450 lines
53 KiB
XML
450 lines
53 KiB
XML
<document id="F1EF3A16CD5035EDA1BC6A47F882F270" ID-DOI="10.5281/zenodo.182961" ID-GBIF-Dataset="6312ffbb-3731-4cbe-9f81-f7a36a027038" ID-ISSN="1175-5326" ID-Zenodo-Dep="182961" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459884825173" checkinUser="plazi" docAuthor="Kirkendall, Lawrence R., Faccoli, Massimo & Ye, Hui" docDate="2008" docId="FB3B083C671AFF9FFF576F8BFA1CF85B" docLanguage="en" docName="zt01819p039.pdf" docOrigin="Zootaxa 1819" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tomicus Latreille, sp. n." docType="key" docVersion="10" lastPageNumber="31" masterDocId="070270446719FF99FFC06C0EFF80FF8B" masterDocTitle="Description of the Yunnan shoot borer, Tomicus yunnanensis Kirkendall & Faccoli sp. n. (Curculionidae, Scolytinae), an unusually aggressive pine shoot beetle from southern China, with a key to the species of Tomicus" masterLastPageNumber="39" masterPageNumber="25" pageNumber="28" updateTime="1698227703634" updateUser="plazi">
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<mods:titleInfo id="359978D07E4A2C3A60D5F5F9CF77972D">
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<mods:title id="53A2EDB30DD1E94C14D3BDAC38723600">Description of the Yunnan shoot borer, Tomicus yunnanensis Kirkendall & Faccoli sp. n. (Curculionidae, Scolytinae), an unusually aggressive pine shoot beetle from southern China, with a key to the species of Tomicus</mods:title>
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<mods:name id="6D3B96AEC18A19DB070120129C07FAF3" type="personal">
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<mods:roleTerm id="DB2E0AE26A8A302F6E832C0AA38CD27A">Author</mods:roleTerm>
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<mods:namePart id="BE71EF57455B91E15ADD1689AFB4AEDE">Kirkendall, Lawrence R.</mods:namePart>
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<mods:name id="82138CF3616C814DCB5F8E4D948712C7" type="personal">
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<mods:roleTerm id="D42815491B5CA277C39620E1A3A1EBDB">Author</mods:roleTerm>
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<mods:namePart id="742C7663E97865AEF5E24730F67F4219">Faccoli, Massimo</mods:namePart>
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</mods:name>
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<mods:name id="2F015580296515F544EE080C32641D12" type="personal">
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<mods:roleTerm id="E081217162B31D55A5565000D4AB5422">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BFD2BD89F639D7E28BA838867D6164BF">Ye, Hui</mods:namePart>
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<mods:typeOfResource id="B23482136F55D505DAD58C8AC9BD87B7">text</mods:typeOfResource>
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<mods:title id="A13FB7F3A98A2039B0778EBC100A1741">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5F124C1E02B6FD0C5A812CD8EB3CF555">
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<mods:date id="DF207CB130E2B3A418705FAA422957AB">2008</mods:date>
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<mods:detail id="FA4CF73238DD3CD8B6ED8BA3E95C1046" type="volume">
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<mods:number id="6684388FED158FA27C14B3CEB3CFA23B">1819</mods:number>
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<mods:classification id="FA432A68FD31DDE023B502E8933BDC2F">journal article</mods:classification>
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<mods:identifier id="422713447BB40B70CEA087824DB706C1" type="DOI">10.5281/zenodo.182961</mods:identifier>
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<mods:identifier id="79367B822578BCF5A86D3AD4A9E065F0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="670659F4CCCCF02B190294E47694784E" type="Zenodo-Dep">182961</mods:identifier>
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<treatment id="FB3B083C671AFF9FFF576F8BFA1CF85B" ID-DOI="http://doi.org/10.5281/zenodo.5657516" ID-Zenodo-Dep="5657516" LSID="urn:lsid:plazi:treatment:FB3B083C671AFF9FFF576F8BFA1CF85B" httpUri="http://treatment.plazi.org/id/FB3B083C671AFF9FFF576F8BFA1CF85B" lastPageId="6" lastPageNumber="31" pageId="3" pageNumber="28">
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<paragraph id="732DB92A671AFF9AFF576F8BFDF3FC14" blockId="3.[151,627,901,927]" box="[151,627,901,927]" pageId="3" pageNumber="28">
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<heading id="28650E46671AFF9AFF576F8BFDF3FC14" bold="true" box="[151,627,901,927]" fontSize="11" level="1" pageId="3" pageNumber="28" reason="1">
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<emphasis id="41E66538671AFF9AFF576F8BFDF3FC14" bold="true" box="[151,627,901,927]" pageId="3" pageNumber="28">
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Key to the species of
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<taxonomicName id="B492C2A9671AFF9AFE5C6F8BFDF3FC14" ID-CoL="8HS72" authority="Latreille" authorityName="Latreille" box="[412,627,901,927]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis id="41E66538671AFF9AFE5C6F8BFE7FFC14" bold="true" box="[412,511,901,927]" italics="true" pageId="3" pageNumber="28">Tomicus</emphasis>
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Latreille
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="732DB92A671AFF9AFF576FDEFAF3FB51" blockId="3.[151,1436,976,1242]" pageId="3" pageNumber="28">
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No key to the six previously published species of
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<taxonomicName id="B492C2A9671AFF9AFD346FDEFCD2FC62" box="[756,850,976,1001]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis id="41E66538671AFF9AFD346FDEFCD2FC62" box="[756,850,976,1001]" italics="true" pageId="3" pageNumber="28">Tomicus</emphasis>
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</taxonomicName>
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exists. The bark beetles killing trees in Yunnan, identified in the literature as
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<emphasis id="41E66538671AFF9AFDC76FF6FD22FB9A" box="[519,674,1016,1041]" italics="true" pageId="3" pageNumber="28">
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<taxonomicName id="B492C2A9671AFF9AFDC76FF6FD1FFB9A" box="[519,671,1016,1041]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="piniperda">T. piniperda</taxonomicName>
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,
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</emphasis>
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are clearly morphologically distinct from that taxon and are described as a new, seventh species below. The following key to all seven species of the genus is based on specimens examined by us.
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<taxonomicName id="B492C2A9671AFF9AFE276846FD26FBEA" box="[487,678,1096,1121]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="puellus">
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<emphasis id="41E66538671AFF9AFE276846FD26FBEA" box="[487,678,1096,1121]" italics="true" pageId="3" pageNumber="28">Tomicus puellus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="B492C2A9671AFF9AFD236846FCC9FBEA" box="[739,841,1096,1121]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="pilifer">
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<emphasis id="41E66538671AFF9AFD236846FCC9FBEA" box="[739,841,1096,1121]" italics="true" pageId="3" pageNumber="28">T. pilifer</emphasis>
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</taxonomicName>
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are rare in Western collections and nowhere well illustrated, and although we could study only a few specimens of these two species, they proved readily identifiable. Many characters used in the key vary intraspecifically and the extreme character states may overlap interspecifically; however, combinations of these characters reliably distinguish otherwise similar species.
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</paragraph>
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<subSubSection id="3B88EAA1671AFF9FFF57691EFA1CF85B" lastPageId="6" lastPageNumber="31" pageId="3" pageNumber="28" type="key">
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<keyStep id="C866A18F671AFF9AFF57691EFA1CFAF1" pageId="3" pageNumber="28">
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<paragraph id="732DB92A671AFF9AFF57691EFA1BFAD9" blockId="3.[151,1436,1295,2002]" pageId="3" pageNumber="28">
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<keyLead id="C8631A1F671AFF9AFF57691EFA1BFAD9" pageId="3" pageNumber="28">
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1.
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<emphasis id="41E66538671AFF9AFF226901FCF7FAA2" bold="true" box="[226,887,1295,1321]" pageId="3" pageNumber="28">Interstria 2 on declivity with rows of small granules</emphasis>
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, not impressed (weakly impressed in some
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<taxonomicName id="B492C2A9671AFF9AFA46691EFEA9FADA" ID-CoL="57D36" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="minor">
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<emphasis id="41E66538671AFF9AFA46691EFEA9FADA" italics="true" pageId="3" pageNumber="28">T. minor</emphasis>
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</taxonomicName>
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) (
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<figureCitation id="EBA9A5AF671AFF9AFE826936FE01FAD9" box="[322,385,1336,1362]" captionStart="FIGURE 1" captionStartId="4.[151,255,1750,1774]" captionTargetBox="[151,1434,683,1728]" captionTargetId="figure@4.[151,1436,683,1729]" captionTargetPageId="4" captionText="FIGURE 1. Eytral declivity of Tomicus species: (a, b) T. puellus (Reitter), (c) T. minor (Hartig), (d) T. pilifer (Spessivtsew)." httpUri="https://zenodo.org/record/182962/files/figure.png" pageId="3" pageNumber="28">Fig. 1</figureCitation>
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)...........................................................................................................................................2
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</keyLead>
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||
</paragraph>
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<paragraph id="732DB92A671AFF9AFF57696EFA1CFAF1" blockId="3.[151,1436,1295,2002]" box="[151,1436,1375,1402]" pageId="3" pageNumber="28">
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<keyLead id="C8631A1F671AFF9AFF57696EFA1CFAF1" box="[151,1436,1375,1402]" pageId="3" pageNumber="28">
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||
-
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||
<emphasis id="41E66538671AFF9AFF226951FDDDFAF2" bold="true" box="[226,605,1375,1401]" pageId="3" pageNumber="28">Interstria 2 devoid of granules</emphasis>
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, clearly impressed (
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<figureCitation id="EBA9A5AF671AFF9AFC89696EFC10FAF1" box="[841,912,1376,1402]" captionStart="FIGURE 2" captionStartId="5.[151,255,1405,1429]" captionTargetBox="[151,1434,354,1380]" captionTargetId="figure@5.[151,1436,354,1381]" captionTargetPageId="5" captionText="FIGURE 2. Eytral declivity of Tomicus species: (a) brevipilosus (Eggers), (b) T. piniperda (Linnaeus), (c) T. destruens (Wollaston), (d) T. yunnanensis sp. n. (holotype)." httpUri="https://zenodo.org/record/182963/files/figure.png" pageId="3" pageNumber="28">Fig. 2</figureCitation>
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) .................................................................. 4
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep id="C866A18F671AFF9AFF576986FA1CF8B9" pageId="3" pageNumber="28">
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<paragraph id="732DB92A671AFF9AFF576986FA1BF991" blockId="3.[151,1436,1295,2002]" pageId="3" pageNumber="28">
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||
<keyLead id="C8631A1F671AFF9AFF576986FA1BF991" pageId="3" pageNumber="28">
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2(1).
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<emphasis id="41E66538671AFF9AFF226989FD1CFA42" bold="true" pageId="3" pageNumber="28">Elytral vestiture consisting of longer, erect interstrial hairs (arising from granules) in uniseriate rows and shorter decumbent hairs</emphasis>
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(ground vestiture), erect hairs longer on declivity (
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<figureCitation id="EBA9A5AF671AFF9AFAC969BEFADBFA41" box="[1289,1371,1456,1482]" captionStart="FIGURE 1" captionStartId="4.[151,255,1750,1774]" captionTargetBox="[151,1434,683,1728]" captionTargetId="figure@4.[151,1436,683,1729]" captionTargetPageId="4" captionText="FIGURE 1. Eytral declivity of Tomicus species: (a, b) T. puellus (Reitter), (c) T. minor (Hartig), (d) T. pilifer (Spessivtsew)." httpUri="https://zenodo.org/record/182962/files/figure.png" pageId="3" pageNumber="28">Figs. 1</figureCitation>
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c, d). Elytral declivity with conspicuous interstrial tubercles in regular uniseriate rows. Larger species, length
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<quantity id="B46A14CF671AFF9AFF236A0EFEF4F991" box="[227,372,1536,1562]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="5.2" metricValueMin="3.1" pageId="3" pageNumber="28" unit="mm" value="4.15" valueMax="5.2" valueMin="3.1">3.1–5.2 mm</quantity>
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. Normal hosts
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<taxonomicName id="B492C2A9671AFF9AFDE36A0EFDE6F992" ID-CoL="6QPY" box="[547,614,1536,1561]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
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<emphasis id="41E66538671AFF9AFDE36A0EFDE6F992" box="[547,614,1536,1561]" italics="true" pageId="3" pageNumber="28">Pinus</emphasis>
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</taxonomicName>
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spp.......................................................................................................3
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</keyLead>
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</paragraph>
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<paragraph id="732DB92A671AFF9AFF576A29FA1CF8B9" blockId="3.[151,1436,1295,2002]" pageId="3" pageNumber="28">
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<keyLead id="C8631A1F671AFF9AFF576A29FA1CF8B9" pageId="3" pageNumber="28">
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<emphasis id="41E66538671AFF9AFF576A29FB0BF9CA" bold="true" box="[151,1163,1575,1601]" pageId="3" pageNumber="28">- Elytral interstrial hairs and ground vestiture equally short, dense, confused</emphasis>
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,
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<emphasis id="41E66538671AFF9AFB586A29FF7DF9E2" bold="true" pageId="3" pageNumber="28">decumbent or nearly so</emphasis>
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, not longer on declivity (
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<figureCitation id="EBA9A5AF671AFF9AFDE76A5EFDF5F9E1" box="[551,629,1616,1642]" captionStart="FIGURE 1" captionStartId="4.[151,255,1750,1774]" captionTargetBox="[151,1434,683,1728]" captionTargetId="figure@4.[151,1436,683,1729]" captionTargetPageId="4" captionText="FIGURE 1. Eytral declivity of Tomicus species: (a, b) T. puellus (Reitter), (c) T. minor (Hartig), (d) T. pilifer (Spessivtsew)." httpUri="https://zenodo.org/record/182962/files/figure.png" pageId="3" pageNumber="28">Figs. 1</figureCitation>
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a, b). Elytral interstriae strongly crenulate; interstrial tubercles large, transversely confluent, confused. Interstrial punctures confused on declivity, only slightly larger than strial punctures; interstrial tubercles inconspicuous on declivity. Smallest species, length
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<quantity id="B46A14CF671AFF9AFACD6AAEFA1CF931" box="[1293,1436,1696,1722]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.5" metricValueMin="2.9" pageId="3" pageNumber="28" unit="mm" value="3.2" valueMax="3.5" valueMin="2.9">2.9–3.5 mm</quantity>
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(ours 3.0–3.2). Maternal gallery monoramous, longitudinal. Distribution: Siberia, Russian Far East and Sakhalin Island. Normal hosts:
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<taxonomicName id="B492C2A9671AFF9AFD976AFEFC8BF882" ID-CoL="4HQ39" box="[599,779,1776,1801]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="jezoensis">
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<emphasis id="41E66538671AFF9AFD976AFEFC8BF882" box="[599,779,1776,1801]" italics="true" pageId="3" pageNumber="28">Picea jezoensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="B492C2A9671AFF9AFCDB6AFEFC27F882" ID-CoL="4HPZH" box="[795,935,1776,1801]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="ajanensis">
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<emphasis id="41E66538671AFF9AFCDB6AFEFC27F882" box="[795,935,1776,1801]" italics="true" pageId="3" pageNumber="28">P. ajanensis</emphasis>
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</taxonomicName>
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,
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<emphasis id="41E66538671AFF9AFC766AFEFAB7F882" box="[950,1335,1776,1801]" italics="true" pageId="3" pageNumber="28">
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<taxonomicName id="B492C2A9671AFF9AFC766AFEFB00F882" ID-CoL="63Z5X" box="[950,1152,1776,1801]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="holophylla">Abies holophylla</taxonomicName>
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,
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<taxonomicName id="B492C2A9671AFF9AFB4C6AFEFAB7F882" ID-CoL="63YW4" box="[1164,1335,1776,1801]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="nephrolepis">A. nephrolepis</taxonomicName>
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</emphasis>
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but also recorded from
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<emphasis id="41E66538671AFF9AFE516B16FB41F8BA" box="[401,1217,1816,1841]" italics="true" pageId="3" pageNumber="28">
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<taxonomicName id="B492C2A9671AFF9AFE516B16FDD3F8BA" ID-CoL="4J29P" box="[401,595,1816,1841]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="koraiensis">Pinus koraiensis</taxonomicName>
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....................................................................................
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</emphasis>
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<taxonomicName id="B492C2A9671AFF9AFB016B16FA1CF8B9" authority="(Reitter)" authorityName="Reitter" baseAuthorityName="Reitter" box="[1217,1436,1816,1842]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="puellus">
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<emphasis id="41E66538671AFF9AFB016B16FAB1F8BA" box="[1217,1329,1816,1841]" italics="true" pageId="3" pageNumber="28">T. puellus</emphasis>
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(Reitter)
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</taxonomicName>
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</keyLead>
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</paragraph>
|
||
</keyStep>
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<keyStep id="C866A18F671AFF9DFF576B4EFA1CFDC9" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28">
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<paragraph id="732DB92A671AFF9AFF576B4EFA1CF859" blockId="3.[151,1436,1295,2002]" pageId="3" pageNumber="28">
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<keyLead id="C8631A1F671AFF9AFF576B4EFA1CF859" pageId="3" pageNumber="28">
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3(2).
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<emphasis id="41E66538671AFF9AFF226B31FEA9F80A" bold="true" pageId="3" pageNumber="28">Interstrial punctures on disc and declivity fine points, difficult to see with normal lighting, not dense</emphasis>
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(
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<figureCitation id="EBA9A5AF671AFF9AFEF96B66FEFCF809" box="[313,380,1896,1922]" captionStart="FIGURE 1" captionStartId="4.[151,255,1750,1774]" captionTargetBox="[151,1434,683,1728]" captionTargetId="figure@4.[151,1436,683,1729]" captionTargetPageId="4" captionText="FIGURE 1. Eytral declivity of Tomicus species: (a, b) T. puellus (Reitter), (c) T. minor (Hartig), (d) T. pilifer (Spessivtsew)." httpUri="https://zenodo.org/record/182962/files/figure.png" pageId="3" pageNumber="28">Fig. 1</figureCitation>
|
||
c). Declivital ground vestiture absent or sparse and difficult to see, inconspicuous. Pronotal punctures sparse, most separated by much more than their diameter; most specimens with a distinct central impunctate longitudinal median strip (
|
||
<figureCitation id="EBA9A5AF671AFF9AFCC46BB6FCCEF859" box="[772,846,1976,2002]" captionStart="FIGURE 6" captionStartId="9.[151,255,1086,1110]" captionTargetBox="[151,1434,153,1026]" captionTargetId="figure@9.[151,1436,153,1027]" captionTargetPageId="9" captionText="FIGURE 6. Pronotum of Tomicus species: (a) T. minor (Hartig), (b) T. pilifer (Spessivtsew) (note difference in density of punctures)." httpUri="https://zenodo.org/record/182967/files/figure.png" pageId="3" pageNumber="28">Fig. 6</figureCitation>
|
||
a). Antennal club pale to medium brown, at most
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="732DB92A671DFF9DFF236C96FA1CFDC9" blockId="4.[151,1437,152,578]" pageId="4" pageNumber="29">
|
||
little darker than funicle (
|
||
<figureCitation id="EBA9A5AF671DFF9DFDCF6C96FDD6FF39" box="[527,598,152,178]" captionStart="FIGURE 3" captionStartId="6.[151,255,1467,1491]" captionTargetBox="[151,1434,309,1439]" captionTargetId="figure@6.[151,1436,309,1441]" captionTargetPageId="6" captionText="FIGURE 3. Antenna of Tomicus species: (a) T. brevipilosus (Eggers), (b) T. minor (Hartig), (c) T. puellus (Reitter), (d) T. pilifer (Spessivtsew)." httpUri="https://zenodo.org/record/182964/files/figure.png" pageId="4" pageNumber="29">Fig. 3</figureCitation>
|
||
b). Larger, length
|
||
<quantity id="B46A14CF671DFF9DFCEA6C96FC38FF39" box="[810,952,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="5.2" metricValueMin="3.2" pageId="4" pageNumber="29" unit="mm" value="4.2" valueMax="5.2" valueMin="3.2">3.2–5.2 mm</quantity>
|
||
(Yunnan,
|
||
<quantity id="B46A14CF671DFF9DFBF26C96FB45FF39" box="[1074,1221,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.6" metricValueMin="4.1" pageId="4" pageNumber="29" unit="mm" value="4.35" valueMax="4.6" valueMin="4.1">4.1–4.6 mm</quantity>
|
||
). Maternal gallery biramous, transverse. Distribution: Europe and Asia. Host-trees: all
|
||
<taxonomicName id="B492C2A9671DFF9DFC3F6CCEFBC2FF52" ID-CoL="6QPY" box="[1023,1090,192,217]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="41E66538671DFF9DFC3F6CCEFBC2FF52" box="[1023,1090,192,217]" italics="true" pageId="4" pageNumber="29">Pinus</emphasis>
|
||
</taxonomicName>
|
||
spp in its range .....................
|
||
<emphasis id="41E66538671DFF9DFF236CE6FB4FFE8A" box="[227,1231,232,257]" italics="true" pageId="4" pageNumber="29">.........................................................................................................................................</emphasis>
|
||
<taxonomicName id="B492C2A9671DFF9DFB146CE6FA1CFE89" ID-CoL="57D36" authority="(Hartig)" authorityName="Hartig" baseAuthorityName="Hartig" box="[1236,1436,232,258]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="41E66538671DFF9DFB146CE6FAB6FE8A" box="[1236,1334,232,257]" italics="true" pageId="4" pageNumber="29">T. minor</emphasis>
|
||
(Hartig)
|
||
</taxonomicName>
|
||
<keyLead id="C8631A1F671DFF9DFF576D1EFA1CFDC9" pageId="4" pageNumber="29">
|
||
-
|
||
<emphasis id="41E66538671DFF9DFF226D01FB04FEA2" bold="true" box="[226,1156,271,297]" pageId="4" pageNumber="29">Interstrial punctures on disc and declivity conspicuous, uniformly dense</emphasis>
|
||
(
|
||
<figureCitation id="EBA9A5AF671DFF9DFB586D1EFB5FFEA1" box="[1176,1247,272,298]" captionStart="FIGURE 1" captionStartId="4.[151,255,1750,1774]" captionTargetBox="[151,1434,683,1728]" captionTargetId="figure@4.[151,1436,683,1729]" captionTargetPageId="4" captionText="FIGURE 1. Eytral declivity of Tomicus species: (a, b) T. puellus (Reitter), (c) T. minor (Hartig), (d) T. pilifer (Spessivtsew)." httpUri="https://zenodo.org/record/182962/files/figure.png" pageId="4" pageNumber="29">Fig. 1</figureCitation>
|
||
d), on declivity only slightly smaller than, or equal to, strial punctures.
|
||
<emphasis id="41E66538671DFF9DFCB86D39FB18FEDA" bold="true" box="[888,1176,311,337]" pageId="4" pageNumber="29">Declivity densely hairy</emphasis>
|
||
due to abundant conspicuous decumbent ground vestiture (but hairs can be completely worn away in older specimens). Pronotal punctures dense, separated on average by about their diameter, no impunctate median strip (
|
||
<figureCitation id="EBA9A5AF671DFF9DFF2C6DBEFEB4FE41" box="[236,308,432,458]" captionStart="FIGURE 6" captionStartId="9.[151,255,1086,1110]" captionTargetBox="[151,1434,153,1026]" captionTargetId="figure@9.[151,1436,153,1027]" captionTargetPageId="9" captionText="FIGURE 6. Pronotum of Tomicus species: (a) T. minor (Hartig), (b) T. pilifer (Spessivtsew) (note difference in density of punctures)." httpUri="https://zenodo.org/record/182967/files/figure.png" pageId="4" pageNumber="29">Fig. 6</figureCitation>
|
||
b). Antennal club brown to dark brown, distinctly darker than funicle (
|
||
<figureCitation id="EBA9A5AF671DFF9DFBBA6DBEFB41FE41" box="[1146,1217,432,458]" captionStart="FIGURE 3" captionStartId="6.[151,255,1467,1491]" captionTargetBox="[151,1434,309,1439]" captionTargetId="figure@6.[151,1436,309,1441]" captionTargetPageId="6" captionText="FIGURE 3. Antenna of Tomicus species: (a) T. brevipilosus (Eggers), (b) T. minor (Hartig), (c) T. puellus (Reitter), (d) T. pilifer (Spessivtsew)." httpUri="https://zenodo.org/record/182964/files/figure.png" pageId="4" pageNumber="29">Fig. 3</figureCitation>
|
||
d). Smaller, length 3.0–
|
||
<quantity id="B46A14CF671DFF9DFED76DD6FEF4FE79" box="[279,372,472,498]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="29" unit="mm" value="4.3">4.3 mm</quantity>
|
||
(3.0–
|
||
<quantity id="B46A14CF671DFF9DFE7C6DD6FD9AFE79" box="[444,538,472,498]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="29" unit="mm" value="3.8">3.8 mm</quantity>
|
||
in the specimens examined). Maternal gallery monoramous, longitudinal. Distribution: Siberia and Russian Far East, Heilongjiang Province in
|
||
<collectingCountry id="0B85F9BA671DFF9DFBCE6E0EFBD8FD91" box="[1038,1112,512,538]" name="China" pageId="4" pageNumber="29">China</collectingCountry>
|
||
. Hosts:
|
||
<taxonomicName id="B492C2A9671DFF9DFB756E0EFAF7FD92" ID-CoL="4J29P" box="[1205,1399,512,537]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="species" species="koraiensis">
|
||
<emphasis id="41E66538671DFF9DFB756E0EFAF7FD92" box="[1205,1399,512,537]" italics="true" pageId="4" pageNumber="29">Pinus koraiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B492C2A9671DFF9DFA466E0EFECEFDCA" ID-CoL="4J22Q" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="species" species="armandii">
|
||
<emphasis id="41E66538671DFF9DFA466E0EFECEFDCA" italics="true" pageId="4" pageNumber="29">P. armandii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="41E66538671DFF9DFE9D6E26FB0EFDCA" box="[349,1166,552,577]" italics="true" pageId="4" pageNumber="29">
|
||
<taxonomicName id="B492C2A9671DFF9DFE9D6E26FD9CFDCA" box="[349,540,552,577]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="species" species="tabulaeformis">P. tabulaeformis</taxonomicName>
|
||
....................................................................................
|
||
</emphasis>
|
||
<taxonomicName id="B492C2A9671DFF9DFB4F6E26FA1CFDC9" ID-CoL="57D3L" authority="(Spessivtsew)" authorityName="Spessivtsew" baseAuthorityName="Spessivtsew" box="[1167,1436,552,578]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="pilifer">
|
||
<emphasis id="41E66538671DFF9DFB4F6E26FB71FDCA" box="[1167,1265,552,577]" italics="true" pageId="4" pageNumber="29">T. pilifer</emphasis>
|
||
(Spessivtsew)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<caption id="27EDE9A2671DFF9DFF576AD8FE9CF89B" httpUri="https://zenodo.org/record/182962/files/figure.png" pageId="4" pageNumber="29" targetBox="[151,1434,683,1728]" targetPageId="4">
|
||
<paragraph id="732DB92A671DFF9DFF576AD8FE9CF89B" blockId="4.[151,1436,1750,1808]" pageId="4" pageNumber="29">
|
||
<emphasis id="41E66538671DFF9DFF576AD8FE99F965" bold="true" box="[151,281,1750,1774]" pageId="4" pageNumber="29">FIGURE 1.</emphasis>
|
||
Eytral declivity of
|
||
<taxonomicName id="B492C2A9671DFF9DFE2B6AD8FDC1F966" ID-CoL="8HS72" box="[491,577,1750,1773]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="41E66538671DFF9DFE2B6AD8FDC1F966" box="[491,577,1750,1773]" italics="true" pageId="4" pageNumber="29">Tomicus</emphasis>
|
||
</taxonomicName>
|
||
species: (a, b)
|
||
<taxonomicName id="B492C2A9671DFF9DFD266AD8FC32F966" authority="Reitter" authorityName="Reitter" box="[742,946,1750,1773]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="puellus">
|
||
<emphasis id="41E66538671DFF9DFD266AD8FCCCF966" box="[742,844,1750,1773]" italics="true" pageId="4" pageNumber="29">T. puellus</emphasis>
|
||
(Reitter)
|
||
</taxonomicName>
|
||
, (c)
|
||
<taxonomicName id="B492C2A9671DFF9DFC226AD8FB1DF966" ID-CoL="57D36" authority="Hartig" authorityName="Hartig" box="[994,1181,1750,1773]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="41E66538671DFF9DFC226AD8FBBAF966" box="[994,1082,1750,1773]" italics="true" pageId="4" pageNumber="29">T. minor</emphasis>
|
||
(Hartig)
|
||
</taxonomicName>
|
||
, (d)
|
||
<taxonomicName id="B492C2A9671DFF9DFB0E6AD8FE98F89B" ID-CoL="57D3L" authority="Spessivtsew" authorityName="Spessivtsew" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="pilifer">
|
||
<emphasis id="41E66538671DFF9DFB0E6AD8FAAAF966" box="[1230,1322,1750,1773]" italics="true" pageId="4" pageNumber="29">T. pilifer</emphasis>
|
||
(Spessivtsew)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<keyStep id="C866A18F671DFF9CFF576B4DFA1CFEA1" lastPageId="5" lastPageNumber="30" pageId="4" pageNumber="29">
|
||
<paragraph id="732DB92A671DFF9DFF576B4DFA1BF85E" blockId="4.[151,1436,1859,2005]" pageId="4" pageNumber="29">
|
||
<keyLead id="C8631A1F671DFF9DFF576B4DFA1BF85E" pageId="4" pageNumber="29">
|
||
4(1).
|
||
<emphasis id="41E66538671DFF9DFF226B4DFC06F8D6" bold="true" box="[226,902,1859,1885]" pageId="4" pageNumber="29">Fine punctures of interstria 2 on declivity uniseriate;</emphasis>
|
||
punctures of striae 1 to 3 on declivity more than twice as large as fine interstrial punctures (
|
||
<figureCitation id="EBA9A5AF671DFF9DFCE96B65FCFEF80E" box="[809,894,1899,1925]" captionStart="FIGURE 2" captionStartId="5.[151,255,1405,1429]" captionTargetBox="[151,1434,354,1380]" captionTargetId="figure@5.[151,1436,354,1381]" captionTargetPageId="5" captionText="FIGURE 2. Eytral declivity of Tomicus species: (a) brevipilosus (Eggers), (b) T. piniperda (Linnaeus), (c) T. destruens (Wollaston), (d) T. yunnanensis sp. n. (holotype)." httpUri="https://zenodo.org/record/182963/files/figure.png" pageId="4" pageNumber="29">Figs. 2</figureCitation>
|
||
a, b). Granules on interstriae 2 and 3 on disc closely spaced, most by a distance equal to 1.5–2.5 punctures of adjacent striae. Antennal club brown, same color as or darker than funicle (
|
||
<figureCitation id="EBA9A5AF671DFF9DFD566BB5FD67F85E" box="[662,743,1979,2005]" captionStart="FIGURE 3" captionStartId="6.[151,255,1467,1491]" captionTargetBox="[151,1434,309,1439]" captionTargetId="figure@6.[151,1436,309,1441]" captionTargetPageId="6" captionText="FIGURE 3. Antenna of Tomicus species: (a) T. brevipilosus (Eggers), (b) T. minor (Hartig), (c) T. puellus (Reitter), (d) T. pilifer (Spessivtsew)." httpUri="https://zenodo.org/record/182964/files/figure.png" pageId="4" pageNumber="29">Figs. 3</figureCitation>
|
||
a, 4c)...................................................................................5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="732DB92A671CFF9CFF576C96FA1CFEA1" blockId="5.[151,1436,151,298]" pageId="5" pageNumber="30">
|
||
<keyLead id="C8631A1F671CFF9CFF576C96FA1CFEA1" pageId="5" pageNumber="30">
|
||
-
|
||
<emphasis id="41E66538671CFF9CFF226C99FBFEFF3A" bold="true" box="[226,1150,151,177]" pageId="5" pageNumber="30">Fine punctures of interstria 2 on declivity confused or appearing biseriate</emphasis>
|
||
, punctures of striae 1 to 3 on declivity less than or equal to twice as large as fine interstrial punctures (
|
||
<figureCitation id="EBA9A5AF671CFF9CFBB86CCEFB4BFF51" box="[1144,1227,192,218]" captionStart="FIGURE 2" captionStartId="5.[151,255,1405,1429]" captionTargetBox="[151,1434,354,1380]" captionTargetId="figure@5.[151,1436,354,1381]" captionTargetPageId="5" captionText="FIGURE 2. Eytral declivity of Tomicus species: (a) brevipilosus (Eggers), (b) T. piniperda (Linnaeus), (c) T. destruens (Wollaston), (d) T. yunnanensis sp. n. (holotype)." httpUri="https://zenodo.org/record/182963/files/figure.png" pageId="5" pageNumber="30">Figs. 2</figureCitation>
|
||
c, d). Granules on interstriae 2 and 3 on disc closely or distantly spaced. Antenna uniformly colored, club yellow to yellow-brown (
|
||
<figureCitation id="EBA9A5AF671CFF9CFEB36D1EFE45FEA1" box="[371,453,272,298]" captionStart="FIGURE 4" captionStartId="7.[151,255,1513,1537]" captionTargetBox="[151,1434,480,1486]" captionTargetId="figure@7.[151,1436,480,1487]" captionTargetPageId="7" captionText="FIGURE 4. Antenna of Tomicus species: (a, b) T. yunnanensis, (c) T. piniperda (Linnaeus), (d) T. destruens (Wollaston)." httpUri="https://zenodo.org/record/182965/files/figure.png" pageId="5" pageNumber="30">Figs. 4</figureCitation>
|
||
a, b, d).........................................................................................................................6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<caption id="27EDE9A2671CFF9CFF576973FD2AFA3C" httpUri="https://zenodo.org/record/182963/files/figure.png" pageId="5" pageNumber="30" targetBox="[151,1434,354,1380]" targetPageId="5">
|
||
<paragraph id="732DB92A671CFF9CFF576973FD2AFA3C" blockId="5.[151,1433,1405,1464]" pageId="5" pageNumber="30">
|
||
<emphasis id="41E66538671CFF9CFF576973FE9AFA1E" bold="true" box="[151,282,1405,1429]" pageId="5" pageNumber="30">FIGURE 2.</emphasis>
|
||
Eytral declivity of
|
||
<taxonomicName id="B492C2A9671CFF9CFE2E6970FDC3FA1E" ID-CoL="8HS72" box="[494,579,1406,1429]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="41E66538671CFF9CFE2E6970FDC3FA1E" box="[494,579,1406,1429]" italics="true" pageId="5" pageNumber="30">Tomicus</emphasis>
|
||
</taxonomicName>
|
||
species: (a)
|
||
<taxonomicName id="B492C2A9671CFF9CFD0F6970FC37FA1F" ID-CoL="57CZJ" authority="Eggers" authorityName="Eggers" box="[719,951,1405,1429]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="brevipilosus">
|
||
<emphasis id="41E66538671CFF9CFD0F6970FCD0FA1E" box="[719,848,1406,1429]" italics="true" pageId="5" pageNumber="30">brevipilosus</emphasis>
|
||
(Eggers)
|
||
</taxonomicName>
|
||
, (b)
|
||
<taxonomicName id="B492C2A9671CFF9CFC296970FB69FA1F" ID-CoL="57D3N" authority="Linnaeus" authorityName="Linnaeus" box="[1001,1257,1405,1429]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="piniperda">
|
||
<emphasis id="41E66538671CFF9CFC296970FBECFA1E" box="[1001,1132,1406,1429]" italics="true" pageId="5" pageNumber="30">T. piniperda</emphasis>
|
||
(Linnaeus)
|
||
</taxonomicName>
|
||
, (c)
|
||
<emphasis id="41E66538671CFF9CFADA6970FE99FA3C" italics="true" pageId="5" pageNumber="30">
|
||
<taxonomicName id="B492C2A9671CFF9CFADA6970FE95FA3C" ID-CoL="57CZZ" authority="Wollaston" authorityName="Wollaston" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="destruens">T. destruens (Wollaston)</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
(d)
|
||
<taxonomicName id="B492C2A9671CFF9CFE8C69AEFE6AFA3C" ID-CoL="4J2L5" box="[332,490,1440,1463]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="30" phylum="Tracheophyta" rank="species" species="yunnanensis" status="sp. nov.">
|
||
<emphasis id="41E66538671CFF9CFE8C69AEFE6AFA3C" box="[332,490,1440,1463]" italics="true" pageId="5" pageNumber="30">T. yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5AD5D843671CFF9CFE3069AEFDACFA33" box="[496,556,1440,1464]" pageId="5" pageNumber="30" rank="species">
|
||
<emphasis id="41E66538671CFF9CFE3069AEFDACFA33" bold="true" box="[496,556,1440,1464]" pageId="5" pageNumber="30">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="732DB92A671CFF9CFF5769E5FA1CF97E" blockId="5.[151,1436,1514,2021]" pageId="5" pageNumber="30">
|
||
<keyLead id="C8631A1F671CFF9CFF5769E5FA1CF97E" pageId="5" pageNumber="30">
|
||
<keyStep id="C866A18F671CFF9CFF5769E5FA1CF97E" pageId="5" pageNumber="30">
|
||
5(4).
|
||
<emphasis id="41E66538671CFF9CFF2269E4FD05F98F" bold="true" box="[226,645,1514,1540]" pageId="5" pageNumber="30">Erect elytral hairs on disc longer</emphasis>
|
||
, about as long as distance between striae; erect hairs on declivity distinctly longer than those on disc (
|
||
<figureCitation id="EBA9A5AF671CFF9CFD5D6A1DFD66F9A6" box="[669,742,1555,1581]" captionStart="FIGURE 2" captionStartId="5.[151,255,1405,1429]" captionTargetBox="[151,1434,354,1380]" captionTargetId="figure@5.[151,1436,354,1381]" captionTargetPageId="5" captionText="FIGURE 2. Eytral declivity of Tomicus species: (a) brevipilosus (Eggers), (b) T. piniperda (Linnaeus), (c) T. destruens (Wollaston), (d) T. yunnanensis sp. n. (holotype)." httpUri="https://zenodo.org/record/182963/files/figure.png" pageId="5" pageNumber="30">Fig. 2</figureCitation>
|
||
b). Antennal club brown, antenna uniformly colored (
|
||
<figureCitation id="EBA9A5AF671CFF9CFAAD6A1DFF74F9DE" captionStart="FIGURE 4" captionStartId="7.[151,255,1513,1537]" captionTargetBox="[151,1434,480,1486]" captionTargetId="figure@7.[151,1436,480,1487]" captionTargetPageId="7" captionText="FIGURE 4. Antenna of Tomicus species: (a, b) T. yunnanensis, (c) T. piniperda (Linnaeus), (d) T. destruens (Wollaston)." httpUri="https://zenodo.org/record/182965/files/figure.png" pageId="5" pageNumber="30">Fig. 4</figureCitation>
|
||
c). Interstria 2 on declivity strongly impressed and concave, with uniseriate, regularly spaced fine punctures (
|
||
<figureCitation id="EBA9A5AF671CFF9CFEA76A6DFE2FF9F6" box="[359,431,1635,1661]" captionStart="FIGURE 2" captionStartId="5.[151,255,1405,1429]" captionTargetBox="[151,1434,354,1380]" captionTargetId="figure@5.[151,1436,354,1381]" captionTargetPageId="5" captionText="FIGURE 2. Eytral declivity of Tomicus species: (a) brevipilosus (Eggers), (b) T. piniperda (Linnaeus), (c) T. destruens (Wollaston), (d) T. yunnanensis sp. n. (holotype)." httpUri="https://zenodo.org/record/182963/files/figure.png" pageId="5" pageNumber="30">Fig. 2</figureCitation>
|
||
b). More slender, elytra 1.7–1.8x longer than wide; larger, length
|
||
<quantity id="B46A14CF671CFF9CFB7F6A6DFACFF9F6" box="[1215,1359,1635,1661]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="5.2" metricValueMin="3.5" pageId="5" pageNumber="30" unit="mm" value="4.35" valueMax="5.2" valueMin="3.5">3.5–5.2 mm</quantity>
|
||
(Jilin:
|
||
<quantity id="B46A14CF671CFF9CFF236A85FEF6F92E" box="[227,374,1675,1701]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.2" metricValueMin="4.4" pageId="5" pageNumber="30" unit="mm" value="4.8" valueMax="5.2" valueMin="4.4">4.4–5.2 mm</quantity>
|
||
). Elytra usually longer than twice width of pronotum. Maternal gallery monoramous, longitudinal. Distribution: Eurasia including
|
||
<collectingCountry id="0B85F9BA671CFF9CFD116ABDFC94F946" box="[721,788,1715,1741]" name="Japan" pageId="5" pageNumber="30">Japan</collectingCountry>
|
||
but not known from Yunnan. Hosts: continental
|
||
<taxonomicName id="B492C2A9671CFF9CFA996ABDFA1CF947" ID-CoL="6QPY" box="[1369,1436,1715,1740]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="30" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="41E66538671CFF9CFA996ABDFA1CF947" box="[1369,1436,1715,1740]" italics="true" pageId="5" pageNumber="30">Pinus</emphasis>
|
||
</taxonomicName>
|
||
spp. and
|
||
<taxonomicName id="B492C2A9671CFF9CFE8C6AD5FE75F97F" ID-CoL="77KSH" box="[332,501,1755,1780]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="30" phylum="Tracheophyta" rank="species" species="pinaster">
|
||
<emphasis id="41E66538671CFF9CFE8C6AD5FE75F97F" box="[332,501,1755,1780]" italics="true" pageId="5" pageNumber="30">Pinus pinaster</emphasis>
|
||
</taxonomicName>
|
||
, not known from
|
||
<emphasis id="41E66538671CFF9CFD076AD5FB05F97F" box="[711,1157,1755,1780]" italics="true" pageId="5" pageNumber="30">
|
||
<taxonomicName id="B492C2A9671CFF9CFD076AD5FCF3F97F" ID-CoL="4J2L5" box="[711,883,1755,1780]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="30" phylum="Tracheophyta" rank="species" species="yunnanensis">P. yunnanensis</taxonomicName>
|
||
.....................................
|
||
</emphasis>
|
||
<taxonomicName id="B492C2A9671CFF9CFB486AD5FA1CF97E" ID-CoL="57D3N" authority="(Linnaeus)" authorityName="Linnaeus" baseAuthorityName="Linnaeus" box="[1160,1436,1755,1781]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="piniperda">
|
||
<emphasis id="41E66538671CFF9CFB486AD5FA96F97F" box="[1160,1302,1755,1780]" italics="true" pageId="5" pageNumber="30">T. piniperda</emphasis>
|
||
(Linnaeus)
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="732DB92A671CFF9FFF576B0CFA1CFF51" blockId="5.[151,1436,1514,2021]" lastBlockId="6.[227,1436,152,218]" lastPageId="6" lastPageNumber="31" pageId="5" pageNumber="30">
|
||
<keyLead id="C8631A1F671CFF9CFF576B0CFA1CF86E" pageId="5" pageNumber="30">
|
||
<emphasis id="41E66538671CFF9CFF576B0CFD02F897" bold="true" box="[151,642,1794,1820]" pageId="5" pageNumber="30">- Erect elytral hairs on disc shorter</emphasis>
|
||
, about 0.5x as long as distance between striae; erect hairs on declivity as long as those on disc (
|
||
<figureCitation id="EBA9A5AF671CFF9CFDFE6B25FD08F8CE" box="[574,648,1835,1861]" captionStart="FIGURE 2" captionStartId="5.[151,255,1405,1429]" captionTargetBox="[151,1434,354,1380]" captionTargetId="figure@5.[151,1436,354,1381]" captionTargetPageId="5" captionText="FIGURE 2. Eytral declivity of Tomicus species: (a) brevipilosus (Eggers), (b) T. piniperda (Linnaeus), (c) T. destruens (Wollaston), (d) T. yunnanensis sp. n. (holotype)." httpUri="https://zenodo.org/record/182963/files/figure.png" pageId="5" pageNumber="30">Fig. 2</figureCitation>
|
||
a). Antennal club brown to dark brown, usually noticeably darker than funicle (
|
||
<figureCitation id="EBA9A5AF671CFF9CFE416B5DFE48F8E6" box="[385,456,1875,1901]" captionStart="FIGURE 3" captionStartId="6.[151,255,1467,1491]" captionTargetBox="[151,1434,309,1439]" captionTargetId="figure@6.[151,1436,309,1441]" captionTargetPageId="6" captionText="FIGURE 3. Antenna of Tomicus species: (a) T. brevipilosus (Eggers), (b) T. minor (Hartig), (c) T. puellus (Reitter), (d) T. pilifer (Spessivtsew)." httpUri="https://zenodo.org/record/182964/files/figure.png" pageId="5" pageNumber="30">Fig. 3</figureCitation>
|
||
a). Interstria 2 on declivity weakly impressed, with punctures very fine, uniseriate, sparse, often widely spaced or even absent on much of declivity (
|
||
<figureCitation id="EBA9A5AF671CFF9CFC366B75FBBFF81E" box="[1014,1087,1915,1941]" captionStart="FIGURE 2" captionStartId="5.[151,255,1405,1429]" captionTargetBox="[151,1434,354,1380]" captionTargetId="figure@5.[151,1436,354,1381]" captionTargetPageId="5" captionText="FIGURE 2. Eytral declivity of Tomicus species: (a) brevipilosus (Eggers), (b) T. piniperda (Linnaeus), (c) T. destruens (Wollaston), (d) T. yunnanensis sp. n. (holotype)." httpUri="https://zenodo.org/record/182963/files/figure.png" pageId="5" pageNumber="30">Fig. 2</figureCitation>
|
||
a). Stouter, elytra 1.6x longer than wide; smaller, length
|
||
<quantity id="B46A14CF671CFF9CFDD66BADFD23F836" box="[534,675,1955,1981]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.4" metricValueMin="3.2" pageId="5" pageNumber="30" unit="mm" value="3.8" valueMax="4.4" valueMin="3.2">3.2–4.4 mm</quantity>
|
||
(Yunnan: 4.0–
|
||
<quantity id="B46A14CF671CFF9CFC8F6BADFC2DF836" box="[847,941,1955,1981]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="30" unit="mm" value="4.4">4.4 mm</quantity>
|
||
). Elytra shorter than twice width of pronotum. Maternal gallery unknown, but probably monoramous and longitudinal. Distribution: Fukien,
|
||
</keyLead>
|
||
Fujian and Yunnan (
|
||
<collectingCountry id="0B85F9BA671FFF9FFE206C96FDACFF39" box="[480,556,152,178]" name="China" pageId="6" pageNumber="31">China</collectingCountry>
|
||
), Assam (
|
||
<collectingCountry id="0B85F9BA671FFF9FFD696C96FD6BFF39" box="[681,747,152,178]" name="India" pageId="6" pageNumber="31">India</collectingCountry>
|
||
),
|
||
<collectingCountry id="0B85F9BA671FFF9FFCC56C96FCCFFF39" box="[773,847,152,178]" name="South Korea" pageId="6" pageNumber="31">Korea</collectingCountry>
|
||
,
|
||
<collectingCountry id="0B85F9BA671FFF9FFCA06C96FC27FF39" box="[864,935,152,178]" name="Japan" pageId="6" pageNumber="31">Japan</collectingCountry>
|
||
. Hosts:
|
||
<taxonomicName id="B492C2A9671FFF9FFBCE6C96FB56FF3A" ID-CoL="4J29P" box="[1038,1238,152,177]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="koraiensis">
|
||
<emphasis id="41E66538671FFF9FFBCE6C96FB56FF3A" box="[1038,1238,152,177]" italics="true" pageId="6" pageNumber="31">Pinus koraiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="41E66538671FFF9FFB2A6C96FAEFFF3A" box="[1258,1391,152,177]" italics="true" pageId="6" pageNumber="31">P. i n s u l a r i s</emphasis>
|
||
,
|
||
<taxonomicName id="B492C2A9671FFF9FFA466C96FED6FF52" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="parvifolia">
|
||
<emphasis id="41E66538671FFF9FFA466C96FED6FF52" italics="true" pageId="6" pageNumber="31">P. parvifolia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="41E66538671FFF9FFEA56CCEFBFDFF52" box="[357,1149,192,217]" italics="true" pageId="6" pageNumber="31">
|
||
<taxonomicName id="B492C2A9671FFF9FFEA56CCEFD92FF52" ID-CoL="4J2L5" box="[357,530,192,217]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="yunnanensis">P. yunnanensis</taxonomicName>
|
||
...................................................................................
|
||
</emphasis>
|
||
<taxonomicName id="B492C2A9671FFF9FFB456CCEFA1CFF51" ID-CoL="57CZJ" authority="(Eggers)" authorityName="Eggers" baseAuthorityName="Eggers" box="[1157,1436,192,218]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="brevipilosus">
|
||
<emphasis id="41E66538671FFF9FFB456CCEFAB0FF52" box="[1157,1328,192,217]" italics="true" pageId="6" pageNumber="31">T. brevipilosus</emphasis>
|
||
(Eggers)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<caption id="27EDE9A2671FFF9FFF5769B5FEFAFA7E" httpUri="https://zenodo.org/record/182964/files/figure.png" pageId="6" pageNumber="31" targetBox="[151,1434,309,1439]" targetPageId="6">
|
||
<paragraph id="732DB92A671FFF9FFF5769B5FEFAFA7E" blockId="6.[151,1437,1467,1525]" pageId="6" pageNumber="31">
|
||
<emphasis id="41E66538671FFF9FFF5769B5FE98FA58" bold="true" box="[151,280,1467,1491]" pageId="6" pageNumber="31">FIGURE 3.</emphasis>
|
||
Antenna of
|
||
<taxonomicName id="B492C2A9671FFF9FFE5C69B5FE71FA59" ID-CoL="8HS72" box="[412,497,1467,1490]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="41E66538671FFF9FFE5C69B5FE71FA59" box="[412,497,1467,1490]" italics="true" pageId="6" pageNumber="31">Tomicus</emphasis>
|
||
</taxonomicName>
|
||
species: (a)
|
||
<taxonomicName id="B492C2A9671FFF9FFDB969B5FCF9FA59" ID-CoL="57CZJ" authority="Eggers" authorityName="Eggers" box="[633,889,1467,1490]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="brevipilosus">
|
||
<emphasis id="41E66538671FFF9FFDB969B5FC95FA59" box="[633,789,1467,1490]" italics="true" pageId="6" pageNumber="31">T. brevipilosus</emphasis>
|
||
(Eggers)
|
||
</taxonomicName>
|
||
, (b)
|
||
<taxonomicName id="B492C2A9671FFF9FFC6869B5FBE1FA59" ID-CoL="57D36" authority="Hartig" authorityName="Hartig" box="[936,1121,1467,1490]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="41E66538671FFF9FFC6869B5FB80FA59" box="[936,1024,1467,1490]" italics="true" pageId="6" pageNumber="31">T. minor</emphasis>
|
||
(Hartig)
|
||
</taxonomicName>
|
||
, (c)
|
||
<taxonomicName id="B492C2A9671FFF9FFB4F69B5FAD8FA59" authority="Reitter" authorityName="Reitter" box="[1167,1368,1467,1490]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="puellus">
|
||
<emphasis id="41E66538671FFF9FFB4F69B5FB75FA59" box="[1167,1269,1467,1490]" italics="true" pageId="6" pageNumber="31">T. puellus</emphasis>
|
||
(Reitter)
|
||
</taxonomicName>
|
||
, (d
|
||
<emphasis id="41E66538671FFF9FFAB969B5FA02FA59" box="[1401,1410,1467,1490]" italics="true" pageId="6" pageNumber="31">)</emphasis>
|
||
<taxonomicName id="B492C2A9671FFF9FFA4869B5FEF6FA7E" ID-CoL="57D3L" authority="(Spessivtsew)" authorityName="Spessivtsew" baseAuthorityName="Spessivtsew" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="pilifer">
|
||
<emphasis id="41E66538671FFF9FFA4869B5FF58FA7E" italics="true" pageId="6" pageNumber="31">T. pilifer</emphasis>
|
||
(Spessivtsew)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<keyStep id="C866A18F671FFF9FFF576A40FA1CF85B" pageId="6" pageNumber="31">
|
||
<paragraph id="732DB92A671FFF9FFF576A40FA1CF80B" blockId="6.[151,1436,1614,2000]" pageId="6" pageNumber="31">
|
||
<keyLead id="C8631A1F671FFF9FFF576A40FA1CF80B" pageId="6" pageNumber="31">
|
||
6(4).
|
||
<emphasis id="41E66538671FFF9FFF226A40FD87F9E3" bold="true" box="[226,519,1614,1640]" pageId="6" pageNumber="31">Mediterranean species.</emphasis>
|
||
Interstria 2 on declivity weakly impressed, punctures dense, confused (
|
||
<figureCitation id="EBA9A5AF671FFF9FFAAD6A40FF74F91B" captionStart="FIGURE 2" captionStartId="5.[151,255,1405,1429]" captionTargetBox="[151,1434,354,1380]" captionTargetId="figure@5.[151,1436,354,1381]" captionTargetPageId="5" captionText="FIGURE 2. Eytral declivity of Tomicus species: (a) brevipilosus (Eggers), (b) T. piniperda (Linnaeus), (c) T. destruens (Wollaston), (d) T. yunnanensis sp. n. (holotype)." httpUri="https://zenodo.org/record/182963/files/figure.png" pageId="6" pageNumber="31">Fig. 2</figureCitation>
|
||
c). Granules of interstriae 2 and 3 on disc closely spaced, most by a distance equal to 1.5–2.5 punctures of adjacent striae; granules of interstriae 1 and 3 on declivity widely spaced, the distance between adjacent granules within a row equal to ca. 2/3 the distance between rows of granules on interstriae 1 and 3. Only base of elytra dark. Protibia with five or six teeth, usually evenly spaced in one cluster (
|
||
<figureCitation id="EBA9A5AF671FFF9FFAAC6AE0FF74F8BB" captionStart="FIGURE 5" captionStartId="8.[151,255,1965,1989]" captionTargetBox="[167,1420,1104,1916]" captionTargetId="figure@8.[151,1436,1091,1939]" captionTargetPageId="8" captionText="FIGURE 5. Protibiae of Tomicus species: T. destruens (Wollaston) (left), T. yunnanensis sp. n. (right)." httpUri="https://zenodo.org/record/182966/files/figure.png" pageId="6" pageNumber="31">Fig. 5</figureCitation>
|
||
). Length
|
||
<quantity id="B46A14CF671FFF9FFEA46B18FE75F8BB" box="[356,501,1814,1840]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.9" metricValueMin="4.1" pageId="6" pageNumber="31" unit="mm" value="4.5" valueMax="4.9" valueMin="4.1">4.1–4.9 mm</quantity>
|
||
. Maternal gallery monoramous, longitudinal. Distribution: Mediterranean basin and Atlantic coastal regions of
|
||
<collectingCountry id="0B85F9BA671FFF9FFD906B30FD16F8D3" box="[592,662,1854,1880]" name="Spain" pageId="6" pageNumber="31">Spain</collectingCountry>
|
||
,
|
||
<collectingCountry id="0B85F9BA671FFF9FFD616B30FC84F8D3" box="[673,772,1854,1880]" name="Portugal" pageId="6" pageNumber="31">Portugal</collectingCountry>
|
||
and North Africa. Hosts: Mediterranean
|
||
<taxonomicName id="B492C2A9671FFF9FFB276B30FAAAF8DC" ID-CoL="6QPY" box="[1255,1322,1854,1879]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="41E66538671FFF9FFB276B30FAAAF8DC" box="[1255,1322,1854,1879]" italics="true" pageId="6" pageNumber="31">Pinus</emphasis>
|
||
</taxonomicName>
|
||
species ..
|
||
<emphasis id="41E66538671FFF9FFF236B68FBFBF8F4" box="[227,1147,1894,1919]" italics="true" pageId="6" pageNumber="31">..............................................................................................................................</emphasis>
|
||
<taxonomicName id="B492C2A9671FFF9FFBBB6B68FA1CF80B" ID-CoL="57CZZ" authority="(Wollaston)" authorityName="Wollaston" baseAuthorityName="Wollaston" box="[1147,1436,1894,1920]" class="Insecta" family="Curculionidae" genus="Tomicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="destruens">
|
||
<emphasis id="41E66538671FFF9FFBBB6B68FA8BF8F4" box="[1147,1291,1894,1919]" italics="true" pageId="6" pageNumber="31">T. destruens</emphasis>
|
||
(Wollaston)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="732DB92A671FFF9FFF576B80FA1CF85B" blockId="6.[151,1436,1614,2000]" pageId="6" pageNumber="31">
|
||
<keyLead id="C8631A1F671FFF9FFF576B80FA1CF85B" pageId="6" pageNumber="31">
|
||
-
|
||
<emphasis id="41E66538671FFF9FFF226B80FE09F823" bold="true" box="[226,393,1934,1960]" pageId="6" pageNumber="31">Asian species</emphasis>
|
||
. Interstria 2 on declivity strongly impressed but broadly convex to flat, punctures more evenly spaced, appearing biseriate or irregularly uniseriate (appearing to zig-zag down declivity) (
|
||
<figureCitation id="EBA9A5AF671FFF9FFAA16BB8FA1CF85B" box="[1377,1436,1974,2000]" captionStart="FIGURE 2" captionStartId="5.[151,255,1405,1429]" captionTargetBox="[151,1434,354,1380]" captionTargetId="figure@5.[151,1436,354,1381]" captionTargetPageId="5" captionText="FIGURE 2. Eytral declivity of Tomicus species: (a) brevipilosus (Eggers), (b) T. piniperda (Linnaeus), (c) T. destruens (Wollaston), (d) T. yunnanensis sp. n. (holotype)." httpUri="https://zenodo.org/record/182963/files/figure.png" pageId="6" pageNumber="31">Figs.</figureCitation>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |