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<document id="992317CD5FF392B88B8B4C26B1F2DB83" ID-DOI="10.11646/zootaxa.2262.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5306371" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629652864432" checkinUser="felipe" docAuthor="Reyes, Javier, Santodomingo, Nadiezhda &amp; Cairns, Stephen" docDate="2009" docId="4B4287D6FF8DDC30A187F9E8FA2FF859" docLanguage="en" docName="zt02262p039.pdf" docOrigin="Zootaxa 2262 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2262.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Heterocyathus antoniae Reyes &amp; Santodomingo &amp; Cairns 2009, new species" docType="treatment" docVersion="4" lastPageNumber="29" masterDocId="B77BFFAEFF97DC2CA110FFBEFFB7FA6D" masterDocTitle="Caryophylliidae (Scleractinia) from the Colombian Caribbean" masterLastPageNumber="39" masterPageNumber="1" pageNumber="27" updateTime="1699089311122" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D971B445CEB4F3ACC162E0CACFDA4752">Caryophylliidae (Scleractinia) from the Colombian Caribbean</mods:title>
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<mods:namePart id="6CD43146B920361429123108333AFB93">Reyes, Javier</mods:namePart>
<mods:affiliation id="D10E77E883FAC8A0A1CCC6540D1CDA35">Instituto de Investigaciones Marinas y Costeras, INVEMAR, Cerro de Punta Betín, AA 1016 Santa Marta, Colombia. E-mail: j _ reyes _ forero @ hotmail. com, nsantodomingo @ gmail. com National Museum of Natural History, Naturalis, PO Box 9517, 2300 RA Leiden, The Netherlands National Museum of Natural History, Washington, D. C. 20013, U. S. A. E-mail: cairnss @ si. edu</mods:affiliation>
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<mods:namePart id="EAD62BF7866F53027F6499D3598B7102">Santodomingo, Nadiezhda</mods:namePart>
<mods:affiliation id="EE21E20D6FC612AB9982E4251F67A484">Instituto de Investigaciones Marinas y Costeras, INVEMAR, Cerro de Punta Betín, AA 1016 Santa Marta, Colombia. E-mail: j _ reyes _ forero @ hotmail. com, nsantodomingo @ gmail. com National Museum of Natural History, Naturalis, PO Box 9517, 2300 RA Leiden, The Netherlands National Museum of Natural History, Washington, D. C. 20013, U. S. A. E-mail: cairnss @ si. edu</mods:affiliation>
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<mods:namePart id="70FDEE4E24273C1118200C57DC09EE81">Cairns, Stephen</mods:namePart>
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<paragraph id="C35436C0FF8DDC36A187F9E8FDE1FC1D" blockId="26.[151,598,1622,1683]" box="[151,598,1622,1648]" pageId="26" pageNumber="27">
<heading id="981C81ACFF8DDC36A187F9E8FDE1FC1D" bold="true" box="[151,598,1622,1648]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="F19FEAD2FF8DDC36A187F9E8FDE1FC1D" bold="true" box="[151,598,1622,1648]" pageId="26" pageNumber="27">
<emphasis id="F19FEAD2FF8DDC36A187F9E8FE08FC1D" bold="true" box="[151,447,1622,1648]" italics="true" pageId="26" pageNumber="27">
<taxonomicName id="04EB4D43FF8DDC36A187F9E8FE0EFC1D" ID-CoL="3L92S" authority="Reyes &amp; Santodomingo &amp; Cairns, 2009" authorityName="Reyes &amp; Santodomingo &amp; Cairns" authorityYear="2009" box="[151,441,1622,1648]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="26" pageNumber="27" phylum="Cnidaria" rank="species" species="antoniae" status="sp. nov.">Heterocyathus antoniae</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="EAAC57A9FF8DDC36A0D6F9E8FDE1FC1D" box="[454,598,1622,1648]" pageId="26" pageNumber="27" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="8BF1654BFF8DDC37A187F9C7FF69F9DA" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="C35436C0FF8DDC36A187F9C7FE97FCFE" blockId="26.[151,598,1622,1683]" box="[151,288,1657,1683]" pageId="26" pageNumber="27">
<figureCitation id="5BD02A45FF8DDC36A187F9C7FE97FCFE" box="[151,288,1657,1683]" captionStart="FIGURE 4" captionStartId="22.[151,256,1591,1615]" captionTargetBox="[263,1334,182,1568]" captionTargetId="figure-278@22.[263,1334,182,1568]" captionTargetPageId="22" captionText="FIGURE 4. A, Lophelia pertusa, INV CNI396, off La Guajira (Honda Bay), colony fragments, x 1; BC, Oxysmilia rotundifolia, INV CNI687, off San Bernardo Islands, calicular and lateral views, both x 1.1; DE, Colangia immersa, INV CNI683, off La Guajira (Punta Gallinas), calicular and lateral views, both x 3; FG, Phyllangia americana americana, INV CNI274, off La Guajira (Caricari Point), colony from calicular and oblique views, both x 2; HI, Rhizosmilia maculata, INV CNI178, off Santa Marta, calicular and lateral views, both x 3.6; JK, Phacelocyathus flos, INV CNI2940, off San Bernardo Islands, calicular and lateral views, both x 3.6; LM, Anomocora fecunda INV CNI2897 off San Bernardo Islands, calicular and lateral views, x 3.2, x 1, respectively; NO, Anomocora prolifera, INV CNI2725, off San Bernardo Islands, calicular view showing rejuvenescence stage and lateral view, both x 4.4; PQ, Anomocora marchadi, INV CNI2303, off La Guajira, calicular and lateral views, both x 3.6; R, Coenosmilia arbuscula, INV CNI855, off San Bernardo Islands, branching pattern, x 1.5; ST, Heterocyathus antoniae, paratype, INV CNI2076, off Manaure, calicular and lateral views, both x 4.5; UW, Heterocyathus antoniae, INV CNI740, holotype, off Santa Marta (Neguanje Bay): U, basal view showing sipunculid holes, V, calicular view, and W, lateral view, all x 4.5. Scale bars for AC, M, R = 1 cm; DL, NQ, SW = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5306379" httpUri="https://zenodo.org/record/5306379/files/figure.png" pageId="26" pageNumber="27">Figs. 4SW</figureCitation>
</paragraph>
<paragraph id="C35436C0FF8DDC37A187F978FB1CFB8A" blockId="26.[151,1436,1734,2031]" lastBlockId="27.[151,1436,152,2034]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="F19FEAD2FF8DDC36A187F978FE81FC8D" bold="true" box="[151,310,1734,1760]" pageId="26" pageNumber="27">Description:</emphasis>
Coralla subcylindrical.
<typeStatus id="1C508862FF8DDC36A34FF978FD78FC8D" box="[607,719,1734,1760]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
is
<quantity id="04139B25FF8DDC36A3E5F978FCFDFC8D" box="[757,842,1734,1760]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="mm" value="10.0">10 mm</quantity>
in GCD, 8.8 mm in LCD and 9.0 mm in height. Corallites initially settled on gastropod shells, which are overgrown during the coral development, and in some specimens the coral skeleton practically covered all the mollusc shell. The
<typeStatus id="1C508862FF8DDC36A552F8AAFB1FFD43" box="[1090,1192,1812,1838]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
base (8.1 x 10.6 mm in BD) displays five holes (sipunculid openings): four of them are aligned with the LCD at the middle part of the base (from the right to the left 0.8, 0.7, 0.2 and 0.6 mm in diameter), while the fifth one is located at the end of a conical shaped pipe (
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diameter) and it is projected out
<quantity id="04139B25FF8DDC36A2A5F836FC4FFDCF" box="[949,1016,1928,1954]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="mm" value="2.0">2 mm</quantity>
and curved 30 from the thecal wall. Internal walls of the holes are covered by a thin, smooth and porcelain-like layer (0.2 mm thick). In the
<typeStatus id="1C508862FF8DDC36A187F86BFF49FD82" box="[151,254,2005,2031]" pageId="26" pageNumber="27" type="paratype">paratype</typeStatus>
specimen only two basal pores were observed. Some sabellid polychaete tubes are attached to the base in the
<typeStatus id="1C508862FF8CDC37A03BFF26FE2EFADF" box="[299,409,152,178]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
specimen. Base is covered by high, conical, almost cylindrical or bent granules perpendicularly oriented. Ridged costae equal in height and width, prominent. Coarse costal granulation develops from secondary accretion of a central rod-shape element, and flanked by two slender spines diagonally oriented against that central element. Lateral spines of C4 flank the C1-C2, and they are twice or three times higher than the external C4 spines. Intercostal furrows are deep and slender, sometimes with thecal pores connecting to the interior of the coralla. Calicular margin is lanceolated: S1 are the most exsert septa projecting up to 1.2 mm from the calicular margin adjacent to S3. Despite S1 are the most exsert septa, the distance from the thecal edge to the septal lobe is slightly longer for the S3 (S1 up to 1.2 mm, S2 up to
<quantity id="04139B25FF8CDC37A440FE18FA20FBAD" box="[1360,1431,422,448]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="28" unit="mm" value="1.0">1 mm</quantity>
, S3 up to 1.3 mm). Septal ornamentation is twice to three times its respective septa width.
</paragraph>
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Septa hexamerally arranged up to five cycles, following a Pourtalès plan (S1=S2=S4&gt;&gt;S3); the fifth cycle incomplete, with two S5 associated to the principal axis. Septal axial edges are smooth and straight; deep and narrow palar notches extend just below the columella. S1 are the only independent septa, reaching the columella. Paliform lobes occur before all septa (P1 to P4), except for the additional S5; P4 composed of up to six palar teeth, gradually smaller while closer to the columella. P3 is composed of one or two lobes, twice the width of S1; P2 is twice to three times the width of P1, and P1 bearing three to four small lobes. Septacolumella junctions are fenestrated. The secondary accretion of septal ornamentation is connected by cylindrical synapticulae to the opposite septal-palar faces near the columella. Septal and palar granulation is laterally fused in carinae, perpendicular to the trabecular axis. Columella is spongy and elliptical (1.8 x 1.1 mm), composed of crispate elements, laterally fused. Fossa is shallow in the
<typeStatus id="1C508862FF8CDC37A572FCEEFB65F907" box="[1122,1234,848,874]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
, but deep in the
<typeStatus id="1C508862FF8CDC37A187FCC8FEB7F9FD" box="[151,256,886,912]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
. Corallum present a white colour, but some specimens exhibit a brown or purple columella and septal bases.
</paragraph>
</subSubSection>
<subSubSection id="8BF1654BFF8CDC37A1D5FC7DFEC1FDC8" pageId="27" pageNumber="28" type="discussion">
<paragraph id="C35436C0FF8CDC37A1D5FC7DFEC1FDC8" blockId="27.[151,1436,152,2034]" pageId="27" pageNumber="28">
<emphasis id="F19FEAD2FF8CDC37A1D5FC7DFEF5F9B0" bold="true" box="[197,322,963,989]" pageId="27" pageNumber="28">Remarks:</emphasis>
The genus
<taxonomicName id="04EB4D43FF8CDC37A0C1FC7AFDC8F9B0" box="[465,639,964,989]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="genus">
<emphasis id="F19FEAD2FF8CDC37A0C1FC7AFDC8F9B0" box="[465,639,964,989]" italics="true" pageId="27" pageNumber="28">Heterocyathus</emphasis>
</taxonomicName>
comprises 16 extant and 5 fossil nominal species (
<bibRefCitation id="A77A4B31FF8CDC37A5F9FC7AFF6FFE69" author="Stolarski, J. &amp; Zibrowius, H. &amp; Loser, H." pageId="27" pageNumber="28" pagination="309 - 330" refId="ref28845" refString="Stolarski, J. Zibrowius, H. &amp; Loser, H. (2001) Antiquity of the scleractinian-sipunculan symbiosis. Acta Palaeontologica Polonica, 46 (3), 309 - 330." type="journal article" year="2001">
Stolarski
<emphasis id="F19FEAD2FF8CDC37A44FFC7AFA2BF9B0" box="[1375,1436,964,989]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2001
</bibRefCitation>
). The extant species were grouped into three valid species by
<bibRefCitation id="A77A4B31FF8CDC37A2DDFC54FAB1FE69" author="Hoeksema, B. W. &amp; Best, M. B." box="[973,1286,1002,1028]" pageId="27" pageNumber="28" pagination="221 - 245" refId="ref26318" refString="Hoeksema, B. W. &amp; Best, M. B. 1991. New observations on scleractinian corals from Indonesia: 2. Sipunculan- associated species belonging to the genera Heterocyathus and Heteropsammia. Zoologische Mededelingen, 65, 221 - 245." type="journal article" year="1991">Hoeksema &amp; Best (1991)</bibRefCitation>
:
<taxonomicName id="04EB4D43FF8CDC37A403FC54FEFCFE46" authority="(Verrill, 1866)" baseAuthorityName="Verrill" baseAuthorityYear="1866" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="sulcatus">
<emphasis id="F19FEAD2FF8CDC37A403FC54FA2BFE6E" box="[1299,1436,1002,1027]" italics="true" pageId="27" pageNumber="28">H. sulcatus</emphasis>
(
<bibRefCitation id="A77A4B31FF8CDC37A18FFBAFFEF5FE46" author="Verrill, A. E." box="[159,322,1041,1067]" pageId="27" pageNumber="28" pagination="17 - 32" refId="ref29213" refString="Verrill, A. E. (1866) Synopsis of the polyps and corals of the North Pacific Explorig Expedition, 1853 - 1856, with descriptions of some additional species from the west coast of North America. Part III. Madreporaria. Proceedings and Communications of the Essex Institute, 5, 17 - 32, pl. 1 - 2." type="journal article" year="1866">Verrill, 1866</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="04EB4D43FF8CDC37A049FBAFFC39FE46" authority="M. Edwards &amp; Haime, 1848" authorityName="M. Edwards &amp; Haime" authorityYear="1848" box="[345,910,1041,1067]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="aequicostatus">
<emphasis id="F19FEAD2FF8CDC37A049FBAFFD90FE47" box="[345,551,1041,1066]" italics="true" pageId="27" pageNumber="28">H. aequicostatus</emphasis>
M.
<bibRefCitation id="A77A4B31FF8CDC37A34DFBAFFC39FE46" author="Milne Edwards, H. &amp; Haime, J." box="[605,910,1041,1067]" pageId="27" pageNumber="28" pagination="209 - 320" refId="ref27026" refString="Milne Edwards, H. &amp; Haime, J. (1848) Recherches sur les polypiers. Memoire 4 (1). Monographie des Astreides. Annales des Sciences Naturelles, 10 (3), 209 - 320, pls. 5 - 9." type="journal article" year="1848">Edwards &amp; Haime, 1848</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="04EB4D43FF8CDC37A2DCFBAFFAAFFE46" authority="Verrill, 1865" authorityName="Verrill" authorityYear="1865" box="[972,1304,1041,1067]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="alternatus">
<emphasis id="F19FEAD2FF8CDC37A2DCFBAFFBD8FE47" box="[972,1135,1041,1066]" italics="true" pageId="27" pageNumber="28">H. alternatus</emphasis>
<bibRefCitation id="A77A4B31FF8CDC37A568FBAFFAAFFE46" author="Verrill, A. E." box="[1144,1304,1041,1067]" pageId="27" pageNumber="28" pagination="145 - 152" refId="ref29160" refString="Verrill, A. E. (1865) Classification of polyps: Extract condensed from a synopsis of the polypi of the North Pacific Exploring Expedition under Captains Ringgold and Rogers, U. S. N. Proceedings and Communications of the Essex Institute, 4, 145 - 152." type="journal article" year="1865">Verrill, 1865</bibRefCitation>
</taxonomicName>
. However,
<bibRefCitation id="A77A4B31FF8CDC37A187FB86FEDAFE3F" author="Zibrowius, H." box="[151,365,1080,1106]" pageId="27" pageNumber="28" pagination="319 - 340" refId="ref29780" refString="Zibrowius, H. (1998) A new type of symbiosis: Heterocyathus japonicus (Cnidaria: Scleractinia) living on Fissidentalium vernedei (Mollusca: Scaphopoda). Zoologische Verhandelingen, 323 (31), 319 - 340." type="journal article" year="1998">Zibrowius (1998)</bibRefCitation>
and
<bibRefCitation id="A77A4B31FF8CDC37A0BAFB86FDD5FE3F" author="Cairns, S. D." box="[426,610,1080,1106]" pageId="27" pageNumber="28" pagination="31 - 167" refId="ref24733" refString="Cairns, S. D. (1999 b) Cnidaria Anthozoa: Deep-water azooxanthellate Scleractinia from Vanuatu, and Wallis and Futuna Islands. Memoires du Museum National d'Histoire Naturelle, 180, 31 - 167." type="journal article" year="1999">Cairns (1999b)</bibRefCitation>
suggest that a more detailed study should be performed in order to clarify the genus taxonomy. Following
<bibRefCitation id="A77A4B31FF8CDC37A375FBE0FC24FE15" author="Hoeksema, B. W. &amp; Best, M. B." box="[613,915,1118,1144]" pageId="27" pageNumber="28" pagination="221 - 245" refId="ref26318" refString="Hoeksema, B. W. &amp; Best, M. B. 1991. New observations on scleractinian corals from Indonesia: 2. Sipunculan- associated species belonging to the genera Heterocyathus and Heteropsammia. Zoologische Mededelingen, 65, 221 - 245." type="journal article" year="1991">Hoeksema &amp; Best (1991)</bibRefCitation>
,
<taxonomicName id="04EB4D43FF8CDC37A2B0FBE1FB10FE15" box="[928,1191,1118,1144]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="F19FEAD2FF8CDC37A2B0FBE1FBFDFE15" box="[928,1098,1119,1144]" italics="true" pageId="27" pageNumber="28">Heterocyathus</emphasis>
species
</taxonomicName>
are differentiated by the septal arrangement, lateral septal projection, and the coloration pattern:
<taxonomicName id="04EB4D43FF8CDC37A53AFB3BFB06FEF3" baseAuthorityName="Verrill" baseAuthorityYear="1866" box="[1066,1201,1157,1182]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="sulcatus">
<emphasis id="F19FEAD2FF8CDC37A53AFB3BFB06FEF3" box="[1066,1201,1157,1182]" italics="true" pageId="27" pageNumber="28">H. sulcatus</emphasis>
</taxonomicName>
is characterized by having a central part of the calices with dark brown or black colour, and short lateral septal projections;
<taxonomicName id="04EB4D43FF8CDC37A46FFB12FE8DFE81" authorityName="M. Edwards &amp; Haime" authorityYear="1848" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="aequicostatus">
<emphasis id="F19FEAD2FF8CDC37A46FFB12FE8DFE81" italics="true" pageId="27" pageNumber="28">H. aequicostatus</emphasis>
</taxonomicName>
presents a compact corallum, white calices, long lateral septal projections and closely packed pali; and
<taxonomicName id="04EB4D43FF8CDC37A013FB47FE2BFF7F" authorityName="Verrill" authorityYear="1865" box="[259,412,1273,1298]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="alternatus">
<emphasis id="F19FEAD2FF8CDC37A013FB47FE2BFF7F" box="[259,412,1273,1298]" italics="true" pageId="27" pageNumber="28">H. alternatus</emphasis>
</taxonomicName>
has spaced S1, white calices, and closely packed pali. Colombian
<taxonomicName id="04EB4D43FF8CDC37A5B2FB47FAFBFF7F" box="[1186,1356,1273,1298]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="genus">
<emphasis id="F19FEAD2FF8CDC37A5B2FB47FAFBFF7F" box="[1186,1356,1273,1298]" italics="true" pageId="27" pageNumber="28">Heterocyathus</emphasis>
</taxonomicName>
shares some characters with these three species: for instance,
<specimenCount id="D5EDFD49FF8CDC37A23FFA9EFBF4FF57" box="[815,1091,1312,1338]" pageId="27" pageNumber="28" type="generic" typeStatus="paratype">one paratype specimen</specimenCount>
(INV CNI2076) has a black columella and long lateral septal projections as
<taxonomicName id="04EB4D43FF8CDC37A3F3FAF9FCDAFF0D" baseAuthorityName="Verrill" baseAuthorityYear="1866" box="[739,877,1351,1376]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="sulcatus">
<emphasis id="F19FEAD2FF8CDC37A3F3FAF9FCDAFF0D" box="[739,877,1351,1376]" italics="true" pageId="27" pageNumber="28">H. sulcatus</emphasis>
</taxonomicName>
; on the contrary, the
<typeStatus id="1C508862FF8CDC37A565FAF8FB68FF0D" box="[1141,1247,1350,1376]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(INV CNI740) presents a completely white corallum, and spaced S1, showing a resemblance to
<taxonomicName id="04EB4D43FF8CDC37A55CFAD3FB50FFEB" authorityName="Verrill" authorityYear="1865" box="[1100,1255,1389,1414]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="alternatus">
<emphasis id="F19FEAD2FF8CDC37A55CFAD3FB50FFEB" box="[1100,1255,1389,1414]" italics="true" pageId="27" pageNumber="28">H. alternatus</emphasis>
</taxonomicName>
.
<taxonomicName id="04EB4D43FF8CDC37A5E6FAD3FA36FFEB" authorityName="Reyes &amp; Santodomingo &amp; Cairns" authorityYear="2009" box="[1270,1409,1389,1414]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="antoniae">
<emphasis id="F19FEAD2FF8CDC37A5E6FAD3FA36FFEB" box="[1270,1409,1389,1414]" italics="true" pageId="27" pageNumber="28">H. antoniae</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="04EB4D43FF8CDC37A041FA2AFE43FFC0" authorityName="Verrill" authorityYear="1865" box="[337,500,1428,1453]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="alternatus">
<emphasis id="F19FEAD2FF8CDC37A041FA2AFE43FFC0" box="[337,500,1428,1453]" italics="true" pageId="27" pageNumber="28">H. alternatus</emphasis>
</taxonomicName>
in size (
<quantity id="04139B25FF8CDC37A371FA2AFD0FFFC3" box="[609,696,1428,1454]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="28" unit="mm" value="10.0">10 mm</quantity>
in GCD, sensu
<bibRefCitation id="A77A4B31FF8CDC37A291FA2AFB9DFFC3" author="Cairns, S. D." box="[897,1066,1428,1454]" pageId="27" pageNumber="28" pagination="31 - 167" refId="ref24733" refString="Cairns, S. D. (1999 b) Cnidaria Anthozoa: Deep-water azooxanthellate Scleractinia from Vanuatu, and Wallis and Futuna Islands. Memoires du Museum National d'Histoire Naturelle, 180, 31 - 167." type="journal article" year="1999">Cairns 1999b</bibRefCitation>
), and having a pointed costae granulation (USNM 90410, Indian Ocean,
<collectingCountry id="BBFC7650FF8CDC37A3BFFA05FCB1FFB8" box="[687,774,1467,1493]" name="Myanmar" pageId="27" pageNumber="28">Burma</collectingCountry>
,
<quantity id="04139B25FF8CDC37A203FA05FCE5FFB8" box="[787,850,1467,1493]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="27" pageNumber="28" unit="m" value="80.0">80 m</quantity>
depth); but it differs from
<taxonomicName id="04EB4D43FF8CDC37A5B7FA05FAFBFFB9" authorityName="Verrill" authorityYear="1865" box="[1191,1356,1467,1492]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="alternatus">
<emphasis id="F19FEAD2FF8CDC37A5B7FA05FAFBFFB9" box="[1191,1356,1467,1492]" italics="true" pageId="27" pageNumber="28">H. alternatus</emphasis>
</taxonomicName>
in the presence of additional pairs of S5 associated to the principal S1, a shallow fossa, slender and crispate columella elements, ridged lobe bases product of the palar granulation fusion, and by its subcylindrical coralla.
<taxonomicName id="04EB4D43FF8CDC37A1E6F991FE0CFC25" authorityName="M. Edwards &amp; Haime" authorityYear="1848" box="[246,443,1583,1608]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="aequicostatus">
<emphasis id="F19FEAD2FF8CDC37A1E6F991FE0CFC25" box="[246,443,1583,1608]" italics="true" pageId="27" pageNumber="28">H. aequicostatus</emphasis>
</taxonomicName>
(USNM 89955, Pacific Ocean,
<collectingCountry id="BBFC7650FF8CDC37A225F990FC0BFC25" box="[821,956,1582,1608]" name="Philippines" pageId="27" pageNumber="28">Philippines</collectingCountry>
,
<quantity id="04139B25FF8CDC37A2D9F990FBB2FC25" box="[969,1029,1582,1608]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="27" pageNumber="28" unit="m" value="33.0">33 m</quantity>
depth) is similar to
<taxonomicName id="04EB4D43FF8CDC37A5E2F991FACAFC25" authorityName="Reyes &amp; Santodomingo &amp; Cairns" authorityYear="2009" box="[1266,1405,1583,1608]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="antoniae">
<emphasis id="F19FEAD2FF8CDC37A5E2F991FACAFC25" box="[1266,1405,1583,1608]" italics="true" pageId="27" pageNumber="28">H. antoniae</emphasis>
</taxonomicName>
in its granular base and its costae extension up to the base edge; but, it is different in the corallum size and its low and rounded costae granulation. The characteristic of having extra pairs of S5 adjacent to the principal S1 system, is shared with
<taxonomicName id="04EB4D43FF8CDC37A0B4F91DFBB8FCD0" authority="(Verrill, 1866) sensu Zibrowius (1998)" authorityName="sensu Zibrowius" authorityYear="1998" baseAuthorityName="Verrill" baseAuthorityYear="1866" box="[420,1039,1699,1725]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="japonicus">
<emphasis id="F19FEAD2FF8CDC37A0B4F91DFD8CFCD1" box="[420,571,1699,1724]" italics="true" pageId="27" pageNumber="28">H. japonicus</emphasis>
(
<bibRefCitation id="A77A4B31FF8CDC37A35BF91DFD53FCD0" author="Verrill, A. E." box="[587,740,1699,1725]" pageId="27" pageNumber="28" pagination="17 - 32" refId="ref29213" refString="Verrill, A. E. (1866) Synopsis of the polyps and corals of the North Pacific Explorig Expedition, 1853 - 1856, with descriptions of some additional species from the west coast of North America. Part III. Madreporaria. Proceedings and Communications of the Essex Institute, 5, 17 - 32, pl. 1 - 2." type="journal article" year="1866">Verrill, 1866</bibRefCitation>
) sensu
<bibRefCitation id="A77A4B31FF8CDC37A22DF91DFBB8FCD0" author="Zibrowius, H." box="[829,1039,1699,1725]" pageId="27" pageNumber="28" pagination="319 - 340" refId="ref29780" refString="Zibrowius, H. (1998) A new type of symbiosis: Heterocyathus japonicus (Cnidaria: Scleractinia) living on Fissidentalium vernedei (Mollusca: Scaphopoda). Zoologische Verhandelingen, 323 (31), 319 - 340." type="journal article" year="1998">Zibrowius (1998)</bibRefCitation>
</taxonomicName>
, but they can be differentiated by the costal length; whilst in
<taxonomicName id="04EB4D43FF8CDC37A0C3F977FDDEFC8F" authorityName="sensu Zibrowius" authorityYear="1998" baseAuthorityName="Verrill" baseAuthorityYear="1866" box="[467,617,1737,1762]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="japonicus">
<emphasis id="F19FEAD2FF8CDC37A0C3F977FDDEFC8F" box="[467,617,1737,1762]" italics="true" pageId="27" pageNumber="28">H. japonicus</emphasis>
</taxonomicName>
the costae extend until the base center of the corallum, in
<taxonomicName id="04EB4D43FF8CDC37A402F977FA2CFC8F" authorityName="Reyes &amp; Santodomingo &amp; Cairns" authorityYear="2009" box="[1298,1435,1737,1762]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="antoniae">
<emphasis id="F19FEAD2FF8CDC37A402F977FA2CFC8F" box="[1298,1435,1737,1762]" italics="true" pageId="27" pageNumber="28">H. antoniae</emphasis>
</taxonomicName>
costae extend only to the base edge.
<taxonomicName id="04EB4D43FF8CDC37A343F94EFD56FD64" authorityName="Reyes &amp; Santodomingo &amp; Cairns" authorityYear="2009" box="[595,737,1776,1801]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="antoniae">
<emphasis id="F19FEAD2FF8CDC37A343F94EFD56FD64" box="[595,737,1776,1801]" italics="true" pageId="27" pageNumber="28">H. antoniae</emphasis>
</taxonomicName>
resembles
<taxonomicName id="04EB4D43FF8CDC37A27AF94EFC45FD64" baseAuthorityName="Verrill" baseAuthorityYear="1866" box="[874,1010,1776,1801]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="sulcatus">
<emphasis id="F19FEAD2FF8CDC37A27AF94EFC45FD64" box="[874,1010,1776,1801]" italics="true" pageId="27" pageNumber="28">H. sulcatus</emphasis>
</taxonomicName>
(USNM 90079,
<collectingCountry id="BBFC7650FF8CDC37A5AEF94EFAFFFD67" box="[1214,1352,1776,1802]" name="Philippines" pageId="27" pageNumber="28">Philippines</collectingCountry>
,
<quantity id="04139B25FF8CDC37A445F94EFA23FD67" box="[1365,1428,1776,1802]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="27" pageNumber="28" unit="m" value="33.0">33 m</quantity>
) because both species have subcylindrical coralla and similar calicular aspect, but
<taxonomicName id="04EB4D43FF8CDC37A54AF8A9FB52FD5D" authorityName="Reyes &amp; Santodomingo &amp; Cairns" authorityYear="2009" box="[1114,1253,1815,1840]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="antoniae">
<emphasis id="F19FEAD2FF8CDC37A54AF8A9FB52FD5D" box="[1114,1253,1815,1840]" italics="true" pageId="27" pageNumber="28">H. antoniae</emphasis>
</taxonomicName>
present mainly white calices and equally wide costae. Thus,
<taxonomicName id="04EB4D43FF8CDC37A3D8F883FCEEFD3B" authorityName="Reyes &amp; Santodomingo &amp; Cairns" authorityYear="2009" box="[712,857,1853,1878]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="antoniae">
<emphasis id="F19FEAD2FF8CDC37A3D8F883FCEEFD3B" box="[712,857,1853,1878]" italics="true" pageId="27" pageNumber="28">H. antoniae</emphasis>
</taxonomicName>
was established as a new species, based on its morphological characteristics and the particular geographic distribution restricted to the northern Caribbean coast of
<collectingCountry id="BBFC7650FF8CDC37A1EAF835FEC5FDC8" box="[250,370,1931,1957]" name="Colombia" pageId="27" pageNumber="28">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="8BF1654BFF8CDC30A1D5F80FFAADFB1E" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="distribution">
<paragraph id="C35436C0FF8CDC30A1D5F80FFAADFB1E" blockId="27.[151,1436,152,2034]" lastBlockId="28.[151,1436,152,564]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="F19FEAD2FF8CDC37A1D5F80FFED0FDA6" bold="true" box="[197,359,1969,1995]" pageId="27" pageNumber="28">Distribution:</emphasis>
<taxonomicName id="04EB4D43FF8CDC37A07FF80FFDC3FDA6" box="[367,628,1969,1995]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="27" pageNumber="28" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="F19FEAD2FF8CDC37A07FF80FFDAEFDA7" box="[367,537,1969,1994]" italics="true" pageId="27" pageNumber="28">Heterocyathus</emphasis>
species
</taxonomicName>
are commonly found on sandy bottoms adjacent to coral reefs of the eastern Pacific and Indian oceans, from
<collectingCountry id="BBFC7650FF8CDC37A360F866FD05FD9F" box="[624,690,2008,2034]" name="Japan" pageId="27" pageNumber="28">Japan</collectingCountry>
to
<collectingCountry id="BBFC7650FF8CDC37A3CAF866FCC2FD9F" box="[730,885,2008,2034]" name="Mozambique" pageId="27" pageNumber="28">Mozambique</collectingCountry>
(
<bibRefCitation id="A77A4B31FF8CDC37A294F866FB14FD9F" author="Hoeksema, B. W. &amp; Best, M. B." box="[900,1187,2008,2034]" pageId="27" pageNumber="28" pagination="221 - 245" refId="ref26318" refString="Hoeksema, B. W. &amp; Best, M. B. 1991. New observations on scleractinian corals from Indonesia: 2. Sipunculan- associated species belonging to the genera Heterocyathus and Heteropsammia. Zoologische Mededelingen, 65, 221 - 245." type="journal article" year="1991">Hoeksema &amp; Best 1991</bibRefCitation>
;
<bibRefCitation id="A77A4B31FF8CDC30A5A0F866FF60FADF" author="Cairns, S. D. &amp; Zibrowius, H." lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" pagination="27 - 243" refId="ref24808" refString="Cairns, S. D. &amp; Zibrowius, H. (1997) Cnidaria Anthozoa: Azooxanthellate Scleractinia from the Philippines and Indonesian regions. Memoires du Museum National d'Histoire Naturelle, 172, 27 - 243, 29 pls." type="journal article" year="1997">Cairns &amp; Zibrowius 1997</bibRefCitation>
;
<bibRefCitation id="A77A4B31FF8BDC30A1F5FF26FE1BFADF" author="Zibrowius, H." box="[229,428,152,178]" pageId="28" pageNumber="29" pagination="319 - 340" refId="ref29780" refString="Zibrowius, H. (1998) A new type of symbiosis: Heterocyathus japonicus (Cnidaria: Scleractinia) living on Fissidentalium vernedei (Mollusca: Scaphopoda). Zoologische Verhandelingen, 323 (31), 319 - 340." type="journal article" year="1998">Zibrowius 1998</bibRefCitation>
;
<bibRefCitation id="A77A4B31FF8BDC30A0AAFF26FDD4FADF" author="Cairns, S. D." box="[442,611,152,178]" pageId="28" pageNumber="29" pagination="31 - 167" refId="ref24733" refString="Cairns, S. D. (1999 b) Cnidaria Anthozoa: Deep-water azooxanthellate Scleractinia from Vanuatu, and Wallis and Futuna Islands. Memoires du Museum National d'Histoire Naturelle, 180, 31 - 167." type="journal article" year="1999">Cairns 1999b</bibRefCitation>
). It is also known from the Gulf of California (
<bibRefCitation id="A77A4B31FF8BDC30A5BBFF26FF6FFAB5" author="Durham, J. W. &amp; Barnard, J. L." pageId="28" pageNumber="29" pagination="1 - 110" refId="ref25593" refString="Durham, J. W. &amp; Barnard, J. L. (1952) Stony corals of the eastern Pacific collected by the Velero III and Velero IV. Allan Hancock Pacific Expeditions, 1 (16), 1 - 110." type="journal article" year="1952">Durham &amp; Barnard 1952</bibRefCitation>
).
<taxonomicName id="04EB4D43FF8BDC30A1FDFF01FE4EFAB5" box="[237,505,190,216]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="28" pageNumber="29" phylum="Cnidaria" rank="species" species="undetermined">
<emphasis id="F19FEAD2FF8BDC30A1FDFF01FE2CFAB5" box="[237,411,191,216]" italics="true" pageId="28" pageNumber="29">Heterocyathus</emphasis>
species
</taxonomicName>
were only known as fossil for the Atlantic Ocean; the youngest record dates from the Miocene (2015 MY, Burdigalian) and it was found at the southwestern
<collectingCountry id="BBFC7650FF8BDC30A594FF5BFB61FA92" box="[1156,1238,229,255]" name="France" pageId="28" pageNumber="29">France</collectingCountry>
(
<bibRefCitation id="A77A4B31FF8BDC30A5F7FF5BFF6FFB4B" author="Stolarski, J. &amp; Zibrowius, H. &amp; Loser, H." pageId="28" pageNumber="29" pagination="309 - 330" refId="ref28845" refString="Stolarski, J. Zibrowius, H. &amp; Loser, H. (2001) Antiquity of the scleractinian-sipunculan symbiosis. Acta Palaeontologica Polonica, 46 (3), 309 - 330." type="journal article" year="2001">
Stolarski
<emphasis id="F19FEAD2FF8BDC30A44EFF5BFA2BFA93" box="[1374,1436,229,254]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
2001
</bibRefCitation>
). The particular distribution of Colombian specimens suggests that the species probably belongs to a relict fauna still present in the Caribbean. In
<collectingCountry id="BBFC7650FF8BDC30A3A7FE8CFC87FB21" box="[695,816,306,332]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
,
<taxonomicName id="04EB4D43FF8BDC30A22DFE8CFC7CFB26" authorityName="Reyes &amp; Santodomingo &amp; Cairns" authorityYear="2009" box="[829,971,306,331]" class="Anthozoa" family="Caryophylliidae" genus="Heterocyathus" kingdom="Animalia" order="Scleractinia" pageId="28" pageNumber="29" phylum="Cnidaria" rank="species" species="antoniae">
<emphasis id="F19FEAD2FF8BDC30A22DFE8CFC7CFB26" box="[829,971,306,331]" italics="true" pageId="28" pageNumber="29">H. antoniae</emphasis>
</taxonomicName>
has been collected in the northeastern Caribbean coast, from Dibulla (
<collectingRegion id="012FF822FF8BDC30A319FEE7FD3EFB1E" box="[521,649,345,371]" country="Colombia" name="La Guajira" pageId="28" pageNumber="29">La Guajira</collectingRegion>
Peninsula) to Santa Marta; between 20 and
<quantity id="04139B25FF8BDC30A59EFEE7FB7DFB1E" box="[1166,1226,345,371]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="28" pageNumber="29" unit="m" value="70.0">70 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="8BF1654BFF8BDC30A1D5FEC1FB7BFBF7" box="[197,1228,383,410]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="C35436C0FF8BDC30A1D5FEC1FB7BFBF7" blockId="28.[151,1436,152,564]" box="[197,1228,383,410]" pageId="28" pageNumber="29">
<emphasis id="F19FEAD2FF8BDC30A1D5FEC1FEE2FBF4" bold="true" box="[197,341,383,409]" pageId="28" pageNumber="29">Etymology:</emphasis>
This species is named after the oldest daughter of J. Reyes, Antonia Reyes.
</paragraph>
</subSubSection>
<subSubSection id="8BF1654BFF8BDC30A1D5FE18FA2FF859" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="C35436C0FF8BDC30A1D5FE18FA2FF859" blockId="28.[151,1436,152,564]" pageId="28" pageNumber="29">
<emphasis id="F19FEAD2FF8BDC30A1D5FE18FEF6FBAD" bold="true" box="[197,321,422,448]" pageId="28" pageNumber="29">Material:</emphasis>
<materialsCitation id="73833C9DFF8BDC30A05BFE18FD34FB8A" collectionCode="GCD" country="Colombia" location="Santa Marta" municipality="Neguanje Bay" pageId="28" pageNumber="29" specimenCode="CNI740, E118" specimenCount="1" typeStatus="holotype">
<typeStatus id="1C508862FF8BDC30A05BFE18FE76FBAD" box="[331,449,422,448]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
, INV
<specimenCode id="934D9EBBFF8BDC30A31EFE18FDC3FBAD" box="[526,628,422,448]" collectionCode="CNI" pageId="28" pageNumber="29">CNI740</specimenCode>
,
<specimenCount id="D5EDFD49FF8BDC30A392FE18FCB8FBAD" box="[642,783,422,448]" pageId="28" pageNumber="29" type="generic">1 specimen</specimenCount>
overgrowing a gastropod shell,
<quantity id="04139B25FF8BDC30A5B4FE18FB4EFBAD" box="[1188,1273,422,448]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="mm" value="10.0">10 mm</quantity>
<collectionCode id="A5FAAE05FF8BDC30A413FE18FAF1FBAD" box="[1283,1350,422,448]" pageId="28" pageNumber="29">GCD</collectionCode>
,
<specimenCode id="934D9EBBFF8BDC30A445FE18FA20FBAD" box="[1365,1431,422,448]" collectionCode="GCD" pageId="28" pageNumber="29">E118</specimenCode>
,
<collectingCountry id="BBFC7650FF8BDC30A187FE73FEB9FB8A" box="[151,270,461,487]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
(
<location id="C634601BFF8BDC30A030FE73FE0DFB8A" LSID="urn:lsid:plazi:treatment:4B4287D6FF8DDC30A187F9E8FA2FF859:C634601BFF8BDC30A030FE73FE0DFB8A" box="[288,442,461,487]" country="Colombia" municipality="Neguanje Bay" name="Santa Marta" pageId="28" pageNumber="29">Santa Marta</location>
,
<collectingMunicipality id="2330ACBAFF8BDC30A0D7FE73FDCEFB8A" box="[455,633,461,487]" pageId="28" pageNumber="29">Neguanje Bay</collectingMunicipality>
)
</materialsCitation>
.
<materialsCitation id="73833C9DFF8BDC30A380FE73FC2BF863" collectionCode="GCD" country="Colombia" location="Manaure" pageId="28" pageNumber="29" specimenCode="CNI2076, E237" specimenCount="1" stateProvince="La Guajira" typeStatus="paratype">
<typeStatus id="1C508862FF8BDC30A380FE73FCBCFB8A" box="[656,779,461,487]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
: INV
<specimenCode id="934D9EBBFF8BDC30A246FE73FC7EFB8A" box="[854,969,461,487]" collectionCode="CNI" pageId="28" pageNumber="29">CNI2076</specimenCode>
,
<specimenCount id="D5EDFD49FF8BDC30A2C7FE73FBD6FB8A" box="[983,1121,461,487]" pageId="28" pageNumber="29" type="generic">1 specimen</specimenCount>
overgrowing a gastropod shell, 9.5 mm
<collectionCode id="A5FAAE05FF8BDC30A054FE4AFE31F863" box="[324,390,500,526]" pageId="28" pageNumber="29">GCD</collectionCode>
,
<specimenCode id="934D9EBBFF8BDC30A082FE4AFE64F863" box="[402,467,500,526]" collectionCode="GCD" pageId="28" pageNumber="29">E237</specimenCode>
,
<collectingCountry id="BBFC7650FF8BDC30A0F1FE4AFDE1F863" box="[481,598,500,526]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
(
<collectingRegion id="012FF822FF8BDC30A377FE4AFD5AF863" box="[615,749,500,526]" country="Colombia" name="La Guajira" pageId="28" pageNumber="29">La Guajira</collectingRegion>
, off
<location id="C634601BFF8BDC30A235FE4AFC23F863" LSID="urn:lsid:plazi:treatment:4B4287D6FF8DDC30A187F9E8FA2FF859:C634601BFF8BDC30A235FE4AFC23F863" box="[805,916,500,526]" country="Colombia" name="Manaure" pageId="28" pageNumber="29" stateProvince="La Guajira">Manaure</location>
)
</materialsCitation>
.
<materialsCitation id="73833C9DFF8BDC30A2B8FE4AFA23F859" collectionCode="CNI" country="Colombia" pageId="28" pageNumber="29" specimenCode="CNI739, CNI738" specimenCount="3" stateProvince="La Guajira">
Other material: INV
<specimenCode id="934D9EBBFF8BDC30A5B4FE4AFAB0F863" box="[1188,1287,500,526]" collectionCode="CNI" pageId="28" pageNumber="29">CNI739</specimenCode>
,
<specimenCount id="D5EDFD49FF8BDC30A404FE4AFA2CF863" box="[1300,1435,500,526]" pageId="28" pageNumber="29" type="generic">1 specimen</specimenCount>
settled
<collectedFrom id="70F7753DFF8BDC30A1FBFDA4FE14F859" box="[235,419,538,564]" pageId="28" pageNumber="29">
on
<taxonomicName id="04EB4D43FF8BDC30A01FFDA4FE14F859" box="[271,419,538,564]" class="Gastropoda" family="Turridae" genus="Polystira" kingdom="Animalia" order="Neogastropoda" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="F19FEAD2FF8BDC30A01FFDA4FECDF85E" box="[271,378,538,563]" italics="true" pageId="28" pageNumber="29">Polystira</emphasis>
sp.
</taxonomicName>
</collectedFrom>
shell, E105; INV
<specimenCode id="934D9EBBFF8BDC30A369FDA4FD6CF859" box="[633,731,538,564]" collectionCode="CNI" pageId="28" pageNumber="29">CNI738</specimenCode>
,
<specimenCount id="D5EDFD49FF8BDC30A3F7FDA4FCC0F859" box="[743,887,538,564]" pageId="28" pageNumber="29" type="generic">2 specimens</specimenCount>
settled on a juvenile
<taxonomicName id="04EB4D43FF8BDC30A560FDA4FAB2F859" box="[1136,1285,538,564]" class="Gastropoda" family="Strombidae" genus="Strombus" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="F19FEAD2FF8BDC30A560FDA4FB6AF85E" box="[1136,1245,538,563]" italics="true" pageId="28" pageNumber="29">Strombus</emphasis>
sp.
</taxonomicName>
shell, E105
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>