499 lines
78 KiB
XML
499 lines
78 KiB
XML
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<mods:title id="98900014DBBB685583CB6644A696F8E9">Leafhoppers of the genus Kybos Fieber, 1866 (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini) in Kyrgyzstan</mods:title>
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<treatment id="039D0A61FF9A201AC894F896AFB3FB28" ID-DOI="http://doi.org/10.5281/zenodo.13621142" ID-Zenodo-Dep="13621142" LSID="urn:lsid:plazi:treatment:039D0A61FF9A201AC894F896AFB3FB28" httpUri="http://treatment.plazi.org/id/039D0A61FF9A201AC894F896AFB3FB28" lastPageId="4" lastPageNumber="93" pageId="1" pageNumber="90">
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<subSubSection id="C32EE8FCFF9A201FC894F896A936F867" box="[151,562,1903,1930]" pageId="1" pageNumber="90" type="nomenclature">
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<paragraph id="8B8BBB77FF9A201FC894F896A936F867" blockId="1.[151,584,1903,1967]" box="[151,562,1903,1930]" pageId="1" pageNumber="90">
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<heading id="D0C30C1BFF9A201FC894F896A936F867" bold="true" box="[151,562,1903,1930]" fontSize="11" level="1" pageId="1" pageNumber="90" reason="1">
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<taxonomicName id="4C34C0F4FF9A201FC894F896A936F867" ID-CoL="9G637" authority="Zachvatkin, 1953" authorityName="Zachvatkin" authorityYear="1953" box="[151,562,1903,1930]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="oshanini">
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||
<emphasis id="B9406765FF9A201FC894F896A936F867" bold="true" box="[151,562,1903,1930]" pageId="1" pageNumber="90">
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<emphasis id="B9406765FF9A201FC894F896AA56F867" bold="true" box="[151,338,1903,1930]" italics="true" pageId="1" pageNumber="90">Kybos oshanini</emphasis>
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Zachvatkin, 1953
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</emphasis>
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</taxonomicName>
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</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C32EE8FCFF9A201FC894F86DA94CF843" box="[151,584,1940,1967]" pageId="1" pageNumber="90" type="description">
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<paragraph id="8B8BBB77FF9A201FC894F86DA94CF843" blockId="1.[151,584,1903,1967]" box="[151,584,1940,1967]" pageId="1" pageNumber="90">
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<figureCitation id="130FA7F2FF9A201FC894F86DABFEF842" box="[151,250,1940,1967]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." figureDoi="http://doi.org/10.5281/zenodo.13330990" httpUri="https://zenodo.org/record/13330990/files/figure.png" pageId="1" pageNumber="90">Figs 5–8</figureCitation>
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,
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<figureCitation id="130FA7F2FF9A201FC905F86DAA4BF843" box="[262,335,1940,1966]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." figureDoi="http://doi.org/10.5281/zenodo.13330992" httpUri="https://zenodo.org/record/13330992/files/figure.png" pageId="1" pageNumber="90">17–32</figureCitation>
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,
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<figureCitation id="130FA7F2FF9A201FC959F86DAAF2F842" box="[346,502,1940,1967]" captionStart="FIGURES 38–63" captionStartId="7.[152,270,1540,1564]" captionTargetBox="[153,1433,184,1509]" captionTargetId="figure-15@7.[151,1436,181,1516]" captionText="FIGURES 38–63. Kybos spp., male genitalia. 38–39, 50, 52, 58—K. montanus; 40–41, 53, 59—K. oshanini; 42–43, 51, 54, 60—K. niveicolor; 44–45, 55, 61—K. auricillatus; 46–47, 56, 62—K. populi; 48–49, 57, 63—K. virgator. 38, 40, 42, 44, 46, 48—aedeagus, lateral view, 39, 41, 43, 45, 47, 49—same, dorsal view, 50–51—style, 52–57—pygofer appendage, 58–63—anal collar appendage." figureDoi="http://doi.org/10.5281/zenodo.13621140" httpUri="https://zenodo.org/record/13621140/files/figure.png" pageId="1" pageNumber="90">40–41, 53, 59</figureCitation>
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, 70–73
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C32EE8FCFF9A201CC894F821A9FCFF20" lastPageId="2" lastPageNumber="91" pageId="1" pageNumber="90" type="reference_group">
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<paragraph id="8B8BBB77FF9A201FC894F821AF15F81C" blockId="1.[151,1041,2008,2033]" box="[151,1041,2008,2033]" pageId="1" pageNumber="90">
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<treatmentCitationGroup id="AB249C59FF9A201FC894F821AF15F81C" box="[151,1041,2008,2033]" pageId="1" pageNumber="90">
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<taxonomicName id="4C34C0F4FF9A201FC894F821A978F81C" ID-CoL="9G634" authority="Zachvatkin, 1953: 237 - 239" authorityName="sensu Zachvatkin" authorityPageNumber="237 - 239" authorityYear="1953" box="[151,636,2008,2033]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="90" phylum="Arthropoda" rank="species" species="mesasiaticus">
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<emphasis id="B9406765FF9A201FC894F821AA5AF81D" box="[151,350,2008,2032]" italics="true" pageId="1" pageNumber="90">Kybos mesasiaticus</emphasis>
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Zachvatkin, 1953: 237–239
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</taxonomicName>
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(synonymy by
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<treatmentCitation id="0A959D66FF9A201FCB19F821AF0EF81C" author="Tishechkin, D. Y." box="[794,1034,2008,2033]" page="1401" pageId="1" pageNumber="90" year="2002">
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||
<bibRefCitation id="EFA5C686FF9A201FCB19F821AF0EF81C" author="Tishechkin, D. Y." box="[794,1034,2008,2033]" pageId="1" pageNumber="90" pagination="1398 - 1401" refId="ref7514" refString="Tishechkin, D. Y. (2002) New data on taxonomy of the genus Kybos (Homoptera, Cicadellidae, Typhlocybinae) from central regions of European Russia. Zoologicheskiy Zhurnal, 81 (11), 1398 - 1401. [in Russian with English summary]" type="journal article" year="2002">Tishechkin, 2002: 1401</bibRefCitation>
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</treatmentCitation>
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)
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8B8BBB77FF99201CC894FF6EA9FDFF42" blockId="2.[151,1028,150,236]" box="[151,761,150,175]" pageId="2" pageNumber="91">
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||
<taxonomicName id="4C34C0F4FF99201CC894FF6EA9ADFF42" ID-CoL="9G634" authority="Zachvatkin, 1953: 208" authorityName="Zachvatkin" authorityPageNumber="208" authorityYear="1953" box="[151,681,150,175]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="subSpecies" species="oshanini" status="syn. nov." subSpecies="mesasiaticus">
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<emphasis id="B9406765FF99201CC894FF6EAABFFF43" box="[151,443,151,175]" italics="true" pageId="2" pageNumber="91">Kybos oshanini mesasiaticus</emphasis>
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Zachvatkin, 1953: 208
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</taxonomicName>
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,
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<emphasis id="B9406765FF99201CCAB7FF6EA9FDFF42" bold="true" box="[692,761,151,175]" pageId="2" pageNumber="91">
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<taxonomicNameLabel id="A273DA1EFF99201CCAB7FF6EA9FDFF42" box="[692,761,151,175]" pageId="2" pageNumber="91">syn. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8B8BBB77FF99201CC894FF4CA9FCFF20" blockId="2.[151,1028,150,236]" box="[151,760,181,205]" pageId="2" pageNumber="91">
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||
<taxonomicName id="4C34C0F4FF99201CC894FF4CA947FF20" ID-CoL="6NQB8" authority="Mitjaev, 1971" authorityName="Mitjaev" authorityYear="1971" box="[151,579,181,205]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="subSpecies" species="mesasiaticus" subSpecies="iliensis">
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<emphasis id="B9406765FF99201CC894FF4CAAA9FF20" box="[151,429,181,205]" italics="true" pageId="2" pageNumber="91">Kybos mesasiaticus iliensis</emphasis>
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Mitjaev, 1971
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</taxonomicName>
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; 122–123,
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<emphasis id="B9406765FF99201CCAB1FF4CA9FCFF20" bold="true" box="[690,760,181,205]" pageId="2" pageNumber="91">syn. n.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EE8FCFF99201CC894FF2CAAE5FCBD" pageId="2" pageNumber="91" type="description">
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<paragraph id="8B8BBB77FF99201CC894FF2CAF00FF01" blockId="2.[151,1028,150,236]" box="[151,1028,212,236]" pageId="2" pageNumber="91">
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<taxonomicName id="4C34C0F4FF99201CC894FF2CA94DFF01" authority="(Mitjaev, 1971)" baseAuthorityName="Mitjaev" baseAuthorityYear="1971" box="[151,585,212,236]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="iliensis" subGenus="Kybos">
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<emphasis id="B9406765FF99201CC894FF2CAAA2FF01" box="[151,422,212,236]" italics="true" pageId="2" pageNumber="91">Empoasca (Kybos) iliensis</emphasis>
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(Mitjaev, 1971)
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</taxonomicName>
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(in Dworakowska, 1976: 446–447),
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<emphasis id="B9406765FF99201CCBBCFF2DAF00FF01" bold="true" box="[959,1028,212,236]" pageId="2" pageNumber="91">syn. n.</emphasis>
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</paragraph>
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<paragraph id="8B8BBB77FF99201CC894FEE3A8E1FE90" blockId="2.[151,1437,282,2001]" pageId="2" pageNumber="91">
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<emphasis id="B9406765FF99201CC894FEE3AA2DFED9" bold="true" box="[151,297,282,308]" pageId="2" pageNumber="91">Description.</emphasis>
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Bright green with lighter spots in fore part of body, old dry specimens slightly yellowish. In darker specimens, forewings sometimes slightly smoky in apical parts, veins of hindwings darker than membrane and visible through folded forewings. Dorsal side of abdomen green (
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<figureCitation id="130FA7F2FF99201CCB71FE9BA8DDFE90" box="[882,985,354,381]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." figureDoi="http://doi.org/10.5281/zenodo.13330990" httpUri="https://zenodo.org/record/13330990/files/figure.png" pageId="2" pageNumber="91">Figs 5–8</figureCitation>
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).
