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<document id="98418AB41FAB83628A563D6654A0632A" ID-CLB-Dataset="8173" ID-DOI="10.5852/ejt.2021.735.1237" ID-GBIF-Dataset="4074b185-a219-4e7e-b1bd-1bc4bd3ef9e7" ID-ISSN="2118-9773" ID-Zenodo-Dep="4542891" ID-ZooBank="8364CCA1-11A1-437F-944D-BA70C690D7F7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613471756163" checkinUser="plazi" docAuthor="Rodríguez-Melgarejo, Maryzender &amp; Chani-Posse, Mariana" docDate="2021" docId="0A25E34BFFC8FFEB940FFBFFFAA8C055" docLanguage="en" docName="ejt-735_15-33.pdf" docOrigin="European Journal of Taxonomy 735" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophionthus Bernhauer 1908" docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="F61C9B33FFCBFFEE964BFF85FF9BC553" masterDocTitle="On the rare Neotropical genus Ophionthus Bernhauer (Coleoptera: Staphylinidae: Staphylininae): redescription of the type species and description of a new species" masterLastPageNumber="33" masterPageNumber="15" pageNumber="18" updateTime="1698929669084" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="07390A63A26678676C28A621912F2B21">On the rare Neotropical genus Ophionthus Bernhauer (Coleoptera: Staphylinidae: Staphylininae): redescription of the type species and description of a new species</mods:title>
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<mods:affiliation id="50587ED55463B18AA145E13485C08A89">Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT CONICET, Mendoza), Casilla de Correo 507, 5500 Mendoza, Argentina. Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Apartado 14 - 0434, Lima 14, Perú. Facultad de Ciencias Exactas y Naturales, Universidad Nacional de Cuyo, Mendoza, Argentina.</mods:affiliation>
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<mods:namePart id="344B339B444D30B889B80D6588038F27">Chani-Posse, Mariana</mods:namePart>
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<mods:affiliation id="B1BE267F28D2DB3C834BB4B50992B8D9">Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Áridas (IADIZA, CCT CONICET, Mendoza), Casilla de Correo 507, 5500 Mendoza, Argentina. Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Apartado 14 - 0434, Lima 14, Perú. Facultad de Ciencias Exactas y Naturales, Universidad Nacional de Cuyo, Mendoza, Argentina.</mods:affiliation>
<mods:nameIdentifier id="FB34504B45C10C41B1CB7CA4DA9B71EE" type="email">mchani@mendoza-conicet.gob.ar</mods:nameIdentifier>
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Genus
<taxonomicName id="458C29DEFFC8FFED94DDFBFCFC6BC1C7" ID-CoL="8XND3" authority="Bernhauer, 1908" authorityName="Bernhauer" authorityYear="1908" box="[662,1008,1145,1172]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B0F88E4FFFC8FFED94DDFBFCFCBEC1C7" bold="true" box="[662,805,1145,1172]" italics="true" pageId="3" pageNumber="18">Ophionthus</emphasis>
<bibRefCitation id="E61D2FACFFC8FFED9566FBFFFC6BC1C7" author="Bernhauer M." box="[813,1008,1146,1172]" pageId="3" pageNumber="18" pagination="283 - 372" refId="ref6978" refString="Bernhauer M. 1908. Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte 74 (1): 283 - 372. Available from https: // www. biodiversitylibrary. org / part / 226088 # / summary [accessed 28 Jan. 2021]." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
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<figureCitation id="1AB74ED8FFC8FFED94ACFB18FCD6C1E4" box="[743,845,1181,1207]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="6.[189,232,1884,1910]" captionStartId-1="8.[189,232,1886,1912]" captionStartId-2="10.[189,232,1916,1942]" captionStartId-3="11.[189,232,1972,1998]" captionStartId-4="12.[189,232,1937,1963]" captionStartId-5="13.[189,232,1815,1841]" captionTargetBox-0="[190,1392,293,1804]" captionTargetBox-1="[199,1394,353,1755]" captionTargetBox-2="[202,1398,532,1882]" captionTargetBox-3="[189,1398,1237,1937]" captionTargetBox-4="[189,1398,1169,1895]" captionTargetBox-5="[230,1362,326,1764]" captionTargetId-0="figure-14@6.[189,1398,293,1811]" captionTargetId-1="figure-15@8.[189,1398,349,1768]" captionTargetId-2="figure-102@10.[189,1398,532,1883]" captionTargetId-3="figure-257@11.[189,1399,1237,1937]" captionTargetId-4="figure-338@12.[189,1398,1168,1896]" captionTargetId-5="figure-11@13.[206,1382,310,1780]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="13" captionText-0="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." captionText-1="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." captionText-2="Fig. 3. Ophionthus asenjoi sp. nov. Holotype, ♀ (IFML). A. Habitus, dorsal view. B. Original labels. C. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm." captionText-3="Fig. 4. Distribution map of Ophionthus serpentinus Bernhauer, 1908 and O. asenjoi sp. nov." captionText-4="Fig. 5. Map of probable localities (correct spelling, see Supp. file 1) for the male holotype of Ophionthus serpentinus Bernhauer, 1908 collected in “Peruvia centralis” by Jelski in 1873." captionText-5="Fig. 6 (continued on next page). Strict consensus of the 28 MP trees obtained showing the systematic placement of Ophionthus Bernhauer, 1908 within the Neotropical lineage of Philonthina Kirby, 1837 (tree length = 579; consistency index (CI) = 0.24; retention index (RI) = 0.71. Black circles: exclusive synapomorphies; open circles, non-exclusive synapomorphies. Numbers above circles are character numbers; numbers below circles represent character states. Support values are indicated in bold below the branches as follows: Bremer/Standard Bootstrap/Jack-knife (cut = 50 for resampling support values)." figureDoi-0="http://doi.org/10.5281/zenodo.4542895" figureDoi-1="http://doi.org/10.5281/zenodo.4542899" figureDoi-2="http://doi.org/10.5281/zenodo.4542901" figureDoi-3="http://doi.org/10.5281/zenodo.4542903" figureDoi-4="http://doi.org/10.5281/zenodo.4542905" figureDoi-5="http://doi.org/10.5281/zenodo.4545915" httpUri-0="https://zenodo.org/record/4542895/files/figure.png" httpUri-1="https://zenodo.org/record/4542899/files/figure.png" httpUri-2="https://zenodo.org/record/4542901/files/figure.png" httpUri-3="https://zenodo.org/record/4542903/files/figure.png" httpUri-4="https://zenodo.org/record/4542905/files/figure.png" httpUri-5="https://zenodo.org/record/4545915/files/figure.png" pageId="3" pageNumber="18">Figs 1–6</figureCitation>
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<treatmentCitationGroup id="A29C7573FFC8FFED96F6FB5AFDC8C1AA" box="[189,595,1247,1273]" pageId="3" pageNumber="18">
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<taxonomicName id="458C29DEFFC8FFED96F6FB5AFDD4C1AA" ID-CoL="8XND3" authority="Bernhauer, 1908: 328" authorityName="Bernhauer" authorityPageNumber="328" authorityYear="1908" box="[189,591,1247,1273]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B0F88E4FFFC8FFED96F6FB5AFEDDC1AA" box="[189,326,1247,1273]" italics="true" pageId="3" pageNumber="18">Ophionthus</emphasis>
<bibRefCitation id="E61D2FACFFC8FFED9706FB5AFDD4C1AA" author="Bernhauer M." box="[333,591,1247,1273]" pageId="3" pageNumber="18" pagination="283 - 372" refId="ref6978" refString="Bernhauer M. 1908. Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte 74 (1): 283 - 372. Available from https: // www. biodiversitylibrary. org / part / 226088 # / summary [accessed 28 Jan. 2021]." type="journal article" year="1908">Bernhauer, 1908: 328</bibRefCitation>
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.
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<taxonomicName id="458C29DEFFC8FFED96F6FAA5FEDDC069" ID-CoL="8XND3" authorityName="Bernhauer" authorityYear="1908" box="[189,326,1312,1338]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">Ophionthus</taxonomicName>
–
</emphasis>
<treatmentCitation id="032D744CFFC8FFED973AFAA5FC8AC069" author="Bernhauer M. &amp; Schubert K." box="[369,785,1312,1338]" page="372" pageId="3" pageNumber="18" year="1914">
<bibRefCitation id="E61D2FACFFC8FFED973AFAA5FC8AC069" author="Bernhauer M. &amp; Schubert K." box="[369,785,1312,1338]" pageId="3" pageNumber="18" pagination="289 - 408" refId="ref7026" refString="Bernhauer M. &amp; Schubert K. 1914. Staphylinidae IV. In: Schenkling S. (ed.) Coleopterorum Catalogus Vol. 57: 289 - 408. W. Junk, Berlin." type="journal article" year="1914">Bernhauer &amp; Schubert 1914: 372</bibRefCitation>
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(catalogue).
<emphasis id="B0F88E4FFFC8FFED95F2FAA4FC48C069" box="[953,979,1313,1338]" italics="true" pageId="3" pageNumber="18">—</emphasis>
<treatmentCitation id="032D744CFFC8FFED95AAFAA4FB29C069" author="Lucas R." box="[993,1202,1312,1339]" page="461" pageId="3" pageNumber="18" year="1920">
<bibRefCitation id="E61D2FACFFC8FFED95AAFAA4FB29C069" author="Lucas R." box="[993,1202,1312,1339]" pageId="3" pageNumber="18" pagination="1 - 5" refId="ref8076" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Archiv fur Naturgeschichte (A) 84 (1 - 5): i-xxxi, 1 - 696. Available from https: // www. zobodat. at / pdf / Archiv-Naturgeschichte _ 84 A _ 1 _ 0001 - 0128. pdf [accessed 5 Feb. 2021]." type="journal article" year="1920">Lucas 1920: 461</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="032D744CFFC8FFED96A1FAC6FD93C00E" author="Blackwelder R. E." box="[234,520,1347,1373]" page="138" pageId="3" pageNumber="18" year="1944">
<bibRefCitation id="E61D2FACFFC8FFED96A1FAC6FD93C00E" author="Blackwelder R. E." box="[234,520,1347,1373]" pageId="3" pageNumber="18" pagination="1 - 188" refId="ref7063" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part. 1. Bulletin of the United States National Museum 185: 1 - 188. https: // doi. org / 10.5479 / si. 03629236.185. i" type="journal article" year="1944">Blackwelder 1944: 138</bibRefCitation>
</treatmentCitation>
(checklist); 1952: 276. —
<treatmentCitation id="032D744CFFC8FFED952AFAC1FBCCC00E" author="Herman L. H." box="[865,1111,1347,1374]" page="2721" pageId="3" pageNumber="18" year="2001">
<bibRefCitation id="E61D2FACFFC8FFED952AFAC1FBCCC00E" author="Herman L. H." box="[865,1111,1347,1374]" pageId="3" pageNumber="18" pagination="2441 - 3020" refId="ref7738" refString="Herman L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. V. Staphylinine Group (Part 2) Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina). Bulletin of the American Museum of Natural History 265: 2441 - 3020. https: // doi. org / 10.1206 / 0003 - 0090.265.1.5" type="journal article" year="2001">Herman 2001: 2721</bibRefCitation>
</treatmentCitation>
(catalogue). — Chani- Posse 2013: 10, 12, 14 (characters, phylogenetic placement); 2014a: 64, 68, 71, 76, 78 (characters, characters in key), 81, fig. 12h (habitus); 2014b: 252 (characters in data matrix). —
<treatmentCitation id="032D744CFFC8FFED92AFFA0FFDB9C09B" author="Chani-Posse M. R. &amp; Solodovnikov A." page="95" pageId="3" pageNumber="18" year="2017">
<bibRefCitation id="E61D2FACFFC8FFED92AFFA0FFDB9C09B" author="Chani-Posse M. R. &amp; Solodovnikov A." pageId="3" pageNumber="18" pagination="83 - 97" refId="ref7516" refString="Chani-Posse M. R. &amp; Solodovnikov A. 2017. Tackling generic limits for the Neotropical Philonthina with a phylogenetic approach: revision of the genera Linoderus Sharp and Odontolinus Sharp (Coleoptera: Staphylinidae). Arthropod Systematics &amp; Phylogeny 75 (1): 83 - 97." type="journal article" year="2017">Chani-Posse &amp; Solodovnikov 2017: 95</bibRefCitation>
</treatmentCitation>
(characters). —
<treatmentCitation id="032D744CFFC8FFED94A5FA28FBD2C09B" ID-DOI="http://doi.org/10.5281/zenodo.5950920" ID-Zenodo-Dep="5950920" author="Chani-Posse M. R. &amp; Newton A. &amp; Kappel Hansen A. &amp; Solodovnikov A." box="[750,1097,1453,1480]" httpUri="http://treatment.plazi.org/id/215187C84C19B37AFF6BF89EF591D59C" page="50" pageId="3" pageNumber="18" year="2018">
<bibRefCitation id="E61D2FACFFC8FFED94A5FA28FBD2C09B" author="Chani-Posse M. R. &amp; Newton A. &amp; Kappel Hansen A. &amp; Solodovnikov A." box="[750,1097,1453,1480]" pageId="3" pageNumber="18" pagination="1 - 95" refId="ref7637" refString="Chani-Posse M. R., Newton A., Kappel Hansen A. &amp; Solodovnikov A. 2018 b. Checklist and taxonomic changes for Central and South American Philonthina (Coleoptera: Staphylinidae). Zootaxa 4449 (1): 1 - 95. https: // doi. org / 10.11646 / zootaxa. 4449.1.1" type="journal article" year="2018">
Chani-Posse
<emphasis id="B0F88E4FFFC8FFED95C6FA2BFC51C094" box="[909,970,1453,1479]" italics="true" pageId="3" pageNumber="18">et al.</emphasis>
2018b: 50
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(checklist). —
<bibRefCitation id="E61D2FACFFC8FFED934EFA28FED0C0B9" author="Asenjo A. &amp; Irmler U. &amp; Klimaszewski J. &amp; Chandler D. S. &amp; Fierros-Lopez H. E. &amp; Vieira J. S." pageId="3" pageNumber="18" pagination="1 - 406" refId="ref6906" refString="Asenjo A., Irmler U., Klimaszewski J., Chandler D. S., Fierros-Lopez H. E. &amp; Vieira J. S. 2019. Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution. Zootaxa 4621 (1): 1 - 406. https: // doi. org / 10.11646 / zootaxa. 4621.1.1" type="journal article" year="2019">
Asenjo
<emphasis id="B0F88E4FFFC8FFED9328FA2BFE9AC0B9" italics="true" pageId="3" pageNumber="18">et al</emphasis>
. 2019
</bibRefCitation>
(catalogue). —
<treatmentCitation id="032D744CFFC8FFED9441FA55FBA0C0B9" author="Chani-Posse M. R. &amp; Ramirez-Salamanca J. M." box="[522,1083,1488,1514]" page="198" pageId="3" pageNumber="18" year="2020">
<bibRefCitation id="E61D2FACFFC8FFED9441FA55FBA0C0B9" author="Chani-Posse M. R. &amp; Ramirez-Salamanca J. M." box="[522,1083,1488,1514]" pageId="3" pageNumber="18" pagination="194 - 217" refId="ref7394" refString="Chani-Posse M. R. &amp; Ramirez-Salamanca J. M. 2020 a. To be biased or to be Neotropical: systematic reassessment of a megadiverse lineage of rove-beetles (Philonthina, Staphylinini, Staphylininae). Cladistics 36 (2): 194 - 217. https: // doi. org / 10.1111 / cla. 12395" type="journal article" year="2020">Chani-Posse &amp; Ramírez-Salamanca 2020a: 198</bibRefCitation>
</treatmentCitation>
, 209–210, 213–214 (taxon sampling, phylogenetic placement); 2020b: 238, 246 (discussion on characters and phylogenetic placement).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CA9601D6FFC8FFED96F6F9DDFC0BC3F2" pageId="3" pageNumber="18" type="type_taxon">
<paragraph id="8233525DFFC8FFED96F6F9DDFEC3C321" blockId="3.[189,344,1624,1650]" box="[189,344,1624,1650]" pageId="3" pageNumber="18">
<heading id="D97BE531FFC8FFED96F6F9DDFEC3C321" bold="true" box="[189,344,1624,1650]" fontSize="11" level="3" pageId="3" pageNumber="18" reason="3">
<emphasis id="B0F88E4FFFC8FFED96F6F9DDFEC3C321" bold="true" box="[189,344,1624,1650]" pageId="3" pageNumber="18">
<typeStatus id="5D37ECFFFFC8FFED96F6F9DDFF61C321" box="[189,250,1624,1650]" pageId="3" pageNumber="18">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8233525DFFC8FFED96F6F903FC0BC3F2" blockId="3.[189,912,1670,1697]" box="[189,912,1670,1697]" pageId="3" pageNumber="18">
<taxonomicName id="458C29DEFFC8FFED96F6F903FD3BC3F2" ID-CoL="49YSH" authority="Bernhauer, 1908" authorityName="Bernhauer" authorityYear="1908" box="[189,672,1670,1697]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="serpentinus">
<emphasis id="B0F88E4FFFC8FFED96F6F903FE4FC3F3" box="[189,468,1670,1696]" italics="true" pageId="3" pageNumber="18">Ophionthus serpentinus</emphasis>
<bibRefCitation id="E61D2FACFFC8FFED9797F903FD3BC3F2" author="Bernhauer M." box="[476,672,1670,1697]" pageId="3" pageNumber="18" pagination="283 - 372" refId="ref6978" refString="Bernhauer M. 1908. Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte 74 (1): 283 - 372. Available from https: // www. biodiversitylibrary. org / part / 226088 # / summary [accessed 28 Jan. 2021]." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
</taxonomicName>
, fixed by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="CA9601D6FFC8FFED96F6F94DFAA8C2CD" pageId="3" pageNumber="18" type="diagnosis">
<paragraph id="8233525DFFC8FFED96F6F94DFEADC3B1" blockId="3.[189,310,1736,1762]" box="[189,310,1736,1762]" pageId="3" pageNumber="18">
<heading id="D97BE531FFC8FFED96F6F94DFEADC3B1" bold="true" box="[189,310,1736,1762]" fontSize="11" level="3" pageId="3" pageNumber="18" reason="3">
<emphasis id="B0F88E4FFFC8FFED96F6F94DFEADC3B1" bold="true" box="[189,310,1736,1762]" pageId="3" pageNumber="18">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8233525DFFC8FFED96F6F973FAA8C2CD" blockId="3.[188,1399,1782,1950]" pageId="3" pageNumber="18">
The genus
<taxonomicName id="458C29DEFFC8FFED9701F973FE48C243" authorityName="Bernhauer" authorityYear="1908" box="[330,467,1782,1808]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B0F88E4FFFC8FFED9701F973FE48C243" box="[330,467,1782,1808]" italics="true" pageId="3" pageNumber="18">Ophionthus</emphasis>
</taxonomicName>
can be differentiated from other genera of Neotropical
<taxonomicName id="458C29DEFFC8FFED92D8F973FA81C243" authorityName="Kirby" authorityYear="1837" box="[1171,1306,1782,1808]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
by the combination of the following characters: head and pronotum broadened anteriad, antennomere 1 at least twice as long as 2 and almost as long as antennomeres 2 and 3 combined, labial palpomere 2 cylindrical, protarsomeres 2–3 with discal setae on ventral surface and without marginal setae, prosternum with sternacostal ridge joining superior line of hypomeron, metatarsomere 1 almost twice as long as 5.
</paragraph>
</subSubSection>
<subSubSection id="CA9601D6FFCFFFEB96F6FE8DFAA8C055" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19" type="description">
<paragraph id="8233525DFFCFFFEA96F6FE8DFEF0C471" blockId="4.[189,363,264,290]" box="[189,363,264,290]" pageId="4" pageNumber="19">
<heading id="D97BE531FFCFFFEA96F6FE8DFEF0C471" bold="true" box="[189,363,264,290]" fontSize="11" level="3" pageId="4" pageNumber="19" reason="3">
<emphasis id="B0F88E4FFFCFFFEA96F6FE8DFEF0C471" bold="true" box="[189,363,264,290]" pageId="4" pageNumber="19">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8233525DFFCFFFEA96F6FEBDFC9DC427" blockId="4.[189,1399,311,373]" pageId="4" pageNumber="19">
DIMENSIONS. BL = 10.5–11.6. Body elongate, more or less parallel-sided, narrowing from seventh abdominal segment to apex (
<figureCitation id="1AB74ED8FFCFFFEA9445FEDFFDEBC427" box="[526,624,346,372]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="4" pageNumber="19">Figs 1A</figureCitation>
,
<figureCitation id="1AB74ED8FFCFFFEA9437FEDFFD11C427" box="[636,650,346,372]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="4" pageNumber="19">2</figureCitation>
A–B, 3A).
</paragraph>
<paragraph id="8233525DFFCFFFEA96F6FE19FF68C74C" blockId="4.[189,1399,412,543]" pageId="4" pageNumber="19">
COLOURATION. Head, thorax, and abdomen shiny black to piceous, except apical half of tergum 7; apical third of sternum 7, and segments 8 and 9 yellow; mandibles, maxillary and labial palps, elytra, and legs dark brown; antennomeres 1–8 or 1–9 dark brown to black, 9–11 or 10–11 yellowish (
<figureCitation id="1AB74ED8FFCFFFEA92F2FE67FA87C4AF" box="[1209,1308,482,508]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="4" pageNumber="19">Figs 1A</figureCitation>
,
<figureCitation id="1AB74ED8FFCFFFEA9362FE67FAACC4AF" box="[1321,1335,482,508]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="4" pageNumber="19">2</figureCitation>
A–B, 3A).
</paragraph>
<paragraph id="8233525DFFCFFFEA96F6FDC2FE19C1AE" blockId="4.[187,1400,583,1278]" pageId="4" pageNumber="19">
HEAD. Subhexagonal and slightly transverse (male) to subquadrate (female) in shape, slightly broadened anteriad, narrower in female (
<figureCitation id="1AB74ED8FFCFFFEA9457FDEFFDE4C7D7" box="[540,639,618,644]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="4" pageNumber="19">Figs 2A</figureCitation>
,
<figureCitation id="1AB74ED8FFCFFFEA94C6FDEFFD29C7D7" box="[653,690,618,644]" captionStart="Fig" captionStartId="10.[189,232,1916,1942]" captionTargetBox="[202,1398,532,1882]" captionTargetId="figure-102@10.[189,1398,532,1883]" captionTargetPageId="10" captionText="Fig. 3. Ophionthus asenjoi sp. nov. Holotype, ♀ (IFML). A. Habitus, dorsal view. B. Original labels. C. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542901" httpUri="https://zenodo.org/record/4542901/files/figure.png" pageId="4" pageNumber="19">3A</figureCitation>
) than male (
<figureCitation id="1AB74ED8FFCFFFEA9503FDEFFC3DC7D7" box="[840,934,618,644]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="4" pageNumber="19">Fig. 1A</figureCitation>
); nuchal ridge present; postmandibular ridge absent; postgenal ridge present; ventral basal ridge present, extending more or less parallel to ventral portion of postoccipital suture; infraorbital ridge present or absent; dorsal and ventral surface of head with scarce setation, microsculpture dense and undulate. Eyes large, moderately convex, occupying apical half of lateral margin of head, shorter than temples seen from above. Gular sutures joined before neck. Frontoclypeus with a short longitudinal groove. Antennae inserted closer to anterior margin of frontoclypeus than to eyes, moderately long, not reaching base of pronotum, increasing gradually in width toward apex; antennomere 1 without apical spine-like seta, distinctly longer than antennomere 2 and almost as long as antennomeres 2 and 3 combined; antennomere 3 slightly longer than antennomere 2; pubescence starting on antennomere 4 to 11. Clypeus entire and fused with frons. Labrum rectangular, distinctly emarginate, sclerotized except apical margin, with several setae and macrosetae at apical margin. Mandible prominent, 1.4× length of head, with setose prostheca well-developed, with a groove on dorso-lateral margin. Maxilla with lacinia elongate and densely setose along internal apical half margin, with galea prominent and densely setose at apex; maxillary palpus long, palpomeres 1 to 4 glabrous with scarce setae at apices; palpomere 1 small; palpomere 2 slightly curved, longest, shorter than 3 and 4 combined; palpomere 4 cylindrical, longer than segment 3. Mentum transverse, with anterior margin slightly emarginate, one seta at each latero-lateral corner. Labial palpus moderately long; palpomere 1 shorter than 2; palpomere 2 shorter than 3; palpomere 3 cylindrical, shorter than 1 and 2 combined.
</paragraph>
<paragraph id="8233525DFFCFFFEA96F6FAA0FC5AC366" blockId="4.[187,1399,1317,1590]" pageId="4" pageNumber="19">
PRONOTUM. Distinctly longer than wide, slightly broadened anteriad, front margin subtruncate, hind margin slightly arcuate, anterior and posterior angles rounded (
<figureCitation id="1AB74ED8FFCFFFEA95EEFACDFB9CC031" box="[933,1031,1352,1378]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="4" pageNumber="19">Figs 1A</figureCitation>
,
<figureCitation id="1AB74ED8FFCFFFEA925FFACDFBA0C031" box="[1044,1083,1352,1378]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="4" pageNumber="19">2A</figureCitation>
,
<figureCitation id="1AB74ED8FFCFFFEA9203FACDFBF4C031" box="[1096,1135,1352,1378]" captionStart="Fig" captionStartId="10.[189,232,1916,1942]" captionTargetBox="[202,1398,532,1882]" captionTargetId="figure-102@10.[189,1398,532,1883]" captionTargetPageId="10" captionText="Fig. 3. Ophionthus asenjoi sp. nov. Holotype, ♀ (IFML). A. Habitus, dorsal view. B. Original labels. C. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542901" httpUri="https://zenodo.org/record/4542901/files/figure.png" pageId="4" pageNumber="19">3A</figureCitation>
); disk with two dorsal rows of punctures sub-parallel to each other; lateral puncture of pronotum bearing long macroseta separated from superior line of pronotal hypomeron by a distance more than three times as large as diameter of puncture; surface with fine and dense microsculpture. Prosternum without distinct midlongitudinal carina; basisternum longer than furcasternum, with transverse carina variable (either complete or rudimentary). Mesoventrite somewhat elongate, with sternopleural suture distinctly oblique; mesoventral intercoxal process rounded, forming an obtuse angle.
</paragraph>
<paragraph id="8233525DFFCFFFEA96F6F9D8FDA0C257" blockId="4.[189,1400,1629,1796]" pageId="4" pageNumber="19">
LEGS. Profemora slightly narrowed toward apex, with lateroventral medial spines; protibiae setose, excavated; protarsomeres 1–4 longer than wide, slightly flattened dorsoventrally and slightly widened apically, with long and curved discal setae (adhesive setae) underneath, without marginal setae; meso- and metatarsomeres 1–2 with strong and distinct marginal setae (
<figureCitation id="1AB74ED8FFCFFFEA95FCF943FB8FC3B3" box="[951,1044,1734,1760]" captionStart="Fig" captionStartId="10.[189,232,1916,1942]" captionTargetBox="[202,1398,532,1882]" captionTargetId="figure-102@10.[189,1398,532,1883]" captionTargetPageId="10" captionText="Fig. 3. Ophionthus asenjoi sp. nov. Holotype, ♀ (IFML). A. Habitus, dorsal view. B. Original labels. C. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542901" httpUri="https://zenodo.org/record/4542901/files/figure.png" pageId="4" pageNumber="19">Fig. 3C</figureCitation>
); metatarsomere 1 moderately longer than 5 (
<figureCitation id="1AB74ED8FFCFFFEA972CF96CFE52C257" box="[359,457,1769,1796]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="4" pageNumber="19">Figs 1A</figureCitation>
,
<figureCitation id="1AB74ED8FFCFFFEA979EF96FFE60C257" box="[469,507,1770,1796]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="4" pageNumber="19">2A</figureCitation>
,
<figureCitation id="1AB74ED8FFCFFFEA9443F96FFDB6C257" box="[520,557,1770,1796]" captionStart="Fig" captionStartId="10.[189,232,1916,1942]" captionTargetBox="[202,1398,532,1882]" captionTargetId="figure-102@10.[189,1398,532,1883]" captionTargetPageId="10" captionText="Fig. 3. Ophionthus asenjoi sp. nov. Holotype, ♀ (IFML). A. Habitus, dorsal view. B. Original labels. C. Metatarsomeres 1 and 2 with strong and distinct marginal setae (arrows). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542901" httpUri="https://zenodo.org/record/4542901/files/figure.png" pageId="4" pageNumber="19">3C</figureCitation>
).
</paragraph>
<paragraph id="8233525DFFCFFFEA96F6F8A9FAFCC23A" blockId="4.[189,1399,1835,1897]" pageId="4" pageNumber="19">ELYTRON. Longer than pronotum, subbasal ridge curved and projected anteriad, punctuation moderately coarse and homogeneously distributed. Hing wing with veins CuA and MP4 fused, vein MP3 present.</paragraph>
<paragraph id="8233525DFFCFFFEB96F6F815FEF6C439" blockId="4.[189,1399,1936,1997]" lastBlockId="5.[189,1398,265,362]" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">ABDOMEN. Tergum I with paired prototergal glands manifested by invaginated capsules with small openings. Terga III–V with anterior basal transverse carina distinct, posterior basal transverse carina always distinct on tergum III, distinct to not distinct on terga IV–V; area between anterior and posterior carinae deeply and sparsely punctate; surface with fine and dense microsculpture of transverse and oblique waves.</paragraph>
<paragraph id="8233525DFFCEFFEB96F6FE14FAD1C72D" blockId="5.[188,1401,401,639]" pageId="5" pageNumber="20">
MALE SEXUAL CHARACTERS. Tergum VIII with apex projected medially, forming an obtuse angle (
<figureCitation id="1AB74ED8FFCEFFEB9346FE14FAFCC4F8" box="[1293,1383,401,427]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="5" pageNumber="20">Fig. 1C</figureCitation>
); sternum VIII slightly emarginate apically (
<figureCitation id="1AB74ED8FFCEFFEB94E0FE31FC93C49D" box="[683,776,436,462]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="5" pageNumber="20">Fig. 1D</figureCitation>
). Genital segment with laterotergal sclerites IX short, not dorsoventrally flattened, with apex rounded and not sexually dimorphic (
<figureCitation id="1AB74ED8FFCEFFEB9224FE52FB48C4A2" box="[1135,1235,471,497]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="5" pageNumber="20">Fig. 2G</figureCitation>
); sternum IX symmetrical, deeply emarginate apically, with several fine and short setae at each side of emargination, basal portion moderately longer than distal portion (
<figureCitation id="1AB74ED8FFCEFFEB9565FD9BFC12C76B" box="[814,905,542,568]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="5" pageNumber="20">Fig. 1E</figureCitation>
); tergum X with apex rounded (
<figureCitation id="1AB74ED8FFCEFFEB9345FD9BFAF3C76B" box="[1294,1384,542,568]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="5" pageNumber="20">Fig. 1F</figureCitation>
). Aedeagus with parameres fused as a short, triangular sclerite, and completely fused to median lobe (
<figureCitation id="1AB74ED8FFCEFFEB968DFDE1FE9CC72D" box="[198,263,612,638]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="5" pageNumber="20">Fig. 1</figureCitation>
G–H); median lobe elongate with apical part narrowed into a rather broad apex (
<figureCitation id="1AB74ED8FFCEFFEB92F1FDE1FB60C72D" box="[1210,1275,612,638]" captionStart="Fig" captionStartId="6.[189,232,1884,1910]" captionTargetBox="[190,1392,293,1804]" captionTargetId="figure-14@6.[189,1398,293,1811]" captionTargetPageId="6" captionText="Fig. 1. Ophionthus serpentinus Bernhauer, 1908. Holotype, ♂ (FMNH). A. Habitus, dorsal view, after Chani-Posse (2014a). B. Original labels. C. Tergum VIII. D. Sternum VIII. E. Sternum IX. F. Tergum X. G. Aedeagus, lateral view. H. Aedeagus, dorsal view. Stippled areas in G represent membranes. Shadded areas in G and H represent internal sclerites Scale bars: A = 5 mm; C–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4542895" httpUri="https://zenodo.org/record/4542895/files/figure.png" pageId="5" pageNumber="20">Fig. 1</figureCitation>
G–H).
</paragraph>
<paragraph id="8233525DFFCEFFEB96F6FD23FAC5C655" blockId="5.[187,1399,677,774]" pageId="5" pageNumber="20">
FEMALE SEXUAL CHARACTERS. Tergum VIII with apex projected medially forming an obtuse angle (
<figureCitation id="1AB74ED8FFCEFFEB968DFD4CFEBBC7B0" box="[198,288,713,739]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="5" pageNumber="20">Fig. 2E</figureCitation>
); sternum VIII with apex rounded (
<figureCitation id="1AB74ED8FFCEFFEB94FAFD4CFC92C7B0" box="[689,777,713,739]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="5" pageNumber="20">Fig. 2F</figureCitation>
); tergum X subtruncate (
<figureCitation id="1AB74ED8FFCEFFEB9268FD4CFBE4C7B0" box="[1059,1151,713,739]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="5" pageNumber="20">Fig. 2G</figureCitation>
); second gonocoxites each with a long macroseta proximally, with minute stylus bearing a long apical macroseta (
<figureCitation id="1AB74ED8FFCEFFEB92B9FD69FACAC655" box="[1266,1361,748,774]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[199,1394,353,1755]" captionTargetId="figure-15@8.[189,1398,349,1768]" captionTargetPageId="8" captionText="Fig. 2. Ophionthus serpentinus Bernhauer, 1908. ♀ (MUSM). A. Habitus, dorsal view.B. Habitus, lateral view. C. Right protarsomeres, ventral view.D. Details of discal setae from right third protarsomere, ventral view. E. Tergum VIII. F. Sternum VIII. G. Lateral tergal sclerites IX and tergum X. H. Gonocoxites of female genital segment. Scale bars: A–B = 5 mm; G–H = 0.5 mm.." figureDoi="http://doi.org/10.5281/zenodo.4542899" httpUri="https://zenodo.org/record/4542899/files/figure.png" pageId="5" pageNumber="20">Fig. 2H</figureCitation>
).
</paragraph>
<paragraph id="8233525DFFCEFFEB96F6FCA8FE15C614" blockId="5.[189,398,813,839]" box="[189,398,813,839]" pageId="5" pageNumber="20">
<heading id="D97BE531FFCEFFEB96F6FCA8FE15C614" bold="true" box="[189,398,813,839]" fontSize="11" level="3" pageId="5" pageNumber="20" reason="3">
<emphasis id="B0F88E4FFFCEFFEB96F6FCA8FE15C614" bold="true" box="[189,398,813,839]" pageId="5" pageNumber="20">Immature stages</emphasis>
</heading>
</paragraph>
<paragraph id="8233525DFFCEFFEB96F6FCD9FEA2C625" blockId="5.[189,313,860,886]" box="[189,313,860,886]" pageId="5" pageNumber="20">Unknown.</paragraph>
<paragraph id="8233525DFFCEFFEB96F6FC18FD99C6E4" blockId="5.[189,514,925,951]" box="[189,514,925,951]" pageId="5" pageNumber="20">
<heading id="D97BE531FFCEFFEB96F6FC18FD99C6E4" bold="true" box="[189,514,925,951]" fontSize="11" level="3" pageId="5" pageNumber="20" reason="3">
<emphasis id="B0F88E4FFFCEFFEB96F6FC18FD99C6E4" bold="true" box="[189,514,925,951]" pageId="5" pageNumber="20">Geographical distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8233525DFFCEFFEB96F6FC49FD39C120" blockId="5.[189,1399,972,1139]" pageId="5" pageNumber="20">
The genus
<taxonomicName id="458C29DEFFCEFFEB970BFC49FE53C6B5" authorityName="Bernhauer" authorityYear="1908" box="[320,456,972,998]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B0F88E4FFFCEFFEB970BFC49FE53C6B5" box="[320,456,972,998]" italics="true" pageId="5" pageNumber="20">Ophionthus</emphasis>
</taxonomicName>
, with two species known at present, has been recorded from the departments of
<collectingRegion id="40489CBFFFCEFFEB96F5FC6AFEA4C15A" box="[190,319,1007,1033]" country="Peru" name="Cajamarca" pageId="5" pageNumber="20">Cajamarca</collectingRegion>
,
<collectingRegion id="40489CBFFFCEFFEB9700FC75FE41C15A" box="[331,474,1007,1034]" country="Peru" name="La Libertad" pageId="5" pageNumber="20">La Libertad</collectingRegion>
, and
<collectingRegion id="40489CBFFFCEFFEB9450FC6AFDC0C15A" box="[539,603,1007,1033]" country="Peru" name="Junin" pageId="5" pageNumber="20">Junín</collectingRegion>
in
<collectingCountry id="FA9B12CDFFCEFFEB94C8FC75FD26C159" box="[643,701,1008,1034]" name="Peru" pageId="5" pageNumber="20">Peru</collectingCountry>
. Based on the information of the specimen labels (locality and habitat),
<taxonomicName id="458C29DEFFCEFFEB971CFB97FE7BC17F" authorityName="Bernhauer" authorityYear="1908" box="[343,480,1042,1068]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B0F88E4FFFCEFFEB971CFB97FE7BC17F" box="[343,480,1042,1068]" italics="true" pageId="5" pageNumber="20">Ophionthus</emphasis>
</taxonomicName>
is distributed mainly in the ecoregion of Peruvian Yungas (
<bibRefCitation id="E61D2FACFFCEFFEB92D2FB97FAF1C17F" author="Olson D. M. &amp; Dinerstein E. &amp; Wikramanayake E. D. &amp; Burgess N. D. &amp; Powell G. V. &amp; Underwood E. C. &amp; D'Amico J. &amp; Itoua I. &amp; Strand H. E. &amp; Morrison J. C. &amp; Loucks C. J. &amp; Allnutt T. F. &amp; Ricketts T. H. &amp; Kura Y. &amp; Lamoreux J. F. &amp; Wettengel W. W. &amp; Hedaeo P. &amp; Kassen K. R." box="[1177,1386,1042,1069]" pageId="5" pageNumber="20" pagination="933 - 938" refId="ref8386" refString="Olson D. M., Dinerstein E., Wikramanayake E. D., Burgess N. D., Powell G. V., Underwood E. C., D'Amico J., Itoua I., Strand H. E., Morrison J. C., Loucks C. J., Allnutt T. F., Ricketts T. H., Kura Y., Lamoreux J. F., Wettengel W. W., Hedaeo P. &amp; Kassen K. R. 2001. Terrestrial ecoregions of the world: a new map of life on earth: a new global map of terrestrial ecoregions provides an innovative tool for conserving biodiversity. BioScience 51 (11): 933 - 938. https: // doi. org / 10.1641 / 0006 - 3568 (2001) 051 [0933: TEOTWA] 2.0. CO; 2" type="journal article" year="2001">
Olson
<emphasis id="B0F88E4FFFCEFFEB92A2FB96FA87C17F" box="[1257,1308,1042,1068]" italics="true" pageId="5" pageNumber="20">et al</emphasis>
. 2001
</bibRefCitation>
). However, according to Morrone´s regionalization (2014) those localities belong to the biogeographical province of Puna (
<figureCitation id="1AB74ED8FFCEFFEB97DEFBDDFE64C120" box="[405,511,1112,1139]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="11.[189,232,1972,1998]" captionStartId-1="12.[189,232,1937,1963]" captionTargetBox-0="[189,1398,1237,1937]" captionTargetBox-1="[189,1398,1169,1895]" captionTargetId-0="figure-257@11.[189,1399,1237,1937]" captionTargetId-1="figure-338@12.[189,1398,1168,1896]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Fig. 4. Distribution map of Ophionthus serpentinus Bernhauer, 1908 and O. asenjoi sp. nov." captionText-1="Fig. 5. Map of probable localities (correct spelling, see Supp. file 1) for the male holotype of Ophionthus serpentinus Bernhauer, 1908 collected in “Peruvia centralis” by Jelski in 1873." figureDoi-0="http://doi.org/10.5281/zenodo.4542903" figureDoi-1="http://doi.org/10.5281/zenodo.4542905" httpUri-0="https://zenodo.org/record/4542903/files/figure.png" httpUri-1="https://zenodo.org/record/4542905/files/figure.png" pageId="5" pageNumber="20">Figs 4–5</figureCitation>
, Supp. file 1).
</paragraph>
<paragraph id="8233525DFFCEFFEB96F6FB1FFEA4C1E7" blockId="5.[189,319,1178,1204]" box="[189,319,1178,1204]" pageId="5" pageNumber="20">
<heading id="D97BE531FFCEFFEB96F6FB1FFEA4C1E7" bold="true" box="[189,319,1178,1204]" fontSize="11" level="3" pageId="5" pageNumber="20" reason="3">
<emphasis id="B0F88E4FFFCEFFEB96F6FB1FFEA4C1E7" bold="true" box="[189,319,1178,1204]" pageId="5" pageNumber="20">Bionomics</emphasis>
</heading>
</paragraph>
<paragraph id="8233525DFFCEFFEB96F6FB4DFAA8C055" blockId="5.[189,1399,1224,1286]" pageId="5" pageNumber="20">
The species of
<taxonomicName id="458C29DEFFCEFFEB9726FB4DFE6DC1B1" authorityName="Bernhauer" authorityYear="1908" box="[365,502,1224,1250]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B0F88E4FFFCEFFEB9726FB4DFE6DC1B1" box="[365,502,1224,1250]" italics="true" pageId="5" pageNumber="20">Ophionthus</emphasis>
</taxonomicName>
are considered general predators as are the other genera of
<taxonomicName id="458C29DEFFCEFFEB92FBFB4DFAA0C1B1" authorityName="Kirby" authorityYear="1837" box="[1200,1339,1224,1250]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
. One specimen of
<taxonomicName id="458C29DEFFCEFFEB9718FB69FE65C055" authorityName="Bernhauer" authorityYear="1908" box="[339,510,1260,1286]" class="Insecta" family="Staphylinidae" genus="Ophionthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="serpentinus">
<emphasis id="B0F88E4FFFCEFFEB9718FB69FE65C055" box="[339,510,1260,1286]" italics="true" pageId="5" pageNumber="20">O. serpentinus</emphasis>
</taxonomicName>
was collected in a clearing in montane forest and riverside shrubbery.
</paragraph>
</subSubSection>
</treatment>
</document>
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