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<mods:title id="B683551006FECE8A5C9CF1D869802132">Cleisostoma mulunense (Orchidaceae), a new species from Guangxi, China</mods:title>
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<mods:namePart id="FADE075CFA7C6377581AC0CE7EDD9997">Qin, Ying</mods:namePart>
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<mods:namePart id="8723D33CD26E34A7A7CE14B69BCF5905">Luo, Liu Juan</mods:namePart>
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<mods:namePart id="E6933AAD7AF8EF6A4076E63945B18BBF">Liu, Yan</mods:namePart>
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<taxonomicName id="4C24F98DFFA1FF8CBBC3C3A4FD0FF8E9" authority="Ying Qin &amp; Yan Liu" authorityName="Ying Qin &amp; Yan Liu" authorityYear="2023" box="[136,648,1780,1807]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="295" phylum="Tracheophyta" rank="species" species="mulunense" status="sp. nov.">
<emphasis id="B9505E1CFFA1FF8CBBC3C3A4FE1CF8E8" bold="true" box="[136,411,1780,1807]" italics="true" pageId="0" pageNumber="295">Cleisostoma mulunense</emphasis>
Ying Qin &amp; Yan Liu
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, 
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(
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&amp; 
<figureCitation id="131F9E8BFFA1FF8CB82CC3A4FCF1F8E9" box="[871,886,1780,1806]" captionStart="FIGURE 2" captionStartId="3.[136,229,1956,1978]" captionTargetBox="[198,1390,245,1931]" captionTargetId="figure-46@3.[198,1390,245,1931]" captionTargetPageId="3" captionText="FIGURE 2. Cleisostoma mulunense. A. Habitat. B. Flower, side view. C. Flower, front view. D. Lip, longitudinal section. E. Dorsal sepal, lateral sepals, petals, lip, column and ovary. F. Callus, side view. G. Callus, front view. H. Lower part of callus. I. Papillae at mouth of spur inside. J. Longitudinal ridge of the disk, side view. K. Longitudinal septum of spur. L. Leaf, cross section. M. Leaves, adaxial view and side view. Photos by Yu-Lan Su (A &amp; B), Yan Liu (C) and Ying Qin (D–M) from living plants of the holotype population." figureDoi="http://doi.org/10.5281/zenodo.10146266" httpUri="https://zenodo.org/record/10146266/files/figure.png" pageId="0" pageNumber="295">2</figureCitation>
)
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:— 
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. 
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: 
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, 
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, elev. ca. 
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(
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!; 
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!)
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.
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Diagnosis:—The new species is close to 
<taxonomicName id="4C24F98DFFA1FF8CB952C2D8FCF3F879" authority="Z. H. Tsi (1983: 76)" authorityName="Z. H. Tsi" authorityPageNumber="76" authorityYear="1983" box="[537,884,1928,1950]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="295" phylum="Tracheophyta" rank="species" species="menghaiense">
<emphasis id="B9505E1CFFA1FF8CB952C2D8FD33F87A" box="[537,692,1928,1949]" italics="true" pageId="0" pageNumber="295">C. menghaiense</emphasis>
Z.H.Tsi (1983: 76)
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, but it is easily distinguished from the latter mainly by its yellowish green to yellow, with purplish brown (vs. pale yellow with purplish red) flowers, suboblong (vs. elliptic) dorsal sepal, broadly oblong and obviously oblique (vs. obovate, slightly oblique) lateral sepals, oblong (vs. obliquely suboblong) petals, lip median lobe with (vs. without) 2 basal backward triangular lobules, cylindroid (vs. somewhat horn-shaped) spur.
</paragraph>
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<emphasis id="B9505E1CFFA0FF8DBBC3C3CDFF7AF955" bold="true" box="[136,253,1693,1715]" pageId="1" pageNumber="296">FIGURE 1.</emphasis>
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<taxonomicName id="4C24F98DFFA0FF8DBA4AC3CDFE60F955" authorityName="Ying Qin &amp; Yan Liu" authorityYear="2023" box="[257,487,1693,1714]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="296" phylum="Tracheophyta" rank="species" species="mulunense">Cleisostoma mulunense</taxonomicName>
.
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A. Habit. B. Flower, obliquely back view. C. Flower, side view. D. Flower, front view. E. Floral bract. F. Dorsal sepal, petal and lateral sepal. G. Column, front view. H. Flower with sepals and petals removed, side view. I. Flower with sepals and petals removed, longitudinal section. J. Anther caps, top view, side view and ventral view. K. Pollinaria. Drawing by Yun-Xi Zhu from a living plant of the holotype population.
</paragraph>
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Herb epiphytic. Stem 2.0– 
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long, 
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in diameter, many-noded. Roots fleshy, 
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long, 
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in diameter. Leaves 6–13, distichous, linear, dorsiventrally compressed but convex between median vein and margin, slightly recurved, thickly fleshy, 
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long, 
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wide, margin entire, apex acute, base with amplexicaul-sheathing; median vein concave adaxially. Inflorescence axillary, 
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long, with 1–3 branches or unbranched, 20–41-flowered; peduncle 1.2–3.0 cm long, 
<quantity id="4CDC2FEBFFA0FF8DB860C28EFC34F81F" box="[811,947,2014,2040]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="1" pageNumber="296" unit="mm" value="1.05" valueMax="1.3" valueMin="0.8">0.8–1.3 mm</quantity>
in diameter, with 1–3 tubular sheaths; rachis slightly zig-zag curved, 
<quantity id="4CDC2FEBFFA0FF8DBAD2CD51FDA2F7FB" box="[409,549,2049,2076]" metricMagnitude="-2" metricUnit="m" metricValue="8.45" metricValueMax="10.1" metricValueMin="6.8" pageId="1" pageNumber="296" unit="cm" value="8.45" valueMax="10.1" valueMin="6.8">6.8–10.1 cm</quantity>
long, 
<quantity id="4CDC2FEBFFA0FF8DB925CD52FD72F7FB" box="[622,757,2050,2076]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="1" pageNumber="296" unit="mm" value="1.05" valueMax="1.3" valueMin="0.8">0.8–1.3 mm</quantity>
in diameter. Floral bracts triangular, acute, 
<quantity id="4CDC2FEBFFA0FF8DBFA8CD51FAECF7FB" box="[1251,1387,2049,2076]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="1" pageNumber="296" unit="mm" value="1.05" valueMax="1.5" valueMin="0.6">0.6–1.5 mm</quantity>
long. Ovary terete, yellowish green, ca. 
<quantity id="4CDC2FEBFFA3FF8EBAB5C5CEFDD5FF5E" box="[510,594,158,185]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="297" unit="mm" value="0.8">0.8 mm</quantity>
in diameter, including pedicel to 
<quantity id="4CDC2FEBFFA3FF8EB8F5C5CEFBBEFF5E" box="[958,1081,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="5.35" metricValueMax="6.6" metricValueMin="4.1" pageId="2" pageNumber="297" unit="cm" value="5.35" valueMax="6.6" valueMin="4.1">4.1–6.6 cm</quantity>
long. Sepals and petals yellowish green abaxially, purplish brown adaxially, margin yellowish green and denticulate, sometimes median vein yellowish green adaxially. Dorsal sepal suboblong, concave, 
<quantity id="4CDC2FEBFFA3FF8EB984C5B6FCDFFEE6" box="[719,856,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="2" pageNumber="297" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4–2.8 mm</quantity>
long, 
<quantity id="4CDC2FEBFFA3FF8EB8EFC5B6FBA9FEE6" box="[932,1070,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="2" pageNumber="297" unit="mm" value="1.7" valueMax="1.9" valueMin="1.5">1.5–1.9 mm</quantity>
wide, apex obtuse; lateral sepals broadly oblong, obviously oblique, 2.8–3.0 mm long, 
<quantity id="4CDC2FEBFFA3FF8EB9B2C45AFC05FEC2" box="[761,898,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="2" pageNumber="297" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
wide, apex obtuse, base adnate to column foot. Petals oblong, 
<quantity id="4CDC2FEBFFA3FF8EBA66C47EFE34FEAE" box="[301,435,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="2" pageNumber="297" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.3–2.7 mm</quantity>
long, 
<quantity id="4CDC2FEBFFA3FF8EBAB2C47EFDF8FEAE" box="[505,639,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="2" pageNumber="297" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4 mm</quantity>
wide, apex obtuse. Lip 3-lobed; lateral lobes broadly triangular, straight, slightly incurved at apex, yellowish green to yellow, 
<quantity id="4CDC2FEBFFA3FF8EB9AFC402FCEBFE8A" box="[740,876,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="2" pageNumber="297" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.1–1.3 mm</quantity>
long, ca. 
<quantity id="4CDC2FEBFFA3FF8EB895C402FBB2FE8A" box="[990,1077,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="297" unit="mm" value="1.9">1.9 mm</quantity>
wide, proximal margin inflexed; median lobe triangular, slightly fleshy, yellowish green to yellow, with purplish brown apex, 
<quantity id="4CDC2FEBFFA3FF8EBFC6C426FA94FE76" box="[1165,1299,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="2" pageNumber="297" unit="mm" value="2.95" valueMax="3.2" valueMin="2.7">2.7–3.2 mm</quantity>
long, 
<quantity id="4CDC2FEBFFA3FF8EBE1CC426FF33FE52" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="2" pageNumber="297" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
wide, base with 2 backward triangular lobules, apex acuminate and upcurved; disk with a longitudinal ridge extending from its center to spur mouth; spur yellowish green to yellow, cylindroid, dorsiventrally compressed, 
<quantity id="4CDC2FEBFFA3FF8EBE1CC4EEFF33FE1A" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="2" pageNumber="297" unit="mm" value="4.05" valueMax="4.2" valueMin="3.9">3.9–4.2 mm</quantity>
long, 
<quantity id="4CDC2FEBFFA3FF8EBBB2C4B2FE07FE1A" box="[249,384,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="2" pageNumber="297" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 mm</quantity>
wide, slightly downcurved, concave on back wall at mouth outside, apex obtuse and with a tiny longitudinal septum inside, papillate at mouth inside, with a callus on back wall of mouth inside; callus nearly ligulatelinear, slightly widened from middle to upper part, papillate in lower part, with a central longitudinal groove. Column erect, stout, 2.0– 
<quantity id="4CDC2FEBFFA3FF8EBA0BC71EFE13FD8E" box="[320,404,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="297" unit="mm" value="2.2">2.2 mm</quantity>
tall, 
<quantity id="4CDC2FEBFFA3FF8EBA81C71EFDD7FD8E" box="[458,592,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="2" pageNumber="297" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
wide, yellowish green with purplish brown, papillate on front surface at base; column foot 
<quantity id="4CDC2FEBFFA3FF8EBA5DC722FE1BFD6A" box="[278,412,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="2" pageNumber="297" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7–1.9 mm</quantity>
long; stigma concave; pollinia 2, spherical, ca. 
<quantity id="4CDC2FEBFFA3FF8EB8E7C722FB86FD6A" box="[940,1025,626,653]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="297" unit="mm" value="0.3">0.3 mm</quantity>
in diameter, each split into 2 subequal hemispheric portions, waxy, dark yellow, with a common stipe and a viscidium; stipe linear, slender, slightly widening near pollinia, ca. 
<quantity id="4CDC2FEBFFA3FF8EBA03C7EAFE19FD32" box="[328,414,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="297" unit="mm" value="1.1">1.1 mm</quantity>
long; viscidium elliptic, ca. 
<quantity id="4CDC2FEBFFA3FF8EB990C7EAFCB7FD32" box="[731,816,698,725]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="297" unit="mm" value="0.2">0.2 mm</quantity>
long; anther cap hemispherical, 0.8–1.0 mm in diameter, with a 
<quantity id="4CDC2FEBFFA3FF8EBB9FC78EFEDCFD1E" box="[212,347,734,761]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="2" pageNumber="297" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.4–0.6 mm</quantity>
long beak. Fruits unknown.
</paragraph>
<paragraph id="8B9B820EFFA3FF8EBBF6C652FD1EFCFB" blockId="2.[136,1452,158,1049]" box="[189,665,770,796]" pageId="2" pageNumber="297">
<emphasis id="B9505E1CFFA3FF8EBBF6C652FEC4FCFB" bold="true" box="[189,323,770,796]" pageId="2" pageNumber="297">Phenology:</emphasis>
—Flowers in July and August.
</paragraph>
</subSubSection>
<subSubSection id="C33ED185FFA3FF8EBBF6C676FBB0FBFF" pageId="2" pageNumber="297" type="etymology">
<paragraph id="8B9B820EFFA3FF8EBBF6C676FA02FCA7" blockId="2.[136,1452,158,1049]" box="[189,1413,806,833]" pageId="2" pageNumber="297">
<emphasis id="B9505E1CFFA3FF8EBBF6C676FECFFCA7" bold="true" box="[189,328,806,832]" pageId="2" pageNumber="297">Etymology:</emphasis>
—The epithet is derived from the 
<typeStatus id="549F3CACFFA3FF8EB98EC677FD72FCA6" box="[709,757,807,833]" pageId="2" pageNumber="297">type</typeStatus>
locality Mulun National Nature Reserve, 
<collectingRegion id="49E04CECFFA3FF8EBF87C676FAA8FCA7" box="[1228,1327,806,832]" country="China" name="Guangxi" pageId="2" pageNumber="297">Guangxi</collectingRegion>
, 
<collectingCountry id="F333C29EFFA3FF8EBE71C676FA06FCA7" box="[1338,1409,806,832]" name="China" pageId="2" pageNumber="297">China</collectingCountry>
.
</paragraph>
<paragraph id="8B9B820EFFA3FF8EBBF6C61AFC4CFC83" blockId="2.[136,1452,158,1049]" box="[189,971,842,869]" pageId="2" pageNumber="297">
<emphasis id="B9505E1CFFA3FF8EBBF6C61AFE12FC83" bold="true" box="[189,405,842,868]" pageId="2" pageNumber="297">Vernacular name:</emphasis>
—ϮḖḆffi± (Chinese pinyin: mù lùn gé jù lán).
</paragraph>
<paragraph id="8B9B820EFFA3FF8EBBF6C63EFBB0FBFF" blockId="2.[136,1452,158,1049]" pageId="2" pageNumber="297">
<emphasis id="B9505E1CFFA3FF8EBBF6C63EFE28FC6F" bold="true" box="[189,431,878,904]" pageId="2" pageNumber="297">Taxonomic notes:—</emphasis>
<taxonomicName id="4C24F98DFFA3FF8EBAE5C63EFD3CFC6F" authorityName="Ying Qin &amp; Yan Liu" authorityYear="2023" box="[430,699,878,904]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="mulunense">
<emphasis id="B9505E1CFFA3FF8EBAE5C63EFD3CFC6F" box="[430,699,878,904]" italics="true" pageId="2" pageNumber="297">Cleisostoma mulunense</emphasis>
</taxonomicName>
is close to 
<taxonomicName id="4C24F98DFFA3FF8EB80DC63FFC7BFC6F" authorityName="Z. H. Tsi" authorityYear="1983" box="[838,1020,878,905]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="menghaiense">
<emphasis id="B9505E1CFFA3FF8EB80DC63FFC7BFC6F" box="[838,1020,878,905]" italics="true" pageId="2" pageNumber="297">C. menghaiense</emphasis>
</taxonomicName>
because of its short stem, linear and dorsiventrally compressed leaves with acute apexes, broadly triangular lateral lobes of lip. However, this new species is easily distinguished from the latter mainly by its yellowish green to yellow, with purplish brown flowers, suboblong dorsal sepal, broadly oblique-oblong lateral sepals, oblong petals, lip median lobe with 2 basal backward triangular lobules, cylindroid spur. Detailed comparisons between them are showed in 
<tableCitation id="C6A6B7B5FFA3FF8EB896C6AEFBB6FBFF" box="[989,1073,1022,1048]" captionStart="TABLE 1" captionStartId="2.[136,224,1092,1116]" captionText="TABLE 1. Morphological comparison between Cleisostoma mulunense and C. menghaiense." pageId="2" pageNumber="297">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5BD286FFA3FF8EBBC3C114FBEBFBBB" box="[136,1132,1092,1116]" pageId="2" pageNumber="297" startId="2.[136,224,1092,1116]" targetType="table">
<paragraph id="8B9B820EFFA3FF8EBBC3C114FBEBFBBB" blockId="2.[136,1132,1092,1116]" box="[136,1132,1092,1116]" pageId="2" pageNumber="297">
<emphasis id="B9505E1CFFA3FF8EBBC3C114FF7CFBBB" bold="true" box="[136,251,1092,1116]" pageId="2" pageNumber="297">TABLE 1.</emphasis>
Morphological comparison between 
<taxonomicName id="4C24F98DFFA3FF8EB9C1C114FC01FBBB" authorityName="Ying Qin &amp; Yan Liu" authorityYear="2023" box="[650,902,1092,1116]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="mulunense">
<emphasis id="B9505E1CFFA3FF8EB9C1C114FC01FBBB" box="[650,902,1092,1116]" italics="true" pageId="2" pageNumber="297">Cleisostoma mulunense</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C24F98DFFA3FF8EB8F1C114FBEFFBBB" authorityName="Z. H. Tsi" authorityYear="1983" box="[954,1128,1092,1116]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="menghaiense">C. menghaiense</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B9B820EFFA3FF8EBBDBC13DFC67F93D" pageId="2" pageNumber="297">
<table id="F92470AEFFA30073BBDBC13DFA10F93D" box="[144,1431,1133,1754]" gridcols="3" gridrows="8" pageId="2" pageNumber="297">
<tr id="3514804CFFA30073BBDBC13DFA10FB64" box="[144,1431,1133,1155]" gridrow="0" pageId="2" pageNumber="297">
<th id="76C5E930FFA30073BBDBC13DFE92FB64" box="[144,277,1133,1155]" gridcol="0" gridrow="0" pageId="2" pageNumber="297">Character</th>
<th id="76C5E930FFA30073BA1AC13DFCF6FB64" box="[337,881,1133,1155]" gridcol="1" gridrow="0" pageId="2" pageNumber="297">
<taxonomicName id="4C24F98DFFA3FF8EBA1AC13DFE53FB65" authorityName="Ying Qin &amp; Yan Liu" authorityYear="2023" box="[337,468,1133,1154]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="mulunense">
<emphasis id="B9505E1CFFA3FF8EBA1AC13DFE53FB65" box="[337,468,1133,1154]" italics="true" pageId="2" pageNumber="297">C. mulunense</emphasis>
</taxonomicName>
</th>
<th id="76C5E930FFA30073B8C0C13DFA10FB64" box="[907,1431,1133,1155]" gridcol="2" gridrow="0" pageId="2" pageNumber="297">
<taxonomicName id="4C24F98DFFA3FF8EB8C0C13DFBA3FB65" authority="Leaves" authorityName="Leaves" box="[907,1060,1133,1154]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="menghaiense">
<emphasis id="B9505E1CFFA3FF8EB8C0C13DFBA3FB65" box="[907,1060,1133,1154]" italics="true" pageId="2" pageNumber="297">C. menghaiense</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3514804CFFA30073BBDBC1C4FA10FB29" box="[144,1431,1172,1230]" gridrow="1" pageId="2" pageNumber="297">
<th id="76C5E930FFA30073BBDBC1C4FE92FB29" box="[144,277,1172,1230]" gridcol="0" gridrow="1" pageId="2" pageNumber="297">Leaves</th>
<td id="76C5E930FFA30073BA1AC1C4FCF6FB29" box="[337,881,1172,1230]" gridcol="1" gridrow="1" pageId="2" pageNumber="297">3.4–6.5 cm × 2.5–5.3 mm, convex between median vein and margin; median vein not raised abaxially</td>
<td id="76C5E930FFA30073B8C0C1C4FA10FB29" box="[907,1431,1172,1230]" gridcol="2" gridrow="1" pageId="2" pageNumber="297">8–14 cm × 4–8 mm, flat between median vein and margin; median vein raised abaxially</td>
</tr>
<tr id="3514804CFFA30073BBDBC1B0FA10FAFD" box="[144,1431,1248,1306]" gridrow="2" pageId="2" pageNumber="297">
<th id="76C5E930FFA30073BBDBC1B0FE92FAFD" box="[144,277,1248,1306]" gridcol="0" gridrow="2" pageId="2" pageNumber="297">Dorsal sepal</th>
<td id="76C5E930FFA30073BA1AC1B0FCF6FAFD" box="[337,881,1248,1306]" gridcol="1" gridrow="2" pageId="2" pageNumber="297">suboblong, 2.4–2.8 × 1.5–1.9 mm, yellowish green with purplish brown</td>
<td id="76C5E930FFA30073B8C0C1B0FA10FAFD" box="[907,1431,1248,1306]" gridcol="2" gridrow="2" pageId="2" pageNumber="297">elliptic, ca. 3.5 × 2 mm, pale yellow with purplish red</td>
</tr>
<tr id="3514804CFFA30073BBDBC07BFA10FA82" box="[144,1431,1323,1381]" gridrow="3" pageId="2" pageNumber="297">
<th id="76C5E930FFA30073BBDBC07BFE92FA82" box="[144,277,1323,1381]" gridcol="0" gridrow="3" pageId="2" pageNumber="297">Lateral sepals</th>
<td id="76C5E930FFA30073BA1AC07BFCF6FA82" box="[337,881,1323,1381]" gridcol="1" gridrow="3" pageId="2" pageNumber="297">broadly oblong, obviously oblique, 2.8–3 × 1.8–2.2 mm, yellowish green with purplish brown</td>
<td id="76C5E930FFA30073B8C0C07BFA10FA82" box="[907,1431,1323,1381]" gridcol="2" gridrow="3" pageId="2" pageNumber="297">obovate, slightly oblique, ca. 3.5 × 2 mm, pale yellow with purplish red</td>
</tr>
<tr id="3514804CFFA30073BBDBC026FA10FA57" box="[144,1431,1398,1456]" gridrow="4" pageId="2" pageNumber="297">
<th id="76C5E930FFA30073BBDBC026FE92FA57" box="[144,277,1398,1456]" gridcol="0" gridrow="4" pageId="2" pageNumber="297">Petals</th>
<td id="76C5E930FFA30073BA1AC026FCF6FA57" box="[337,881,1398,1456]" gridcol="1" gridrow="4" pageId="2" pageNumber="297">oblong, 2.3–2.7 × 1.2–1.4 mm, yellowish green with purplish brown</td>
<td id="76C5E930FFA30073B8C0C026FA10FA57" box="[907,1431,1398,1456]" gridcol="2" gridrow="4" pageId="2" pageNumber="297">obliquely suboblong, ca. 3.0 × 1.5 mm, pale yellow with purplish red</td>
</tr>
<tr id="3514804CFFA30073BBDBC092FA10F9A3" box="[144,1431,1474,1604]" gridrow="5" pageId="2" pageNumber="297">
<th id="76C5E930FFA30073BBDBC092FE92F9A3" box="[144,277,1474,1604]" gridcol="0" gridrow="5" pageId="2" pageNumber="297">Lip</th>
<td id="76C5E930FFA30073BA1AC092FCF6F9A3" box="[337,881,1474,1604]" gridcol="1" gridrow="5" pageId="2" pageNumber="297">median lobe with 2 basal backward triangular lobules, yellowish green to yellow, with purplish brown, apex acuminate; lateral lobes yellowish green to yellow, proximal margin inflexed</td>
<td id="76C5E930FFA30073B8C0C092FA10F9A3" box="[907,1431,1474,1604]" gridcol="2" gridrow="5" pageId="2" pageNumber="297">median lobe without 2 basal backward triangular lobules, purplish red, apex acute; lateral lobes purplish red, proximal margin not inflexed</td>
</tr>
<tr id="3514804CFFA30073BBDBC305FA10F954" box="[144,1431,1621,1715]" gridrow="6" pageId="2" pageNumber="297">
<th id="76C5E930FFA30073BBDBC305FE92F954" box="[144,277,1621,1715]" gridcol="0" gridrow="6" pageId="2" pageNumber="297">Spur</th>
<td id="76C5E930FFA30073BA1AC305FCF6F954" box="[337,881,1621,1715]" gridcol="1" gridrow="6" pageId="2" pageNumber="297">cylindroid, yellowish green to yellow; longitudinal septum tiny; callus nearly ligulate-linear, slightly widened toward upper part</td>
<td id="76C5E930FFA30073B8C0C305FA10F954" box="[907,1431,1621,1715]" gridcol="2" gridrow="6" pageId="2" pageNumber="297">somewhat horn-shaped, purplish red; longitudinal septum obvious; callus 3-lobed</td>
</tr>
<tr id="3514804CFFA30073BBDBC394FA10F93D" box="[144,1431,1732,1754]" gridrow="7" pageId="2" pageNumber="297">
<th id="76C5E930FFA30073BBDBC394FE92F93D" box="[144,277,1732,1754]" gridcol="0" gridrow="7" pageId="2" pageNumber="297">Pollinia</th>
<td id="76C5E930FFA30073BA1AC394FCF6F93D" box="[337,881,1732,1754]" gridcol="1" gridrow="7" pageId="2" pageNumber="297">spherical</td>
<td id="76C5E930FFA30073B8C0C394FA10F93D" box="[907,1431,1732,1754]" gridcol="2" gridrow="7" pageId="2" pageNumber="297">pyriform</td>
</tr>
</table>
</paragraph>
<subSubSection id="C33ED185FFA3FF88BBF6C25AFC6BF89D" lastPageId="4" lastPageNumber="299" pageId="2" pageNumber="297" type="distribution">
<paragraph id="8B9B820EFFA3FF8EBBF6C25AFD5DF852" blockId="2.[136,1452,1802,2081]" pageId="2" pageNumber="297">
<emphasis id="B9505E1CFFA3FF8EBBF6C25AFE6AF8C3" bold="true" box="[189,493,1802,1828]" pageId="2" pageNumber="297">Distribution and ecology:</emphasis>
— 
<taxonomicName id="4C24F98DFFA3FF8EB942C25AFC95F8C3" authorityName="Ying Qin &amp; Yan Liu" authorityYear="2023" box="[521,786,1802,1828]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="mulunense">
<emphasis id="B9505E1CFFA3FF8EB942C25AFC95F8C3" box="[521,786,1802,1828]" italics="true" pageId="2" pageNumber="297">Cleisostoma mulunense</emphasis>
</taxonomicName>
is known only from the limestone areas of Mulun National Nature Reserve, 
<collectingRegion id="49E04CECFFA3FF8EBA0BC27EFE24F8AF" box="[320,419,1838,1864]" country="China" name="Guangxi" pageId="2" pageNumber="297">Guangxi</collectingRegion>
, 
<collectingCountry id="F333C29EFFA3FF8EBAE6C27EFE73F8AF" box="[429,500,1838,1864]" name="China" pageId="2" pageNumber="297">China</collectingCountry>
. This new species is an epiphytic plant on trees with an elevation of ca. 
<specimenCount id="9D224987FFA3FF8EBFB4C27EFACCF8AE" box="[1279,1355,1838,1865]" count="983" pageId="2" pageNumber="297" type="generic">
<quantity id="4CDC2FEBFFA3FF8EBFB4C27EFAC0F8AE" box="[1279,1351,1838,1865]" metricMagnitude="2" metricUnit="m" metricValue="9.83" pageId="2" pageNumber="297" unit="m" value="983.0">983 m</quantity>
.
</specimenCount>
And it is companied by 
<taxonomicName id="4C24F98DFFA3FF8EBA66C202FDABF88B" box="[301,556,1874,1900]" class="Magnoliopsida" family="Juglandaceae" genus="Platycarya" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="strobilacea">
<emphasis id="B9505E1CFFA3FF8EBA66C202FDABF88B" box="[301,556,1874,1900]" italics="true" pageId="2" pageNumber="297">Platycarya strobilacea</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EB971C202FC92F88B" box="[570,789,1874,1900]" class="Magnoliopsida" family="Rosaceae" genus="Eriobotrya" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="seguinii">
<emphasis id="B9505E1CFFA3FF8EB971C202FC92F88B" box="[570,789,1874,1900]" italics="true" pageId="2" pageNumber="297">Eriobotrya seguinii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EB869C202FC77F88B" box="[802,1008,1874,1900]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="sycopseoides">
<emphasis id="B9505E1CFFA3FF8EB869C202FC77F88B" box="[802,1008,1874,1900]" italics="true" pageId="2" pageNumber="297">Acer sycopseoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EB8B5C202FB6DF88B" box="[1022,1258,1874,1900]" class="Pinopsida" family="Cupressaceae" genus="Calocedrus" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="rupestris">
<emphasis id="B9505E1CFFA3FF8EB8B5C202FB6DF88B" box="[1022,1258,1874,1900]" italics="true" pageId="2" pageNumber="297">Calocedrus rupestris</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EBFB3C202FEA0F877" class="Magnoliopsida" family="Sapotaceae" genus="Sinosideroxylon" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="pedunculatum">
<emphasis id="B9505E1CFFA3FF8EBFB3C202FEA0F877" italics="true" pageId="2" pageNumber="297">Sinosideroxylon pedunculatum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EBA7CC227FDA8F877" box="[311,559,1910,1937]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="phillyreoides">
<emphasis id="B9505E1CFFA3FF8EBA7CC227FDA8F877" box="[311,559,1910,1937]" italics="true" pageId="2" pageNumber="297">Quercus phillyreoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EB90BC226FD54F877" box="[576,723,1910,1936]" class="Magnoliopsida" family="Violaceae" genus="Viola" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="shiweii">
<emphasis id="B9505E1CFFA3FF8EB90BC226FD54F877" box="[576,723,1910,1936]" italics="true" pageId="2" pageNumber="297">Viola shiweii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EB9AFC226FC09F877" box="[740,910,1910,1936]" class="Magnoliopsida" family="Apocynaceae" genus="Alyxia" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="sinensis">
<emphasis id="B9505E1CFFA3FF8EB9AFC226FC09F877" box="[740,910,1910,1936]" italics="true" pageId="2" pageNumber="297">Alyxia sinensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EB8D5C226FB34F877" box="[926,1203,1910,1936]" class="Liliopsida" family="Orchidaceae" genus="Cymbidium" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="floribundum">
<emphasis id="B9505E1CFFA3FF8EB8D5C226FB34F877" box="[926,1203,1910,1936]" italics="true" pageId="2" pageNumber="297">Cymbidium floribundum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EBF8FC227FA23F877" box="[1220,1444,1910,1936]" class="Liliopsida" family="Orchidaceae" genus="Luisia" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="appressifolia">
<emphasis id="B9505E1CFFA3FF8EBF8FC227FA23F877" box="[1220,1444,1910,1936]" italics="true" pageId="2" pageNumber="297">Luisia appressifolia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EBBC3C2CAFE46F853" box="[136,449,1946,1972]" class="Liliopsida" family="Orchidaceae" genus="Paphiopedilum" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="micranthum">
<emphasis id="B9505E1CFFA3FF8EBBC3C2CAFE46F853" box="[136,449,1946,1972]" italics="true" pageId="2" pageNumber="297">Paphiopedilum micranthum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C24F98DFFA3FF8EBA84C2CAFD23F853" box="[463,676,1946,1972]" class="Liliopsida" family="Orchidaceae" genus="Pholidota" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="longipes">
<emphasis id="B9505E1CFFA3FF8EBA84C2CAFD23F853" box="[463,676,1946,1972]" italics="true" pageId="2" pageNumber="297">Pholidota longipes</emphasis>
</taxonomicName>
, etc.
</paragraph>
<paragraph id="8B9B820EFFA3FF8FBBF6C2EEFB8DFF3B" blockId="2.[136,1452,1802,2081]" lastBlockId="3.[136,1451,158,221]" lastPageId="3" lastPageNumber="298" pageId="2" pageNumber="297">
<emphasis id="B9505E1CFFA3FF8EBBF6C2EEFE3EF83F" bold="true" box="[189,441,1982,2008]" pageId="2" pageNumber="297">Conservation status:</emphasis>
—After many field surveys, only one population of 
<taxonomicName id="4C24F98DFFA3FF8EBF61C2EEFABEF83F" authorityName="Ying Qin &amp; Yan Liu" authorityYear="2023" box="[1066,1337,1982,2008]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="mulunense">
<emphasis id="B9505E1CFFA3FF8EBF61C2EEFABEF83F" box="[1066,1337,1982,2008]" italics="true" pageId="2" pageNumber="297">Cleisostoma mulunense</emphasis>
</taxonomicName>
has been discovered with 73 mature individuals in Mulun National Nature Reserve. Its estimated area of occupancy is less than 
<quantity id="4CDC2FEBFFA3FF8EBBC3CD56FF48F7C7" box="[136,207,2054,2080]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="297" unit="km" value="10.0">10 km</quantity>
<superScript id="7C512F46FFA3FF8EBB84CD56FF50F7F3" attach="left" box="[207,215,2054,2068]" fontSize="6" pageId="2" pageNumber="297">2</superScript>
. And this species is mainly threatened by natural droughts, which cause death to some individuals. According to Guidelines for using the IUCN Red List Categories and Criteria (
<bibRefCitation id="EFB5FFFFFFA2FF8FB835C5CEFA18FF5F" author="IUCN Standards &amp; Petitions Committee" box="[894,1439,158,184]" pageId="3" pageNumber="298" refId="ref2962" refString="IUCN Standards and Petitions Committee (2022) Guidelines for using the IUCN Red List Categories and Criteria. Version 15.1. Prepared by the Standards and Petitions Committee. [https: // www. iucnredlist. org / documents / RedListGuidelines. pdf]" type="url" year="2022">IUCN Standards and Petitions Committee, 2022</bibRefCitation>
), this species should be assessed as Critically Endangered [CR B2ab(V); C2a(ii)].
</paragraph>
<caption id="DF5BD286FFA2FF8FBBC3C2F4FA9EF7C1" ID-DOI="http://doi.org/10.5281/zenodo.10146266" ID-Zenodo-Dep="10146266" httpUri="https://zenodo.org/record/10146266/files/figure.png" pageId="3" pageNumber="298" startId="3.[136,229,1956,1978]" targetBox="[198,1390,245,1931]" targetPageId="3" targetType="figure">
<paragraph id="8B9B820EFFA2FF8FBBC3C2F4FA9EF7C1" blockId="3.[136,1452,1956,2086]" pageId="3" pageNumber="298">
<emphasis id="B9505E1CFFA2FF8FBBC3C2F4FF7AF85E" bold="true" box="[136,253,1956,1978]" pageId="3" pageNumber="298">FIGURE 2.</emphasis>
<emphasis id="B9505E1CFFA2FF8FBA49C2F4FE6BF85E" box="[258,492,1956,1977]" italics="true" pageId="3" pageNumber="298">
<taxonomicName id="4C24F98DFFA2FF8FBA49C2F4FE6FF85E" authorityName="Ying Qin &amp; Yan Liu" authorityYear="2023" box="[258,488,1956,1977]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="298" phylum="Tracheophyta" rank="species" species="mulunense">Cleisostoma mulunense</taxonomicName>
.
</emphasis>
A. Habitat. B. Flower, side view. C. Flower, front view. D. Lip, longitudinal section. E. Dorsal sepal, lateral sepals, petals, lip, column and ovary. F. Callus, side view. G. Callus, front view. H. Lower part of callus. I. Papillae at mouth of spur inside. J. Longitudinal ridge of the disk, side view. K. Longitudinal septum of spur. L. Leaf, cross section. M. Leaves, adaxial view and side view. Photos by Yu-Lan Su (A &amp; B), Yan Liu (C) and Ying Qin (D–M) from living plants of the holotype population.
</paragraph>
</caption>
<caption id="DF5BD286FFA5FF88BBC3C3DBFBA5F90E" ID-DOI="http://doi.org/10.5281/zenodo.10146268" ID-Zenodo-Dep="10146268" httpUri="https://zenodo.org/record/10146268/files/figure.png" pageId="4" pageNumber="299" startId="4.[136,229,1675,1697]" targetBox="[158,1430,190,1651]" targetPageId="4" targetType="figure">
<paragraph id="8B9B820EFFA5FF88BBC3C3DBFBA5F90E" blockId="4.[136,1452,1675,1769]" pageId="4" pageNumber="299">
<emphasis id="B9505E1CFFA5FF88BBC3C3DBFF7AF947" bold="true" box="[136,253,1675,1697]" pageId="4" pageNumber="299">FIGURE 3.</emphasis>
<emphasis id="B9505E1CFFA5FF88BA49C3DBFD84F947" box="[258,515,1675,1696]" italics="true" pageId="4" pageNumber="299">
<taxonomicName id="4C24F98DFFA5FF88BA49C3DBFE78F947" box="[258,511,1675,1696]" class="Liliopsida" family="Orchidaceae" genus="Cleisostoma" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="menghaiense">Cleisostoma menghaiense</taxonomicName>
.
</emphasis>
A. Habit. B. Inflorescence. C. Flower, side view. D. Flower, front view. E. Flower with sepals and petals removed, side view. F. Leaves, adaxial view and side view. G. Leaf, cross section. H. Callus, front view. I. Callus, side view. J. Lip, longitudinal section. Photos by Ying Qin from a living plant in Mulun National Nature Reserve.
</paragraph>
</caption>
<paragraph id="8B9B820EFFA5FF88BBF6C26BFA2CF8B1" blockId="4.[136,1451,1851,1914]" box="[189,1451,1851,1878]" pageId="4" pageNumber="299">
<materialsCitation id="3B4C8853FFA5FF88BBF6C26BFA2CF8B1" box="[189,1451,1851,1878]" country="China" county="Huanjiang County" location="Mulun National Nature" pageId="4" pageNumber="299" specimenCount="2" stateProvince="Guangxi">
<emphasis id="B9505E1CFFA5FF88BBF6C26BFD4BF8B2" bold="true" box="[189,716,1851,1877]" pageId="4" pageNumber="299">
<typeStatus id="549F3CACFFA5FF88BBF6C26BFDBBF8B2" box="[189,572,1851,1877]" isOtherTypeStatus="true" pageId="4" pageNumber="299">Additional specimens examined</typeStatus>
(
<typeStatus id="549F3CACFFA5FF88B906C26BFD38F8B2" box="[589,703,1851,1877]" pageId="4" pageNumber="299" type="paratype">paratype</typeStatus>
):
</emphasis>
— 
<collectingCountry id="F333C29EFFA5FF88B9A1C26CFCC2F8B1" box="[746,837,1852,1878]" name="China" pageId="4" pageNumber="299">CHINA</collectingCountry>
. 
<collectingRegion id="49E04CECFFA5FF88B81AC26CFC31F8B1" box="[849,950,1852,1878]" country="China" name="Guangxi" pageId="4" pageNumber="299">Guangxi</collectingRegion>
: 
<collectingCounty id="62FAFA82FFA5FF88B888C26CFB12F8B1" box="[963,1173,1852,1878]" pageId="4" pageNumber="299">Huanjiang County</collectingCounty>
, 
<location id="8EFBD4D5FFA5FF88BFE9C26CFA2CF8B1" LSID="urn:lsid:plazi:treatment:038D3318FFA1FF88BBC3C3A4FC6BF89D:8EFBD4D5FFA5FF88BFE9C26CFA2CF8B1" box="[1186,1451,1852,1878]" country="China" county="Huanjiang County" name="Mulun National Nature" pageId="4" pageNumber="299" stateProvince="Guangxi">Mulun National Nature</location>
</materialsCitation>
</paragraph>
<paragraph id="8B9B820EFFA5FF88BBC3C230FC6BF89D" blockId="4.[136,1451,1851,1914]" box="[136,1004,1888,1914]" pageId="4" pageNumber="299">
<materialsCitation id="3B4C8853FFA5FF88BBC3C230FC6FF89D" box="[136,1000,1888,1914]" collectingDate="2023-07-27" collectionCode="IBK" elevation="983" pageId="4" pageNumber="299" specimenCode="GXQY20230727001" specimenCount="1">
Reserve, elev. ca. 
<quantity id="4CDC2FEBFFA5FF88BA19C230FE1CF89D" box="[338,411,1888,1914]" metricMagnitude="2" metricUnit="m" metricValue="9.83" pageId="4" pageNumber="299" unit="m" value="983.0">
<elevation id="0009653DFFA5FF88BA19C230FE1CF89D" box="[338,411,1888,1914]" metricMagnitude="2" metricUnit="m" metricValue="9.83" pageId="4" pageNumber="299" unit="m" value="983.0">983 m</elevation>
</quantity>
, 
<date id="FF9AA4CEFFA5FF88BAECC230FDBDF89D" box="[423,570,1888,1914]" pageId="4" pageNumber="299" value="2023-07-27">
<collectingDate id="EFDE5D26FFA5FF88BAECC230FDBDF89D" box="[423,570,1888,1914]" pageId="4" pageNumber="299" value="2023-07-27">27 July 2023</collectingDate>
</date>
, 
<emphasis id="B9505E1CFFA5FF88B90EC231FC15F89D" box="[581,914,1888,1914]" italics="true" pageId="4" pageNumber="299">
Ying Qin 
<specimenCode id="DB822A75FFA5FF88B9E4C230FC15F89D" box="[687,914,1888,1914]" collectionCode="GXQY" pageId="4" pageNumber="299">GXQY20230727001</specimenCode>
</emphasis>
(
<collectionCode id="ED351ACBFFA5FF88B8E9C230FC5BF89D" box="[930,988,1888,1914]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13150" lsid="urn:lsid:biocol.org:col:13150" name="Guangxi Institute of Botany" pageId="4" pageNumber="299" type="Herbarium">IBK</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>