279 lines
36 KiB
XML
279 lines
36 KiB
XML
<document id="5651656DFCA48635E412334A96C5A727" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="87CE867CA29D4545A74141B1A13EB254" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus complicatus Sharp 1882" docType="treatment" docVersion="5" lastPageNumber="75" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="72" updateTime="1701314941532" updateUser="plazi">
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<mods:title id="A33237833228A998E33CDC3739DADDC9">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="29B06C62EAA140CEBD1F9C6AB2FD8747">Bistroem, Olof</mods:namePart>
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<mods:namePart id="D67E14A4CF07A4C6E100596A72A0E885">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="E208E35505F365CED2267D8A893D7170">Bergsten, Johannes</mods:namePart>
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<mods:date id="B9081F426DD530DF5673680F57A9E4FE">2015</mods:date>
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<mods:number id="3D2B5785A655C92F2E5FADC861730788">542</mods:number>
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<mods:identifier id="E0D2EDD7C7F57507841AA558056A290E" type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
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<treatment id="87CE867CA29D4545A74141B1A13EB254" ID-GBIF-Taxon="127862738" LSID="urn:lsid:plazi:treatment:87CE867CA29D4545A74141B1A13EB254" httpUri="http://treatment.plazi.org/id/87CE867CA29D4545A74141B1A13EB254" lastPageId="74" lastPageNumber="75" pageId="71" pageNumber="72">
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<subSubSection id="FA2D2048870CF3749C8123B867079F50" pageId="71" pageNumber="72" type="multiple">
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<paragraph id="6A8D2470961830D0D8CB7C5DC7DA9FF3" pageId="71" pageNumber="72">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="2517ED4805B5AD00DD997ACCC64D086B" pageId="71" pageNumber="72" type="nomenclature">
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<paragraph id="46D55EFA8C47A5251F9F57EF7CC9CE3A" pageId="71" pageNumber="72">
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<taxonomicName id="A24D27B19F00F386457184DF2C8926F9" ID-CoL="3RM69" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus Sharp, 1882</taxonomicName>
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Figs 59-60, 255, 414, 541
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</paragraph>
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</subSubSection>
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<subSubSection id="3E66141214E0B5FD30AAB1BDB423BC3D" pageId="71" pageNumber="72" type="reference_group">
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<paragraph id="5B68CC73C1A5846474CA041DB771D235" pageId="71" pageNumber="72">
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<taxonomicName id="AFA9EC1BB794EAE1F82B7E78DF908CF7" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
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<bibRefCitation id="58FA23C08DE920A7D0AC03FF8348E4FB" author="Sharp, D" journalOrPublisher="The Scientific Transactions of the Royal Dublin Society" pageId="263" pageNumber="264" pagination="179 - 1003" title="On aquatic carnivorous Coleoptera or Dytiscidae." volume="2" year="1882">Sharp 1882</bibRefCitation>
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: 308, 309 (original description, faunistics, discussion);
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<bibRefCitation id="451C26AAF2BCBF8CC87A3CABD4777E30" author="Kolbe, HJ" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="259" pageNumber="260" pagination="383 - 427" title="Ueber die madagascarischen Dytisciden des Koenigl. Entomologischen Museum zu Berlin." volume="49" year="1883">Kolbe1883</bibRefCitation>
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: 402 (description, faunistics); v. d.
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<bibRefCitation id="10B3CD8082362BF80C922C4BD2661A81" author="Branden, C van den" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="255" pageNumber="256" pagination="1 - 118" title="Catalogue des coleopteres carnassiers aquatiques (Haliplidae, Amphizoidae, Pelobiidae et Dytiscidae)." volume="28" year="1885">Branden 1885</bibRefCitation>
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: 20 (catalogue, faunistics);
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<bibRefCitation id="D932E750AA0F2DAE16F1A4AB7E4F7F3C" author="Regimbart, M" journalOrPublisher="Memoires de la Societe entomologique de Belgique" pageId="262" pageNumber="263" pagination="1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. En contribution a la faune entomologique du Congo." volume="4" year="1895">
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<normalizedToken id="C25D3898A34CEA524565ADC68D32F06B" originalValue="Régimbart">Regimbart</normalizedToken>
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||
1895
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||
</bibRefCitation>
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||
: 138 (description, faunistics);
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||
<bibRefCitation id="8A662E40BF97F2CC4B0EE64091F2FE5A" author="Zimmermann, A" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="263" pageNumber="264" pagination="68 - 249" title="Die Schwimmkaefer des Deutschen Entomologischen Museums in Berlin-Dahlem." volume="83" year="1919">Zimmermann 1919</bibRefCitation>
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||
: 122 (faunistics);
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<bibRefCitation id="7EB7FCBE86F1BE41C5CD861C47216740" author="Zimmermann, A" journalOrPublisher="Junk, Berlin" pageId="263" pageNumber="264" title="Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Vol. 4 pars 71. W." year="1920 a">Zimmermann 1920a</bibRefCitation>
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:17 (catalogue, faunistics);
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<bibRefCitation id="0A1401723A1B4C25568AF512458DF7FE" pageId="71" pageNumber="72">Bertrand 1928a</bibRefCitation>
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: 184, 185 (juvenile faunistics);
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<bibRefCitation id="BC5B06DDACD351150E375D10F3FA53FC" author="Bertrand, H" journalOrPublisher="Faune des Colonies Francaises" pageId="253" pageNumber="254" pagination="361 - 387" title="Description sommaire de quelques larves de dytiscides de Madagascar." volume="2" year="1928 c">Bertrand 1928c</bibRefCitation>
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: 365 (juvenile description);
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<bibRefCitation id="686A23AC6A4AB97A118EC9B1A9B4C51F" author="Bertrand, H" journalOrPublisher="Exploration du Parc National Albert. Mission H. Damas (1935 - 1936)" pageId="253" pageNumber="254" pagination="1 - 27" title="Larves d'hydrocanthares (Dytiscidae et Gyrinidae)." volume="17" year="1948">Bertrand 1948</bibRefCitation>
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: 12: (juvenile faunistics);
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<bibRefCitation id="3CB5E8C2E9AD8DC69B57D0FB7D6B4C5E" author="Bertrand, H" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" pageId="253" pageNumber="254" pagination="283 - 299" title="Larves d'hydrocanthares. La reserve naturelle integrale du Mont Nimba." volume="2" year="1954">Bertrand 1954</bibRefCitation>
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: 284 (juvenile discussion, faunistics);
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<bibRefCitation id="7A7861F43683A0A75BBF3157AE5546D1" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot 1946c</bibRefCitation>
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||
: 279, 281, 283 (discussion, description, faunistics);
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<bibRefCitation id="987CFE76A9DB6C28A1CC19BEE4AAAC2D" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
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: 570, 573, 574 (description, faunistics);
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<bibRefCitation id="DE6CB6069154C2D2FBD71112C6702BB9" author="Bertrand, H" journalOrPublisher="Bulletin de l'Institut Francais d'Afrique Noire (ser. A)" pageId="253" pageNumber="254" pagination="389 - 466" title="Contribution a l'etude des premiers etats des coleopteres aquatiques de la region ethiopienne (5 e note). Familles: Haliplidae, Dytiscidae, Gyrinidae (Hydrocanthares)." volume="25" year="1963">Bertrand 1963</bibRefCitation>
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: 411 (discussion, faunistics);
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<bibRefCitation id="C5C4FFBEE0F339AF3C28B1B49DA4ACB5" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="285 - 294" title="New and little known species of African Laccophilus (Coleoptera: Dytiscidae)." volume="33" year="1970">Omer-Cooper 1970</bibRefCitation>
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: 290, 291 (description); Bertrand and Legros 1971: 245 (faunistics, biology);
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<bibRefCitation id="AD4D6FA34CD470570E9EAE4A0CABF733" author="Bameul, F" journalOrPublisher="Nouvelle Revue d'Entomologie (N. S.)" pageId="252" pageNumber="253" pagination="87 - 103" title="Haliplidae, Noteridae, Dytiscidae et Gyrinidae captures dans les iles Mascareignes et a Madagascar par Yves Gomy (Coleoptera, Hydradephaga)." volume="1" year="1984">Bameul 1984</bibRefCitation>
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||
: 94 (faunistics);
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<bibRefCitation id="AC360FE567A59BDA8C69D4B90E1F8A6C" author="Rocchi, S" journalOrPublisher="Frustula Entomologica (n. s.)" pageId="262" pageNumber="263" pagination="71 - 89" title="Contributo alla conoscenza degli Haliplidae e dei Dytiscidae del Madagascar con descrizione di due nuove specie (Coleoptera)." volume="14" year="1991">Rocchi 1991</bibRefCitation>
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: 80, 86 (faunistics);
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<bibRefCitation id="91B5CAD6A300F9B17AE30452BCEEE57D" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 242 (catalogue, faunistics);
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<bibRefCitation id="5E4A7A8B4E900F9A22714C35A1CCC7A2" author="Pederzani, F" journalOrPublisher="Atti Dell'Accademia Roveretana degli Agiati, Contributi della Classe di Scienze Matematiche, Fisiche e Naturali" pageId="261" pageNumber="262" pagination="81 - 100" title="On some Madagascan Hydradephaga of the Natural History Museum, London, with the description of five new species of Dytiscidae (ColeopteraHaliplidae, Dytiscidae, Noteridae)." volume="8 B" year="2009">Pederzani and Rocchi 2009</bibRefCitation>
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||
: 95 (faunistics, list);
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||
<bibRefCitation id="C3897342F310DD67EA7CA6EA82FC2CE1" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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||
: 210 (catalogue, faunistics).
|
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</paragraph>
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</subSubSection>
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||
<subSubSection id="77829D69A61B65C427820827F3690C27" pageId="71" pageNumber="72" type="type locality">
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<paragraph id="68B5746A33D432F1C838B1D01E121BE0" pageId="71" pageNumber="72">Type locality.</paragraph>
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<paragraph id="1FA17F1E3B57BC70CEBB052C8AD321F7" pageId="71" pageNumber="72">Madagascar.</paragraph>
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</subSubSection>
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<subSubSection id="DAE416FC9B0ED2A6C2DF4546D979945A" pageId="72" pageNumber="73" type="type material">
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<paragraph id="32EC92F9C9CF756E8A8AFF8CB0298EE3" pageId="72" pageNumber="73">
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<pageBreakToken id="523F2B1AF9CDBAA455290F9B5A2563BC" pageId="72" pageNumber="73" start="start">Type</pageBreakToken>
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material studied
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</paragraph>
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<paragraph id="0F5A29C12268A9A687307343D3DC4B39" pageId="72" pageNumber="73">
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(2 exs.).
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<taxonomicName id="751F3FA54DC430BDD2B963BB4151F823" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
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: Lectotype (by present designation) male: "Type / Madagascar / Sharp Coll. 1905-313 / Madag. /
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<taxonomicName id="7679D6567C6B9A7A56D59067257BE2DF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus rivulosus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="rivulosus">Laccophilus rivulosus</taxonomicName>
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Klug /Type 574
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<taxonomicName id="ADBFB6CC480A495E65B0C23B14A013B4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
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sp. n. Madagascar" (BMNH). - Paralectotype: female: "574 / Cotype / Madagascar / Sharp Coll. 1905-313 / Madagascar /
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<taxonomicName id="D463CF903438B938FFC75F844C44938F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
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Shp Co-type" (1 ex. BMNH).
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</paragraph>
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</subSubSection>
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<subSubSection id="BB0B09739663A4C7EC2D763EF836F6E7" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73" type="materials_examined">
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<paragraph id="3F33C304218E69488A88D1214848DAA6" pageId="72" pageNumber="73">Additional material studied</paragraph>
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<paragraph id="164168B2E6A00E9DD69D95D3DDB52776" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
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(390 exs.). Madagascar: "E-Mad., Morarano, N. Morananga 900 m asl, 13.1. 1995
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<normalizedToken id="3F513BEB1F3645FEA0742DADAB28BC87" originalValue="Janák”">Janak"</normalizedToken>
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(9 exs. NMW; habitus in Fig. 414); "Moramanga env. 10-18.12. 1997
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<normalizedToken id="7CFAF351164D02B684715D3AA456462C" originalValue="Pacholátko”">Pacholatko"</normalizedToken>
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(1 ex. NHMB); "E-Mad. Ampamoho nr Andilamena 1200-1300 m asl 18-20.1. 1995 Dunay &
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<normalizedToken id="8752C079B5288B97F2247D941C8C9B25" originalValue="Janák”">Janak"</normalizedToken>
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(20 exs. NMW); "Ambatombe nr Andilamena 900 m asl, 17.1. 1995 Dunay &
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<normalizedToken id="CF9EA7AA1EECE3F96164C593803BB623" originalValue="Janák”">Janak"</normalizedToken>
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(10 exs. NMW, 10 exs. MZH, 1 ex. NHRS); "E-Mad. Andranokobaka, N Moramanga 800 m asl 13.1. 1995 Dunay &
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<normalizedToken id="55340054A6DC87F3A40F085370D1BAD7" originalValue="Janák”">Janak"</normalizedToken>
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(21 exs. NMW); "W-Mad. Manindaray W Sakahara 700-800 m asl 30.1. 1995 Dunay &
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<normalizedToken id="8F3296595BB518428CFDF5BBA1577518" originalValue="Janák”">Janak"</normalizedToken>
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(1 ex. NMW); "5 km S Ampamoho 950-1000 m asl 18-20.1. 1995 Dunay &
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<normalizedToken id="BB393855480643309C04680A0804B511" originalValue="Janák”">Janak"</normalizedToken>
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(3 exs. NMW); "E Sakahara 30.1. 1995 Manindray 700-800 m Dunay &
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<normalizedToken id="112432EFE78D6A832EEA3E5FE4DF0867" originalValue="Janák”">Janak"</normalizedToken>
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(1 ex. NMW); "Mad. Centr. Antananarive 18-19.1. 1993
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<normalizedToken id="5BC6391A51908665B76244451D535067" originalValue="Janák”">Janak"</normalizedToken>
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(1 ex. NMW); "Envir. de Tananarive 7. 1970 Pederzani" (1 ex. AMGS, 2 exs. NHMB); "Prov. Tananarive, env. de Arivonimamo 22.7. 1970 Pederzani" (1 ex. AMGS); "Tan. Manjakatompo 3.1. 1958 Keiser" (1 ex. NHMB); "Mandrare Bas., Loc.
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<normalizedToken id="924F51CF9F151D7A7F0D5B300A1E65EE" originalValue="prés">pres</normalizedToken>
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Andaza, affl. non
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<normalizedToken id="0E8F5730C17CBB91F37F05A1927E366A" originalValue="nommé">nomme</normalizedToken>
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, Riv.,
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<geoCoordinate id="1B204229C5F569D7398C35DC9CF1B3E2" direction="east" orientation="longitude" precision="15" value="46.568054">46°34'05"E</geoCoordinate>
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,
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<geoCoordinate id="3E9A07CA923F69838D25E807D0FE817F" direction="south" orientation="latitude" precision="15" value="-24.054443">24°03'16"S</geoCoordinate>
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, Alt. 315 m 26.4. 1995 Elouard & Pilaka" (1 ex. NMW); "Prov. Tamatava, 3.3 km N Ambabasoratra 31.8. 1962 Cashatt" (1 ex. USNM); "Mad-est 1100-1200 m, NP Ranomafana / Vohiparara 21-24.1. 1993
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<normalizedToken id="712AF02C4F92829C6AF621BDBE0D423C" originalValue="Janák”">Janak"</normalizedToken>
|
||
(2 exs. NMW); "Prov. Fianarantsoa 7 km W Ranomafana, 1100 m 8-21-10. 1988 Steiner W.E. / At black light in montane rainforest" (3 exs. USNM, 1 ex. MZH); "Prov. Fianarantsoa 7 km W Ranomafana, 1000 m 23-28.2. 1990 Steiner W.E." (1 ex. USNM); "Ankaratra (Antananarivo) Res. Manjakatompo 6.10. 2001 / Gerecke et Goldsmith leg. / Helocrene at left border of affluent to Lac Froid, 1700 m asl" (2 exs. BMNH); "Manjakatompo 5.10. 1989 Bartolozzi & Taiti leg. / m 1700 Stazione pescicoltura /
|
||
<taxonomicName id="605F7EDAD528D1BC51A14EB1773BF50C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
|
||
Sharp det. Rocchi 1989"(3 exs. CSR); "Antranovy (Antananarive) 13 km W from Arivonimana, 13.7. 2001 helocrene in rice field exp. N / 1480 m asl / Gerecke et Goldsmith leg." (9 exs. BMNH, 2 exs. MZH); "Tsimelahy (Tuelar) 5.9. 2001, Riv. Antarantsa downstr. piceine naturelle / 200 m asl / Gerecke et Goldsmith leg." (1 ex. BMNH); "Foret de
|
||
<normalizedToken id="C427B2DAA2D9EF34504969C5C7397850" originalValue="l’Est">l'Est</normalizedToken>
|
||
Perinet-Anosibe 11-12. 1959" (1 ex. BMNH); "Tan. Madag, Ampefy, Lac Kavitaha wi 25.III. 58 Keiser" (1 ex. NHMB); "Tam. Perinet 3.12. 1957 Keiser" (1 ex. NHMB); "La Mandraka, ex. coll. Breuning" (1 ex. MRAC); "Suberbieville, ex. coll. Breuning" (5 exs. MRAC); "Tananarive 22-29.i. 1972 Hecq" (1 ex. MRAC); "Tananarive, at light 12.12. 1955 E. Mac (?) Callan" (1 ex. AMGS); "Tananarivo Friedrichs /
|
||
<taxonomicName id="C0588169EECA6FCC1A0CB96F59CB5AF9" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
|
||
Sharp det. Brancucci" (1 ex. ZMHB); "Fianarantsoa Prov., Foret
|
||
<normalizedToken id="123F78B0CF62FA65BEF0C0838073A0D7" originalValue="d’Antsirakanbiaty">d'Antsirakanbiaty</normalizedToken>
|
||
, 7.6 km 285˚ WNW Itremo, elev. 1550 m 22-26.1.2003 /
|
||
<geoCoordinate id="A9E0399D1D3B38EC5A9201701500F1E9" direction="south" orientation="latitude" precision="15" value="-20.593334">20°35'36"S</geoCoordinate>
|
||
<geoCoordinate id="42C7A836A8600875733A6866173E78C9" direction="east" orientation="longitude" precision="15" value="46.56333">046°33'48"E</geoCoordinate>
|
||
, collected at light in montane rainforest, Fischer, Griswold & al. leg." (2 exs. CAS); "Fianarantsoa Ranomafana National Park, Talatakely res. lab. area, black light mercury vapour / light, elev. 940 m
|
||
<geoCoordinate id="0E765008050A75DB3A55F5CA5BAE7A02" direction="south" orientation="latitude" precision="1" value="-21.248196">21°14'53,5"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="136BDFD4C862EED23C91222D962744A9" direction="east" orientation="longitude" precision="1" value="47.42664">47°25'35,9"E</geoCoordinate>
|
||
, 31.10-20.11.1998 Lee & Ribardo leg." (1 ex. CAS); "Fian Isalo Menamaty R., degraded river with lots of vegetation used by women to wash clothes in, 11.5. 2006, N-22°33.001, E45°24.074, 757 m Bergsten et al." (12 exs. NHRS, 5 exs. MZH); "Fian
|
||
<pageBreakToken id="C4610202F6AC70CE0E925B59E011B62F" pageId="73" pageNumber="74" start="start">Andringitra</pageBreakToken>
|
||
Zomandao R. bridge on road to the camp belamba, vegetation rich edges along the river N-22°6.225, E46°55.244, 8-9.5.2006, 1421 m Bergsten et al." (13 exs. NHRS); "Fian. Col. des Tapias Rte Tana-Fianarantsoa, vegetation- rich pond with fish 6.5. 2006 N-20°46.376, E47°10.749, 1718 m Bergsten et al." (9 exs. NHRS); "Mangoky Bas, Loc. Andringitra, Camp B, Zomandao River
|
||
<geoCoordinate id="F201E9BE8B1C35933CCE802B36EAFA6C" direction="east" orientation="longitude" precision="15" value="46.896114">46°53'46"E</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="009BD425D2066C6922EAB8F0667B687A" direction="south" orientation="latitude" precision="15" value="-22.12">22°07'12"S</geoCoordinate>
|
||
1600 m asl 30.11. 1993 leg. ORSTOM" (1 ex. NHRS); "Andringitra Rambavy R. (Cascade)
|
||
<geoCoordinate id="F858DF237C714D2479A4FFFCEDCD16CB" direction="north" orientation="latitude" precision="55" value="22.153">N-22.153</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="7F5FD3C4D565718768913311C7CB12F3" direction="east" orientation="longitude" precision="5555" value="46.9">E 46.9</geoCoordinate>
|
||
, 1979 m, 8.5. 2006 Bergsten et al. / BMNH(E) <794244> DNA Voucher" (1 ex. NHRS); "Ft Dauphin, rice paddies
|
||
<geoCoordinate id="1DC131F6A4AED1AB57251B5D592B3B53" direction="north" orientation="latitude" precision="55" value="24.824">N-24.824</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="69EAF227DCC1DD56A5D484CF8B9A0FD0" direction="east" orientation="longitude" precision="55" value="46.866">E 46.866</geoCoordinate>
|
||
, 34.44 m 19.5. 2006 Bergsten at al/BMNH(E)(E) <794247> DNA voucher" (1 ex. NHRS); "TOLI Zombitse, N-22 64 Andramomena R. pool,
|
||
<geoCoordinate id="E4D95070AB6949515B806849156FD67D" direction="north" orientation="latitude" precision="555" value="22.64">N-22.64</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E6708062B0A6E0D85B7DF50EC017B67B" direction="east" orientation="longitude" precision="55" value="44.864">E44.864</geoCoordinate>
|
||
: 577 m, 15.5.2006 leg. Bergsten" (1 ex. NHRS); "Mangoro Bas Loc. Ankirihitra Tsarantanana River
|
||
<geoCoordinate id="69689334F18DD1A84A5B02916FF235BD" direction="east" orientation="longitude" precision="15" value="47.29361">47°17'37"E</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="0E00A3D744FAAB3890FC0FFF817B52CE" direction="south" orientation="latitude" precision="15" value="-19.383333">19°23'00"S</geoCoordinate>
|
||
, 11.1. 1997 Elouard leg." (1 ex. NHRS); "Mad-est. 1100-1200 m P.N. Ranomafana / Vohipara 21-24.1. 1993 Dunay &
|
||
<normalizedToken id="C79E7D23DCD598F74300698852F2B8A5" originalValue="Janák">Janak</normalizedToken>
|
||
leg." (1 ex. NHRS); "Mah/Tol. Melaky/Menabe Ambojihanahary NP,
|
||
<geoCoordinate id="74B1054575195AAC0F99D5ABF7081C59" direction="south" orientation="latitude" precision="1" value="-18.26849">S18.26849</geoCoordinate>
|
||
.
|
||
<geoCoordinate id="5BA85A0763158CD95D6903D67741B405" direction="east" orientation="longitude" precision="1" value="45.46346">E045.46346</geoCoordinate>
|
||
, 906 m.a.o., 19.12. 2009 water net, field Bergsten et al. / NHRS-JLKB 000000723" (1 ex. NHRS);
|
||
<normalizedToken id="8381931A3A0CB9D44C33736D335CEC40" originalValue="“Madagascar”">"Madagascar"</normalizedToken>
|
||
(1 ex. USNM, 2 exs. ZMHB).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67DC4DC62C55BB587387D4DA350646A4" pageId="73" pageNumber="74" type="specimen with uncertain locality">
|
||
<paragraph id="973329BCA61C676FB4E78AED0E8EB3C7" pageId="73" pageNumber="74">Specimen with uncertain locality</paragraph>
|
||
<paragraph id="BB50E103A55B5A568657EAA0097877B0" pageId="73" pageNumber="74">
|
||
(2 exs.). "Dirjo 24" (1 ex. NMW);
|
||
<normalizedToken id="A00E02D597E38C8E9F3F24C9BF800014" originalValue="“Mexique”">"Mexique"</normalizedToken>
|
||
(1 ex. MHNG).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="102F468F3BAC9CC5C81014BB8137C1D1" pageId="73" pageNumber="74" type="diagnosis">
|
||
<paragraph id="913276231F5D247901A56B010938FB05" pageId="73" pageNumber="74">Diagnosis.</paragraph>
|
||
<paragraph id="F6166DB9AA5679D474ADB055F2186383" pageId="73" pageNumber="74">
|
||
<taxonomicName id="2687119CA4D43F69647EB60CC1255D4B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
|
||
, distributed solely on Madagascar, resembles most of three species on mainland of Africa, viz.
|
||
<taxonomicName id="DFC0A6F7BB8D5E01206AAD8E024F6C93" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus simplicistriatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="simplicistriatus">Laccophilus simplicistriatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="F8BFADA5A81DFD51322BB5D50BBEE685" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
and
|
||
<taxonomicName id="58E794F9E87604856327E8ADF0F6B0A5" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus propinquus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="propinquus">Laccophilus propinquus</taxonomicName>
|
||
. In general
|
||
<taxonomicName id="3DFE9673FE16335F08183F873407AEC2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
|
||
is somewhat larger than the three other species. Elytral colour pattern in
|
||
<taxonomicName id="8088473DC7958D442C4481FDD5E58DEB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
|
||
is uniform and more evenly distributed than in
|
||
<taxonomicName id="BF97A03444F090129EA86972F3E781B6" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
and in
|
||
<taxonomicName id="830AA388C57ECDD5D01C00BBD1816ADB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus propinquus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="propinquus">Laccophilus propinquus</taxonomicName>
|
||
, in which a vague and irregular, dark, longitudinal marking can be discerned discally on each elytron. Elytral colour pattern in
|
||
<taxonomicName id="A34606F2BE9CC040B16386E56D2B1E2D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus simplicistriatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="simplicistriatus">Laccophilus simplicistriatus</taxonomicName>
|
||
is fairly uniform and exhibits only slight variation. In
|
||
<taxonomicName id="855F8978CA16ED1000365C06FF88C2D4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
|
||
penis in lateral view is not expanded posterior to apex while in
|
||
<taxonomicName id="FAB8DBCE8B32B22CB242BF2B03B07EEF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus simplicistriatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="simplicistriatus">Laccophilus simplicistriatus</taxonomicName>
|
||
corresponding feature is clearly expanded on each side. From
|
||
<taxonomicName id="D3154243798450AB6D156454A7BFE97D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
the species is distinguished by the tip of penis, which is clearly curved
|
||
<normalizedToken id="9E21587F2BBC3ED73A45830D61900DEF" originalValue="”upwards”">"upwards"</normalizedToken>
|
||
while tip of
|
||
<taxonomicName id="70205742D0630DA63A032986C48FF89E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
|
||
penis is almost straight. Regarding
|
||
<taxonomicName id="E9649572221E7FDCE50171F3BBA26328" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus propinquus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="propinquus">Laccophilus propinquus</taxonomicName>
|
||
,
|
||
<taxonomicName id="8FBE905B4DE4612DB570EE30DA4FAC90" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
|
||
is separated from it as
|
||
<taxonomicName id="DF3C02A7D7577AEF6FDBD888210F5229" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
|
||
is but tip of penis vague.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2E7418F9DEE08408CFB0EC598E11D0B2" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="description">
|
||
<paragraph id="05D8F1479D0AD15C733648F3E830415A" pageId="73" pageNumber="74">Description.</paragraph>
|
||
<paragraph id="1C200D180BEF58785618AA55BDC140A4" pageId="73" pageNumber="74">Body length 3.9-4.6 mm, width 2.2-2.6 mm. Pale ferrugineous, with blackish ferrugineous to dark ferrugineous, dense irrorations on elytra (Fig. 414). Dorsal colour pattern of body quite stable; rarely elytral irrorations laterally and at base slightly reduced.</paragraph>
|
||
<paragraph id="AA51A05139AC07EDE7E59F8D49F4793B" pageId="73" pageNumber="74">Head: Pale ferrugineous, posteriorly, narrowly dark ferrugineous. Submat, reticulation in part double; large meshes often weakly developed and hardly visible. At eyes, with fine and irregular punctures, extending medially towards midhead.</paragraph>
|
||
<paragraph id="2F3C6BD29037696DB54B0193C03851C7" pageId="73" pageNumber="74">Pronotum: Pale ferrugineous, anteriorly and posteriorly in middle with vague, darkened areas. Reticulation quite dense, double. Large meshes weakly developed; one mesh contains between three and six fine meshes. At margins with fine, irregular punctures, which are rather indistinct at posterior margin.</paragraph>
|
||
<paragraph id="5AB33BD76E0083DEF7FA9F5AEA26DE5D" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
|
||
Elytra: Pale ferrugineous, with dense blackish ferrugineous to dark ferrugineous irrorations (Fig. 414). Submat, with quite dense reticulation. Reticulation indistinctly
|
||
<pageBreakToken id="6D4EFF0C6863AD671996652455BF4C60" pageId="74" pageNumber="75" start="start">double</pageBreakToken>
|
||
; indistinct fragments of larger meshes extensively discernible. Each elytron provided with three, longitudinal, rather vague areas of fine and sparse punctures.
|
||
</paragraph>
|
||
<paragraph id="59C90B5654F6326A16BB3402D0FD4DDB" pageId="74" pageNumber="75">Ventral aspect: Pale ferrugineous to ferrugineous, abdomen in posterior part blackish to dark ferrugineus. Rather shiny to submat, very finely reticulated. Almost impunctate; fine punctures discernible on apical ventrite. Apical ventrite as in Fig. 59. Basal ventrites provided with fine, slightly curved, striae. Metacoxal plate with some very shallow, almost transversely located furrows. Prosternal process, slender, extended and pointed.</paragraph>
|
||
<paragraph id="34F1654E76A2F678EF16DE8D2FC04949" pageId="74" pageNumber="75">Legs: Pro- and mesotarsus somewhat enlarged; provided with suckers.</paragraph>
|
||
<paragraph id="C12C1F716B9374348F96FFCE990252D7" pageId="74" pageNumber="75">Male genitalia: Penis in lateral aspect evenly curved, broad; extreme apex curved upwards and sharp (Fig. 255).</paragraph>
|
||
<paragraph id="228D2606B901C12EE3AAA15BD8683417" pageId="74" pageNumber="75">Female: Pro- and mesotarsus slender. Apical ventrite as in Fig. 60. Rarely microsculpture of body is strongly developed and such a specimen is clearly matter than ordinary female specimens.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08830CB511C67AB0FA9AE5AC71260726" pageId="74" pageNumber="75" type="distribution">
|
||
<paragraph id="DA14B2D5B4CAC07D6DF02BA36F382A4A" pageId="74" pageNumber="75">Distribution.</paragraph>
|
||
<paragraph id="B1B8D3698045A28B75F3604033E490FE" pageId="74" pageNumber="75">Madagascar (Fig. 541).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="35BAD5D996D7A6A8593F7394BAA5B775" pageId="74" pageNumber="75" type="collecting circumstances">
|
||
<paragraph id="9A377455B3694B18246325E51353BD97" pageId="74" pageNumber="75">Collecting circumstances.</paragraph>
|
||
<paragraph id="59AE0CF588F2AB2AD6A2A1FE0EB06E65" pageId="74" pageNumber="75">
|
||
In literature, detailed documentation is not available. From collecting labels appear that
|
||
<taxonomicName id="7123B28ACB03AAB970BB99BF5A4238DD" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus complicatus" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="complicatus">Laccophilus complicatus</taxonomicName>
|
||
has been sampled in areas between 700-1700 m a.s.l. At least once collected in a degraded river with lots of vegetation, used to wash clothes in. Obviously it is attracted by light and sampled in a montane rain forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |