257 lines
28 KiB
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257 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1182.97346" ID-Pensoft-Pub="1313-2970-1182-165" ID-Pensoft-UUID="DCECB4AAD9E95ADB80188D91F386DD1C" ID-ZooBank="86F5AC45D35E4DDFAB042BCBDEEE8407" ModsDocID="1313-2970-1182-165" checkinTime="1697273694970" checkinUser="pensoft" docAuthor="Choi, Yeon-Jae, Park, Sun-Jae, Lee, Seung-Gyu & Park, Jong-Seok" docDate="2023" docId="C69925DF89585287953CD03ACA412693" docLanguage="en" docName="ZooKeys 1182: 165-181" docOrigin="ZooKeys 1182" docPubDate="2023-10-13" docSource="http://dx.doi.org/10.3897/zookeys.1182.97346" docTitle="Bryaxis nemorosus Choi, Park, Lee & Park 2023, sp. nov." docType="treatment" docUuid="812B1845-BAEB-4E11-A8BD-7F51B922433F" docUuidSource="ZooBank" docVersion="1" id="DCECB4AAD9E95ADB80188D91F386DD1C" lastPageNumber="165" masterDocId="DCECB4AAD9E95ADB80188D91F386DD1C" masterDocTitle="Five new species of Bryaxis Kugelann (Coleoptera, Staphylinidae, Pselaphinae) from Korea and a nomenclatural note on Bryaxis mahunkai Loebl" masterLastPageNumber="181" masterPageNumber="165" pageNumber="165" updateTime="1697273694970" updateUser="pensoft">
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<mods:mods id="182E0E059AA4ED9AC4E1D9D2E1059D78" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="900830568929F3AD64E138656BB4E9D4">
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<mods:title id="83976B8DC9C472A9F99A41FE491B6340">Five new species of Bryaxis Kugelann (Coleoptera, Staphylinidae, Pselaphinae) from Korea and a nomenclatural note on Bryaxis mahunkai Loebl</mods:title>
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<mods:name id="BF5C9C46441CBE64F47A5BC50DAD5796" type="personal">
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<mods:roleTerm id="738783309221DECAA32FE759799D12E4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E234AF5F6DCE32E39501905F14B038E6">Choi, Yeon-Jae</mods:namePart>
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<mods:nameIdentifier id="F466556180B37CE68AA77548144C1538" type="ORCID">https://orcid.org/0000-0003-0172-1128</mods:nameIdentifier>
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<mods:affiliation id="6A01BEDA4392EA3448D4376BB4BA7E35">Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju 28644, Republic of Korea</mods:affiliation>
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</mods:name>
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<mods:name id="5F28549D939DB0F0E6A8313DA2FDCD61" type="personal">
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<mods:roleTerm id="CA5756B1CAB0F1099131F9DE43C3E80D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7B6CDD3EB12BED14EBEAD58366C628FE">Park, Sun-Jae</mods:namePart>
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<mods:nameIdentifier id="B8118C26C2D199B8FC0924B8626A9B64" type="ORCID">https://orcid.org/0000-0003-3759-6517</mods:nameIdentifier>
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<mods:affiliation id="C1CB1DE9FE2CFED9BFA812F771C93C86">Climate Change and Environmental Biology Research Division, National Institute of Biological Resources, Incheon 22689, Republic of Korea</mods:affiliation>
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</mods:name>
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<mods:name id="CB739C186F55CA21ECC24584F6EF3E9B" type="personal">
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<mods:role id="DAD1BAF2AF2C5B1A7C16CB8E4F061CFE">
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<mods:roleTerm id="F643A63428B50D0F803499D89A963691">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6380205C59D1B6A914690551A6A21599">Lee, Seung-Gyu</mods:namePart>
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<mods:nameIdentifier id="93BDFBD70D9387B83A61FA334087806D" type="ORCID">https://orcid.org/0000-0001-5576-2799</mods:nameIdentifier>
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<mods:affiliation id="D8C0A3EB3FDD659FB0B7FF92C3E4AAB8">Climate Change and Environmental Biology Research Division, National Institute of Biological Resources, Incheon 22689, Republic of Korea</mods:affiliation>
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</mods:name>
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<mods:name id="25A6A152F316A74827DC71789F096343" type="personal">
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</mods:role>
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<mods:namePart id="A9FE051A95FB0C972ECAA35606771177">Park, Jong-Seok</mods:namePart>
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<mods:nameIdentifier id="B411B46C5AB8822C24312D9F21DD44C2" type="ORCID">https://orcid.org/0000-0003-2063-2534</mods:nameIdentifier>
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<mods:affiliation id="2B090D8D1B54B03612A0DDA1C8CC01F1">Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju 28644, Republic of Korea</mods:affiliation>
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<mods:nameIdentifier id="DCE93ACFE589BC393BA544BFA168182C" type="email">jpark16@cbnu.ac.kr</mods:nameIdentifier>
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<mods:typeOfResource id="134E0D6A6E913C63E74CA37D1FF8683F">text</mods:typeOfResource>
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<mods:titleInfo id="069886A31500B46A4AEC84E7372EC3B9">
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<mods:title id="C370873BC6E6A98ECB9FB460B3688072">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="C23FF7C75FD43CA5F2A54FAB043DE7BE">
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<mods:date id="7E4DDCC2931DE9E0750282D119380E0F">2023</mods:date>
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<mods:number id="AC8578EAB157F65EE39FF55B60B0C50E">2023-10-13</mods:number>
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<mods:detail id="D2F17E1B9CF8F00D6916BBA879351540" type="volume">
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<mods:number id="F3D6E018490D493CBA3E7F3DAC4B1AC9">1182</mods:number>
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<mods:start id="B205B85CEE2F43F45AF05DC8BF9CF5FC">165</mods:start>
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<mods:end id="02E3F553AF12652FB821597F43614EA0">181</mods:end>
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<mods:url id="4D02F0C72D1E522F38B0CB60D14264F2">http://dx.doi.org/10.3897/zookeys.1182.97346</mods:url>
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<mods:classification id="4FCBD9FC5906AE415933A806CCBA8163">journal article</mods:classification>
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<mods:identifier id="10BB9263A2EB3856F4F14CA1174E373C" type="DOI">http://dx.doi.org/10.3897/zookeys.1182.97346</mods:identifier>
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<mods:identifier id="6CB1FF99D360F8D320C719F028104E2D" type="Pensoft-Pub">1313-2970-1182-165</mods:identifier>
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<mods:identifier id="E64CC2731180594A39E432067BA9812F" type="ZooBank">86F5AC45D35E4DDFAB042BCBDEEE8407</mods:identifier>
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<treatment id="C69925DF89585287953CD03ACA412693" LSID="urn:lsid:zoobank.org:act:812B1845-BAEB-4E11-A8BD-7F51B922433F" httpUri="http://treatment.plazi.org/id/C69925DF89585287953CD03ACA412693" lastPageNumber="165" pageId="0" pageNumber="165">
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<subSubSection id="E06BFF202BD10A22FBD24A11532C0FB3" pageId="0" pageNumber="165" type="nomenclature">
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<paragraph id="849FCE789A1362B4289611E526E2049D" pageId="0" pageNumber="165">
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<taxonomicName id="FF3F35266BB297952FDF70AC17BBC608" LSID="https://zoobank.org/812B1845-BAEB-4E11-A8BD-7F51B922433F" authority="Choi, Park, Lee & Park" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis nemorosus" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="nemorosus" status="sp. nov.">Bryaxis nemorosus Choi, Park, Lee & Park</taxonomicName>
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<taxonomicNameLabel id="D20C50A19BC92656A10BEB6571958FFF" pageId="0" pageNumber="165">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="5D4495A0FFC94ABEC11CC25312426CEF" pageId="0" pageNumber="165" type="description">
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<paragraph id="2A9EC23D4BB32D8A82C69A541F7FC18D" pageId="0" pageNumber="165">
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<figureCitation id="C3A56129D9841265DE5EEBB774C3C83F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">Figs 8</figureCitation>
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<figureCitation id="F6199D112E0C08DAD4E62A85B2D3514F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diagnostic characters of Bryaxis girinensis sp. nov. (A, C, E) and B. nemorosus sp. nov. (B, D, F). A, B maxillary palpi C, D protibiae E, F heads. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure9" httpUri="https://binary.pensoft.net/fig/920952" pageId="0" pageNumber="165">, 9B, D, F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C46B6BCB01548353EE581365C9C92BF9" pageId="0" pageNumber="165" type="type material">
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<paragraph id="E88911C3F92C909C85AA2ED19515D90D" pageId="0" pageNumber="165">Type materials</paragraph>
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<paragraph id="468042CBFC9A7F70CD0BB815BC072CE7" pageId="0" pageNumber="165">
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<emphasis id="7E76613B10590D3C2CE16FFBD66A4447" bold="true" pageId="0" pageNumber="165">
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(
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<emphasis id="9B0980D4408CCD82F3E942424E7C89C9" italics="true" pageId="0" pageNumber="165">N</emphasis>
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= 5, 4
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</emphasis>
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♂♂,
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<emphasis id="BBE28B7C9EE8607039D18D24F25008A0" bold="true" pageId="0" pageNumber="165">1</emphasis>
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♀)
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<emphasis id="1969BA4F3BE39D58E6F2B284B9A27B7B" bold="true" pageId="0" pageNumber="165">
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.
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<emphasis id="A3D3B1C488B6EC25CCFC3A95009FC0AB" italics="true" pageId="0" pageNumber="165">Holotype male</emphasis>
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.
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</emphasis>
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"Korea: Jeonnam Prov. Mt. Doksil, Gageo island. Gageodo-gil, Heuksan-myeon, Sinan-gun, 13 Jul 2021,
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<geoCoordinate id="C43E7D126F34D4A7AC8D8186F209BB59" degrees="34" direction="north" minutes="05" orientation="latitude" precision="1" seconds="06.1" value="34.085026">34°05'06.1"N</geoCoordinate>
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,
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<geoCoordinate id="E8327AA47D5C1E0610D9FA7C96E4A6DF" degrees="125" direction="east" minutes="06" orientation="longitude" precision="1" seconds="17.4" value="125.104836">125°06'17.4"E</geoCoordinate>
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, 468 m, sifting leaf & soil litter, J.-W. Seo" (NIBR).
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<emphasis id="C24B06CAC3BE835E55ECCE3E302E10BF" bold="true" pageId="0" pageNumber="165">
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<emphasis id="76E7A61374FAB15264F5B17F7D359D3D" italics="true" pageId="0" pageNumber="165">Paratype</emphasis>
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.
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</emphasis>
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1♂ (CBNUIC, slide mounted). "Korea: Jeonnam Prov. Mt. Doksil, Gageo island. Gageodo-gil, Heuksan-myeon, Sinan-gun, 13 Jul 2021,
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<geoCoordinate id="5B247AE5537C77C144E7A78EBF043C06" degrees="34" direction="north" minutes="05" orientation="latitude" precision="1" seconds="06.1" value="34.085026">34°05'06.1"N</geoCoordinate>
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,
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<geoCoordinate id="1B80DC092F8DA416C2E98B1B70FA6D10" degrees="125" direction="east" minutes="06" orientation="longitude" precision="1" seconds="17.4" value="125.104836">125°06'17.4"E</geoCoordinate>
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, 468 m, sifting leaf & soil litter, J.-W. Seo". 1♂ (CBNUIC, dried). "Korea: Jeonnam Prov. Mt. Doksil, Gageo island. Gageodo-gil, Heuksan-myeon, Sinan-gun, 8 Jul 2020,
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<geoCoordinate id="0AB947DE04FD71F2A30AC36CE6C8B721" degrees="34" direction="north" minutes="05" orientation="latitude" precision="1" seconds="35.0" value="34.093056">34°05'35.0"N</geoCoordinate>
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,
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<geoCoordinate id="5E332F59713253B7E5E786377CB31787" degrees="125" direction="east" minutes="06" orientation="longitude" precision="1" seconds="25.0" value="125.10694">125°06'25.0"E</geoCoordinate>
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, 590 m, sifting leaf & soil litter, T.-Y. Jang". 1♂ (CBNUIC, dried). "Korea: Jeonnam Prov. Mt. Doksil, Gageo island. Gageodo-gil, Heuksan-myeon, Sinan-gun, 8 Jul 2020,
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<geoCoordinate id="C0A074194796BFDF67F431F2B1A00991" degrees="34" direction="north" minutes="04" orientation="latitude" precision="1" seconds="40.0" value="34.077778">34°04'40.0"N</geoCoordinate>
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,
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<geoCoordinate id="06F81B962BFF6B711A9519C512665B13" degrees="125" direction="east" minutes="06" orientation="longitude" precision="1" seconds="23.0" value="125.106384">125°06'23.0"E</geoCoordinate>
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, 540 m, sifting leaf & soil litter, U.-J. Byeon, T.-Y. Jang". 1♀ (CBNUIC, dried). "Korea: Jeonnam Prov. Mt. Doksil, Gageo island. Gageodo-gil, Heuksan-myeon, Sinan-gun, 7 Jul 2020,
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<geoCoordinate id="C99CEBFE0174AC91210CB92EECBA8803" degrees="34" direction="north" minutes="04" orientation="latitude" precision="1" seconds="34.7" value="34.076305">34°04'34.7"N</geoCoordinate>
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,
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<geoCoordinate id="074C1EDCEA9E19A9589FCCB1D1A837C1" degrees="125" direction="east" minutes="06" orientation="longitude" precision="1" seconds="28.8" value="125.108">125°06'28.8"E</geoCoordinate>
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, 534 m, sifting leaf & soil litter, T.-Y. Jang".
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</paragraph>
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</subSubSection>
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<subSubSection id="578F82F046B71AF1B6826F5D54910F19" pageId="0" pageNumber="165" type="diagnosis">
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<paragraph id="B96F879BF7FD05BD305AC224E185F52D" pageId="0" pageNumber="165">Diagnosis.</paragraph>
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<paragraph id="39A18CB4D3AA06AEB6D845D84C6DB865" pageId="0" pageNumber="165">
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Elongated head with small eyes situated on mid-length of head (Fig.
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<figureCitation id="AED28EA4E4B4345CB7637F8AAF1DBA63" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">8E</figureCitation>
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, arrow).
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</paragraph>
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<caption id="71A51AA5A8AE0BAE7F03C8E56D1539BB" doi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165" start="Figure 8" startId="F8">
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<paragraph id="7BC9F94FE13EBDD6B71621326BF661C3" pageId="0" pageNumber="165">
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<emphasis id="73DB9C3224D2EA38FA95793FAB791B21" bold="true" pageId="0" pageNumber="165">Figure 8.</emphasis>
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<taxonomicName id="98AE8A67EEB198CC2951549D35A36AD7" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis nemorosus" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="nemorosus">
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<emphasis id="4F08BA1FA8AC05EE0EAD368DF74731B6" italics="true" pageId="0" pageNumber="165">Bryaxis nemorosus</emphasis>
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</taxonomicName>
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Choi, Park, Lee & Park, sp. nov. male (
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<emphasis id="44DBF3980FAE3F9B1286E0A71CE86D5B" bold="true" pageId="0" pageNumber="165">A, B, E-I</emphasis>
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), female (
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<emphasis id="2AAB7993228AB44DA16F2859A7BF8CC9" bold="true" pageId="0" pageNumber="165">C, D</emphasis>
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).
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<emphasis id="CD01B93429F07F6A77651E4B685A4DB7" bold="true" pageId="0" pageNumber="165">A, C</emphasis>
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dorsal habitus
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<emphasis id="955BB56EFC973AA4DB467689D9D98835" bold="true" pageId="0" pageNumber="165">B, D</emphasis>
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ventral habitus
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<emphasis id="D9E8408D753105BED50AEB3D15B1EBCB" bold="true" pageId="0" pageNumber="165">E</emphasis>
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head
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<emphasis id="AB6E73AE2289D6B062B6A6CF72C63031" bold="true" pageId="0" pageNumber="165">F</emphasis>
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antenna
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<emphasis id="0871FDE05564E32DFCCCDC30A40A8ADE" bold="true" pageId="0" pageNumber="165">G</emphasis>
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fore leg
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<emphasis id="985C301D0E7A1902BE8D53C3738FF517" bold="true" pageId="0" pageNumber="165">H</emphasis>
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hind leg
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<emphasis id="EEB25F978E050925F7BB6B5A2A2DE054" bold="true" pageId="0" pageNumber="165">I</emphasis>
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aedeagus. Scale bars: 1 mm (
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<emphasis id="9B6434042DC09F6946928828CF30112E" bold="true" pageId="0" pageNumber="165">A-D</emphasis>
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); 0.1 mm (
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<emphasis id="FAE4BBD8BD4E7F0F9A3841B5AEB47B26" bold="true" pageId="0" pageNumber="165">E, I</emphasis>
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); 0.5 mm (
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<emphasis id="CD9FACA8D05EE2D3A6E49D2A009621F4" bold="true" pageId="0" pageNumber="165">F-H</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="1BDA0A204D1B6D18665B887E1D2B373C" pageId="0" pageNumber="165" type="description">
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<paragraph id="D02B8A4210A06B4838AD6562681CE5C8" pageId="0" pageNumber="165">Description.</paragraph>
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<paragraph id="C01371DEAE8BADA31A30DBF7FD217DAE" pageId="0" pageNumber="165">
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Body reddish brown, antennae, maxillary palpi, legs slightly lighter than body, length 1.57-1.58 mm, maximum width 0.61-0.67 mm (Fig.
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<figureCitation id="8F5E507A27F53B042D2C5D9C3572FEFE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">8A-D</figureCitation>
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). Setae on body yellowish. Head 1.15 times as long as wide (Fig.
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<figureCitation id="F85CF712F54FD65586C8B873502B52E5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">8E</figureCitation>
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). Frons with U-shaped impression between antennal tubercles; frontal foveae absent; frontal rostrum distinct anteriorly. Vertex weakly convex; longitudinal carina absent; vertexal foveae small. Eyes reduced with 9 facets. Maxillary palpi moderately developed; palpomeres II smooth; III with tubercles; palpomere IV 0.24-0.28 mm long and about 3.06-3.24 times as long as wide, subcylindrical pseudosegment at apex. Antennae about 0.61-0.68 mm long; scapes subcylindrical and elongated, 2-2.18 times as long as pedicels, without modification; pedicels subcylindrical, 1.38-1.40 times as long as wide; antennomere III 1.67-1.72 times as long as wide; IV-VII subequal in length; VIII subglobose as long as wide; IX 0.82-0.86 times as long as wide; X transverse, 0.64-0.65; XI largest, pointed at apex, 1.76-1.84 times as long as wide (Fig.
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<figureCitation id="8816416DE5FB81C1C80CDC61E2FDDD53" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">8F</figureCitation>
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). Pronotum 0.85-0.89 times as long as wide and widest at basal 2/3, lateral antebasal foveae connected by antebasal sulcus. Elytra slightly convex, 0.87 times as long as wide and widest at basal 1/3, each elytron with two basal foveae and subhumeral fovea. Legs slender; protibiae without spine (Fig.
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<figureCitation id="CB0ACDF027AAE9D38D1F7AA33BDF73A2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">8G</figureCitation>
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); metatibiae with spine on apical (Fig.
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<figureCitation id="3C1D8C2D3EBAE91E2612794787AB2EC4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">8H</figureCitation>
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, arrow). Aedeagus robust, 0.53 mm long and 1.79 times as long as wide; penis bulbous and dorsal diaphragm small, transversely ovoid; parameres symmetrical, each bearing two setae; endophallus comprising simple strut basally and two symmetrical struts curved along with parameres, thickened basally (Fig.
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<figureCitation id="04C51E8AD83223155DD34C2064C301EC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">8I</figureCitation>
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).
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</paragraph>
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<paragraph id="C29D67CF25180B294D242500843D78CF" pageId="0" pageNumber="165">
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<emphasis id="13E74177DE338B6758B2954A1F2B82DB" bold="true" pageId="0" pageNumber="165">
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<emphasis id="A1AA43F9267EE412829EAB3750FCBB11" italics="true" pageId="0" pageNumber="165">Sexual dimorphism</emphasis>
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.
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</emphasis>
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Female metatibial spines absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="BB72BE8941D64C0C291CCD57DB0F2E4F" pageId="0" pageNumber="165" type="remarks">
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<paragraph id="5DE077EF7CB44FFC24FB972470EE3C4B" pageId="0" pageNumber="165">Remarks.</paragraph>
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<paragraph id="AA939679B0F2EC83FC33C1B56FD6D183" pageId="0" pageNumber="165">
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Adults of this species are similar to that of
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<taxonomicName id="45C5A0A71196EE6FEDECADA92D14C58B" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis girinensis" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="girinensis">
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<emphasis id="6D14192B8AEFE5F785D5B3EDCF77E2D0" italics="true" pageId="0" pageNumber="165">Bryaxis girinensis</emphasis>
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</taxonomicName>
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Choi, Park, Lee & Park sp. nov. in the shape of antennomeres IV-XI (Figs
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<figureCitation id="1FA15494F02608CACF5038ED82B580F0" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bryaxis girinensis Choi, Park, Lee & Park, sp. nov. A dorsal habitus B ventral habitus C head D antenna E fore leg F hind leg G aedeagus. Scale bars: 1 mm (A, B); 0.1 mm (C, G); 0.5 mm (D-F)." figureDoi="10.3897/zookeys.1182.97346.figure7" httpUri="https://binary.pensoft.net/fig/920950" pageId="0" pageNumber="165">7D</figureCitation>
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,
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<figureCitation id="566420C67FDFFEF27D0EC638E73059BF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">8F</figureCitation>
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). However, they can be distinguished by having angular tempora much longer than the eyes (Fig.
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<figureCitation id="9C1F793F030349104D4FB325F91E721D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diagnostic characters of Bryaxis girinensis sp. nov. (A, C, E) and B. nemorosus sp. nov. (B, D, F). A, B maxillary palpi C, D protibiae E, F heads. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure9" httpUri="https://binary.pensoft.net/fig/920952" pageId="0" pageNumber="165">9F</figureCitation>
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), smooth maxillary palpomere II (Fig.
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<figureCitation id="AB474E4B9D5821C2D54B7AEEDC0CFC18" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diagnostic characters of Bryaxis girinensis sp. nov. (A, C, E) and B. nemorosus sp. nov. (B, D, F). A, B maxillary palpi C, D protibiae E, F heads. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure9" httpUri="https://binary.pensoft.net/fig/920952" pageId="0" pageNumber="165">9B</figureCitation>
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), unadorned antennal scapes and pedicels (Fig.
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<figureCitation id="08D1F1F9FE2EEC31BAD4D55365FB8C44" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">8F</figureCitation>
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), slender protibiae without a spine (Fig.
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<figureCitation id="A86D74C190FC25CB6BAB288A3D81FBD1" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diagnostic characters of Bryaxis girinensis sp. nov. (A, C, E) and B. nemorosus sp. nov. (B, D, F). A, B maxillary palpi C, D protibiae E, F heads. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure9" httpUri="https://binary.pensoft.net/fig/920952" pageId="0" pageNumber="165">9D</figureCitation>
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), and an endophallus composed of three long struts (Fig.
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<figureCitation id="23D53B1A8DEF6651A880919C26F41650" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure8" httpUri="https://binary.pensoft.net/fig/920951" pageId="0" pageNumber="165">8I</figureCitation>
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).
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</paragraph>
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<caption id="C54F4C895990E44294F1DB6187693063" doi="10.3897/zookeys.1182.97346.figure9" httpUri="https://binary.pensoft.net/fig/920952" pageId="0" pageNumber="165" start="Figure 9" startId="F9">
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<paragraph id="5A9E4CFF3D88C59256A8E13ABA6D5C98" pageId="0" pageNumber="165">
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<emphasis id="A3B6B521B4E243B48122C0599E50CE8B" bold="true" pageId="0" pageNumber="165">Figure 9.</emphasis>
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Diagnostic characters of
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<taxonomicName id="CAD2965AE695F249C3346FED50B04BED" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis girinensis" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="girinensis">
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<emphasis id="73AC00E83AE08E7F750AA5F2933C9801" italics="true" pageId="0" pageNumber="165">Bryaxis girinensis</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis id="6C1E8B7AAA8D5AAC7C38B64616389A4A" bold="true" pageId="0" pageNumber="165">A, C, E</emphasis>
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) and
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<taxonomicName id="49ADF49B672D405409B161BB3166BD92" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="B. nemorosus" order="Coleoptera" pageId="0" pageNumber="165" rank="species" species="nemorosus">
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<emphasis id="EAA79066DEF99B7B5A8895AAE754FB4D" italics="true" pageId="0" pageNumber="165">B. nemorosus</emphasis>
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|
</taxonomicName>
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sp. nov. (
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<emphasis id="7A048DF7352C3528E8F40A3629EC6F07" bold="true" pageId="0" pageNumber="165">B, D, F</emphasis>
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|
).
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<emphasis id="899AB385A3C3F56A70C4967A9AE543A2" bold="true" pageId="0" pageNumber="165">A, B</emphasis>
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|
maxillary palpi
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|
<emphasis id="3EC510AD4723E40D3460E95565BC20EC" bold="true" pageId="0" pageNumber="165">C, D</emphasis>
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|
protibiae
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|
<emphasis id="F3A2749AA08B7CFFCE942FD2C81F2734" bold="true" pageId="0" pageNumber="165">E, F</emphasis>
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|
heads. Scale bars: 0.1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
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|
<subSubSection id="5FB26ADB2CF3B8397EDCBBCBDF0A092B" pageId="0" pageNumber="165" type="etymology">
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<paragraph id="2DE3D250CE7CFDD69B81F7EEBDBEFD45" pageId="0" pageNumber="165">Etymology.</paragraph>
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|
<paragraph id="F77915CB2F59372437FE703C27300DE4" pageId="0" pageNumber="165">
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|
The specific epithet is the Latin word
|
|
<taxonomicName id="6414290F19A51005DD0DCD69DD270D25" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis nemorosus" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="nemorosus">
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<emphasis id="A50BA80CE2FE42C29A8EB9E1F15B2C57" italics="true" pageId="0" pageNumber="165">Bryaxis nemorosus</emphasis>
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|
</taxonomicName>
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|
("wooded, shady", masculine) that refers to habitat where the types collected.
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|
</paragraph>
|
|
</subSubSection>
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|
<subSubSection id="13AD0CEE33866346FDF80EC2963FF2DC" pageId="0" pageNumber="165" type="habitat">
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|
<paragraph id="D0E00EEF257A70D3161A2628BC8C9C66" pageId="0" pageNumber="165">Habitat.</paragraph>
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<paragraph id="CC832634B74FDCDBB355C8F56988B129" pageId="0" pageNumber="165">Specimens of this species were collected by sifting leaf and soil litter in wet forest with dense canopy, which is located on an island.</paragraph>
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</subSubSection>
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|
<subSubSection id="909C67C71115218CCDD7221536CCDC30" pageId="0" pageNumber="165" type="distribution">
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|
<paragraph id="F143FFFF26CC3BA4C1F0309729D83576" pageId="0" pageNumber="165">Distribution.</paragraph>
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<paragraph id="75DDC66A85D0175CE4B71189C641AC01" pageId="0" pageNumber="165">Korea (Gageo island, Sinan-gun, Jeollanam-do).</paragraph>
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</subSubSection>
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</treatment>
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</document> |