136 lines
19 KiB
XML
136 lines
19 KiB
XML
<document id="06F52CB6503ACCD86D31D6D481EAF05C" ID-CLB-Dataset="42355" ID-DOI="10.11646/zootaxa.3873.5.7" ID-GBIF-Dataset="71209bd4-3468-45b3-9bda-3c3a31bbde76" ID-ISSN="1175-5326" ID-Zenodo-Dep="252165" ID-ZooBank="F51524C8-B5B3-4DDC-B861-01754495DD75" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460940005797" checkinUser="plazi" docAuthor="Żabka, Marek & Patoleta, Barbara M." docDate="2014" docId="039087C2FFCF9B4CFF7C6E99FC0D9FEA" docLanguage="en" docName="zt03873p589.pdf" docOrigin="Zootaxa 3873 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helpis wanlessi Żabka & Patoleta, 2014, sp. nov." docType="treatment" docVersion="15" lastPageNumber="580" masterDocId="FFA9FFBAFFC69B45FFEB6C4EFFB2997D" masterDocTitle="New species of Helpis Simon, 1901 from Australia (Araneae: Salticidae), with a new definition of the genus" masterLastPageNumber="589" masterPageNumber="571" pageNumber="580" updateTime="1698584017404" updateUser="ExternalLinkService">
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<mods:title id="A44B46848A668D17FF0C9698E472C4D4">New species of Helpis Simon, 1901 from Australia (Araneae: Salticidae), with a new definition of the genus</mods:title>
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<mods:namePart id="7B371578ACFABD0A4A9F29B90E828E6E">Żabka, Marek</mods:namePart>
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<mods:namePart id="AE4BFB4EE8628C30037E816EED7281BE">Patoleta, Barbara M.</mods:namePart>
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<mods:title id="2B4650599E0DB1A2FC26319813017E26">Zootaxa</mods:title>
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<mods:part id="CB6BEC633060F9516C7ED114B682ABA8">
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<mods:date id="505700F4974F8B45B2529B6B114FC4BC">2014</mods:date>
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<mods:number id="F0D49510EF2544B57C3011DAC08845AD">3873</mods:number>
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<mods:number id="EA6481F725FD4D330B973B223FADED0F">5</mods:number>
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<mods:identifier id="CCFE265EE54B275DA6259B086F168748" type="DOI">10.11646/zootaxa.3873.5.7</mods:identifier>
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<treatment id="039087C2FFCF9B4CFF7C6E99FC0D9FEA" ID-DOI="http://doi.org/10.5281/zenodo.4685403" ID-GBIF-Taxon="119590914" ID-Zenodo-Dep="4685403" LSID="urn:lsid:plazi:treatment:039087C2FFCF9B4CFF7C6E99FC0D9FEA" httpUri="http://treatment.plazi.org/id/039087C2FFCF9B4CFF7C6E99FC0D9FEA" lastPageNumber="580" pageId="9" pageNumber="580">
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<subSubSection id="C323655FFFCF9B4CFF7C6E99FE8C9A6E" pageId="9" pageNumber="580" type="nomenclature">
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<paragraph id="8B8636D4FFCF9B4CFF7C6E99FE0A9B8F" blockId="9.[151,440,727,787]" box="[151,440,727,754]" pageId="9" pageNumber="580">
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<heading id="D0CE81B8FFCF9B4CFF7C6E99FE0A9B8F" bold="true" box="[151,440,727,754]" fontSize="11" level="1" pageId="9" pageNumber="580" reason="1">
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<emphasis id="B94DEAC6FFCF9B4CFF7C6E99FE0A9B8F" bold="true" box="[151,440,727,754]" pageId="9" pageNumber="580">
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<taxonomicName id="4C394D57FFCF9B4CFF7C6E99FEE19B8C" ID-CoL="6LMRC" box="[151,339,727,753]" class="Arachnida" family="Salticidae" genus="Helpis" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="wanlessi" status="sp. nov.">
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<emphasis id="B94DEAC6FFCF9B4CFF7C6E99FEE19B8C" bold="true" box="[151,339,727,753]" italics="true" pageId="9" pageNumber="580">Helpis wanlessi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27E57BDFFCF9B4CFEB16E96FE0A9B8F" box="[346,440,728,754]" pageId="9" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8636D4FFCF9B4CFF7C6EB4FE8C9A6E" blockId="9.[151,440,727,787]" box="[151,318,762,787]" pageId="9" pageNumber="580">
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<figureCitation id="13022A51FFCF9B4CFF7C6EB4FEA59A6E" box="[151,279,762,787]" captionStart="FIGURES 42 – 50" captionStartId="10.[151,264,1912,1934]" captionTargetBox="[214,1366,225,1861]" captionTargetId="figure@10.[187,1400,193,1890]" captionTargetPageId="10" captionText="FIGURES 42 – 50. Helpis wanlessi sp. nov. 42 female holotype, dorsal view; 43 same, ventral view; 44 same, frontal view; 45 same, endites, labium and chelicerae; 46 same, cheliceral dentition; 47 same, lateral view, 48 – 49 same, epigyne; 50 same, vulva; (ag: accessory gland, cd: copulatory duct, co: copulatory opening, mg: median guide, p: pouch, s: spermatheca). Scale: 42 – 47 = 1 mm, 49 = as in Fig." httpUri="https://zenodo.org/record/252171/files/figure.png" pageId="9" pageNumber="580">Figs 42–50</figureCitation>
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,
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<figureCitation id="13022A51FFCF9B4CFEC96EB4FE8C9A6E" box="[290,318,762,787]" captionStart="FIGURE 86" captionStartId="17.[151,250,745,767]" captionTargetBox="[255,1357,193,728]" captionTargetId="figure@17.[227,1357,193,728]" captionTargetPageId="17" captionText="FIGURE 86. Distribution records of Helpis merriwa (square) and H. wanlessi (triangle)." httpUri="https://zenodo.org/record/252177/files/figure.png" pageId="9" pageNumber="580">86</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C323655FFFCF9B4CFF7C6F0CFCAA9ADE" pageId="9" pageNumber="580" type="materials_examined">
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<paragraph id="8B8636D4FFCF9B4CFF7C6F0CFCAA9ADE" blockId="9.[151,1437,834,1687]" pageId="9" pageNumber="580">
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<emphasis id="B94DEAC6FFCF9B4CFF7C6F0CFEF09A26" bold="true" box="[151,322,834,859]" pageId="9" pageNumber="580">
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<typeStatus id="54828876FFCF9B4CFF7C6F0CFF609A26" box="[151,210,834,859]" pageId="9" pageNumber="580">Type</typeStatus>
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material
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</emphasis>
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.
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<collectingCountry id="F32E7644FFCF9B4CFEBE6F0DFE459A26" box="[341,503,835,859]" name="Australia" pageId="9" pageNumber="580">AUSTRALIA</collectingCountry>
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, New South
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<collectingCountry id="F32E7644FFCF9B4CFD7D6F0CFD6C9A26" box="[662,734,834,859]" name="United Kingdom" pageId="9" pageNumber="580">Wales</collectingCountry>
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: ♂
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<typeStatus id="54828876FFCF9B4CFCE56F0CFCC19A26" box="[782,883,834,859]" pageId="9" pageNumber="580" type="holotype">holotype</typeStatus>
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, Warrumbungles National Park, John Renshaw Parkway,
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<quantity id="4CC19B31FFCF9B4CFEED6F29FEE69A02" box="[262,340,870,895]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="580" unit="km" value="1.9">1.9 km</quantity>
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W of Camp Wambelong,
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<geoCoordinate id="EE0D5013FFCF9B4CFD966F28FCB19A02" box="[637,771,870,895]" direction="south" orientation="latitude" pageId="9" pageNumber="580" precision="15" value="-31.275555">31º16'32" S</geoCoordinate>
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,
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<geoCoordinate id="EE0D5013FFCF9B4CFCFB6F28FC149A03" box="[784,934,870,895]" direction="east" orientation="longitude" pageId="9" pageNumber="580" precision="15" value="148.96028">148º57'37" E</geoCoordinate>
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, [-31.2756, 148.9603], under rocks,
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<date id="FF871014FFCF9B4CFAA86F29FF609ADE" pageId="9" pageNumber="580" value="2001-11-10">10 Nov. 2001</date>
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, M. Gray, G. Milledge & H. Smith, AMS KS75206.
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</paragraph>
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</subSubSection>
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<subSubSection id="C323655FFFCF9B4CFF2C6FE0FED79A96" pageId="9" pageNumber="580" type="etymology">
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<paragraph id="8B8636D4FFCF9B4CFF2C6FE0FED79A96" blockId="9.[151,1437,834,1687]" pageId="9" pageNumber="580">
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<emphasis id="B94DEAC6FFCF9B4CFF2C6FE0FEF49ABA" bold="true" box="[199,326,942,967]" pageId="9" pageNumber="580">Etymology</emphasis>
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. In honour of the British arachnologist Mr Fred R. Wanless, in recognition of his excellent work on salticid taxonomy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C323655FFFCF9B4CFF2C6FB8FC329D4E" pageId="9" pageNumber="580" type="diagnosis">
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<paragraph id="8B8636D4FFCF9B4CFF2C6FB8FC329D4E" blockId="9.[151,1437,834,1687]" pageId="9" pageNumber="580">
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<emphasis id="B94DEAC6FFCF9B4CFF2C6FB8FE889D72" bold="true" box="[199,314,1014,1039]" pageId="9" pageNumber="580">Diagnosis</emphasis>
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. In comparison with other relatives, median guides wider, copulatory openings oriented laterally (the only case), insemination ducts relatively shorter (
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<figureCitation id="13022A51FFCF9B4CFD1A6854FCC79D4E" box="[753,885,1050,1075]" captionStart="FIGURES 42 – 50" captionStartId="10.[151,264,1912,1934]" captionTargetBox="[214,1366,225,1861]" captionTargetId="figure@10.[187,1400,193,1890]" captionTargetPageId="10" captionText="FIGURES 42 – 50. Helpis wanlessi sp. nov. 42 female holotype, dorsal view; 43 same, ventral view; 44 same, frontal view; 45 same, endites, labium and chelicerae; 46 same, cheliceral dentition; 47 same, lateral view, 48 – 49 same, epigyne; 50 same, vulva; (ag: accessory gland, cd: copulatory duct, co: copulatory opening, mg: median guide, p: pouch, s: spermatheca). Scale: 42 – 47 = 1 mm, 49 = as in Fig." httpUri="https://zenodo.org/record/252171/files/figure.png" pageId="9" pageNumber="580">Figs 49–50</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C323655FFFCF9B4CFF2C6870FE379F0E" pageId="9" pageNumber="580" type="description">
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<paragraph id="8B8636D4FFCF9B4CFF2C6870FE579F32" blockId="9.[151,1437,834,1687]" pageId="9" pageNumber="580">
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<emphasis id="B94DEAC6FFCF9B4CFF2C6870FEE39D2A" bold="true" box="[199,337,1086,1111]" pageId="9" pageNumber="580">Description</emphasis>
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. Female
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<typeStatus id="54828876FFCF9B4CFE2F6870FD989D2A" box="[452,554,1086,1111]" pageId="9" pageNumber="580" type="holotype">holotype</typeStatus>
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. Cephalothorax pear-shaped, greyish brown covered with brown hairs and sparse chestnut setae (
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<figureCitation id="13022A51FFCF9B4CFE7C682CFE5F9D06" box="[407,493,1122,1147]" captionStart="FIGURES 42 – 50" captionStartId="10.[151,264,1912,1934]" captionTargetBox="[214,1366,225,1861]" captionTargetId="figure@10.[187,1400,193,1890]" captionTargetPageId="10" captionText="FIGURES 42 – 50. Helpis wanlessi sp. nov. 42 female holotype, dorsal view; 43 same, ventral view; 44 same, frontal view; 45 same, endites, labium and chelicerae; 46 same, cheliceral dentition; 47 same, lateral view, 48 – 49 same, epigyne; 50 same, vulva; (ag: accessory gland, cd: copulatory duct, co: copulatory opening, mg: median guide, p: pouch, s: spermatheca). Scale: 42 – 47 = 1 mm, 49 = as in Fig." httpUri="https://zenodo.org/record/252171/files/figure.png" pageId="9" pageNumber="580">Fig. 42</figureCitation>
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). Eyes surrounded in dark brown. Thorax with middle yellowish stripe. Abdomen with a grey pattern on light background. Spinnerets whitish, but the anterior ones with wide grey band. Clypeus grey brown, narrow (about 16% of AME diameter), with some protruding bristles below and between AME (
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<figureCitation id="13022A51FFCF9B4CFA8468E4FF059D9A" captionStart="FIGURES 42 – 50" captionStartId="10.[151,264,1912,1934]" captionTargetBox="[214,1366,225,1861]" captionTargetId="figure@10.[187,1400,193,1890]" captionTargetPageId="10" captionText="FIGURES 42 – 50. Helpis wanlessi sp. nov. 42 female holotype, dorsal view; 43 same, ventral view; 44 same, frontal view; 45 same, endites, labium and chelicerae; 46 same, cheliceral dentition; 47 same, lateral view, 48 – 49 same, epigyne; 50 same, vulva; (ag: accessory gland, cd: copulatory duct, co: copulatory opening, mg: median guide, p: pouch, s: spermatheca). Scale: 42 – 47 = 1 mm, 49 = as in Fig." httpUri="https://zenodo.org/record/252171/files/figure.png" pageId="9" pageNumber="580">Fig. 44</figureCitation>
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). Chelicerae grey brown, with rugose frontal surfaces, promargin and retromargin with four and five teeth, respectively (
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<figureCitation id="13022A51FFCF9B4CFEDA68BCFE389C76" box="[305,394,1266,1291]" captionStart="FIGURES 42 – 50" captionStartId="10.[151,264,1912,1934]" captionTargetBox="[214,1366,225,1861]" captionTargetId="figure@10.[187,1400,193,1890]" captionTargetPageId="10" captionText="FIGURES 42 – 50. Helpis wanlessi sp. nov. 42 female holotype, dorsal view; 43 same, ventral view; 44 same, frontal view; 45 same, endites, labium and chelicerae; 46 same, cheliceral dentition; 47 same, lateral view, 48 – 49 same, epigyne; 50 same, vulva; (ag: accessory gland, cd: copulatory duct, co: copulatory opening, mg: median guide, p: pouch, s: spermatheca). Scale: 42 – 47 = 1 mm, 49 = as in Fig." httpUri="https://zenodo.org/record/252171/files/figure.png" pageId="9" pageNumber="580">Fig. 46</figureCitation>
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). Pedipalps greyish brown, metatarsus with a dorsal and a retrolateral spine. Endites and labium basally greyish brown with lighter distal halves (
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<figureCitation id="13022A51FFCF9B4CFCFE6958FCD89C52" box="[789,874,1302,1327]" captionStart="FIGURES 42 – 50" captionStartId="10.[151,264,1912,1934]" captionTargetBox="[214,1366,225,1861]" captionTargetId="figure@10.[187,1400,193,1890]" captionTargetPageId="10" captionText="FIGURES 42 – 50. Helpis wanlessi sp. nov. 42 female holotype, dorsal view; 43 same, ventral view; 44 same, frontal view; 45 same, endites, labium and chelicerae; 46 same, cheliceral dentition; 47 same, lateral view, 48 – 49 same, epigyne; 50 same, vulva; (ag: accessory gland, cd: copulatory duct, co: copulatory opening, mg: median guide, p: pouch, s: spermatheca). Scale: 42 – 47 = 1 mm, 49 = as in Fig." httpUri="https://zenodo.org/record/252171/files/figure.png" pageId="9" pageNumber="580">Fig. 45</figureCitation>
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). Sternum yellowish, with darker margins. Venter whitish with grey pattern (
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<figureCitation id="13022A51FFCF9B4CFE2B6974FDA79C2E" box="[448,533,1338,1363]" captionStart="FIGURES 42 – 50" captionStartId="10.[151,264,1912,1934]" captionTargetBox="[214,1366,225,1861]" captionTargetId="figure@10.[187,1400,193,1890]" captionTargetPageId="10" captionText="FIGURES 42 – 50. Helpis wanlessi sp. nov. 42 female holotype, dorsal view; 43 same, ventral view; 44 same, frontal view; 45 same, endites, labium and chelicerae; 46 same, cheliceral dentition; 47 same, lateral view, 48 – 49 same, epigyne; 50 same, vulva; (ag: accessory gland, cd: copulatory duct, co: copulatory opening, mg: median guide, p: pouch, s: spermatheca). Scale: 42 – 47 = 1 mm, 49 = as in Fig." httpUri="https://zenodo.org/record/252171/files/figure.png" pageId="9" pageNumber="580">Fig. 43</figureCitation>
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). Legs I the longest and more robust than others, greyish brown, only coxae and femora pale yellow, darker laterally. Ventral spination of first legs: tibia 2-2-2-2, metatarsus 2-2. Legs II more delicate, with similar colouration and spination. Legs III–IV lighter, with darker bands. All femora with dorsal spines: first femora 1-1-2, the others 1-1-3. Epigyne with clearly visible posterior pouch (
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<figureCitation id="13022A51FFCF9B4CFB4469E8FAB59CC2" box="[1199,1287,1446,1471]" captionStart="FIGURES 42 – 50" captionStartId="10.[151,264,1912,1934]" captionTargetBox="[214,1366,225,1861]" captionTargetId="figure@10.[187,1400,193,1890]" captionTargetPageId="10" captionText="FIGURES 42 – 50. Helpis wanlessi sp. nov. 42 female holotype, dorsal view; 43 same, ventral view; 44 same, frontal view; 45 same, endites, labium and chelicerae; 46 same, cheliceral dentition; 47 same, lateral view, 48 – 49 same, epigyne; 50 same, vulva; (ag: accessory gland, cd: copulatory duct, co: copulatory opening, mg: median guide, p: pouch, s: spermatheca). Scale: 42 – 47 = 1 mm, 49 = as in Fig." httpUri="https://zenodo.org/record/252171/files/figure.png" pageId="9" pageNumber="580">Fig. 49</figureCitation>
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). Copulatory ducts short, slightly oblique, spermathecae pear-shaped, accessory glands short (
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<figureCitation id="13022A51FFCF9B4CFBD86984FB389C9E" box="[1075,1162,1482,1507]" captionStart="FIGURES 42 – 50" captionStartId="10.[151,264,1912,1934]" captionTargetBox="[214,1366,225,1861]" captionTargetId="figure@10.[187,1400,193,1890]" captionTargetPageId="10" captionText="FIGURES 42 – 50. Helpis wanlessi sp. nov. 42 female holotype, dorsal view; 43 same, ventral view; 44 same, frontal view; 45 same, endites, labium and chelicerae; 46 same, cheliceral dentition; 47 same, lateral view, 48 – 49 same, epigyne; 50 same, vulva; (ag: accessory gland, cd: copulatory duct, co: copulatory opening, mg: median guide, p: pouch, s: spermatheca). Scale: 42 – 47 = 1 mm, 49 = as in Fig." httpUri="https://zenodo.org/record/252171/files/figure.png" pageId="9" pageNumber="580">Fig. 50</figureCitation>
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). Dimensions: CL 2.81,
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<collectingCountry id="F32E7644FFCF9B4CFF7C69A1FF0C9F7A" box="[151,190,1519,1543]" name="Switzerland" pageId="9" pageNumber="580">CH</collectingCountry>
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1.45, CW 2.22, AL 3.32, AW 2.03, EFL 1.24, AEW 1.87, PEW 1.53, leg I 7.30 (2.14+1.32+1.85+1.31+0.68), leg II 6.12 (1.74+1.09+1.49+1.20+0.60), leg III 6.22 (1.71+0.95+1.44+1.46+0.66), leg IV 7.83 (2.20+0.78+2.03+1.86+0.70).
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</paragraph>
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<paragraph id="8B8636D4FFCF9B4CFF2C6A14FE379F0E" blockId="9.[151,1437,834,1687]" box="[199,389,1626,1651]" pageId="9" pageNumber="580">Male. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="C323655FFFCF9B4CFF2C6A30FC0D9FEA" box="[199,959,1662,1687]" pageId="9" pageNumber="580" type="distribution">
|
||
<paragraph id="8B8636D4FFCF9B4CFF2C6A30FC0D9FEA" blockId="9.[151,1437,834,1687]" box="[199,959,1662,1687]" pageId="9" pageNumber="580">
|
||
<emphasis id="B94DEAC6FFCF9B4CFF2C6A30FEEB9FEA" bold="true" box="[199,345,1662,1687]" pageId="9" pageNumber="580">Distribution</emphasis>
|
||
. Known only from the
|
||
<typeStatus id="54828876FFCF9B4CFD8B6A31FD229FEA" box="[608,656,1663,1687]" pageId="9" pageNumber="580">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13022A51FFCF9B4CFD116A30FCFF9FEA" box="[762,845,1662,1687]" captionStart="FIGURE 86" captionStartId="17.[151,250,745,767]" captionTargetBox="[255,1357,193,728]" captionTargetId="figure@17.[227,1357,193,728]" captionTargetPageId="17" captionText="FIGURE 86. Distribution records of Helpis merriwa (square) and H. wanlessi (triangle)." httpUri="https://zenodo.org/record/252177/files/figure.png" pageId="9" pageNumber="580">Fig. 86</figureCitation>
|
||
, triangle).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |