417 lines
66 KiB
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417 lines
66 KiB
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<mods:title id="42EE0918E82509F846794199314FA504">On the rare Neotropical genus Ophionthus Bernhauer (Coleoptera: Staphylinidae: Staphylininae): redescription of the type species and description of a new species</mods:title>
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<mods:namePart id="F4E1ADA06785FA70C7D0F6248DAD4209">Rodríguez-Melgarejo, Maryzender</mods:namePart>
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<mods:affiliation id="1231FBD28946E0910CE6B8B4B080E83A">Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT CONICET, Mendoza), Casilla de Correo 507, 5500 Mendoza, Argentina. Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Apartado 14 - 0434, Lima 14, Perú. Facultad de Ciencias Exactas y Naturales, Universidad Nacional de Cuyo, Mendoza, Argentina.</mods:affiliation>
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<mods:namePart id="203EB49D3AB0FE041218F0963330A68A">Chani-Posse, Mariana</mods:namePart>
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<mods:affiliation id="8B610C34C17F8A0F9A6F4B329969D153">Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT CONICET, Mendoza), Casilla de Correo 507, 5500 Mendoza, Argentina. Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Apartado 14 - 0434, Lima 14, Perú. Facultad de Ciencias Exactas y Naturales, Universidad Nacional de Cuyo, Mendoza, Argentina.</mods:affiliation>
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<heading id="D97BE531FFCEFFEB9468FAC6FB8BC00D" box="[547,1040,1347,1374]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="20" reason="2">
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<taxonomicName id="458C29DEFFCEFFEB9468FAC6FB8BC00D" ID-CoL="49YSH" authority="Bernhauer, 1908" authorityName="Bernhauer" authorityYear="1908" box="[547,1040,1347,1374]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="serpentinus">
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<emphasis id="B0F88E4FFFCEFFEB9468FAC6FCDDC00D" bold="true" box="[547,838,1347,1374]" italics="true" pageId="5" pageNumber="20">Ophionthus serpentinus</emphasis>
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<bibRefCitation id="E61D2FACFFCEFFEB9505FAC1FB8BC00D" author="Bernhauer M." box="[846,1040,1348,1374]" pageId="5" pageNumber="20" pagination="283 - 372" refId="ref6978" refString="Bernhauer M. 1908. Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte 74 (1): 283 - 372. Available from https: // www. biodiversitylibrary. org / part / 226088 # / summary [accessed 28 Jan. 2021]." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
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<paragraph id="8233525DFFCEFFEB9482FAE2FCF1C0D2" blockId="5.[547,1040,1347,1409]" box="[713,874,1383,1409]" pageId="5" pageNumber="20">
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<figureCitation id="1AB74ED8FFCEFFEB9482FAE2FCA9C0D2" box="[713,818,1383,1409]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[189,232,1884,1910]" captionStartId-1="8.[189,232,1886,1912]" captionTargetBox-0="[190,1392,293,1804]" captionTargetBox-1="[199,1394,353,1755]" captionTargetId-0="figure-14@6.[189,1398,293,1811]" captionTargetId-1="figure-15@8.[189,1398,349,1768]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." captionText-1="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi-0="http://doi.org/10.5281/zenodo.4542895" figureDoi-1="http://doi.org/10.5281/zenodo.4542899" httpUri-0="https://zenodo.org/record/4542895/files/figure.png" httpUri-1="https://zenodo.org/record/4542899/files/figure.png" pageId="5" pageNumber="20">Figs 1–2</figureCitation>
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,
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<figureCitation id="1AB74ED8FFCEFFEB9575FAE2FCF1C0D2" box="[830,874,1383,1409]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="11.[189,232,1972,1998]" captionStartId-1="12.[189,232,1937,1963]" captionStartId-2="13.[189,232,1815,1841]" captionTargetBox-0="[189,1398,1237,1937]" captionTargetBox-1="[189,1398,1169,1895]" captionTargetBox-2="[230,1362,326,1764]" captionTargetId-0="figure-257@11.[189,1399,1237,1937]" captionTargetId-1="figure-338@12.[189,1398,1168,1896]" captionTargetId-2="figure-11@13.[206,1382,310,1780]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="Fig. 4. Distribution map of Ophionthus serpentinus Bernhauer, 1908 and O. asenjoi sp. nov." captionText-1="Fig. 5. Map of probable localities (correct spelling, see Supp. file 1) for the male holotype of Ophionthus serpentinus Bernhauer, 1908 collected in “Peruvia centralis” by Jelski in 1873." captionText-2="Fig. 6 (continued on next page). Strict consensus of the 28 MP trees obtained showing the systematic placement of Ophionthus Bernhauer, 1908 within the Neotropical lineage of Philonthina Kirby, 1837 (tree length = 579; consistency index (CI) = 0.24; retention index (RI) = 0.71. Black circles: exclusive synapomorphies; open circles, non-exclusive synapomorphies. Numbers above circles are character numbers; numbers below circles represent character states. Support values are indicated in bold below the branches as follows: Bremer/Standard Bootstrap/Jack-knife (cut = 50 for resampling support values)." figureDoi-0="http://doi.org/10.5281/zenodo.4542903" figureDoi-1="http://doi.org/10.5281/zenodo.4542905" figureDoi-2="http://doi.org/10.5281/zenodo.4545915" httpUri-0="https://zenodo.org/record/4542903/files/figure.png" httpUri-1="https://zenodo.org/record/4542905/files/figure.png" httpUri-2="https://zenodo.org/record/4545915/files/figure.png" pageId="5" pageNumber="20">4–6</figureCitation>
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<paragraph id="8233525DFFCEFFEB96F6FA2DFD7AC090" blockId="5.[189,737,1448,1475]" box="[189,737,1448,1475]" pageId="5" pageNumber="20">
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<treatmentCitation id="032D744CFFCEFFEB96F6FA2DFD45C090" author="Bernhauer M." box="[189,734,1448,1475]" page="329" pageId="5" pageNumber="20" year="1908">
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<taxonomicName id="458C29DEFFCEFFEB96F6FA2DFD45C090" ID-CoL="49YSH" authority="Bernhauer, 1908: 329" authorityName="Bernhauer" authorityPageNumber="329" authorityYear="1908" box="[189,734,1448,1475]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="serpentinus">
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<emphasis id="B0F88E4FFFCEFFEB96F6FA2DFE4FC091" box="[189,468,1448,1474]" italics="true" pageId="5" pageNumber="20">Ophionthus serpentinus</emphasis>