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</paragraph>
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<paragraph id="8B8BBB77FF99201CC8C4FE7FABC0FDB8" blockId="2.[151,1437,282,2001]" pageId="2" pageNumber="91">
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Length of dorsal apodemes same or slightly greater than their width at base, their inner margins parallel or slightly divergent, usually separated by rather narrow notch (
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<figureCitation id="130FA7F2FF99201CCB49FE53A8CBFE29" box="[842,975,426,452]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." figureDoi="http://doi.org/10.5281/zenodo.13330992" httpUri="https://zenodo.org/record/13330992/files/figure.png" pageId="2" pageNumber="91">Figs 17–24</figureCitation>
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). Ventral apodemes 1.8–2.5 times longer than their width at base, parallel-sided, touching with inner margins or separated by narrow notch (
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<figureCitation id="130FA7F2FF99201CCD08FE37AE95FE04" box="[1291,1425,462,489]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." figureDoi="http://doi.org/10.5281/zenodo.13330992" httpUri="https://zenodo.org/record/13330992/files/figure.png" pageId="2" pageNumber="91">Figs 25–32</figureCitation>
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). Aedeagus without lateral processes (
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<figureCitation id="130FA7F2FF99201CCA3FFE0BA9C7FDE1" box="[572,707,498,524]" captionStart="FIGURES 38–63" captionStartId="7.[152,270,1540,1564]" captionTargetBox="[153,1433,184,1509]" captionTargetId="figure-15@7.[151,1436,181,1516]" captionText="FIGURES 38–63. Kybos spp., male genitalia. 38–39, 50, 52, 58—K. montanus; 40–41, 53, 59—K. oshanini; 42–43, 51, 54, 60—K. niveicolor; 44–45, 55, 61—K. auricillatus; 46–47, 56, 62—K. populi; 48–49, 57, 63—K. virgator. 38, 40, 42, 44, 46, 48—aedeagus, lateral view, 39, 41, 43, 45, 47, 49—same, dorsal view, 50–51—style, 52–57—pygofer appendage, 58–63—anal collar appendage." figureDoi="http://doi.org/10.5281/zenodo.13621140" httpUri="https://zenodo.org/record/13621140/files/figure.png" pageId="2" pageNumber="91">Figs 40–41</figureCitation>
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). Style same as in
|
||
<taxonomicName id="4C34C0F4FF99201CCBA3FE0AAF35FDE1" authorityName="Mitjaev" authorityYear="1971" box="[928,1073,499,524]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B9406765FF99201CCBA3FE0AAF35FDE1" box="[928,1073,499,524]" italics="true" pageId="2" pageNumber="91">K. montanus</emphasis>
|
||
</taxonomicName>
|
||
, with numerous long setae and serrated falcate apical part. Pygofer appendages narrow (
|
||
<figureCitation id="130FA7F2FF99201CCB2EFDEFA887FDDC" box="[813,899,534,561]" captionStart="FIGURES 38–63" captionStartId="7.[152,270,1540,1564]" captionTargetBox="[153,1433,184,1509]" captionTargetId="figure-15@7.[151,1436,181,1516]" captionText="FIGURES 38–63. Kybos spp., male genitalia. 38–39, 50, 52, 58—K. montanus; 40–41, 53, 59—K. oshanini; 42–43, 51, 54, 60—K. niveicolor; 44–45, 55, 61—K. auricillatus; 46–47, 56, 62—K. populi; 48–49, 57, 63—K. virgator. 38, 40, 42, 44, 46, 48—aedeagus, lateral view, 39, 41, 43, 45, 47, 49—same, dorsal view, 50–51—style, 52–57—pygofer appendage, 58–63—anal collar appendage." figureDoi="http://doi.org/10.5281/zenodo.13621140" httpUri="https://zenodo.org/record/13621140/files/figure.png" pageId="2" pageNumber="91">Fig. 53</figureCitation>
|
||
). Anal collar appendages narrow, falcate (
|
||
<figureCitation id="130FA7F2FF99201CCD6CFDEFABB3FDB8" captionStart="FIGURES 38–63" captionStartId="7.[152,270,1540,1564]" captionTargetBox="[153,1433,184,1509]" captionTargetId="figure-15@7.[151,1436,181,1516]" captionText="FIGURES 38–63. Kybos spp., male genitalia. 38–39, 50, 52, 58—K. montanus; 40–41, 53, 59—K. oshanini; 42–43, 51, 54, 60—K. niveicolor; 44–45, 55, 61—K. auricillatus; 46–47, 56, 62—K. populi; 48–49, 57, 63—K. virgator. 38, 40, 42, 44, 46, 48—aedeagus, lateral view, 39, 41, 43, 45, 47, 49—same, dorsal view, 50–51—style, 52–57—pygofer appendage, 58–63—anal collar appendage." figureDoi="http://doi.org/10.5281/zenodo.13621140" httpUri="https://zenodo.org/record/13621140/files/figure.png" pageId="2" pageNumber="91">Fig. 59</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8BBB77FF99201CC8C4FDA7AAE5FCBD" blockId="2.[151,1437,282,2001]" pageId="2" pageNumber="91">
|
||
Males with very long ventral apodemes (
|
||
<figureCitation id="130FA7F2FF99201CCA97FDA7A810FD94" box="[660,788,606,633]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." figureDoi="http://doi.org/10.5281/zenodo.13330992" httpUri="https://zenodo.org/record/13330992/files/figure.png" pageId="2" pageNumber="91">Figs 25–27</figureCitation>
|
||
) were found only in a population on the northern edge of Fergana Valley near Shekaftar (=Shakaptar) Village,
|
||
<quantity id="4CCC1692FF99201CCAEAFD7BA834FD71" box="[745,816,642,669]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="91" unit="km" value="45.0">45 km</quantity>
|
||
southwest from Kerben. However, in the same locality on the same group of trees, males with the same coloration but with shorter apodemes were found (
|
||
<figureCitation id="130FA7F2FF99201CCD1DFD5FAE71FD2D" box="[1310,1397,678,704]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." figureDoi="http://doi.org/10.5281/zenodo.13330992" httpUri="https://zenodo.org/record/13330992/files/figure.png" pageId="2" pageNumber="91">Fig. 28</figureCitation>
|
||
). It is likely, that males with different lengths of apodemes may predominate in different populations; this was the reason for the description of the form with shorter apodemes as a subspecies,
|
||
<taxonomicName id="4C34C0F4FF99201CCBFAFD16AFFBFCE5" authorityName="Mitjaev" authorityYear="1971" box="[1017,1279,750,776]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="subSpecies" species="mesasiaticus" subSpecies="iliensis">
|
||
<emphasis id="B9406765FF99201CCBFAFD16AFFBFCE5" box="[1017,1279,750,776]" italics="true" pageId="2" pageNumber="91">K. mesasiaticus iliensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130FA7F2FF99201CCD0DFD17AE8BFCE5" box="[1294,1423,750,776]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." figureDoi="http://doi.org/10.5281/zenodo.13330992" httpUri="https://zenodo.org/record/13330992/files/figure.png" pageId="2" pageNumber="91">Figs 23, 31</figureCitation>
|
||
). However, since both forms are sometimes found in the same sample, it is clear that such differences are the result of intrapopulation variability.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32EE8FCFF99201CC8C4FCA3AFA0FA81" pageId="2" pageNumber="91" type="biology_ecology">
|
||
<paragraph id="8B8BBB77FF99201CC8C4FCA3AFD6FC75" blockId="2.[151,1437,282,2001]" pageId="2" pageNumber="91">
|
||
<emphasis id="B9406765FF99201CC8C4FCA3AA15FC99" bold="true" box="[199,273,858,884]" pageId="2" pageNumber="91">Hosts.</emphasis>
|
||
We collected
|
||
<taxonomicName id="4C34C0F4FF99201CC9A9FCA2A92CFC99" authorityName="Zachvatkin" authorityYear="1953" box="[426,552,858,884]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF99201CC9A9FCA2A92CFC99" box="[426,552,858,884]" italics="true" pageId="2" pageNumber="91">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
from different species of
|
||
<taxonomicName id="4C34C0F4FF99201CCB42FCA3A87DFC99" box="[833,889,858,884]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="91" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9406765FF99201CCB42FCA3A87DFC99" box="[833,889,858,884]" italics="true" pageId="2" pageNumber="91">Salix</emphasis>
|
||
</taxonomicName>
|
||
from the section
|
||
<taxonomicName id="4C34C0F4FF99201CCC36FCA3AF76FC99" authorityName="Linnaeus" authorityYear="1758" box="[1077,1138,858,884]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="91" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9406765FF99201CCC36FCA3AF76FC99" box="[1077,1138,858,884]" italics="true" pageId="2" pageNumber="91">Helix</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F323FBE7FF99201CCC91FCA2AE11FC98" box="[1170,1301,859,885]" name="Kyrgyzstan" pageId="2" pageNumber="91">Kyrgyzstan</collectingCountry>
|
||
,
|
||
<bibRefCitation id="EFA5C686FF99201CCD1DFCA3ABE3FC74" author="Zachvatkin, A. A." pageId="2" pageNumber="91" pagination="205 - 223" refId="ref7639" refString="Zachvatkin, A. A. (1953 a) Biological and taxonomic notes on leafhoppers of Central Russian fauna. In: Smirnov, E. S. (Ed.), Sbornik nauchnykh rabot. Moscow University, Moscow, pp. 205 - 223. [in Russian]" type="book chapter" year="1953">Zachvatkin (1953a</bibRefCitation>
|
||
, b) collected it from
|
||
<taxonomicName id="4C34C0F4FF99201CC9D2FC86A959FC75" box="[465,605,895,921]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="91" phylum="Tracheophyta" rank="species" species="songorica">
|
||
<emphasis id="B9406765FF99201CC9D2FC86A959FC75" box="[465,605,895,921]" italics="true" pageId="2" pageNumber="91">S. songorica</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F323FBE7FF99201CCA82FC87A9FBFC75" box="[641,767,894,920]" name="Uzbekistan" pageId="2" pageNumber="91">Uzbekistan</collectingCountry>
|
||
and from
|
||
<taxonomicName id="4C34C0F4FF99201CCB70FC86A8F2FC75" baseAuthorityName="Zachvatkin" baseAuthorityYear="1953" box="[883,1014,894,921]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="91" phylum="Tracheophyta" rank="species" species="viminalis">
|
||
<emphasis id="B9406765FF99201CCB70FC86A8F2FC75" box="[883,1014,894,921]" italics="true" pageId="2" pageNumber="91">S. viminalis</emphasis>
|
||
</taxonomicName>
|
||
in Moscow Oblast.
|
||
</paragraph>
|
||
<paragraph id="8B8BBB77FF99201CC8C4FC5BAAC9FBE9" blockId="2.[151,1437,282,2001]" pageId="2" pageNumber="91">
|
||
<emphasis id="B9406765FF99201CC8C4FC5BAA72FC51" bold="true" box="[199,374,930,956]" pageId="2" pageNumber="91">Calling signal.</emphasis>
|
||
Signals of
|
||
<specimenCount id="9D3270FEFF99201CCA03FC5BA97CFC51" box="[512,632,930,956]" count="5" pageId="2" pageNumber="91" type="male">five males</specimenCount>
|
||
from
|
||
<collectingCountry id="F323FBE7FF99201CCAC7FC5AA841FC50" box="[708,837,931,957]" name="Kyrgyzstan" pageId="2" pageNumber="91">Kyrgyzstan</collectingCountry>
|
||
(the bank of the Taldy-Sai River
|
||
<quantity id="4CCC1692FF99201CCCD6FC5BAE16FC51" box="[1237,1298,930,956]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="91" unit="km" value="3.0">3 km</quantity>
|
||
down from Shekaftar (=Shakaptar) Village,
|
||
<quantity id="4CCC1692FF99201CCA00FC3FA94FFC0D" box="[515,587,966,993]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="91" unit="km" value="45.0">45 km</quantity>
|
||
southwest from Kerben Town, on
|
||
<taxonomicName id="4C34C0F4FF99201CCBD0FC3FAF0FFC0D" box="[979,1035,966,992]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="91" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9406765FF99201CCBD0FC3FAF0FFC0D" box="[979,1035,966,992]" italics="true" pageId="2" pageNumber="91">Salix</emphasis>
|
||
</taxonomicName>
|
||
sect.
|
||
<taxonomicName id="4C34C0F4FF99201CCC4EFC3FAF8EFC0D" authorityName="Linnaeus" authorityYear="1758" box="[1101,1162,966,992]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="91" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9406765FF99201CCC4EFC3FAF8EFC0D" box="[1101,1162,966,992]" italics="true" pageId="2" pageNumber="91">Helix</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="FF8A9DB7FF99201CCC9AFC3EAE21FC0D" box="[1177,1317,966,993]" pageId="2" pageNumber="91" value="2023-07-07">7. VII. 2023</date>
|
||
, recording at 22
|
||
<superScript id="7C41163FFF99201CC8D6FC13ABF6FBE9" attach="right" box="[213,242,1002,1028]" fontSize="6" pageId="2" pageNumber="91">oC</superScript>
|
||
) were investigated.
|
||
</paragraph>
|
||
<paragraph id="8B8BBB77FF99201CC8C4FBF7A90FFB55" blockId="2.[151,1437,282,2001]" pageId="2" pageNumber="91">
|
||
Calling signal is a phrase consisting of 2–5 syllables, following each other with a period of 360–760 ms; syllable repetition period usually increases towards the end of a phrase (
|
||
<figureCitation id="130FA7F2FF99201CCB62FBCBA8E0FBA1" box="[865,996,1074,1100]" captionStart="FIGURES 64–78" captionStartId="9.[152,270,1943,1967]" captionTargetBox="[179,1400,189,1917]" captionTargetId="figure-15@9.[175,1411,181,1919]" captionTargetPageId="9" captionText="FIGURES 64–78. Oscillograms of male calling signals of Kybos spp. 64–69—K. montanus; 70–73—K. oshanini; 74–76—K. niveicolor; 77–78—K. populi. Faster oscillograms of the parts of signals indicated as “66”, “68”, “72–73”, “76”, and “78” are given under the same numbers." figureDoi="http://doi.org/10.5281/zenodo.13330994" httpUri="https://zenodo.org/record/13330994/files/figure.png" pageId="2" pageNumber="91">Figs 70–71</figureCitation>
|
||
). Each syllable consists of several low-amplitude pulses following with a period of 80–140 ms, and one higher-amplitude pulse (
|
||
<figureCitation id="130FA7F2FF99201CCC99FBAFAFF4FB9D" box="[1178,1264,1110,1136]" captionStart="FIGURES 64–78" captionStartId="9.[152,270,1943,1967]" captionTargetBox="[179,1400,189,1917]" captionTargetId="figure-15@9.[175,1411,181,1919]" captionTargetPageId="9" captionText="FIGURES 64–78. Oscillograms of male calling signals of Kybos spp. 64–69—K. montanus; 70–73—K. oshanini; 74–76—K. niveicolor; 77–78—K. populi. Faster oscillograms of the parts of signals indicated as “66”, “68”, “72–73”, “76”, and “78” are given under the same numbers." figureDoi="http://doi.org/10.5281/zenodo.13330994" httpUri="https://zenodo.org/record/13330994/files/figure.png" pageId="2" pageNumber="91">Fig. 72</figureCitation>
|
||
). Occasionally, low-amplitude pulses are divided in two and an additional low-amplitude short pulse follows immediately before the high-amplitude one (
|
||
<figureCitation id="130FA7F2FF99201CC9A9FB67AAFBFB55" box="[426,511,1182,1208]" captionStart="FIGURES 64–78" captionStartId="9.[152,270,1943,1967]" captionTargetBox="[179,1400,189,1917]" captionTargetId="figure-15@9.[175,1411,181,1919]" captionTargetPageId="9" captionText="FIGURES 64–78. Oscillograms of male calling signals of Kybos spp. 64–69—K. montanus; 70–73—K. oshanini; 74–76—K. niveicolor; 77–78—K. populi. Faster oscillograms of the parts of signals indicated as “66”, “68”, “72–73”, “76”, and “78” are given under the same numbers." figureDoi="http://doi.org/10.5281/zenodo.13330994" httpUri="https://zenodo.org/record/13330994/files/figure.png" pageId="2" pageNumber="91">Fig. 73</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8BBB77FF99201CC8C4FB3BAFA0FA81" blockId="2.[151,1437,282,2001]" pageId="2" pageNumber="91">
|
||
The signal
|
||
<taxonomicName id="4C34C0F4FF99201CC942FB3AAAC4FB31" authorityName="Zachvatkin" authorityYear="1953" box="[321,448,1218,1244]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF99201CC942FB3AAAC4FB31" box="[321,448,1218,1244]" italics="true" pageId="2" pageNumber="91">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
is very similar to the calling signal of
|
||
<taxonomicName id="4C34C0F4FF99201CCB66FB3AA8F5FB31" authorityName="Mitjaev" authorityYear="1971" box="[869,1009,1219,1244]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B9406765FF99201CCB66FB3AA8F5FB31" box="[869,1009,1219,1244]" italics="true" pageId="2" pageNumber="91">K. montanus</emphasis>
|
||
</taxonomicName>
|
||
. The most noticeable difference is that low-amplitude pulses in the beginning of the syllable are always present in the signals of
|
||
<taxonomicName id="4C34C0F4FF99201CCC73FB1EAFEBFAED" authorityName="Zachvatkin" authorityYear="1953" box="[1136,1263,1254,1280]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF99201CCC73FB1EAFEBFAED" box="[1136,1263,1254,1280]" italics="true" pageId="2" pageNumber="91">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
, whereas in the signals of
|
||
<taxonomicName id="4C34C0F4FF99201CC908FAF2AA9DFAC9" authorityName="Mitjaev" authorityYear="1971" box="[267,409,1291,1316]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B9406765FF99201CC908FAF2AA9DFAC9" box="[267,409,1291,1316]" italics="true" pageId="2" pageNumber="91">K. montanus</emphasis>
|
||
</taxonomicName>
|
||
they are fully or partially reduced. Also, the higher-amplitude component of syllable in the signal of
|
||
<taxonomicName id="4C34C0F4FF99201CC903FAD6AA8AFAA5" authorityName="Mitjaev" authorityYear="1971" box="[256,398,1327,1352]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B9406765FF99201CC903FAD6AA8AFAA5" box="[256,398,1327,1352]" italics="true" pageId="2" pageNumber="91">K. montanus</emphasis>
|
||
</taxonomicName>
|
||
is longer, has a flat leading edge, and sometimes consists of several partially merged pulses. In
|
||
<taxonomicName id="4C34C0F4FF99201CC8B5FAAAAA32FA81" authorityName="Zachvatkin" authorityYear="1953" box="[182,310,1362,1388]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF99201CC8B5FAAAAA32FA81" box="[182,310,1362,1388]" italics="true" pageId="2" pageNumber="91">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
it is shorter, with a steep leading edge, and consists of only one or two pulses.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32EE8FCFF99201CC8C4FA8FAFA7FA10" pageId="2" pageNumber="91" type="distribution">
|
||
<paragraph id="8B8BBB77FF99201CC8C4FA8FAFA7FA10" blockId="2.[151,1437,282,2001]" pageId="2" pageNumber="91">
|
||
<emphasis id="B9406765FF99201CC8C4FA8FAA65FA7D" bold="true" box="[199,353,1398,1424]" pageId="2" pageNumber="91">Distribution.</emphasis>
|
||
Widely distributed on the plains and arid low mountains of Central Asia along rivers and in oases; does not penetrate into the mountains above the zone of arid shrubs. In European
|
||
<collectingCountry id="F323FBE7FF99201CCC74FA63AFC6FA59" box="[1143,1218,1434,1460]" name="Russia" pageId="2" pageNumber="91">Russia</collectingCountry>
|
||
it is known only in the enclave of steppe vegetation in the Oka River Valley,
|
||
<collectingRegion id="49F07595FF99201CCB20FA46A8DCFA35" box="[803,984,1470,1497]" country="Russia" name="Moscow" pageId="2" pageNumber="91">Moscow Oblast</collectingRegion>
|
||
, Serpukhov Region, where a significant number of steppe insect species are found that do not penetrate farther to the north and west.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32EE8FCFF99201AC8C4F9FFAFB3FB28" lastPageId="4" lastPageNumber="93" pageId="2" pageNumber="91" type="discussion">
|
||
<paragraph id="8B8BBB77FF99201CC8C4F9FFA928F83D" blockId="2.[151,1437,282,2001]" pageId="2" pageNumber="91">
|
||
<emphasis id="B9406765FF99201CC8C4F9FFAA3FF9CD" bold="true" box="[199,315,1542,1568]" pageId="2" pageNumber="91">Remarks.</emphasis>
|
||
The description of
|
||
<taxonomicName id="4C34C0F4FF99201CCA1EF9FEA99BF9CD" authorityName="Zachvatkin" authorityYear="1953" box="[541,671,1542,1568]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF99201CCA1EF9FEA99BF9CD" box="[541,671,1542,1568]" italics="true" pageId="2" pageNumber="91">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
is based on specimens collected in the environs of Luzhki Village, Serpukhov Region,
|
||
<collectingRegion id="49F07595FF99201CC97FF9D2A92BF9A9" box="[380,559,1578,1605]" country="Russia" name="Moscow" pageId="2" pageNumber="91">Moscow Oblast</collectingRegion>
|
||
from
|
||
<taxonomicName id="4C34C0F4FF99201CCA7AF9D3AF10F9A8" authority="(Zachvatkin, 1953 a)" baseAuthorityName="Zachvatkin" baseAuthorityYear="1953" box="[633,1044,1578,1605]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="91" phylum="Tracheophyta" rank="species" species="viminalis">
|
||
<emphasis id="B9406765FF99201CCA7AF9D3A826F9A9" box="[633,802,1578,1604]" italics="true" pageId="2" pageNumber="91">Salix viminalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA5C686FF99201CCB30F9D3AF08F9A8" author="Zachvatkin, A. A." box="[819,1036,1578,1605]" pageId="2" pageNumber="91" pagination="205 - 223" refId="ref7639" refString="Zachvatkin, A. A. (1953 a) Biological and taxonomic notes on leafhoppers of Central Russian fauna. In: Smirnov, E. S. (Ed.), Sbornik nauchnykh rabot. Moscow University, Moscow, pp. 205 - 223. [in Russian]" type="book chapter" year="1953">Zachvatkin, 1953a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. It is indicated that this species is widespread and forms several races, one of which,
|
||
<taxonomicName id="4C34C0F4FF99201CCAEBF9B6AF00F985" authorityName="Zachvatkin" authorityYear="1953" box="[744,1028,1614,1640]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="subSpecies" species="oshanini" subSpecies="mesasiaticus">
|
||
<emphasis id="B9406765FF99201CCAEBF9B6AF00F985" box="[744,1028,1614,1640]" italics="true" pageId="2" pageNumber="91">K. oshanini mesasiaticus</emphasis>
|
||
</taxonomicName>
|
||
, is common in the oasis regions of Central Asia on willows and differs from the nominotypical one in its brighter coloration and somewhat smaller size. Type specimens are absent in the collection of the Zoological Museum of
|
||
<collectingRegion id="49F07595FF99201CCC00F96EAF67F95C" box="[1027,1123,1687,1713]" country="Russia" name="Moscow" pageId="2" pageNumber="91">Moscow</collectingRegion>
|
||
University. However, in the materials collected by Zachvatkin and his students, a series of specimens with the label “
|
||
<collectingRegion id="49F07595FF99201CCC6CF943AE21F939" box="[1135,1317,1722,1748]" country="Russia" name="Moscow" pageId="2" pageNumber="91">Moscow Oblast</collectingRegion>
|
||
, Prioksko-Terrasny Nature Reserve,
|
||
<date id="FF8A9DB7FF99201CC9C7F927A967F915" box="[452,611,1758,1785]" pageId="2" pageNumber="91" value="1946-07-10">10. VII. 1946</date>
|
||
,
|
||
<taxonomicName id="4C34C0F4FF99201CCA72F927A81EF915" baseAuthorityName="Zachvatkin" baseAuthorityYear="1953" box="[625,794,1758,1784]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="91" phylum="Tracheophyta" rank="species" species="viminalis">
|
||
<emphasis id="B9406765FF99201CCA72F927A81EF915" box="[625,794,1758,1784]" italics="true" pageId="2" pageNumber="91">Salix viminalis</emphasis>
|
||
</taxonomicName>
|
||
” was found. Since these specimens fully correspond to the original description, and Luzhki Village is located on the bank of the Oka River close to the Prioksko-Terrasny Nature Reserve boundary, we previously suggested that they represent
|
||
<typeStatus id="548F05D5FF99201CCBA8F8DEAF1CF8AC" box="[939,1048,1831,1857]" pageId="2" pageNumber="91" type="topotype">topotypes</typeStatus>
|
||
of this species (
|
||
<bibRefCitation id="EFA5C686FF99201CCCCAF8DFAE95F8AD" author="Tishechkin, D. Y." box="[1225,1425,1830,1856]" pageId="2" pageNumber="91" pagination="1398 - 1401" refId="ref7514" refString="Tishechkin, D. Y. (2002) New data on taxonomy of the genus Kybos (Homoptera, Cicadellidae, Typhlocybinae) from central regions of European Russia. Zoologicheskiy Zhurnal, 81 (11), 1398 - 1401. [in Russian with English summary]" type="journal article" year="2002">Tishechkin, 2002</bibRefCitation>
|
||
). It should be added that
|
||
<taxonomicName id="4C34C0F4FF99201CC9A1F8B3A94DF889" baseAuthorityName="Zachvatkin" baseAuthorityYear="1953" box="[418,585,1866,1892]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="91" phylum="Tracheophyta" rank="species" species="viminalis">
|
||
<emphasis id="B9406765FF99201CC9A1F8B3A94DF889" box="[418,585,1866,1892]" italics="true" pageId="2" pageNumber="91">Salix viminalis</emphasis>
|
||
</taxonomicName>
|
||
grows in this area only on alluvial sands on the banks of the Oka River, so these specimens cannot be collected in other parts of the reserve, where forest communities predominate. Thus, identification of
|
||
<taxonomicName id="4C34C0F4FF99201CC952F86AAAD5F841" authorityName="Zachvatkin" authorityYear="1953" box="[337,465,1938,1964]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="91" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF99201CC952F86AAAD5F841" box="[337,465,1938,1964]" italics="true" pageId="2" pageNumber="91">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
from European
|
||
<collectingCountry id="F323FBE7FF99201CCA85F86BA9D5F841" box="[646,721,1938,1964]" name="Russia" pageId="2" pageNumber="91">Russia</collectingCountry>
|
||
is based on these specimens, which are illustrated in
|
||
<bibRefCitation id="EFA5C686FF99201CCD22F86BABE6F83D" author="Tishechkin, D. Y." pageId="2" pageNumber="91" pagination="1398 - 1401" refId="ref7514" refString="Tishechkin, D. Y. (2002) New data on taxonomy of the genus Kybos (Homoptera, Cicadellidae, Typhlocybinae) from central regions of European Russia. Zoologicheskiy Zhurnal, 81 (11), 1398 - 1401. [in Russian with English summary]" type="journal article" year="2002">Tishechkin (2002)</bibRefCitation>
|
||
and shown on
|
||
<figureCitation id="130FA7F2FF99201CC988F84FAAD6F83D" box="[395,466,1974,2000]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." figureDoi="http://doi.org/10.5281/zenodo.13330990" httpUri="https://zenodo.org/record/13330990/files/figure.png" pageId="2" pageNumber="91">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="130FA7F2FF99201CC9DCF84FA92CF83D" box="[479,552,1974,2000]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." figureDoi="http://doi.org/10.5281/zenodo.13330992" httpUri="https://zenodo.org/record/13330992/files/figure.png" pageId="2" pageNumber="91">24, 32</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF4BEBFFFF98201DC89BF8B9AAD9F805" ID-DOI="http://doi.org/10.5281/zenodo.13330990" ID-Zenodo-Dep="13330990" httpUri="https://zenodo.org/record/13330990/files/figure.png" pageId="3" pageNumber="92" startId="3.[152,270,1856,1880]" targetBox="[254,1332,185,1824]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8B8BBB77FF98201DC89BF8B9AAD9F805" blockId="3.[151,1437,1856,2024]" pageId="3" pageNumber="92">
|
||
<emphasis id="B9406765FF98201DC89BF8B9AA4BF8B5" bold="true" box="[152,335,1856,1880]" pageId="3" pageNumber="92">FIGURES 1–12.</emphasis>
|
||
<taxonomicName id="4C34C0F4FF98201DC955F8B9AACDF8B5" box="[342,457,1856,1880]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B9406765FF98201DC955F8B9AA91F8B5" box="[342,405,1856,1880]" italics="true" pageId="3" pageNumber="92">Kybos</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, dorsal habitus. 1–4—
|
||
<taxonomicName id="4C34C0F4FF98201DCAB2F8B8A836F8B5" authorityName="Mitjaev" authorityYear="1971" box="[689,818,1857,1880]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B9406765FF98201DCAB2F8B8A836F8B5" box="[689,818,1857,1880]" italics="true" pageId="3" pageNumber="92">K. montanus</emphasis>
|
||
</taxonomicName>
|
||
(1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—
|
||
<taxonomicName id="4C34C0F4FF98201DCAE2F89CA85CF891" authorityName="Zachvatkin" authorityYear="1953" box="[737,856,1892,1916]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF98201DCAE2F89CA85CF891" box="[737,856,1892,1916]" italics="true" pageId="3" pageNumber="92">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
(5—specimen from the type locality of
|
||
<taxonomicName id="4C34C0F4FF98201DCD07F89CAE99F891" baseAuthorityName="Zakhvatkin" baseAuthorityYear="1953" box="[1284,1437,1893,1916]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="mesasiatica">
|
||
<emphasis id="B9406765FF98201DCD07F89CAE99F891" box="[1284,1437,1893,1916]" italics="true" pageId="3" pageNumber="92">K. mesasiatica</emphasis>
|
||
</taxonomicName>
|
||
in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of
|
||
<taxonomicName id="4C34C0F4FF98201DCAB4F854A82AF829" authorityName="Zachvatkin" authorityYear="1953" box="[695,814,1964,1988]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF98201DCAB4F854A82AF829" box="[695,814,1964,1988]" italics="true" pageId="3" pageNumber="92">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
in Moscow Oblast); 9—
|
||
<taxonomicName id="4C34C0F4FF98201DCC35F854AFB9F829" authorityName="Zachvatkin" authorityYear="1953" box="[1078,1213,1964,1988]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="niveicolor">
|
||
<emphasis id="B9406765FF98201DCC35F854AFB9F829" box="[1078,1213,1964,1988]" italics="true" pageId="3" pageNumber="92">K. niveicolor</emphasis>
|
||
</taxonomicName>
|
||
; 10—
|
||
<taxonomicName id="4C34C0F4FF98201DCD03F854AE91F829" authorityName="Dlabola" authorityYear="1963" box="[1280,1429,1964,1988]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="auricillatus">
|
||
<emphasis id="B9406765FF98201DCD03F854AE91F829" box="[1280,1429,1964,1988]" italics="true" pageId="3" pageNumber="92">K. auricillatus</emphasis>
|
||
</taxonomicName>
|
||
; 11—
|
||
<taxonomicName id="4C34C0F4FF98201DC8CAF828AA23F805" baseAuthorityName="Edwards" baseAuthorityYear="1908" box="[201,295,2000,2024]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="populi">
|
||
<emphasis id="B9406765FF98201DC8CAF828AA23F805" box="[201,295,2000,2024]" italics="true" pageId="3" pageNumber="92">K. populi</emphasis>
|
||
</taxonomicName>
|
||
; 12—
|
||
<taxonomicName id="4C34C0F4FF98201DC964F828AAD3F805" baseAuthorityName="Ribaut" baseAuthorityYear="1933" box="[359,471,2001,2024]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="virgator">
|
||
<emphasis id="B9406765FF98201DC964F828AAD3F805" box="[359,471,2001,2024]" italics="true" pageId="3" pageNumber="92">K. virgator</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8BBB77FF9F201AC8C4FF6EA992FE3C" blockId="4.[151,1437,150,1221]" pageId="4" pageNumber="93">
|
||
In another article from the same collection,
|
||
<bibRefCitation id="EFA5C686FF9F201ACAA8FF6EA88CFF5C" author="Zachvatkin, A. A." box="[683,904,150,177]" pageId="4" pageNumber="93" pagination="237 - 245" refId="ref7692" refString="Zachvatkin, A. A. (1953 b) On the fauna of Eupterygidae (Homoptera, Cicadina) of Central Asia. In: Smirnov, E. S. (Ed.), Sbornik nauchnykh rabot. Moscow University, Moscow, pp. 237 - 245. [in Russian]" type="book chapter" year="1953">Zachvatkin (1953b)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C34C0F4FF9F201ACBFCFF6EAFAAFF5D" authorityName="sensu Zachvatkin" authorityYear="1953" box="[1023,1198,151,176]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="mesasiaticus">
|
||
<emphasis id="B9406765FF9F201ACBFCFF6EAFAAFF5D" box="[1023,1198,151,176]" italics="true" pageId="4" pageNumber="93">K. mesasiaticus</emphasis>
|
||
</taxonomicName>
|
||
from Yargak Village, Khatyrchi Region, Navoi Oblast (later belonged to Samarkand Oblast),
|
||
<collectingCountry id="F323FBE7FF9F201ACBA0FF42AF25FF38" box="[931,1057,187,213]" name="Uzbekistan" pageId="4" pageNumber="93">Uzbekistan</collectingCountry>
|
||
based on the large series, collected at
|
||
<date id="FF8A9DB7FF9F201AC8B7FF26AA22FF14" box="[180,294,223,249]" pageId="4" pageNumber="93" value="1928-07">July 1928</date>
|
||
; in the original description, it is stated that this species is widespread in the oasis regions of Central Asia (
|
||
<bibRefCitation id="EFA5C686FF9F201AC8D9FEFAAAB6FEF0" author="Zachvatkin, A. A." box="[218,434,258,285]" pageId="4" pageNumber="93" pagination="237 - 245" refId="ref7692" refString="Zachvatkin, A. A. (1953 b) On the fauna of Eupterygidae (Homoptera, Cicadina) of Central Asia. In: Smirnov, E. S. (Ed.), Sbornik nauchnykh rabot. Moscow University, Moscow, pp. 237 - 245. [in Russian]" type="book chapter" year="1953">Zachvatkin, 1953b</bibRefCitation>
|
||
). Only a series of
|
||
<specimenCount id="9D3270FEFF9F201ACA82FEFAA9FFFEF0" box="[641,763,259,285]" count="21" pageId="4" pageNumber="93" type="female">21 females</specimenCount>
|
||
with labels “Bukhara Oblast (sic!), env. Khatyrchi, Yargak,
|
||
<date id="FF8A9DB7FF9F201AC894FEDEAA3CFEAC" box="[151,312,295,321]" pageId="4" pageNumber="93" value="1928-08-14">14. VIII. 1928</date>
|
||
, Zachvatkin”, which are
|
||
<typeStatus id="548F05D5FF9F201ACA4EFEDEA9BEFEAC" box="[589,698,295,321]" pageId="4" pageNumber="93" type="topotype">topotypes</typeStatus>
|
||
of this taxon (
|
||
<figureCitation id="130FA7F2FF9F201ACB5AFEDEA89FFEAC" box="[857,923,294,321]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." figureDoi="http://doi.org/10.5281/zenodo.13330990" httpUri="https://zenodo.org/record/13330990/files/figure.png" pageId="4" pageNumber="93">Fig. 5</figureCitation>
|
||
) was found in the collection of the Zoological Museum of Moscow University. Our specimens from
|
||
<collectingCountry id="F323FBE7FF9F201ACB05FEB2A883FE88" box="[774,903,331,357]" name="Kyrgyzstan" pageId="4" pageNumber="93">Kyrgyzstan</collectingCountry>
|
||
do not differ from them in appearance; in the shape of genitalia and apodemes they are similar to the specimen of
|
||
<taxonomicName id="4C34C0F4FF9F201ACB88FE96AF3EFE65" authorityName="sensu Zachvatkin" authorityYear="1953" box="[907,1082,367,392]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="mesasiaticus">
|
||
<emphasis id="B9406765FF9F201ACB88FE96AF3EFE65" box="[907,1082,367,392]" italics="true" pageId="4" pageNumber="93">K. mesasiaticus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F323FBE7FF9F201ACC78FE96AE00FE64" box="[1147,1284,367,393]" name="Afghanistan" pageId="4" pageNumber="93">Afghanistan</collectingCountry>
|
||
illustrated by Dworakowska (1976: 445, figs 308–313). As a result, our specimens from
|
||
<collectingCountry id="F323FBE7FF9F201ACBD5FE6AAF53FE40" box="[982,1111,403,429]" name="Kyrgyzstan" pageId="4" pageNumber="93">Kyrgyzstan</collectingCountry>
|
||
can be confidently identified as
|
||
<taxonomicName id="4C34C0F4FF9F201AC8B5FE4EA996FE3C" authority="sensu Zachvatkin (1953 b)" authorityName="sensu Zachvatkin" authorityYear="1953" box="[182,658,438,465]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="mesasiaticus">
|
||
<emphasis id="B9406765FF9F201AC8B5FE4EAA61FE3D" box="[182,357,439,464]" italics="true" pageId="4" pageNumber="93">K. mesasiaticus</emphasis>
|
||
sensu
|
||
<bibRefCitation id="EFA5C686FF9F201AC9B1FE4EA996FE3C" author="Zachvatkin, A. A." box="[434,658,438,465]" pageId="4" pageNumber="93" pagination="237 - 245" refId="ref7692" refString="Zachvatkin, A. A. (1953 b) On the fauna of Eupterygidae (Homoptera, Cicadina) of Central Asia. In: Smirnov, E. S. (Ed.), Sbornik nauchnykh rabot. Moscow University, Moscow, pp. 237 - 245. [in Russian]" type="book chapter" year="1953">Zachvatkin (1953b)</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8BBB77FF9F201AC8C4FE22A9BFFD1C" blockId="4.[151,1437,150,1221]" pageId="4" pageNumber="93">
|
||
Comparison of specimens from Central Asia (
|
||
<figureCitation id="130FA7F2FF9F201ACAD4FE22A83AFE18" box="[727,830,474,501]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." figureDoi="http://doi.org/10.5281/zenodo.13330990" httpUri="https://zenodo.org/record/13330990/files/figure.png" pageId="4" pageNumber="93">Figs 5–7</figureCitation>
|
||
,
|
||
<figureCitation id="130FA7F2FF9F201ACB4FFE22A8EEFE18" box="[844,1002,474,501]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." figureDoi="http://doi.org/10.5281/zenodo.13330992" httpUri="https://zenodo.org/record/13330992/files/figure.png" pageId="4" pageNumber="93">17–23, 25–31</figureCitation>
|
||
) and
|
||
<collectingRegion id="49F07595FF9F201ACC2EFE22AFE4FE18" box="[1069,1248,475,501]" country="Russia" name="Moscow" pageId="4" pageNumber="93">Moscow Oblast</collectingRegion>
|
||
(
|
||
<figureCitation id="130FA7F2FF9F201ACCF2FE22AE3FFE18" box="[1265,1339,475,501]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." figureDoi="http://doi.org/10.5281/zenodo.13330990" httpUri="https://zenodo.org/record/13330990/files/figure.png" pageId="4" pageNumber="93">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="130FA7F2FF9F201ACD49FE22AE90FE18" box="[1354,1428,475,501]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." figureDoi="http://doi.org/10.5281/zenodo.13330992" httpUri="https://zenodo.org/record/13330992/files/figure.png" pageId="4" pageNumber="93">24, 32</figureCitation>
|
||
) showed, that they belong to the same species. Certain differences in the shape of the abdominal apodemes, as mentioned above, are the result of intrapopulation variability. The description of
|
||
<taxonomicName id="4C34C0F4FF9F201ACC28FDDAAFD8FDD1" authorityName="sensu Zachvatkin" authorityYear="1953" box="[1067,1244,547,572]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="mesasiaticus">
|
||
<emphasis id="B9406765FF9F201ACC28FDDAAFD8FDD1" box="[1067,1244,547,572]" italics="true" pageId="4" pageNumber="93">K. mesasiaticus</emphasis>
|
||
</taxonomicName>
|
||
indicates a body length of 3.78–4.22 mm for females (data on males are not given;
|
||
<bibRefCitation id="EFA5C686FF9F201ACB86FDBEAF5FFD8C" author="Zachvatkin, A. A." box="[901,1115,582,609]" pageId="4" pageNumber="93" pagination="237 - 245" refId="ref7692" refString="Zachvatkin, A. A. (1953 b) On the fauna of Eupterygidae (Homoptera, Cicadina) of Central Asia. In: Smirnov, E. S. (Ed.), Sbornik nauchnykh rabot. Moscow University, Moscow, pp. 237 - 245. [in Russian]" type="book chapter" year="1953">Zachvatkin, 1953b</bibRefCitation>
|
||
), while the description of
|
||
<taxonomicName id="4C34C0F4FF9F201ACD80FDBEABF3FD68" authorityName="Zachvatkin" authorityYear="1953" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF9F201ACD80FDBEABF3FD68" italics="true" pageId="4" pageNumber="93">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
indicates a body length of 3.79–4.22 mm (sex is not specified;
|
||
<bibRefCitation id="EFA5C686FF9F201ACBD1FD92AFACFD68" author="Zachvatkin, A. A." box="[978,1192,618,645]" pageId="4" pageNumber="93" pagination="205 - 223" refId="ref7639" refString="Zachvatkin, A. A. (1953 a) Biological and taxonomic notes on leafhoppers of Central Russian fauna. In: Smirnov, E. S. (Ed.), Sbornik nauchnykh rabot. Moscow University, Moscow, pp. 205 - 223. [in Russian]" type="book chapter" year="1953">Zachvatkin, 1953a</bibRefCitation>
|
||
). This contradicts the statement in
|
||
<bibRefCitation id="EFA5C686FF9F201AC92AFD76A902FD44" author="Zachvatkin, A. A." box="[297,518,654,681]" pageId="4" pageNumber="93" pagination="205 - 223" refId="ref7639" refString="Zachvatkin, A. A. (1953 a) Biological and taxonomic notes on leafhoppers of Central Russian fauna. In: Smirnov, E. S. (Ed.), Sbornik nauchnykh rabot. Moscow University, Moscow, pp. 205 - 223. [in Russian]" type="book chapter" year="1953">Zachvatkin (1953a)</bibRefCitation>
|
||
that these taxa differ in size. Such inconsistencies in these articles are due to the fact that they were published after the death of Prof. Zachvatkin in a collection of works, the manuscripts of which, in an unfinished form, were found in his archive.
|
||
</paragraph>
|
||
<paragraph id="8B8BBB77FF9F201AC8C4FD02AF19FC24" blockId="4.[151,1437,150,1221]" pageId="4" pageNumber="93">
|
||
Previously, as a first reviser, we chose for this species the valid name
|
||
<taxonomicName id="4C34C0F4FF9F201ACBE5FD02AF6CFCF8" authorityName="Zachvatkin" authorityYear="1953" box="[998,1128,763,789]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF9F201ACBE5FD02AF6CFCF8" box="[998,1128,763,789]" italics="true" pageId="4" pageNumber="93">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
and, following
|
||
<bibRefCitation id="EFA5C686FF9F201ACD1DFD02ABF4FCD4" author="Zachvatkin, A. A." pageId="4" pageNumber="93" pagination="205 - 223" refId="ref7639" refString="Zachvatkin, A. A. (1953 a) Biological and taxonomic notes on leafhoppers of Central Russian fauna. In: Smirnov, E. S. (Ed.), Sbornik nauchnykh rabot. Moscow University, Moscow, pp. 205 - 223. [in Russian]" type="book chapter" year="1953">Zachvatkin (1953a)</bibRefCitation>
|
||
, left the Central Asian taxon in a subspecies rank,
|
||
<taxonomicName id="4C34C0F4FF9F201ACB3FFCE6AE3EFCD4" authority="(Tishechkin, 2002)" baseAuthorityName="Tishechkin" baseAuthorityYear="2002" box="[828,1338,799,825]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="oshanini" subSpecies="mesasiaticus">
|
||
<emphasis id="B9406765FF9F201ACB3FFCE6AF53FCD5" box="[828,1111,799,825]" italics="true" pageId="4" pageNumber="93">K. oshanini mesasiaticus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA5C686FF9F201ACC6BFCE6AE36FCD4" author="Tishechkin, D. Y." box="[1128,1330,799,825]" pageId="4" pageNumber="93" pagination="1398 - 1401" refId="ref7514" refString="Tishechkin, D. Y. (2002) New data on taxonomy of the genus Kybos (Homoptera, Cicadellidae, Typhlocybinae) from central regions of European Russia. Zoologicheskiy Zhurnal, 81 (11), 1398 - 1401. [in Russian with English summary]" type="journal article" year="2002">Tishechkin, 2002</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Despite this, afterwards,
|
||
<taxonomicName id="4C34C0F4FF9F201AC951FCBAA906FCB1" authorityName="sensu Zachvatkin" authorityYear="1953" box="[338,514,835,860]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="mesasiaticus">
|
||
<emphasis id="B9406765FF9F201AC951FCBAA906FCB1" box="[338,514,835,860]" italics="true" pageId="4" pageNumber="93">K. mesasiaticus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C34C0F4FF9F201ACA3AFCBAA9BEFCB0" authorityName="Zachvatkin" authorityYear="1953" box="[569,698,835,861]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF9F201ACA3AFCBAA9BEFCB0" box="[569,698,835,861]" italics="true" pageId="4" pageNumber="93">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
were considered as separate species by
|
||
<bibRefCitation id="EFA5C686FF9F201ACC7EFCBAAE24FCB0" author="Mitjaev, I. D." box="[1149,1312,835,861]" pageId="4" pageNumber="93" pagination="1 - 170" refId="ref7271" refString="Mitjaev, I. D. (2002) Fauna, Ecology, and Zoogeography of leafhoppers (Homoptera, Cicadinea) of Kazakhstan. Tethys Entomological Research, 5, 1 - 170. [in Russian]" type="journal article" year="2002">Mitjaev (2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA5C686FF9F201ACD28FCBBAE61FCB0" author="Mitjaev, I. D." box="[1323,1381,834,861]" pageId="4" pageNumber="93" pagination="43 - 81" refId="ref7310" refString="Mitjaev, I. D. (2015) Leafhoppers (Homoptera, Cicadinea) of Kazakhstan, annotated check-list of species. Selevinia, 23, 43 - 81. [in Russian with English summary]" type="journal article" year="2015">2015</bibRefCitation>
|
||
) and
|
||
<bibRefCitation id="EFA5C686FF9F201AC894FC9EAA83FC6C" author="Dmitriev, D. A. & Anufriev, G. A. & Bartlett, C. R. & Blanco-Rodriguez, E. & Borodin, O. I. & Cao, Y. - H. & Deitz, L. L. & Dietrich, C. H. & Dmitrieva, M. O. & El-Sonbati, S. A. & Evangelista de Souza, O. & Gjonov, I. V. & Goncalves, A. C. & Hendrix, S. & McKamey, S. & Kohler, M. & Kunz, G. & Malenovsky, I. & Morris, B. O. & Novoselova, M. & Pinedo-Escatel, J. A. & Rakitov, R. A. & Rothschild, M. J. & Sanborn, A. F. & Takiya, D. M. & Wallace, M. S. & Zahniser, J. N." box="[151,391,871,897]" pageId="4" pageNumber="93" refId="ref6962" refString="Dmitriev, D. A., Anufriev, G. A., Bartlett, C. R., Blanco-Rodriguez, E., Borodin, O. I., Cao, Y. - H., Deitz, L. L., Dietrich, C. H., Dmitrieva, M. O., El-Sonbati, S. A., Evangelista de Souza, O., Gjonov, I. V., Goncalves, A. C., Hendrix, S., McKamey, S., Kohler, M., Kunz, G., Malenovsky, I., Morris, B. O., Novoselova, M., Pinedo-Escatel, J. A., Rakitov, R. A., Rothschild, M. J., Sanborn, A. F., Takiya, D. M., Wallace, M. S. & Zahniser, J. N. (2022 onward). Kybos Fieber, 1866. World Auchenorrhyncha Database. TaxonPages. Available from: https: // hoppers. speciesfile. org / otus / 43322 / overview (accessed 9 March 2024)" type="journal volume" year="2022">
|
||
Dmitriev
|
||
<emphasis id="B9406765FF9F201AC907FC9EAA39FC6C" box="[260,317,871,897]" italics="true" pageId="4" pageNumber="93">et al.</emphasis>
|
||
(2022
|
||
</bibRefCitation>
|
||
onward). Presently, after studying additional materials from different regions of
|
||
<collectingCountry id="F323FBE7FF9F201ACD19FC9EAE99FC6C" box="[1306,1437,871,897]" name="Kazakhstan" pageId="4" pageNumber="93">Kazakhstan</collectingCountry>
|
||
and
|
||
<collectingCountry id="F323FBE7FF9F201AC8C4FC72AA4EFC48" box="[199,330,907,933]" name="Kyrgyzstan" pageId="4" pageNumber="93">Kyrgyzstan</collectingCountry>
|
||
, we found no significant differences between the European and Central Asian taxa. In this regard, we establish synonymy
|
||
<taxonomicName id="4C34C0F4FF9F201AC9A0FC56A988FC24" authorityName="Zachvatkin" authorityYear="1953" box="[419,652,943,969]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="oshanini" subSpecies="oshanini">
|
||
<emphasis id="B9406765FF9F201AC9A0FC56A988FC24" box="[419,652,943,969]" italics="true" pageId="4" pageNumber="93">K. oshanini oshanini</emphasis>
|
||
</taxonomicName>
|
||
=
|
||
<taxonomicName id="4C34C0F4FF9F201ACAA9FC56A8C5FC25" baseAuthorityName="Tishechkin" baseAuthorityYear="2002" box="[682,961,943,969]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="oshanini" status="syn. n." subSpecies="mesasiaticus">
|
||
<emphasis id="B9406765FF9F201ACAA9FC56A8C5FC25" box="[682,961,943,969]" italics="true" pageId="4" pageNumber="93">K. oshanini mesasiaticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9406765FF9F201ACBCCFC56AF19FC24" bold="true" box="[975,1053,943,969]" pageId="4" pageNumber="93">
|
||
<taxonomicNameLabel id="A273DA1EFF9F201ACBCCFC56AF19FC24" box="[975,1053,943,969]" pageId="4" pageNumber="93">syn. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8BBB77FF9F201AC8C4FC2AAFB3FB28" blockId="4.[151,1437,150,1221]" pageId="4" pageNumber="93">
|
||
<taxonomicName id="4C34C0F4FF9F201AC8C4FC2AAACBFC00" authorityName="Mitjaev" authorityYear="1971" box="[199,463,979,1005]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="mesasiaticus" subSpecies="iliensis">
|
||
<emphasis id="B9406765FF9F201AC8C4FC2AAACBFC00" box="[199,463,979,1005]" italics="true" pageId="4" pageNumber="93">K. mesasiaticus iliensis</emphasis>
|
||
</taxonomicName>
|
||
was briefly described in a key (Mitjaev, 1971) from the region south of the Balkhash Lake and from the middle part of the Ili River valley. According to the drawings, it differs from the nominotypical subspecies in having shorter abdominal apodemes. Indeed, the male we collected in the Charyn River valley near the confluence with the Ili has short apodemes and fully corresponds to the drawings in Mitjaev (1971) (
|
||
<figureCitation id="130FA7F2FF9F201ACCF0FBC6AE3FFBB4" box="[1267,1339,1087,1113]" captionStart="FIGURES 1–12" captionStartId="3.[152,270,1856,1880]" captionTargetBox="[254,1332,185,1824]" captionTargetId="figure-15@3.[248,1338,179,1830]" captionTargetPageId="3" captionText="FIGURES 1–12. Kybos spp., dorsal habitus. 1–4—K. montanus (1–2—specimens from Inner Tien Shan, 3—specimen from West Tien Shan, 4—specimen from Alai Range); 5–8—K. oshanini (5—specimen from the type locality of K. mesasiatica in Uzbekistan, 6—specimen from Shekaftar, the northern edge of the Fergana Valley, 7—specimen from the Ili River basin, Kazakhstan, 8—specimen from the type locality of K. oshanini in Moscow Oblast); 9—K. niveicolor; 10—K. auricillatus; 11—K. populi; 12—K. virgator." figureDoi="http://doi.org/10.5281/zenodo.13330990" httpUri="https://zenodo.org/record/13330990/files/figure.png" pageId="4" pageNumber="93">Figs 7</figureCitation>
|
||
,
|
||
<figureCitation id="130FA7F2FF9F201ACD44FBC6AE8BFBB4" box="[1351,1423,1087,1113]" captionStart="FIGURES 13–37" captionStartId="5.[152,270,1804,1828]" captionTargetBox="[216,1373,187,1774]" captionTargetId="figure-15@5.[212,1376,181,1780]" captionTargetPageId="5" captionText="FIGURES 13–37. Kybos spp., abdominal apodemes. 13–16—K. montanus (13–14—dorsal apodemes, 15–16—ventral apodemes); 17–32—K. oshanini (17–24—dorsal apodemes, 25–32—ventral apodemes; 17–20, 25–28—specimens from Shekaftar, the northern edge of the Fergana Valley, 21–22, 29–30—specimens from the same locality in West Tien Shan, 23, 31—specimen from the Ili River basin, Kazakhstan, 24, 32—specimen from the type locality of K. oshanini in Moscow Oblast); 33—K. niveicolor, dorsal and ventral apodemes; 34–35—K. auricillatus (34—dorsal apodemes, 35—ventral apodemes); 36— K. populi, dorsal and ventral apodemes; 37—K. virgator, same." figureDoi="http://doi.org/10.5281/zenodo.13330992" httpUri="https://zenodo.org/record/13330992/files/figure.png" pageId="4" pageNumber="93">23, 31</figureCitation>
|
||
). Apparently, based on these differences, Dworakowska (1976) raised this taxon to the species rank,
|
||
<taxonomicName id="4C34C0F4FF9F201ACCCBFB9AABE2FB4C" baseAuthorityName="Mitjaev" baseAuthorityYear="1971" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="iliensis" subGenus="Kybos">
|
||
<emphasis id="B9406765FF9F201ACCCBFB9AABE2FB4C" italics="true" pageId="4" pageNumber="93">Empoasca (Kybos) iliensis</emphasis>
|
||
</taxonomicName>
|
||
. However, since the shape of apodemes in
|
||
<taxonomicName id="4C34C0F4FF9F201ACAC7FB7EA840FB4C" authorityName="Zachvatkin" authorityYear="1953" box="[708,836,1159,1185]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF9F201ACAC7FB7EA840FB4C" box="[708,836,1159,1185]" italics="true" pageId="4" pageNumber="93">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
can vary within the same population, we establish the synonymy
|
||
<taxonomicName id="4C34C0F4FF9F201AC910FB52AA90FB28" authorityName="Zachvatkin" authorityYear="1953" box="[275,404,1195,1221]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="oshanini">
|
||
<emphasis id="B9406765FF9F201AC910FB52AA90FB28" box="[275,404,1195,1221]" italics="true" pageId="4" pageNumber="93">K. oshanini</emphasis>
|
||
</taxonomicName>
|
||
=
|
||
<taxonomicName id="4C34C0F4FF9F201AC9B1FB52A9BCFB28" authorityName="Mitjaev" authorityYear="1971" box="[434,696,1195,1221]" class="Insecta" family="Cicadellidae" genus="Kybos" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="mesasiaticus" status="syn. n." subSpecies="iliensis">
|
||
<emphasis id="B9406765FF9F201AC9B1FB52A9BCFB28" box="[434,696,1195,1221]" italics="true" pageId="4" pageNumber="93">K. mesasiaticus iliensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9406765FF9F201ACAC5FB52A82EFB28" bold="true" box="[710,810,1195,1221]" pageId="4" pageNumber="93">
|
||
<taxonomicNameLabel id="A273DA1EFF9F201ACAC5FB52A817FB28" box="[710,787,1195,1221]" pageId="4" pageNumber="93">syn. n.</taxonomicNameLabel>
|
||
=
|
||
</emphasis>
|
||
<taxonomicName id="4C34C0F4FF9F201ACB32FB52AF5FFB28" baseAuthorityName="Mitjaev" baseAuthorityYear="1971" box="[817,1115,1195,1221]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="iliensis" status="syn. nov." subGenus="Kybos">
|
||
<emphasis id="B9406765FF9F201ACB32FB52AF5FFB28" box="[817,1115,1195,1221]" italics="true" pageId="4" pageNumber="93">Empoasca (Kybos) iliensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9406765FF9F201ACC69FB52AFB3FB28" bold="true" box="[1130,1207,1195,1221]" pageId="4" pageNumber="93">
|
||
<taxonomicNameLabel id="A273DA1EFF9F201ACC69FB52AFB3FB28" box="[1130,1207,1195,1221]" pageId="4" pageNumber="93">syn. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |