T Dikow 
<taxonomicName box="[209,272,253,273]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="183" phylum="Tracheophyta" rank="genus">
<emphasis box="[209,272,253,273]" italics="true" pageId="12" pageNumber="183">Acacia</emphasis>
</taxonomicName>
savannah’; 1 
<emphasis box="[389,403,253,272]" italics="true" pageId="12" pageNumber="183">ơ</emphasis>
‘S Africa: N 
<collectingRegion box="[519,565,252,272]" country="South Africa" name="Western Cape" pageId="12" pageNumber="183">Cape</collectingRegion>
/ Witsand Farm, 
<geoCoordinate box="[715,785,250,273]" degrees="28" direction="south" minutes="32" orientation="latitude" pageId="12" pageNumber="183" precision="925" value="-28.533333">28°32'S</geoCoordinate>
/ 
<geoCoordinate box="[796,871,250,273]" degrees="22" direction="east" minutes="30" orientation="longitude" pageId="12" pageNumber="183" precision="925" value="22.5">22°30'E</geoCoordinate>
. 
<date box="[878,982,252,272]" pageId="12" pageNumber="183" value="1979-02-02" valueMax="1979-02-04" valueMin="1979-02-02">2–4.ii.1979</date>
/ B/ Lamoral, I Bampton / J. Barnley. Malaise tr’; 4 
<emphasis box="[482,539,276,296]" italics="true" pageId="12" pageNumber="183">ơ 4^‘</emphasis>
South Africa: N 
<collectingRegion box="[691,738,276,296]" country="South Africa" name="Western Cape" pageId="12" pageNumber="183">Cape</collectingRegion>
/ Witsand Nature Reserve / 
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/ 
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J Londt &amp; T Dikow / 
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<taxonomicName box="[713,776,301,321]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="183" phylum="Tracheophyta" rank="genus">
<emphasis box="[713,776,301,321]" italics="true" pageId="12" pageNumber="183">Acacia</emphasis>
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/ savannah &amp; white dune area’; 2 
<emphasis italics="true" pageId="12" pageNumber="183">ơ 1^‘</emphasis>
South Africa: N 
<collectingRegion box="[339,386,324,344]" country="South Africa" name="Western Cape" pageId="12" pageNumber="183">Cape</collectingRegion>
/ Witsand Nature Reserve / 
<geoCoordinate box="[649,760,322,345]" degrees="28" direction="south" minutes="33.673" orientation="latitude" pageId="12" pageNumber="183" precision="1" value="-28.561216">28°33.673'S</geoCoordinate>
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/ 
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J Londt &amp; T Dikow / 
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<emphasis box="[366,429,349,369]" italics="true" pageId="12" pageNumber="183">Acacia</emphasis>
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/ savannah. Red sandy ridge’; 6 
<emphasis box="[721,821,348,368]" italics="true" pageId="12" pageNumber="183">ơ 1^same</emphasis>
data as 
<typeStatus box="[894,976,348,368]" pageId="12" pageNumber="183">holotype</typeStatus>
.
</paragraph>
<paragraph blockId="12.[144,1104,204,991]" pageId="12" pageNumber="183">
Other material examined: I have seen the following specimen, identified as 
<taxonomicName baseAuthorityName="Macquart" baseAuthorityYear="1838" box="[809,883,381,401]" class="Insecta" family="Asilidae" genus="Laphria" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="183" phylum="Arthropoda" rank="species" species="testacea">
<emphasis box="[809,883,381,401]" italics="true" pageId="12" pageNumber="183">testacea</emphasis>
</taxonomicName>
by Engel, which appears to belong to 
<taxonomicName authorityName="Londt" authorityYear="2008" box="[257,320,405,425]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="183" phylum="Arthropoda" rank="species" species="effrena">
<emphasis box="[257,320,405,425]" italics="true" pageId="12" pageNumber="183">effrena</emphasis>
</taxonomicName>
. Because the locality is so far removed from the Northern Cape records, I refrain from including the specimen in the 
<typeStatus box="[417,456,428,448]" pageId="12" pageNumber="183">type</typeStatus>
series. 
<collectingCountry box="[526,653,428,448]" name="Zimbabwe" pageId="12" pageNumber="183">ZIMBABWE</collectingCountry>
: 1 
<emphasis box="[673,687,429,448]" italics="true" pageId="12" pageNumber="183">ơ</emphasis>
‘Victoria Falls [
<geoCoordinate box="[835,907,426,449]" degrees="17" direction="south" minutes="55" orientation="latitude" pageId="12" pageNumber="183" precision="925" value="-17.916666">17°55'S</geoCoordinate>
: 
<geoCoordinate box="[911,989,426,449]" degrees="25" direction="east" minutes="51" orientation="longitude" pageId="12" pageNumber="183" precision="925" value="25.85">25°51'E</geoCoordinate>
], / 
<date box="[1017,1092,428,448]" pageId="12" pageNumber="183" value="1920-01-04">4.i.1920</date>
/ Rhodesia / Museum’, Pres. by / Imp. Inst. Ent. / Brit. Mus. / 1930-298.’, ‘ 
<taxonomicName baseAuthorityName="Macquart" baseAuthorityYear="1838" box="[794,966,453,473]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="183" phylum="Arthropoda" rank="species" species="testacea">
<emphasis box="[794,966,453,473]" italics="true" pageId="12" pageNumber="183">Trichardis testacea</emphasis>
</taxonomicName>
/ Merh. / Dr. E. O. Engel det.’ (BMNH).
</paragraph>
<paragraph blockId="12.[144,1104,204,991]" pageId="12" pageNumber="183">
Distribution and biology: The species is a southern African endemic, being found in the 
<collectingRegion box="[144,417,542,567]" country="South Africa" name="Northern Cape" pageId="12" pageNumber="183">Northern Cape Province</collectingRegion>
of 
<collectingCountry box="[456,599,542,567]" name="South Africa" pageId="12" pageNumber="183">South Africa</collectingCountry>
and in western 
<collectingCountry box="[778,898,542,567]" name="Zimbabwe" pageId="12" pageNumber="183">Zimbabwe</collectingCountry>
(
<figureCitation box="[914,999,542,567]" captionStart="Fig 59" captionStartId="55.[183,213,872,892]" captionTargetBox="[287,1039,203,852]" captionTargetId="figure-10@55.[285,1041,203,852]" captionTargetPageId="55" captionText="Fig 59. Distributions of southern African Trichardis species: – T. cribrata (Loew, 1858), 9 – T. effrena sp. n., 1974, • – T. picta Hermann, 1906." figureDoi="http://doi.org/10.5281/zenodo.7917407" httpUri="https://zenodo.org/record/7917407/files/figure.png" pageId="12" pageNumber="183">Fig. 59</figureCitation>
). While I hesitate to give 
<typeStatus box="[320,368,574,599]" pageId="12" pageNumber="183">type</typeStatus>
status to the single Zimbabwean specimen because of its isolated position relative to the other records, I am fairly confident that the specimen is correctly labelled and identified. 
<collectingCountry box="[405,515,638,663]" name="Botswana" pageId="12" pageNumber="183">Botswana</collectingCountry>
is generally poorly sampled and so this kind of apparently disjuct distributional pattern should not cause undue concern. Other asilid species have been shown to have a similar distributional pattern. For example Londt (2004) demonstrated that 
<taxonomicName authority="(Engel, 1932)" baseAuthorityName="Engel" baseAuthorityYear="1932" box="[353,753,734,759]" class="Insecta" family="Asilidae" genus="Laphystotes" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="183" phylum="Arthropoda" rank="species" species="albicans">
<emphasis box="[353,590,734,758]" italics="true" pageId="12" pageNumber="183">Laphystotes albicans</emphasis>
(Engel, 1932)
</taxonomicName>
is similarly distributed. Adults of the new species are active during summer and have been collected between January and March (Table 1). This species is associated with open 
<taxonomicName box="[792,870,798,822]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="183" phylum="Tracheophyta" rank="genus">
<emphasis box="[792,870,798,822]" italics="true" pageId="12" pageNumber="183">Acacia</emphasis>
</taxonomicName>
savannah and mixed woodland.All the specimens captured at Witsand Nat. Res. were found resting on sandy pathways.
</paragraph>
<paragraph blockId="12.[144,1104,204,991]" pageId="12" pageNumber="183">
Similar species: Although sharing a number of characters with 
<taxonomicName authorityName="Londt" authorityYear="2008" box="[877,952,902,926]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="183" phylum="Arthropoda" rank="species" species="glabra">
<emphasis box="[877,952,902,926]" italics="true" pageId="12" pageNumber="183">glabra</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Londt" authorityYear="2008" box="[1013,1097,902,926]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="183" phylum="Arthropoda" rank="species" species="mellina">
<emphasis box="[1013,1097,902,926]" italics="true" pageId="12" pageNumber="183">mellina</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Londt" authorityYear="2008" box="[144,222,934,958]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="183" phylum="Arthropoda" rank="species" species="effrena">
<emphasis box="[144,222,934,958]" italics="true" pageId="12" pageNumber="183">effrena</emphasis>
</taxonomicName>
is a distinctive species in that it displays a remarkable reduction in wing venation and has a distinctive male genital form.
</paragraph>
</subSubSection>
</treatment>
</document>