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<bibRefCitation id="E61D2FACFFCEFFEB9797FA2DFD45C090" author="Bernhauer M." box="[476,734,1448,1475]" pageId="5" pageNumber="20" pagination="283 - 372" refId="ref6978" refString="Bernhauer M. 1908. Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte 74 (1): 283 - 372. Available from https: // www. biodiversitylibrary. org / part / 226088 # / summary [accessed 28 Jan. 2021]." type="journal article" year="1908">Bernhauer, 1908: 329</bibRefCitation>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="8233525DFFCEFFEB96F6FA6FFB56C3E6" blockId="5.[189,1399,1514,1717]" pageId="5" pageNumber="20">
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<taxonomicName id="458C29DEFFCEFFEB96F6FA6FFE48C357" ID-CoL="49YSH" authorityName="Bernhauer" authorityYear="1908" box="[189,467,1514,1540]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="serpentinus">
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<emphasis id="B0F88E4FFFCEFFEB96F6FA6FFE48C357" box="[189,467,1514,1540]" italics="true" pageId="5" pageNumber="20">Ophionthus serpentinus</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="032D744CFFCEFFEB97A6FA6FFCF7C357" author="Bernhauer M. & Schubert K." box="[493,876,1514,1540]" page="372" pageId="5" pageNumber="20" year="1914">
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<bibRefCitation id="E61D2FACFFCEFFEB97A6FA6FFCF7C357" author="Bernhauer M. & Schubert K." box="[493,876,1514,1540]" pageId="5" pageNumber="20" pagination="289 - 408" refId="ref7026" refString="Bernhauer M. & Schubert K. 1914. Staphylinidae IV. In: Schenkling S. (ed.) Coleopterorum Catalogus Vol. 57: 289 - 408. W. Junk, Berlin." type="journal article" year="1914">Bernhauer & Schubert 1914: 372</bibRefCitation>
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</treatmentCitation>
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||
(catalogue). —
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<treatmentCitation id="032D744CFFCEFFEB926EFA6FFB7DC357" author="Lucas R." box="[1061,1254,1514,1540]" page="461" pageId="5" pageNumber="20" year="1920">
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<bibRefCitation id="E61D2FACFFCEFFEB926EFA6FFB7DC357" author="Lucas R." box="[1061,1254,1514,1540]" pageId="5" pageNumber="20" pagination="1 - 5" refId="ref8076" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Archiv fur Naturgeschichte (A) 84 (1 - 5): i-xxxi, 1 - 696. Available from https: // www. zobodat. at / pdf / Archiv-Naturgeschichte _ 84 A _ 1 _ 0001 - 0128. pdf [accessed 5 Feb. 2021]." type="journal article" year="1920">Lucas 1920: 461</bibRefCitation>
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</treatmentCitation>
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||
(catalogue). —
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||
<treatmentCitation id="032D744CFFCEFFEB975EF988FDAEC374" author="Blackwelder R. E." box="[277,565,1549,1575]" page="138" pageId="5" pageNumber="20" year="1944">
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<bibRefCitation id="E61D2FACFFCEFFEB975EF988FDAEC374" author="Blackwelder R. E." box="[277,565,1549,1575]" pageId="5" pageNumber="20" pagination="1 - 188" refId="ref7063" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part. 1. Bulletin of the United States National Museum 185: 1 - 188. https: // doi. org / 10.5479 / si. 03629236.185. i" type="journal article" year="1944">Blackwelder 1944: 138</bibRefCitation>
|
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</treatmentCitation>
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||
(checklist); 1952: 276 (
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<typeStatus id="5D37ECFFFFCEFFEB9521F988FC05C374" box="[874,926,1549,1575]" pageId="5" pageNumber="20">type</typeStatus>
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species:
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<taxonomicName id="458C29DEFFCEFFEB9251F98BFB57C374" ID-CoL="49YSH" authorityName="Bernhauer" authorityYear="1908" box="[1050,1228,1550,1576]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="serpentinus">
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<emphasis id="B0F88E4FFFCEFFEB9251F98BFB57C374" box="[1050,1228,1550,1576]" italics="true" pageId="5" pageNumber="20">O. serpentinus</emphasis>
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</taxonomicName>
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). —
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<treatmentCitation id="032D744CFFCEFFEB9353F98BFEE8C319" author="Herman L. H." page="2721" pageId="5" pageNumber="20" year="2001">
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<bibRefCitation id="E61D2FACFFCEFFEB9353F98BFEE8C319" author="Herman L. H." pageId="5" pageNumber="20" pagination="2441 - 3020" refId="ref7738" refString="Herman L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. V. Staphylinine Group (Part 2) Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina). Bulletin of the American Museum of Natural History 265: 2441 - 3020. https: // doi. org / 10.1206 / 0003 - 0090.265.1.5" type="journal article" year="2001">Herman 2001: 2721</bibRefCitation>
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</treatmentCitation>
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(catalogue). —
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<treatmentCitation id="032D744CFFCEFFEB9471F9B5FCCBC319" author="Chani-Posse M. R." box="[570,848,1584,1610]" page="81" pageId="5" pageNumber="20" year="2014">
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<bibRefCitation id="E61D2FACFFCEFFEB9471F9B5FCCBC319" author="Chani-Posse M. R." box="[570,848,1584,1610]" pageId="5" pageNumber="20" pagination="62 - 86" refId="ref7271" refString="Chani-Posse M. R. 2014 a. An illustrated key to the New World genera of Philonthina Kirby (Coleoptera: Staphylinidae), with morphological, taxonomical and distributional notes. Zootaxa 3755 (1): 62 - 86. https: // doi. org / 10.11646 / zootaxa. 3755.1.3" type="journal article" year="2014">Chani-Posse 2014a: 81</bibRefCitation>
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</treatmentCitation>
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, fig. 12h (habitus); 2014b: 252 (characters in data matrix). —
|
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<treatmentCitation id="032D744CFFCEFFEB97FCF9D6FC8CC33D" author="Chani-Posse M. R. & Newton A. & Kappel Hansen A. & Solodovnikov A." box="[439,791,1619,1646]" page="51" pageId="5" pageNumber="20" year="2018">
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<bibRefCitation id="E61D2FACFFCEFFEB97FCF9D6FC8CC33D" author="Chani-Posse M. R. & Newton A. & Kappel Hansen A. & Solodovnikov A." box="[439,791,1619,1646]" pageId="5" pageNumber="20" pagination="1 - 95" refId="ref7637" refString="Chani-Posse M. R., Newton A., Kappel Hansen A. & Solodovnikov A. 2018 b. Checklist and taxonomic changes for Central and South American Philonthina (Coleoptera: Staphylinidae). Zootaxa 4449 (1): 1 - 95. https: // doi. org / 10.11646 / zootaxa. 4449.1.1" type="journal article" year="2018">
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Chani-Posse
|
||
<emphasis id="B0F88E4FFFCEFFEB941FF9D1FD10C33E" box="[596,651,1619,1645]" italics="true" pageId="5" pageNumber="20">et al</emphasis>
|
||
. 2018b: 51
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(checklist). —
|
||
<bibRefCitation id="E61D2FACFFCEFFEB959DF9D6FB23C33D" author="Asenjo A. & Irmler U. & Klimaszewski J. & Chandler D. S. & Fierros-Lopez H. E. & Vieira J. S." box="[982,1208,1619,1646]" pageId="5" pageNumber="20" pagination="1 - 406" refId="ref6906" refString="Asenjo A., Irmler U., Klimaszewski J., Chandler D. S., Fierros-Lopez H. E. & Vieira J. S. 2019. Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution. Zootaxa 4621 (1): 1 - 406. https: // doi. org / 10.11646 / zootaxa. 4621.1.1" type="journal article" year="2019">
|
||
Asenjo
|
||
<emphasis id="B0F88E4FFFCEFFEB927FF9D1FBE8C33E" box="[1076,1139,1619,1645]" italics="true" pageId="5" pageNumber="20">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
(catalogue). —
|
||
<treatmentCitation id="032D744CFFCEFFEB96A1F9F2FC85C3C2" author="Chani-Posse M. R. & Ramirez-Salamanca J. M." box="[234,798,1655,1681]" page="198" pageId="5" pageNumber="20" year="2020">
|
||
<bibRefCitation id="E61D2FACFFCEFFEB96A1F9F2FC85C3C2" author="Chani-Posse M. R. & Ramirez-Salamanca J. M." box="[234,798,1655,1681]" pageId="5" pageNumber="20" pagination="194 - 217" refId="ref7394" refString="Chani-Posse M. R. & Ramirez-Salamanca J. M. 2020 a. To be biased or to be Neotropical: systematic reassessment of a megadiverse lineage of rove-beetles (Philonthina, Staphylinini, Staphylininae). Cladistics 36 (2): 194 - 217. https: // doi. org / 10.1111 / cla. 12395" type="journal article" year="2020">Chani-Posse & Ramírez-Salamanca 2020a: 198</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 209–210, 213–214 (taxon sampling, phylogenetic placement); 2020b: 238, 246 (discussion on characters and phylogenetic placement).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CA9601D6FFCEFFE996F6F95EFD82C709" lastPageId="7" lastPageNumber="22" pageId="5" pageNumber="20" type="materials_examined">
|
||
<paragraph id="8233525DFFCEFFEB96F6F95EFE31C3A6" blockId="5.[189,426,1755,1781]" box="[189,426,1755,1781]" pageId="5" pageNumber="20">
|
||
<heading id="D97BE531FFCEFFEB96F6F95EFE31C3A6" bold="true" box="[189,426,1755,1781]" fontSize="11" level="3" pageId="5" pageNumber="20" reason="3">
|
||
<emphasis id="B0F88E4FFFCEFFEB96F6F95EFE31C3A6" bold="true" box="[189,426,1755,1781]" pageId="5" pageNumber="20">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCEFFEB96A1F88FFEC7C277" blockId="5.[189,1399,1802,2004]" box="[234,348,1802,1828]" pageId="5" pageNumber="20">
|
||
<emphasis id="B0F88E4FFFCEFFEB96A1F88FFEC7C277" bold="true" box="[234,348,1802,1828]" pageId="5" pageNumber="20">
|
||
<typeStatus id="5D37ECFFFFCEFFEB96A1F88FFEC7C277" box="[234,348,1802,1828]" pageId="5" pageNumber="20" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCEFFEB96F6F8A8FE59C287" blockId="5.[189,1399,1802,2004]" pageId="5" pageNumber="20">
|
||
<materialsCitation id="32E45800FFCEFFEB96F6F8A8FE59C287" ID-GBIF-Occurrence="3037423302" collectionCode="FMNH" collectorName="C. Jelski & M. Bernhauer" country="Peru" location="PERU" pageId="5" pageNumber="20" specimenCount="1" typeStatus="typus">
|
||
<emphasis id="B0F88E4FFFCEFFEB96F6F8A8FE96C214" bold="true" box="[189,269,1837,1863]" pageId="5" pageNumber="20">
|
||
<collectingCountry id="FA9B12CDFFCEFFEB96F6F8A8FE96C214" box="[189,269,1837,1863]" name="Peru" pageId="5" pageNumber="20">PERU</collectingCountry>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="948A99D4FFCEFFEB9766F8AEFED8C214" box="[301,323,1835,1863]" pageId="5" pageNumber="20" type="male">♂</specimenCount>
|
||
[specimen glued to the tip of a triangle of white card]; “
|
||
<taxonomicName id="458C29DEFFCEFFEB9248F8A8FB07C214" box="[1027,1180,1837,1863]" class="Insecta" family="Staphylinidae" genus="Xanthopygus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B0F88E4FFFCEFFEB9248F8A8FB07C214" box="[1027,1180,1837,1863]" italics="true" pageId="5" pageNumber="20">Xanthopygus</emphasis>
|
||
</taxonomicName>
|
||
sp. / Perú central 1873. / C. Jelski [handwriting on white label]”, “Peruvia centr. / lg. Jelski 1873 / ded. [latin
|
||
<emphasis id="B0F88E4FFFCEFFEB936AF8D5FAC7C239" box="[1313,1372,1872,1898]" italics="true" pageId="5" pageNumber="20">dedit</emphasis>
|
||
= gave] Skalitzky. [Bernhauer’s handwriting on white label]”, “
|
||
<taxonomicName id="458C29DEFFCEFFEB95E9F8F6FBB0C2DE" authorityName="Bernhauer" authorityYear="1908" box="[930,1067,1907,1933]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B0F88E4FFFCEFFEB95E9F8F6FBB0C2DE" box="[930,1067,1907,1933]" italics="true" pageId="5" pageNumber="20">Ophionthus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="458C29DEFFCEFFEB920CF8F1FB55C2DE" authorityName="Bernhauer" authorityYear="1908" box="[1095,1230,1908,1933]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="serpentinus">
|
||
<emphasis id="B0F88E4FFFCEFFEB920CF8F1FB55C2DE" box="[1095,1230,1908,1933]" italics="true" pageId="5" pageNumber="20">serpentinus</emphasis>
|
||
</taxonomicName>
|
||
/ Brh.
|
||
<typeStatus id="5D37ECFFFFCEFFEB9363F8F1FAE8C2DD" box="[1320,1395,1908,1934]" pageId="5" pageNumber="20">Typus</typeStatus>
|
||
. [Bernhauer’s handwriting on brown label]”, “Chicago NHMus / M. Bernhauer / Collection [printed on white label]”;
|
||
<collectionCode id="E49DCA98FFCEFFEB972EF83FFE25C287" box="[357,446,1978,2004]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="20" type="Museum">FMNH</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="D6F302D5FFCDFFE896F6F8D9FB1CC2B2" ID-DOI="http://doi.org/10.5281/zenodo.4542895" ID-Zenodo-Dep="4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="6" pageNumber="21" startId="6.[189,232,1884,1910]" targetBox="[190,1392,293,1804]" targetPageId="6">
|
||
<paragraph id="8233525DFFCDFFE896F6F8D9FB1CC2B2" blockId="6.[187,1399,1882,2017]" pageId="6" pageNumber="21">
|
||
<emphasis id="B0F88E4FFFCDFFE896F6F8D9FE97C225" bold="true" box="[189,268,1884,1910]" pageId="6" pageNumber="21">Fig. 1.</emphasis>
|
||
<taxonomicName id="458C29DEFFCDFFE8975EF8D9FD60C224" authority="Bernhauer, 1908" authorityName="Bernhauer" authorityYear="1908" box="[277,763,1884,1911]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="serpentinus">
|
||
<emphasis id="B0F88E4FFFCDFFE8975EF8D9FDB6C225" box="[277,557,1884,1910]" italics="true" pageId="6" pageNumber="21">Ophionthus serpentinus</emphasis>
|
||
<bibRefCitation id="E61D2FACFFCDFFE8947FF8D9FD60C224" author="Bernhauer M." box="[564,763,1884,1911]" pageId="6" pageNumber="21" pagination="283 - 372" refId="ref6978" refString="Bernhauer M. 1908. Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte 74 (1): 283 - 372. Available from https: // www. biodiversitylibrary. org / part / 226088 # / summary [accessed 28 Jan. 2021]." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
|
||
</taxonomicName>
|
||
. Holotype, ♂ (FMNH).
|
||
<emphasis id="B0F88E4FFFCDFFE89251F8D9FBB4C225" bold="true" box="[1050,1071,1884,1910]" pageId="6" pageNumber="21">A</emphasis>
|
||
. Habitus, dorsal view, after
|
||
<bibRefCitation id="E61D2FACFFCDFFE896F5F805FE2EC2C9" author="Chani-Posse M. R." box="[190,437,1920,1946]" pageId="6" pageNumber="21" pagination="62 - 86" refId="ref7271" refString="Chani-Posse M. R. 2014 a. An illustrated key to the New World genera of Philonthina Kirby (Coleoptera: Staphylinidae), with morphological, taxonomical and distributional notes. Zootaxa 3755 (1): 62 - 86. https: // doi. org / 10.11646 / zootaxa. 3755.1.3" type="journal article" year="2014">Chani-Posse (2014a)</bibRefCitation>
|
||
.
|
||
<emphasis id="B0F88E4FFFCDFFE897F6F805FE4AC2C9" bold="true" box="[445,465,1920,1946]" pageId="6" pageNumber="21">B</emphasis>
|
||
. Original labels.
|
||
<emphasis id="B0F88E4FFFCDFFE894DCF8FAFD37C2CA" bold="true" box="[663,684,1919,1945]" pageId="6" pageNumber="21">C</emphasis>
|
||
. Tergum VIII.
|
||
<emphasis id="B0F88E4FFFCDFFE89513F805FCF6C2C9" bold="true" box="[856,877,1920,1946]" pageId="6" pageNumber="21">D</emphasis>
|
||
. Sternum VIII.
|
||
<emphasis id="B0F88E4FFFCDFFE89268F805FBACC2C9" bold="true" box="[1059,1079,1920,1946]" pageId="6" pageNumber="21">E</emphasis>
|
||
. Sternum IX.
|
||
<emphasis id="B0F88E4FFFCDFFE89291F805FB77C2C9" bold="true" box="[1242,1260,1920,1946]" pageId="6" pageNumber="21">F</emphasis>
|
||
. Tergum X.
|
||
<emphasis id="B0F88E4FFFCDFFE896F0F826FF49C2EE" bold="true" box="[187,210,1955,1981]" pageId="6" pageNumber="21">G</emphasis>
|
||
. Aedeagus, lateral view.
|
||
<emphasis id="B0F88E4FFFCDFFE897B8F826FD91C2EE" bold="true" box="[499,522,1955,1981]" pageId="6" pageNumber="21">H</emphasis>
|
||
. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8233525DFFCCFFE996A1FE8DFEEAC471" blockId="7.[189,1399,264,397]" box="[234,369,264,290]" pageId="7" pageNumber="22">
|
||
<emphasis id="B0F88E4FFFCCFFE996A1FE8DFEEAC471" bold="true" box="[234,369,264,290]" pageId="7" pageNumber="22">Comments</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6FEA9FEA1C4DE" blockId="7.[189,1399,264,397]" pageId="7" pageNumber="22">
|
||
<bibRefCitation id="E61D2FACFFCCFFE996F6FEA9FE0BC415" author="Bernhauer M." box="[189,400,300,326]" pageId="7" pageNumber="22" pagination="283 - 372" refId="ref6978" refString="Bernhauer M. 1908. Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte 74 (1): 283 - 372. Available from https: // www. biodiversitylibrary. org / part / 226088 # / summary [accessed 28 Jan. 2021]." type="journal article" year="1908">Bernhauer (1908)</bibRefCitation>
|
||
stated that Skalitzky gave him a unique specimen of
|
||
<taxonomicName id="458C29DEFFCCFFE99251FEA9FB53C416" authorityName="Bernhauer" authorityYear="1908" box="[1050,1224,300,326]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="22" phylum="Arthropoda" rank="species" species="serpentinus">
|
||
<emphasis id="B0F88E4FFFCCFFE99251FEA9FB53C416" box="[1050,1224,300,326]" italics="true" pageId="7" pageNumber="22">O. serpentinus</emphasis>
|
||
</taxonomicName>
|
||
; the specimen mentioned above agrees with the original description and is considered to be the
|
||
<typeStatus id="5D37ECFFFFCCFFE992D0FECAFA9EC43A" box="[1179,1285,335,361]" pageId="7" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
, fixed by monotypy.
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996A1FE36FE3DC49E" blockId="7.[189,1399,435,603]" box="[234,422,435,461]" pageId="7" pageNumber="22">
|
||
<emphasis id="B0F88E4FFFCCFFE996A1FE36FE3DC49E" bold="true" box="[234,422,435,461]" pageId="7" pageNumber="22">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6FE52FD82C709" blockId="7.[189,1399,435,603]" pageId="7" pageNumber="22">
|
||
<materialsCitation id="32E45800FFCCFFE996F6FE52FD8EC709" ID-GBIF-Occurrence="3037423303" collectedFrom="a clearing in montane forest and riverside shrubbery" collectingDate="1952-12-12" collectionCode="MUSM" collectorName="Koepcke & M. Koepcke & Coll. Koepcke" country="Peru" elevation="1800" latitude="-7.6666665" location="Llaguen" longLatPrecision="1303" longitude="-78.666664" municipality="Quebrada Borgapampa" pageId="7" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="La Libertad">
|
||
<collectingCountry id="FA9B12CDFFCCFFE996F6FE52FE93C4A2" box="[189,264,471,497]" name="Peru" pageId="7" pageNumber="22">PERU</collectingCountry>
|
||
–
|
||
<emphasis id="B0F88E4FFFCCFFE99763FE52FE59C4A3" bold="true" box="[296,450,470,497]" pageId="7" pageNumber="22">
|
||
<collectingRegion id="40489CBFFFCCFFE99763FE52FE59C4A3" box="[296,450,470,497]" country="Peru" name="La Libertad" pageId="7" pageNumber="22">La Libertad</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="948A99D4FFCCFFE99794FE52FD90C4A2" box="[479,523,471,497]" pageId="7" pageNumber="22" type="female">1 ♀</specimenCount>
|
||
[specimen glued to the tip of a triangle of white card]; LL [La Libertad],
|
||
<collectingMunicipality id="6257C827FFCCFFE996F6FE7FFE54C747" box="[189,463,506,532]" pageId="7" pageNumber="22">Quebrada Borgapampa</collectingMunicipality>
|
||
, Hda. [Hacienda]
|
||
<location id="87530486FFCCFFE994EDFE7FFC91C747" LSID="urn:lsid:plazi:treatment:0A25E34BFFCEFFE79468FAC6FCF4C7A4:87530486FFCCFFE994EDFE7FFC91C747" box="[678,778,506,532]" country="Peru" latitude="-7.6666665" longLatPrecision="1303" longitude="-78.666664" municipality="Quebrada Borgapampa" name="Llaguen" pageId="7" pageNumber="22" stateProvince="La Libertad">Llaguén</location>
|
||
; ca
|
||
<geoCoordinate id="E7B8349AFFCCFFE99572FE7DFC0AC747" box="[825,913,504,532]" degrees="7" direction="south" minutes="40" orientation="latitude" pageId="7" pageNumber="22" precision="925" value="-7.6666665">7°40′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E7B8349AFFCCFFE995D4FE7DFB8AC747" box="[927,1041,504,532]" degrees="78" direction="west" minutes="40" orientation="longitude" pageId="7" pageNumber="22" precision="925" value="-78.666664">78°40′ W</geoCoordinate>
|
||
; ca
|
||
<quantity id="4574FFB8FFCCFFE9920BFE7FFB02C747" box="[1088,1177,506,532]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="22" unit="m" value="1800.0">
|
||
<elevation id="09A1B56EFFCCFFE9920BFE7FFB02C747" box="[1088,1177,506,532]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="22" unit="m" value="1800.0">1800 m</elevation>
|
||
</quantity>
|
||
[m a.s.l.];
|
||
<date id="F632749DFFCCFFE99351FE7FFF60C76B" pageId="7" pageNumber="22" value="1952-12-12">
|
||
<collectingDate id="E6768D75FFCCFFE99351FE7FFF60C76B" pageId="7" pageNumber="22" value="1952-12-12">12 Dec. 1952</collectingDate>
|
||
</date>
|
||
; H-W.
|
||
<collectorName id="2F79378BFFCCFFE99710FD98FE58C764" box="[347,451,541,567]" pageId="7" pageNumber="22">Koepcke</collectorName>
|
||
&
|
||
<collectorName id="2F79378BFFCCFFE997BDFD9BFD16C764" box="[502,653,541,568]" pageId="7" pageNumber="22">M. Koepcke</collectorName>
|
||
leg.;
|
||
<collectedFrom id="319011A0FFCCFFE99497FD9BFAE9C764" box="[732,1394,541,568]" pageId="7" pageNumber="22">a clearing in montane forest and riverside shrubbery</collectedFrom>
|
||
;
|
||
<collectorName id="2F79378BFFCCFFE996F6FDC5FEFDC709" box="[189,358,576,602]" pageId="7" pageNumber="22">Coll. Koepcke</collectorName>
|
||
#665;
|
||
<collectionCode id="E49DCA98FFCCFFE997F3FDC5FD8EC709" box="[440,533,576,602]" pageId="7" pageNumber="22">MUSM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CA9601D6FFCCFFE996F6FD04FB73C642" pageId="7" pageNumber="22" type="diagnosis">
|
||
<paragraph id="8233525DFFCCFFE996F6FD04FEADC7C8" blockId="7.[189,310,641,667]" box="[189,310,641,667]" pageId="7" pageNumber="22">
|
||
<heading id="D97BE531FFCCFFE996F6FD04FEADC7C8" bold="true" box="[189,310,641,667]" fontSize="11" level="3" pageId="7" pageNumber="22" reason="3">
|
||
<emphasis id="B0F88E4FFFCCFFE996F6FD04FEADC7C8" bold="true" box="[189,310,641,667]" pageId="7" pageNumber="22">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6FD35FB73C642" blockId="7.[188,1398,688,785]" pageId="7" pageNumber="22">
|
||
<taxonomicName id="458C29DEFFCCFFE996F6FD35FE48C799" authorityName="Bernhauer" authorityYear="1908" box="[189,467,688,714]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="22" phylum="Arthropoda" rank="species" species="serpentinus">
|
||
<emphasis id="B0F88E4FFFCCFFE996F6FD35FE48C799" box="[189,467,688,714]" italics="true" pageId="7" pageNumber="22">Ophionthus serpentinus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="458C29DEFFCCFFE99420FD34FD78C799" authority="Rodríguez-Melgarejo & Chani-Posse, 2021" authorityName="Rodríguez-Melgarejo & Chani-Posse" authorityYear="2021" box="[619,739,689,715]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="22" phylum="Arthropoda" rank="species" species="asenjoi" status="sp. nov.">
|
||
<emphasis id="B0F88E4FFFCCFFE99420FD34FD78C799" box="[619,739,689,715]" italics="true" pageId="7" pageNumber="22">O. asenjoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="ABCB3334FFCCFFE994A1FD34FCDFC798" box="[746,836,689,715]" pageId="7" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having the head with two pairs of interocular punctures, antennae with the last two antennomeres much lighter than the preceding segments (
|
||
<figureCitation id="1AB74ED8FFCCFFE9935BFD56FAEBC7BD" box="[1296,1392,723,750]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="7" pageNumber="22">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="1AB74ED8FFCCFFE996F7FD72FF51C642" box="[188,202,759,785]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="7" pageNumber="22">2</figureCitation>
|
||
A–B) and the abdominal terga III–V each with a distinct posterior basal transverse carina.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CA9601D6FFCCFFE796F6FCBDFCF4C7A4" lastPageId="9" lastPageNumber="24" pageId="7" pageNumber="22" type="description">
|
||
<paragraph id="8233525DFFCCFFE996F6FCBDFEF0C601" blockId="7.[189,363,824,850]" box="[189,363,824,850]" pageId="7" pageNumber="22">
|
||
<heading id="D97BE531FFCCFFE996F6FCBDFEF0C601" bold="true" box="[189,363,824,850]" fontSize="11" level="3" pageId="7" pageNumber="22" reason="3">
|
||
<emphasis id="B0F88E4FFFCCFFE996F6FCBDFEF0C601" bold="true" box="[189,363,824,850]" pageId="7" pageNumber="22">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6FCE2FD52C6D2" blockId="7.[189,713,871,897]" box="[189,713,871,897]" pageId="7" pageNumber="22">
|
||
DIMENSIONS. BL= 11.3–1.6 (
|
||
<figureCitation id="1AB74ED8FFCCFFE99448FCE2FDFEC6D2" box="[515,613,871,897]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="7" pageNumber="22">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="1AB74ED8FFCCFFE9943AFCE2FD1BC6D2" box="[625,640,871,897]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="7" pageNumber="22">2</figureCitation>
|
||
A–B).
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6FC2DFB77C15A" blockId="7.[189,1399,936,1033]" pageId="7" pageNumber="22">
|
||
COLOURATION. Head, thorax, and abdomen shiny black, except apical half of tergum VII, apical third of sternum VII, and segments VIII and IX yellow; mandibles, maxillary and labial palps, elytra, and legs dark brown; antennomeres 1–9 dark brown to black and 10–11 yellowish (
|
||
<figureCitation id="1AB74ED8FFCCFFE9926CFC6BFB13C15A" box="[1063,1160,1006,1033]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="7" pageNumber="22">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="1AB74ED8FFCCFFE992DEFC6AFB3FC15A" box="[1173,1188,1007,1033]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="7" pageNumber="22">2</figureCitation>
|
||
A–B).
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6FBB5FC63C030" blockId="7.[189,1401,1072,1380]" pageId="7" pageNumber="22">
|
||
HEAD. As wide as long (female) (
|
||
<figureCitation id="1AB74ED8FFCCFFE9940CFBB5FD3FC119" box="[583,676,1072,1098]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="7" pageNumber="22">Fig. 2A</figureCitation>
|
||
) to wider than long (male) (
|
||
<figureCitation id="1AB74ED8FFCCFFE995B9FBB5FBD5C119" box="[1010,1102,1072,1098]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="7" pageNumber="22">Fig. 1A</figureCitation>
|
||
) (HW/HL= 1.0–1.2) and moderately wider than pronotum (HW/PW = 1.2–1.4). Eyes moderately shorter than temples (YL/TL = 0.5–0.6) seen from above. Epicranium with 2 pairs of interocular punctures; each side of vertex with 4–5 postocular punctures. Antennae with antennomere 1 at least 2× as long as antennomere 2 (A1/A2 = 2.1–2.6) and almost as long as antennomeres 2 and 3 combined (A1/(A2+A3) = 0.9–1.1), antennomere 2 moderately shorter than antennomere 3 (A2/A3 = 0.8). Labial palpus with palpomere 2 1.4× as long as palpomere 1 and 0.6× as long as palpomere 3. Maxillary palpus with palpomere 1 distinctly short; palpomere 2 curved, 2× as long as palpomere 3; palpomere 4 slightly longer than palpomere 2, 2× as long as palpomere 3. Neck about 0.4× as wide as head at widest point.
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6FA09FDFEC09A" blockId="7.[189,1398,1419,1481]" pageId="7" pageNumber="22">PRONOTUM. Longer than wide (PL/PW = 1.2–1.3), with two dorsal rows of 4 punctures each on disk of pronotum, 3–4 sublateral punctures.</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6FA75FC48C359" blockId="7.[189,979,1519,1546]" box="[189,979,1519,1546]" pageId="7" pageNumber="22">
|
||
LEGS. First metatarsomere longer than fifth (S1/S5 = 1.8) (
|
||
<figureCitation id="1AB74ED8FFCCFFE9952CFA75FC5DC359" box="[871,966,1520,1546]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="7" pageNumber="22">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6F9B7FA8EC31F" blockId="7.[189,1301,1585,1612]" box="[189,1301,1585,1612]" pageId="7" pageNumber="22">ELYTRON. Longer than wide (EL/EW = 1.8), at sides about 1.6× as long as elytron along suture.</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6F9F6FB3FC3DF" blockId="7.[189,1188,1650,1677]" box="[189,1188,1650,1677]" pageId="7" pageNumber="22">ABDOMEN. Abdominal terga III–V each with posterior basal transverse carina distinct.</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6F931FB2FC2F2" blockId="7.[189,1399,1716,1954]" pageId="7" pageNumber="22">
|
||
MALE SEXUAL CHARACTERS. Tergum VIII with hind margin projected, forming an obtuse angle (
|
||
<figureCitation id="1AB74ED8FFCCFFE99347F931FAF3C39D" box="[1292,1384,1716,1742]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="7" pageNumber="22">Fig. 1C</figureCitation>
|
||
). Sternum VIII slightly emarginate apically at middle, with numerous long setae on apical area and small setae on basal area (
|
||
<figureCitation id="1AB74ED8FFCCFFE997E5F97FFD96C247" box="[430,525,1786,1812]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="7" pageNumber="22">Fig. 1D</figureCitation>
|
||
). Sternum IX symmetrical, narrowed in the middle of its lateral margins, deeply emarginate apically, with several fine and short setae at each side of emargination (
|
||
<figureCitation id="1AB74ED8FFCCFFE99347F898FAF3C26B" box="[1292,1384,1821,1848]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="7" pageNumber="22">Fig. 1E</figureCitation>
|
||
). Tergum X with apex rounded, with numerous setae at apex (
|
||
<figureCitation id="1AB74ED8FFCCFFE995E6F8C4FB91C208" box="[941,1034,1857,1883]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="7" pageNumber="22">Fig. 1F</figureCitation>
|
||
). Aedeagus with median lobe distinctly broadened at middle and gradually narrowed towards a rather truncate apex seen from above, paramere triangular with base concave (
|
||
<figureCitation id="1AB74ED8FFCCFFE994D9F802FD48C2F2" box="[658,723,1927,1953]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="7" pageNumber="22">Fig. 1</figureCitation>
|
||
G–H), internal sclerites as in
|
||
<figureCitation id="1AB74ED8FFCCFFE99264F802FBF4C2F2" box="[1071,1135,1927,1953]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="7" pageNumber="22">Fig. 1</figureCitation>
|
||
G–H.
|
||
</paragraph>
|
||
<paragraph id="8233525DFFCCFFE996F6F84CFCD2C2B0" blockId="7.[189,841,1992,2019]" box="[189,841,1992,2019]" pageId="7" pageNumber="22">FEMALE SEXUAL CHARACTERS. As described for the genus.</paragraph>
|
||
<caption id="D6F302D5FFC3FFE696F6F8DBFC50C2B0" ID-DOI="http://doi.org/10.5281/zenodo.4542899" ID-Zenodo-Dep="4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="8" pageNumber="23" startId="8.[189,232,1886,1912]" targetBox="[199,1394,353,1755]" targetPageId="8">
|
||
<paragraph id="8233525DFFC3FFE696F6F8DBFC50C2B0" blockId="8.[186,1398,1886,2019]" pageId="8" pageNumber="23">
|
||
<emphasis id="B0F88E4FFFC3FFE696F6F8DBFE93C22B" bold="true" box="[189,264,1886,1912]" pageId="8" pageNumber="23">Fig. 2.</emphasis>
|
||
<taxonomicName id="458C29DEFFC3FFE69747F8DBFD77C22A" authority="Bernhauer, 1908" authorityName="Bernhauer" authorityYear="1908" box="[268,748,1886,1913]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="23" phylum="Arthropoda" rank="species" species="serpentinus">
|
||
<emphasis id="B0F88E4FFFC3FFE69747F8DBFDBAC22B" box="[268,545,1886,1912]" italics="true" pageId="8" pageNumber="23">Ophionthus serpentinus</emphasis>
|
||
<bibRefCitation id="E61D2FACFFC3FFE69463F8DBFD77C22A" author="Bernhauer M." box="[552,748,1886,1913]" pageId="8" pageNumber="23" pagination="283 - 372" refId="ref6978" refString="Bernhauer M. 1908. Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte 74 (1): 283 - 372. Available from https: // www. biodiversitylibrary. org / part / 226088 # / summary [accessed 28 Jan. 2021]." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
|
||
</taxonomicName>
|
||
. ♀ (MUSM).
|
||
<emphasis id="B0F88E4FFFC3FFE695C0F8DBFC3BC22B" bold="true" box="[907,928,1886,1912]" pageId="8" pageNumber="23">A</emphasis>
|
||
. Habitus, dorsal view.
|
||
<emphasis id="B0F88E4FFFC3FFE692E8F8DAFB2CC22A" bold="true" box="[1187,1207,1887,1913]" pageId="8" pageNumber="23">B</emphasis>
|
||
. Habitus, lateral view.
|
||
<emphasis id="B0F88E4FFFC3FFE6974BF804FE8EC2C8" bold="true" box="[256,277,1921,1947]" pageId="8" pageNumber="23">C</emphasis>
|
||
. Right protarsomeres, ventral view.
|
||
<emphasis id="B0F88E4FFFC3FFE694E7F807FD5AC2CF" bold="true" box="[684,705,1922,1948]" pageId="8" pageNumber="23">D</emphasis>
|
||
. Details of discal setae from right third protarsomere, ventral view.
|
||
<emphasis id="B0F88E4FFFC3FFE69748F820FE8CC2EC" bold="true" box="[259,279,1957,1983]" pageId="8" pageNumber="23">E</emphasis>
|
||
. Tergum VIII.
|
||
<emphasis id="B0F88E4FFFC3FFE6978CF820FE42C2EC" bold="true" box="[455,473,1957,1983]" pageId="8" pageNumber="23">F</emphasis>
|
||
. Sternum VIII.
|
||
<emphasis id="B0F88E4FFFC3FFE694DFF820FD30C2EC" bold="true" box="[660,683,1957,1983]" pageId="8" pageNumber="23">G</emphasis>
|
||
. Lateral tergal sclerites IX and tergum X.
|
||
<emphasis id="B0F88E4FFFC3FFE692D6F820FB2FC2EC" bold="true" box="[1181,1204,1957,1983]" pageId="8" pageNumber="23">H</emphasis>
|
||
. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm..
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8233525DFFC2FFE796F6FE8DFD24C471" blockId="9.[189,703,264,291]" box="[189,703,264,291]" pageId="9" pageNumber="24">
|
||
<heading id="D97BE531FFC2FFE796F6FE8DFD24C471" bold="true" box="[189,703,264,291]" fontSize="11" level="3" pageId="9" pageNumber="24" reason="3">
|
||
<emphasis id="B0F88E4FFFC2FFE796F6FE8DFD24C471" bold="true" box="[189,703,264,291]" pageId="9" pageNumber="24">Geographical distribution and bionomics</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8233525DFFC2FFE796F6FEB2FCF4C7A4" blockId="9.[189,1400,311,760]" pageId="9" pageNumber="24">
|
||
The male
|
||
<typeStatus id="5D37ECFFFFC2FFE7977EFEB2FE07C402" box="[309,412,311,337]" pageId="9" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="458C29DEFFC2FFE79783FEB2FDEFC402" authorityName="Bernhauer" authorityYear="1908" box="[456,628,311,337]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="serpentinus">
|
||
<emphasis id="B0F88E4FFFC2FFE79783FEB2FDEFC402" box="[456,628,311,337]" italics="true" pageId="9" pageNumber="24">O. serpentinus</emphasis>
|
||
</taxonomicName>
|
||
was collected in 1873 by Jelski in “Peruvia centralis” (
|
||
<figureCitation id="1AB74ED8FFC2FFE79340FEB2FAFCC402" box="[1291,1383,311,337]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="9" pageNumber="24">Fig. 1B</figureCitation>
|
||
). Jelski travelled and collected in the central Andes of
|
||
<collectingCountry id="FA9B12CDFFC2FFE79579FEDEFCF3C426" box="[818,872,347,373]" name="Peru" pageId="9" pageNumber="24">Peru</collectingCountry>
|
||
during that year (
|
||
<bibRefCitation id="E61D2FACFFC2FFE79270FEDFFB48C426" author="Solsky S." box="[1083,1235,346,373]" pageId="9" pageNumber="24" pagination="3 - 26" refId="ref8721" refString="Solsky S. 1875. Materiaux pour l'entomographie de l'Amerique du Sud. Staphylinides recueillis par MM. C. Jelsky et le Baron de Nolcken dans le Perou et la Nouvelle Grenade. Article III. Horae Societatis Entomologicae Rossicae 11 (1): 3 - 26." type="journal article" year="1875">Solsky 1875</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E61D2FACFFC2FFE79294FEDFFF67C4CB" author="Taczanowski L." pageId="9" pageNumber="24" pagination="501 - 565" refId="ref8868" refString="Taczanowski L. 1875. Liste des oiseaux recueillis par M. Constantin Jelski dans la partie centrale du Perou occidental. Proceedings of the Scientific Meetings of the Zoological Society of London 1874 (3): 501 - 565. https: // doi. org / 10.1111 / j. 1096 - 3642.1874. tb 02510. x" type="journal article" year="1875">Taczanowski 1875</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E61D2FACFFC2FFE79741FEF8FE55C4CB" author="Mlikovsky J." box="[266,462,381,408]" pageId="9" pageNumber="24" pagination="17 - 180" refId="ref8246" refString="Mlikovsky J. 2009. Types of birds in the collections of the Museum and Institute of Zoology, Polish Academy of Sciences, Warszawa, Poland. Part 3: South American birds. Journal of the National Museum (Prague), Natural History Series 178 (5): 17 - 180. Available from https: // publikace. nm. cz / en / file / 6 a 5 c 9 f 94784 eaa 047577335 ba 8553 b 02 / 16385 / 178 - 2009 - 05 - Mlikovsky. pdf [accessed 5 Feb. 2021]." type="journal article" year="2009">Mlíkovský 2009</bibRefCitation>
|
||
) and the localities he visited are summarized in the Supp. file 1 and
|
||
<figureCitation id="1AB74ED8FFC2FFE7934CFEF8FAE9C4CB" box="[1287,1394,381,408]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="11.[189,232,1972,1998]" captionStartId-1="12.[189,232,1937,1963]" captionTargetBox-0="[189,1398,1237,1937]" captionTargetBox-1="[189,1398,1169,1895]" captionTargetId-0="figure-257@11.[189,1399,1237,1937]" captionTargetId-1="figure-338@12.[189,1398,1168,1896]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Fig. 4. Distribution map of Ophionthus serpentinus Bernhauer, 1908 and O. asenjoi sp. nov." captionText-1="Fig. 5. Map of probable localities (correct spelling, see Supp. file 1) for the male holotype of Ophionthus serpentinus Bernhauer, 1908 collected in “Peruvia centralis” by Jelski in 1873." figureDoi-0="http://doi.org/10.5281/zenodo.4542903" figureDoi-1="http://doi.org/10.5281/zenodo.4542905" httpUri-0="https://zenodo.org/record/4542903/files/figure.png" httpUri-1="https://zenodo.org/record/4542905/files/figure.png" pageId="9" pageNumber="24">Figs 4–5</figureCitation>
|
||
. Based on these analyses, the type locality “Peruvia centralis” is located in the department of
|
||
<collectingRegion id="40489CBFFFC2FFE7934FFE24FADCC4E8" box="[1284,1351,417,443]" country="Peru" name="Junin" pageId="9" pageNumber="24">Junín</collectingRegion>
|
||
, but we do not have more information (e.g., day and/or month of collection) to improve the accuracy of this type locality (
|
||
<figureCitation id="1AB74ED8FFC2FFE7972BFE62FE20C752" box="[352,443,487,513]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="9" pageNumber="24">Fig. 1B</figureCitation>
|
||
). Additionally, the new locality provided by the female specimen from Hacienda Llaguén, ca
|
||
<geoCoordinate id="E7B8349AFFC2FFE79719FD8CFE37C777" box="[338,428,521,549]" degrees="7" direction="south" minutes="40" orientation="latitude" pageId="9" pageNumber="24" precision="925" value="-7.6666665">7°40′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E7B8349AFFC2FFE797F7FD8CFDB5C776" box="[444,558,521,549]" degrees="78" direction="west" minutes="40" orientation="longitude" pageId="9" pageNumber="24" precision="925" value="-78.666664">78°40′ W</geoCoordinate>
|
||
,
|
||
<quantity id="4574FFB8FFC2FFE79477FD8FFD0CC776" box="[572,663,522,549]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="24" unit="m" value="1800.0">1800 m</quantity>
|
||
[m a.s.l.] (
|
||
<bibRefCitation id="E61D2FACFFC2FFE79556FD8FFC56C777" author="Koepcke H. W." box="[797,973,522,548]" pageId="9" pageNumber="24" refId="ref7874" refString="Koepcke H. W. 1982. Erlauterung der Beobachtungs- und Sammler-Nummern zu den in den Jahren 1949 bis 1974 in Peru durchgefuhrten biologischen Studien von Hans-Wilhelm Koepcke und Maria Koepcke. Hamburg. Availabe from https: // issuu. com / ifrankej / docs / koepcke _ h _ w _ 1982 _ erl __ uterung [accessed 5 Feb. 2021]." type="url" year="1982">Koepcke 1982</bibRefCitation>
|
||
) is located in the department of
|
||
<collectingRegion id="40489CBFFFC2FFE79313FD8EFEBBC714" country="Peru" name="La Libertad" pageId="9" pageNumber="24">La Libertad</collectingRegion>
|
||
and close to the Forest of Llaguén (
|
||
<geoCoordinate id="E7B8349AFFC2FFE79484FDA9FCB2C71B" box="[719,809,556,584]" degrees="7" direction="south" minutes="42" orientation="latitude" pageId="9" pageNumber="24" precision="925" value="-7.7">7°42′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E7B8349AFFC2FFE7957CFDA9FC33C71B" box="[823,936,556,584]" degrees="78" direction="west" minutes="44" orientation="longitude" pageId="9" pageNumber="24" precision="925" value="-78.73333">78°44′ W</geoCoordinate>
|
||
,
|
||
<quantity id="4574FFB8FFC2FFE795FEFDA8FB94C71B" box="[949,1039,557,584]" metricMagnitude="3" metricUnit="m" metricValue="2.642" pageId="9" pageNumber="24" unit="m" value="2642.0">2642 m</quantity>
|
||
[m a.s.l.]), which is described as a dry cloud forest, with high human disturbance and used for grazing cattle and as a source of fuel (
|
||
<bibRefCitation id="E61D2FACFFC2FFE7968DFDF1FEEEC7DD" author="Valencia N." box="[198,373,628,654]" pageId="9" pageNumber="24" refId="ref8987" refString="Valencia N. 1990. Ecology of the Forests on the Western Slopes of the Peruvian Andes. PhD thesis, University of Aberdeen, Aberdeen. Available from https: // abdn. alma. exlibrisgroup. com / view / delivery / 44 ABE _ INST / 12152267510005941 [accessed 5 Feb. 2021]." type="url" year="1990">Valencia 1990</bibRefCitation>
|
||
). Hacienda Llaguén is placed in the ecoregion of Peruvian Yungas (
|
||
<bibRefCitation id="E61D2FACFFC2FFE792D5FDF1FAF5C7DD" author="Olson D. M. & Dinerstein E. & Wikramanayake E. D. & Burgess N. D. & Powell G. V. & Underwood E. C. & D'Amico J. & Itoua I. & Strand H. E. & Morrison J. C. & Loucks C. J. & Allnutt T. F. & Ricketts T. H. & Kura Y. & Lamoreux J. F. & Wettengel W. W. & Hedaeo P. & Kassen K. R." box="[1182,1390,628,654]" pageId="9" pageNumber="24" pagination="933 - 938" refId="ref8386" refString="Olson D. M., Dinerstein E., Wikramanayake E. D., Burgess N. D., Powell G. V., Underwood E. C., D'Amico J., Itoua I., Strand H. E., Morrison J. C., Loucks C. J., Allnutt T. F., Ricketts T. H., Kura Y., Lamoreux J. F., Wettengel W. W., Hedaeo P. & Kassen K. R. 2001. Terrestrial ecoregions of the world: a new map of life on earth: a new global map of terrestrial ecoregions provides an innovative tool for conserving biodiversity. BioScience 51 (11): 933 - 938. https: // doi. org / 10.1641 / 0006 - 3568 (2001) 051 [0933: TEOTWA] 2.0. CO; 2" type="journal article" year="2001">
|
||
Olson
|
||
<emphasis id="B0F88E4FFFC2FFE792A4FDF0FAB9C7DD" box="[1263,1314,628,654]" italics="true" pageId="9" pageNumber="24">et al</emphasis>
|
||
. 2001
|
||
</bibRefCitation>
|
||
) or in the Puna province (
|
||
<bibRefCitation id="E61D2FACFFC2FFE797AAFD12FD16C7E2" author="Morrone J. J." box="[481,653,663,689]" pageId="9" pageNumber="24" pagination="1 - 110" refId="ref8350" refString="Morrone J. J. 2014. Biogeographical regionalisation of the Neotropical region. Zootaxa 3782 (1): 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">Morrone 2014</bibRefCitation>
|
||
) (
|
||
<figureCitation id="1AB74ED8FFC2FFE794EDFD12FD76C7E2" box="[678,749,663,689]" captionStart="Fig" captionStartId="11.[189,232,1972,1998]" captionTargetBox="[189,1398,1237,1937]" captionTargetId="figure-257@11.[189,1399,1237,1937]" captionTargetPageId="11" captionText="Fig. 4. Distribution map of Ophionthus serpentinus Bernhauer, 1908 and O. asenjoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4542903" httpUri="https://zenodo.org/record/4542903/files/figure.png" pageId="9" pageNumber="24">Fig. 4</figureCitation>
|
||
) and nowadays it is placed in the ecosystem of Andean shrublands (
|
||
<bibRefCitation id="E61D2FACFFC2FFE79706FD3FFD9BC787" author="MINAM" box="[333,512,698,724]" pageId="9" pageNumber="24" refId="ref8217" refString="MINAM [Ministerio del Ambiente del Peru]. 2019. Mapa nacional de ecosistemas del Peru. Available from mapa-nacional-de-ecosistemas-del-peru [accessed 3 Jun. 2020]." type="book" year="2019">MINAM 2019</bibRefCitation>
|
||
). According to label information, the female specimen was collected in an open area within a montane forest and riverside shrubbery.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |