385 lines
57 KiB
XML
385 lines
57 KiB
XML
<document id="180DD30F36D6A39B66DC3A43053E9AFF" ID-DOI="10.5281/zenodo.181452" ID-GBIF-Dataset="0e6e9d41-cb9a-4ec3-a951-bf5f2eb06577" ID-ISSN="1175-5326" ID-Zenodo-Dep="181452" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459833984607" checkinUser="plazi" docAuthor="Naruse, Tohru, Ng, Peter K. L. & Guinot, Danièle" docDate="2008" docId="9C7287B0994BFFBDC195D5F9FC09FBB2" docLanguage="en" docName="zt01739p040.pdf" docOrigin="Zootaxa 1739" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Guaplax denticulata Naruse, Ng & Guinot, 2008, spec. nov." docType="treatment" docVersion="10" lastPageNumber="38" masterDocId="604BFFC89947FFACC102D56EFFB9FFEB" masterDocTitle="Two new genera and two new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on Cancrocaeca Ng, 1991" masterLastPageNumber="40" masterPageNumber="21" pageNumber="33" updateTime="1698223242837" updateUser="plazi">
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<mods:mods id="6175BC8E9171E7667929228E778320C7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F7D0C58E285F3BA375B2E037166242AE">
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<mods:title id="CB7E152D6FEAF01C0885F7214847A201">Two new genera and two new species of troglobitic false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae) from Indonesia, with notes on Cancrocaeca Ng, 1991</mods:title>
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</mods:titleInfo>
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<mods:name id="99271CA7CC85F0E67649794CF9BAE3B2" type="personal">
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<mods:role id="AE661DCA01B7FAD170032EEBD7F59BF9">
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<mods:roleTerm id="9DDA65A42ED372CDDBFB17FADBEC2E94">Author</mods:roleTerm>
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<mods:namePart id="833DF9153CA8552463E60C16D19444A6">Naruse, Tohru</mods:namePart>
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<mods:name id="3164D1021A80ABB2A5DC61747671D778" type="personal">
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<mods:roleTerm id="A2236DB216072BE6A2907BB4278CD62B">Author</mods:roleTerm>
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<mods:namePart id="4D18B4A0870061C6EA5286DD102DC440">Ng, Peter K. L.</mods:namePart>
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<mods:name id="F5E3DF0BBC3D971690AFDE934B04EAC0" type="personal">
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<mods:roleTerm id="F6C3C68DB7CF750F68C2D0DD688CD7DE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A18B567A010973E658F10F2BF572517C">Guinot, Danièle</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="755F5A62E63B06F25DD59A6C0CDC03AE">text</mods:typeOfResource>
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<mods:titleInfo id="BD5EE1A5673D01C06197B44C6285706D">
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<mods:title id="FE8AD376E5E76A67B6F77DE9F18CE089">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="794B0F6C9DAA4F5139187FD9DF713DA2">
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<mods:date id="9A265B111804895E1EDE0927E77F0C3D">2008</mods:date>
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<mods:detail id="82C1759866C936AE6F0C4DEB9C2222F7" type="volume">
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<mods:number id="3C6103644B388B46DE239529B1710F5A">1739</mods:number>
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<mods:classification id="57886EE06F7F22C0C8399914A9F354E3">journal article</mods:classification>
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<mods:identifier id="73820375760A45E9780AD286B7C30045" type="DOI">10.5281/zenodo.181452</mods:identifier>
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<mods:identifier id="0D60FEDFDD169892E0DAF7EF0C23D13A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2F816E2585875B72217849E1EFC7F963" type="Zenodo-Dep">181452</mods:identifier>
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<treatment id="9C7287B0994BFFBDC195D5F9FC09FBB2" ID-DOI="http://doi.org/10.5281/zenodo.6233828" ID-GBIF-Taxon="119358432" ID-Zenodo-Dep="6233828" LSID="urn:lsid:zoobank.org:act:CFB6634F-5F25-422B-992A-1AAC555A7426" httpUri="http://treatment.plazi.org/id/9C7287B0994BFFBDC195D5F9FC09FBB2" lastPageId="17" lastPageNumber="38" pageId="12" pageNumber="33">
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<subSubSection id="5CC1652D994BFFA0C195D5F9FEA1FF3F" pageId="12" pageNumber="33" type="nomenclature">
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<paragraph id="146436A6994BFFA0C195D5F9FDB5FF5A" blockId="12.[151,524,151,212]" box="[151,524,151,177]" pageId="12" pageNumber="33">
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<heading id="4F2C81CA994BFFA0C195D5F9FDB5FF5A" bold="true" box="[151,524,151,177]" fontSize="11" level="1" pageId="12" pageNumber="33" reason="1">
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<emphasis id="26AFEAB4994BFFA0C195D5F9FDB5FF5A" bold="true" box="[151,524,151,177]" pageId="12" pageNumber="33">
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<taxonomicName id="D3DB4D25994BFFA0C195D5F9FE34FF5A" ID-CoL="3HJ7R" box="[151,397,151,177]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
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<emphasis id="26AFEAB4994BFFA0C195D5F9FE34FF5A" bold="true" box="[151,397,151,177]" italics="true" pageId="12" pageNumber="33">Guaplax denticulata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3D9C57CF994BFFA0C096D5F9FDB5FF5A" box="[404,524,151,177]" pageId="12" pageNumber="33" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="146436A6994BFFA0C195D5D4FEA1FF3F" blockId="12.[151,524,151,212]" box="[151,280,186,212]" pageId="12" pageNumber="33">
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(
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<figureCitation id="8CE02A23994BFFA0C19DD5D4FEB6FF3F" box="[159,271,186,212]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="12.[151,255,1678,1702]" captionStartId-1="13.[151,255,1905,1929]" captionStartId-2="14.[151,255,1571,1595]" captionTargetBox-0="[156,1408,881,1640]" captionTargetBox-1="[167,1372,191,1865]" captionTargetBox-2="[160,1401,194,1531]" captionTargetId-0="figure@12.[151,1436,863,1651]" captionTargetId-1="figure@13.[164,1400,146,1866]" captionTargetId-2="figure@14.[151,1436,194,1547]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 7. Guaplax denticulata spec. nov.: a, habitus; b, carapace, frontal view. a, male holotype, MZB Cru 1654 (holotype), 2.7 × 3.1 mm; b, female paratype, MNHN-B 3061, 3.4 × 4.1 mm. Scale bars, 1 mm." captionText-1="FIGURE 8. Guaplax denticulata spec. nov.: a, right third maxilliped; b, male left chela; c, left P 3; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view; g, left G 2; h, G 2 on abdomen, inner view. a, female paratype, MNHN-B 3061, 3.4 × 4.1 mm; b – h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e – g, 0.5 mm; b – d, h, 1 mm." captionText-2="FIGURE 9. Guaplax denticulata spec. nov.: a, thoracic sternum of female; b – d, female abdomen-pleotelson, outer side (b and d) and inner surface (c); e – h, second to fifth left pleopods, anterior view. Female paratype, MNHN-B 3061, 3.4 × 4.1 mm. Numbers and “ V ” refer to thoracic sternites and vulvae, respectively. Scale bars, 1 mm." httpUri-0="https://zenodo.org/record/181459/files/figure.png" httpUri-1="https://zenodo.org/record/181460/files/figure.png" httpUri-2="https://zenodo.org/record/181461/files/figure.png" pageId="12" pageNumber="33">Figs. 7–9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="5CC1652D994BFFA0C195D464FDD9FE77" pageId="12" pageNumber="33" type="materials_examined">
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<paragraph id="146436A6994BFFA0C195D464FC62FEA7" blockId="12.[151,1437,266,812]" pageId="12" pageNumber="33">
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<emphasis id="26AFEAB4994BFFA0C195D464FDB2FECF" bold="true" box="[151,523,266,292]" pageId="12" pageNumber="33">
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Material examined.
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<typeStatus id="CB608804994BFFA0C090D464FDBFFECF" box="[402,518,266,292]" pageId="12" pageNumber="33" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male, 2.7 ×
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<quantity id="D3239B43994BFFA0C3A2D464FD44FECF" box="[672,765,266,292]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="12" pageNumber="33" unit="mm" value="3.1">3.1 mm</quantity>
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,
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<collectionCode id="72CAAE63994BFFA0C20AD464FCF1FECF" box="[776,840,266,292]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="12" pageNumber="33">MZB</collectionCode>
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Cru 1654, Gua Laowacling, Tabalar ulu, Kalimantan Timur,
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<collectingCountry id="6CCC7636994BFFA0C01AD45CFE34FEA7" box="[280,397,306,332]" name="Indonesia" pageId="12" pageNumber="33">Indonesia</collectingCountry>
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, coll. C. Rahmadi & L. Deharveng,
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<date id="60651066994BFFA0C236D45CFC6FFEA7" box="[820,982,306,332]" pageId="12" pageNumber="33" value="2004-08-11">11 Aug. 2004</date>
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.
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</paragraph>
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<paragraph id="146436A6994BFFA0C1C7D434FDD9FE77" blockId="12.[151,1437,266,812]" pageId="12" pageNumber="33">
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<emphasis id="26AFEAB4994BFFA0C1C7D434FEF3FE9F" bold="true" box="[197,330,346,372]" pageId="12" pageNumber="33">
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<typeStatus id="CB608804994BFFA0C1C7D434FEFDFE9F" box="[197,324,346,372]" pageId="12" pageNumber="33" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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One male, 2.9 ×
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<quantity id="D3239B43994BFFA0C322D434FD39FE9F" box="[544,640,346,372]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="33" unit="mm" value="3.4">3.4 mm</quantity>
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,
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<collectionCode id="72CAAE63994BFFA0C38FD434FD7FFE9F" box="[653,710,346,372]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="33">ZRC</collectionCode>
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2007.0120, data same as
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<typeStatus id="CB608804994BFFA0C502D434FBD3FE9F" box="[1024,1130,346,372]" pageId="12" pageNumber="33" type="holotype">holotype</typeStatus>
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;
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<specimenCount id="02DDFD2F994BFFA0C57AD434FB5AFE9F" box="[1144,1251,346,372]" pageId="12" pageNumber="33" type="female">1 female</specimenCount>
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, 3.4 ×
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<quantity id="D3239B43994BFFA0C43AD434FA21FE9F" box="[1336,1432,346,372]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="12" pageNumber="33" unit="mm" value="4.1">4.1 mm</quantity>
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, MNHN-B3061, data same as
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<typeStatus id="CB608804994BFFA0C0F0D4ECFDE5FE77" box="[498,604,386,412]" pageId="12" pageNumber="33" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5CC1652D994BFFA3C1C7D4C4FBA7FCD9" lastPageId="15" lastPageNumber="36" pageId="12" pageNumber="33" type="description">
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<paragraph id="146436A6994BFFA0C1C7D4C4FC13FCC7" blockId="12.[151,1437,266,812]" pageId="12" pageNumber="33">
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<emphasis id="26AFEAB4994BFFA0C1C7D4C4FEE4FE2F" bold="true" box="[197,349,426,452]" pageId="12" pageNumber="33">Description.</emphasis>
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Carapace (
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<figureCitation id="8CE02A23994BFFA0C0E3D4C4FD9FFE2F" box="[481,550,426,452]" captionStart="FIGURE 7" captionStartId="12.[151,255,1678,1702]" captionTargetBox="[156,1408,881,1640]" captionTargetId="figure@12.[151,1436,863,1651]" captionTargetPageId="12" captionText="FIGURE 7. Guaplax denticulata spec. nov.: a, habitus; b, carapace, frontal view. a, male holotype, MZB Cru 1654 (holotype), 2.7 × 3.1 mm; b, female paratype, MNHN-B 3061, 3.4 × 4.1 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181459/files/figure.png" pageId="12" pageNumber="33">Fig. 7</figureCitation>
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) subcircular, slightly wider than long, CW 1.17–1.20 times (n = 3) CL; dorsal surface flat, surrounded by continuous rim, regions clearly separated by grooves, cervical groove branching near distal end, branches confluent with rim; rostrum separated from carapace by rim, trilobate, median tooth slightly longer than, and placed inferior to, lateral teeth, teeth proximally fused; anterior margin of cephalothorax produced anteriorly in lateral view; anterolateral margin with 4 rounded teeth on dorsal rim, first to third teeth equidistantly placed, fourth tooth just behind third tooth, side wall below posterolateral margin gently swollen laterally, with two flat, triangular teeth. Antennular fossae very shallow, medially separated by thin, very low septum; a rounded tubercle on outwards of eye. Eye (
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<figureCitation id="8CE02A23994BFFA0C2BCD7ACFBBCFD37" box="[958,1029,706,732]" captionStart="FIGURE 7" captionStartId="12.[151,255,1678,1702]" captionTargetBox="[156,1408,881,1640]" captionTargetId="figure@12.[151,1436,863,1651]" captionTargetPageId="12" captionText="FIGURE 7. Guaplax denticulata spec. nov.: a, habitus; b, carapace, frontal view. a, male holotype, MZB Cru 1654 (holotype), 2.7 × 3.1 mm; b, female paratype, MNHN-B 3061, 3.4 × 4.1 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181459/files/figure.png" pageId="12" pageNumber="33">Fig. 7</figureCitation>
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) slightly visible from dorsal view, base of ocular peduncle fused to orbit, immovable, with conical tip, cornea without trace of black pigment. Epistome with posterior margin straight, cristate, slightly upturned.
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</paragraph>
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<caption id="40A4662E994BFFA0C195D3E0FB37F923" httpUri="https://zenodo.org/record/181459/files/figure.png" pageId="12" pageNumber="33" targetBox="[156,1408,881,1640]" targetPageId="12">
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<paragraph id="146436A6994BFFA0C195D3E0FB37F923" blockId="12.[151,1436,1678,1736]" pageId="12" pageNumber="33">
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<emphasis id="26AFEAB4994BFFA0C195D3E0FEA5F94D" bold="true" box="[151,284,1678,1702]" pageId="12" pageNumber="33">FIGURE 7.</emphasis>
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<taxonomicName id="D3DB4D25994BFFA0C024D3E1FDBAF94D" box="[294,515,1679,1702]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
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<emphasis id="26AFEAB4994BFFA0C024D3E1FDBAF94D" box="[294,515,1679,1702]" italics="true" pageId="12" pageNumber="33">Guaplax denticulata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3D9C57CF994BFFA0C30CD3E0FDC7F94D" box="[526,638,1678,1702]" pageId="12" pageNumber="33" rank="species">
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<emphasis id="26AFEAB4994BFFA0C30CD3E0FDCEF94D" bold="true" box="[526,631,1678,1702]" pageId="12" pageNumber="33">spec. nov</emphasis>
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.
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</taxonomicNameLabel>
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: a, habitus; b, carapace, frontal view. a, male holotype, MZB Cru 1654 (holotype), 2.7 × 3.1 mm; b, female paratype, MNHN-B3061, 3.4 × 4.1 mm. Scale bars, 1 mm.
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</paragraph>
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</caption>
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<paragraph id="146436A6994BFFA0C1C7D392FCB7F865" blockId="12.[151,1436,1788,1934]" pageId="12" pageNumber="33">
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Third maxillipeds (
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<figureCitation id="8CE02A23994BFFA0C0AFD392FE4FF8FD" box="[429,502,1788,1814]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[167,1372,191,1865]" captionTargetId="figure@13.[164,1400,146,1866]" captionTargetPageId="13" captionText="FIGURE 8. Guaplax denticulata spec. nov.: a, right third maxilliped; b, male left chela; c, left P 3; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view; g, left G 2; h, G 2 on abdomen, inner view. a, female paratype, MNHN-B 3061, 3.4 × 4.1 mm; b – h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e – g, 0.5 mm; b – d, h, 1 mm." httpUri="https://zenodo.org/record/181460/files/figure.png" pageId="12" pageNumber="33">Fig. 8</figureCitation>
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a) narrow, forming wide gape between endopods; exopod long, reaching just proximal to distal end of merus, with cylindrical flagellum; ischium short, with wide distal inner extension; merus ovoid, midlength about 1.5 times length of ischium; dactylus slender, slightly longer than propodus, distal end reaching distal inner part of ischium
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<emphasis id="26AFEAB4994BFFA0C3BCD21AFCB1F866" box="[702,776,1908,1933]" italics="true" pageId="12" pageNumber="33">in situ</emphasis>
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.
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</paragraph>
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<caption id="40A4662E994AFFA1C195D21FFAD3F826" httpUri="https://zenodo.org/record/181460/files/figure.png" pageId="13" pageNumber="34" targetBox="[167,1372,191,1865]" targetPageId="13">
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||
<paragraph id="146436A6994AFFA1C195D21FFAD3F826" blockId="13.[151,1436,1905,1997]" pageId="13" pageNumber="34">
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<emphasis id="26AFEAB4994AFFA1C195D21FFEA1F862" bold="true" box="[151,280,1905,1929]" pageId="13" pageNumber="34">FIGURE 8.</emphasis>
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<taxonomicName id="D3DB4D25994AFFA1C01DD21FFE40F863" box="[287,505,1905,1928]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
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<emphasis id="26AFEAB4994AFFA1C01DD21FFE40F863" box="[287,505,1905,1928]" italics="true" pageId="13" pageNumber="34">Guaplax denticulata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3D9C57CF994AFFA1C0FDD21FFDD5F863" box="[511,620,1905,1929]" pageId="13" pageNumber="34" rank="species">
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||
<emphasis id="26AFEAB4994AFFA1C0FDD21FFDDCF862" bold="true" box="[511,613,1905,1929]" pageId="13" pageNumber="34">spec. nov</emphasis>
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.
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</taxonomicNameLabel>
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: a, right third maxilliped; b, male left chela; c, left P3; d, male abdomen-pleotelson; e, left G1, ventral view; f, left G1, dorsal view; g, left G2; h, G2 on abdomen, inner view. a, female paratype, MNHN-B3061, 3.4 × 4.1 mm; b–h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e–g, 0.5 mm; b–d, h, 1 mm.
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</paragraph>
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</caption>
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<caption id="40A4662E9949FFA2C195D34DFB23F96B" httpUri="https://zenodo.org/record/181461/files/figure.png" pageId="14" pageNumber="35" targetBox="[160,1401,194,1531]" targetPageId="14">
|
||
<paragraph id="146436A69949FFA2C195D34DFB23F96B" blockId="14.[151,1436,1571,1664]" pageId="14" pageNumber="35">
|
||
<emphasis id="26AFEAB49949FFA2C195D34DFEA0F9D0" bold="true" box="[151,281,1571,1595]" pageId="14" pageNumber="35">FIGURE 9.</emphasis>
|
||
<taxonomicName id="D3DB4D259949FFA2C023D34DFE42F9D1" box="[289,507,1571,1594]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
|
||
<emphasis id="26AFEAB49949FFA2C023D34DFE42F9D1" box="[289,507,1571,1594]" italics="true" pageId="14" pageNumber="35">Guaplax denticulata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="26AFEAB49949FFA2C301D34DFDC8F9D0" bold="true" box="[515,625,1571,1595]" pageId="14" pageNumber="35">
|
||
<taxonomicNameLabel id="3D9C57CF9949FFA2C301D34DFDC8F9D0" box="[515,625,1571,1595]" pageId="14" pageNumber="35" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: a, thoracic sternum of female; b–d, female abdomen-pleotelson, outer side (b and d) and inner surface (c); e–h, second to fifth left pleopods, anterior view. Female paratype, MNHN-B3061, 3.4 × 4.1 mm. Numbers and “V” refer to thoracic sternites and vulvae, respectively. Scale bars, 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="146436A69949FFA2C1C7D3DDFBC9F87E" blockId="14.[151,1437,1715,2021]" pageId="14" pageNumber="35">
|
||
Female with circular vulvae (
|
||
<figureCitation id="8CE02A239949FFA2C322D3DDFDD1F926" box="[544,616,1715,1741]" captionStart="FIGURE 9" captionStartId="14.[151,255,1571,1595]" captionTargetBox="[160,1401,194,1531]" captionTargetId="figure@14.[151,1436,194,1547]" captionTargetPageId="14" captionText="FIGURE 9. Guaplax denticulata spec. nov.: a, thoracic sternum of female; b – d, female abdomen-pleotelson, outer side (b and d) and inner surface (c); e – h, second to fifth left pleopods, anterior view. Female paratype, MNHN-B 3061, 3.4 × 4.1 mm. Numbers and “ V ” refer to thoracic sternites and vulvae, respectively. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181461/files/figure.png" pageId="14" pageNumber="35">Fig. 9</figureCitation>
|
||
a), vulvae not raised, located on imaginary line joining posterior ends of P2 coxae on medial fused plate of thoracic sternum. Chelipeds symmetrical in both males and females, slightly larger in males; merus curving inwards, with flat ventral surface, ventral margins rounded, ventral outer margin without tooth; carpus slightly shorter than dorsal length of palm, outer surface rounded, without inner angle; palm of chela (
|
||
<figureCitation id="8CE02A239949FFA2C0D5D23DFDA5F886" box="[471,540,1875,1901]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[167,1372,191,1865]" captionTargetId="figure@13.[164,1400,146,1866]" captionTargetPageId="13" captionText="FIGURE 8. Guaplax denticulata spec. nov.: a, right third maxilliped; b, male left chela; c, left P 3; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view; g, left G 2; h, G 2 on abdomen, inner view. a, female paratype, MNHN-B 3061, 3.4 × 4.1 mm; b – h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e – g, 0.5 mm; b – d, h, 1 mm." httpUri="https://zenodo.org/record/181460/files/figure.png" pageId="14" pageNumber="35">Fig. 8</figureCitation>
|
||
b) with swollen outer surface, as long as dactylus; fingers straight, distal part slightly incurving backwards, with crossing acute tips, cutting edges with tiny teeth.
|
||
</paragraph>
|
||
<paragraph id="146436A69949FFA3C1C7D2CDFB81FEE9" blockId="14.[151,1437,1715,2021]" lastBlockId="15.[151,1437,152,1738]" lastPageId="15" lastPageNumber="36" pageId="14" pageNumber="35">
|
||
Ambulatory legs (
|
||
<figureCitation id="8CE02A239949FFA2C09AD2CDFE50F856" box="[408,489,1955,1981]" captionStart="FIGURE 7" captionStartId="12.[151,255,1678,1702]" captionTargetBox="[156,1408,881,1640]" captionTargetId="figure@12.[151,1436,863,1651]" captionTargetPageId="12" captionText="FIGURE 7. Guaplax denticulata spec. nov.: a, habitus; b, carapace, frontal view. a, male holotype, MZB Cru 1654 (holotype), 2.7 × 3.1 mm; b, female paratype, MNHN-B 3061, 3.4 × 4.1 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181459/files/figure.png" pageId="14" pageNumber="35">Figs. 7</figureCitation>
|
||
a, 8c) thin, moderately long; P3 and P4 subequal in length, longer than P2, P5 not clearly shorter than others; meri with elliptical cross-section, with constant diameter throughout length, P3 and P4 meri shorter than carapace length and width of dorsal rim, with rounded thin projection on distal anterior angle; propodi shorter than respective meri; dactyli clearly shorter than propodi, slightly incurved, inner margin lined with short setae, ending in sharp spine, without subterminal tooth.
|
||
</paragraph>
|
||
<paragraph id="146436A69948FFA3C1C7D47EFAA0FD81" blockId="15.[151,1437,152,1738]" pageId="15" pageNumber="36">
|
||
Male abdomen-pleotelson (
|
||
<figureCitation id="8CE02A239948FFA3C308D47EFDE8FEC1" box="[522,593,272,298]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[167,1372,191,1865]" captionTargetId="figure@13.[164,1400,146,1866]" captionTargetPageId="13" captionText="FIGURE 8. Guaplax denticulata spec. nov.: a, right third maxilliped; b, male left chela; c, left P 3; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view; g, left G 2; h, G 2 on abdomen, inner view. a, female paratype, MNHN-B 3061, 3.4 × 4.1 mm; b – h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e – g, 0.5 mm; b – d, h, 1 mm." httpUri="https://zenodo.org/record/181460/files/figure.png" pageId="15" pageNumber="36">Fig. 8</figureCitation>
|
||
d) 4 segmented, third to fifth abdominal segments fused; first segment with proximal part placed below posterior part of carapace, disto-lateral angles forming cylindrical lobe, lobe extending beyond distal margin of median part of first segment; second segment short, narrower than first and fused segments; fused segments with narrow proximal margins, slightly wider than distal margins, lateral margins of proximal half convex, distal half narrowed, slightly concave; pleotelson tong-shaped, with convex proximal margin distinctly narrower than half width of abdomen. G1 (
|
||
<figureCitation id="8CE02A239948FFA3C2C8D4B6FBA9FE19" box="[970,1040,472,498]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[167,1372,191,1865]" captionTargetId="figure@13.[164,1400,146,1866]" captionTargetPageId="13" captionText="FIGURE 8. Guaplax denticulata spec. nov.: a, right third maxilliped; b, male left chela; c, left P 3; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view; g, left G 2; h, G 2 on abdomen, inner view. a, female paratype, MNHN-B 3061, 3.4 × 4.1 mm; b – h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e – g, 0.5 mm; b – d, h, 1 mm." httpUri="https://zenodo.org/record/181460/files/figure.png" pageId="15" pageNumber="36">Fig. 8</figureCitation>
|
||
e, f) almost straight, tapering simply, dorsal side of inner margin of distal part lined with long, stiff setae; suture running from dorsal side of proximal part to outer margin of distal part. G2 (
|
||
<figureCitation id="8CE02A239948FFA3C3E4D746FC97FDA9" box="[742,814,552,578]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[167,1372,191,1865]" captionTargetId="figure@13.[164,1400,146,1866]" captionTargetPageId="13" captionText="FIGURE 8. Guaplax denticulata spec. nov.: a, right third maxilliped; b, male left chela; c, left P 3; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view; g, left G 2; h, G 2 on abdomen, inner view. a, female paratype, MNHN-B 3061, 3.4 × 4.1 mm; b – h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e – g, 0.5 mm; b – d, h, 1 mm." httpUri="https://zenodo.org/record/181460/files/figure.png" pageId="15" pageNumber="36">
|
||
Fig.
|
||
<quantity id="D3239B439948FFA3C222D746FC97FDA9" box="[800,814,552,578]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="15" pageNumber="36" unit="g" value="8.0">8</quantity>
|
||
</figureCitation>
|
||
g) short, with dilated base, proximal outer part protruded from lateral margin of second abdominal segment, partially visible in dorsal view (
|
||
<figureCitation id="8CE02A239948FFA3C5B6D73EFB40FD81" box="[1204,1273,592,618]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[167,1372,191,1865]" captionTargetId="figure@13.[164,1400,146,1866]" captionTargetPageId="13" captionText="FIGURE 8. Guaplax denticulata spec. nov.: a, right third maxilliped; b, male left chela; c, left P 3; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view; g, left G 2; h, G 2 on abdomen, inner view. a, female paratype, MNHN-B 3061, 3.4 × 4.1 mm; b – h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e – g, 0.5 mm; b – d, h, 1 mm." httpUri="https://zenodo.org/record/181460/files/figure.png" pageId="15" pageNumber="36">Fig. 8</figureCitation>
|
||
h).
|
||
</paragraph>
|
||
<paragraph id="146436A69948FFA3C1C7D716FBA7FCD9" blockId="15.[151,1437,152,1738]" pageId="15" pageNumber="36">
|
||
Female abdomen-pleotelson (
|
||
<figureCitation id="8CE02A239948FFA3C325D716FDC9FD79" box="[551,624,632,658]" captionStart="FIGURE 9" captionStartId="14.[151,255,1571,1595]" captionTargetBox="[160,1401,194,1531]" captionTargetId="figure@14.[151,1436,194,1547]" captionTargetPageId="14" captionText="FIGURE 9. Guaplax denticulata spec. nov.: a, thoracic sternum of female; b – d, female abdomen-pleotelson, outer side (b and d) and inner surface (c); e – h, second to fifth left pleopods, anterior view. Female paratype, MNHN-B 3061, 3.4 × 4.1 mm. Numbers and “ V ” refer to thoracic sternites and vulvae, respectively. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181461/files/figure.png" pageId="15" pageNumber="36">Fig. 9</figureCitation>
|
||
d) rounded, second segment to pleotelson fused, first segment with proximal part placed below posterior margin of carapace, disto-lateral angles forming rounded teeth; second segment short, boundaries between second and third and between fifth and pleotelson demarcated only by shallow sutures externally; inner surface of abdomen covered with thin membrane, forming pouch; pleopods present on second to fifth segments, biramous, short, contained inside pouch.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5CC1652D9948FFA3C1C7D651FDDAFC69" pageId="15" pageNumber="36" type="etymology">
|
||
<paragraph id="146436A69948FFA3C1C7D651FDDAFC69" blockId="15.[151,1437,152,1738]" pageId="15" pageNumber="36">
|
||
<emphasis id="26AFEAB49948FFA3C1C7D651FEEBFCB2" bold="true" box="[197,338,831,857]" pageId="15" pageNumber="36">Etymology.</emphasis>
|
||
From the Latin
|
||
<emphasis id="26AFEAB49948FFA3C310D62EFD18FCB2" box="[530,673,832,857]" italics="true" pageId="15" pageNumber="36">denticulatus</emphasis>
|
||
meaning “bearing small teeth”, alluding to the teeth on the carapace. The name is used as an adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5CC1652D9948FFBCC1C7D6E1FB3EFF53" lastPageId="16" lastPageNumber="37" pageId="15" pageNumber="36" type="discussion">
|
||
<paragraph id="146436A69948FFA3C1C7D6E1FBCDFB01" blockId="15.[151,1437,152,1738]" pageId="15" pageNumber="36">
|
||
<emphasis id="26AFEAB49948FFA3C1C7D6E1FE87FC42" bold="true" box="[197,318,911,937]" pageId="15" pageNumber="36">Remarks.</emphasis>
|
||
Apart from the generic characters separating
|
||
<taxonomicName id="D3DB4D259948FFA3C275D6FEFC65FC42" box="[887,988,912,937]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49948FFA3C275D6FEFC65FC42" box="[887,988,912,937]" italics="true" pageId="15" pageNumber="36">Guaplax</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="D3DB4D259948FFA3C52FD6FEFB2AFC42" box="[1069,1171,912,937]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26AFEAB49948FFA3C52FD6FEFB2AFC42" box="[1069,1171,912,937]" italics="true" pageId="15" pageNumber="36">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D3DB4D259948FFA3C5A5D6FEFA25FC42" box="[1191,1436,912,937]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
|
||
<emphasis id="26AFEAB49948FFA3C5A5D6FEFA25FC42" box="[1191,1436,912,937]" italics="true" pageId="15" pageNumber="36">Guaplax denticulata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="26AFEAB49948FFA3C195D6D9FEB6FC3A" bold="true" box="[151,271,951,977]" pageId="15" pageNumber="36">
|
||
<taxonomicNameLabel id="3D9C57CF9948FFA3C195D6D9FEB6FC3A" box="[151,271,951,977]" pageId="15" pageNumber="36" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be immediately differentiated from
|
||
<taxonomicName id="D3DB4D259948FFA3C3E8D6D6FCE1FC3A" box="[746,856,952,977]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="ensifer" status="sp. nov.">
|
||
<emphasis id="26AFEAB49948FFA3C3E8D6D6FCE1FC3A" box="[746,856,952,977]" italics="true" pageId="15" pageNumber="36">S. ensifer</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="26AFEAB49948FFA3C262D6D9FC61FC3A" bold="true" box="[864,984,951,977]" pageId="15" pageNumber="36">
|
||
<taxonomicNameLabel id="3D9C57CF9948FFA3C262D6D9FC61FC3A" box="[864,984,951,977]" pageId="15" pageNumber="36" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by possessing moderately long ambulatory legs (
|
||
<figureCitation id="8CE02A239948FFA3C020D68EFECCFC11" box="[290,373,992,1018]" captionStart="FIGURE 7" captionStartId="12.[151,255,1678,1702]" captionTargetBox="[156,1408,881,1640]" captionTargetId="figure@12.[151,1436,863,1651]" captionTargetPageId="12" captionText="FIGURE 7. Guaplax denticulata spec. nov.: a, habitus; b, carapace, frontal view. a, male holotype, MZB Cru 1654 (holotype), 2.7 × 3.1 mm; b, female paratype, MNHN-B 3061, 3.4 × 4.1 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181459/files/figure.png" pageId="15" pageNumber="36">Figs. 7</figureCitation>
|
||
a, 8c) (relatively much longer in
|
||
<taxonomicName id="D3DB4D259948FFA3C3F4D68EFCDBFC12" box="[758,866,992,1017]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="ensifer">
|
||
<emphasis id="26AFEAB49948FFA3C3F4D68EFCDBFC12" box="[758,866,992,1017]" italics="true" pageId="15" pageNumber="36">S. ensifer</emphasis>
|
||
</taxonomicName>
|
||
, fig. 3, 5a) and the presence of anterolateral and posterolateral teeth on the carapace (
|
||
<figureCitation id="8CE02A239948FFA3C34ED166FD2DFBC9" box="[588,660,1032,1058]" captionStart="FIGURE 7" captionStartId="12.[151,255,1678,1702]" captionTargetBox="[156,1408,881,1640]" captionTargetId="figure@12.[151,1436,863,1651]" captionTargetPageId="12" captionText="FIGURE 7. Guaplax denticulata spec. nov.: a, habitus; b, carapace, frontal view. a, male holotype, MZB Cru 1654 (holotype), 2.7 × 3.1 mm; b, female paratype, MNHN-B 3061, 3.4 × 4.1 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181459/files/figure.png" pageId="15" pageNumber="36">Fig. 7</figureCitation>
|
||
a, b) (absent in
|
||
<taxonomicName id="D3DB4D259948FFA3C252D166FC07FBCA" box="[848,958,1032,1057]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="ensifer">
|
||
<emphasis id="26AFEAB49948FFA3C252D166FC07FBCA" box="[848,958,1032,1057]" italics="true" pageId="15" pageNumber="36">S. ensifer</emphasis>
|
||
</taxonomicName>
|
||
, fig. 4a, b).
|
||
<taxonomicName id="D3DB4D259948FFA3C556D166FAFCFBCA" box="[1108,1349,1032,1057]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="denticulata">
|
||
<emphasis id="26AFEAB49948FFA3C556D166FAFCFBCA" box="[1108,1349,1032,1057]" italics="true" pageId="15" pageNumber="36">Guaplax denticulata</emphasis>
|
||
</taxonomicName>
|
||
can be also distinguished from
|
||
<taxonomicName id="D3DB4D259948FFA3C0B5D15EFD9EFBA2" box="[439,551,1072,1097]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="ensifer">
|
||
<emphasis id="26AFEAB49948FFA3C0B5D15EFD9EFBA2" box="[439,551,1072,1097]" italics="true" pageId="15" pageNumber="36">S. ensifer</emphasis>
|
||
</taxonomicName>
|
||
by the relatively shorter fingers of the chelae, with the immovable finger slightly longer than the length of the palm (
|
||
<figureCitation id="8CE02A239948FFA3C39CD136FD5DFB99" box="[670,740,1112,1138]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[167,1372,191,1865]" captionTargetId="figure@13.[164,1400,146,1866]" captionTargetPageId="13" captionText="FIGURE 8. Guaplax denticulata spec. nov.: a, right third maxilliped; b, male left chela; c, left P 3; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view; g, left G 2; h, G 2 on abdomen, inner view. a, female paratype, MNHN-B 3061, 3.4 × 4.1 mm; b – h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e – g, 0.5 mm; b – d, h, 1 mm." httpUri="https://zenodo.org/record/181460/files/figure.png" pageId="15" pageNumber="36">Fig. 8</figureCitation>
|
||
b) (clearly longer in
|
||
<taxonomicName id="D3DB4D259948FFA3C2D8D136FBF1FB9A" box="[986,1096,1112,1137]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="ensifer">
|
||
<emphasis id="26AFEAB49948FFA3C2D8D136FBF1FB9A" box="[986,1096,1112,1137]" italics="true" pageId="15" pageNumber="36">S. ensifer</emphasis>
|
||
</taxonomicName>
|
||
, fig. 5b, c), the absence of a subdistal tooth on the ventral outer margin of the cheliped merus (present in
|
||
<taxonomicName id="D3DB4D259948FFA3C513D1EEFBC7FB72" box="[1041,1150,1152,1177]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="ensifer">
|
||
<emphasis id="26AFEAB49948FFA3C513D1EEFBC7FB72" box="[1041,1150,1152,1177]" italics="true" pageId="15" pageNumber="36">S. ensifer</emphasis>
|
||
</taxonomicName>
|
||
), a long male pleotelson having convex proximal margin (
|
||
<figureCitation id="8CE02A239948FFA3C323D1C6FDDEFB29" box="[545,615,1192,1218]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[167,1372,191,1865]" captionTargetId="figure@13.[164,1400,146,1866]" captionTargetPageId="13" captionText="FIGURE 8. Guaplax denticulata spec. nov.: a, right third maxilliped; b, male left chela; c, left P 3; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view; g, left G 2; h, G 2 on abdomen, inner view. a, female paratype, MNHN-B 3061, 3.4 × 4.1 mm; b – h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e – g, 0.5 mm; b – d, h, 1 mm." httpUri="https://zenodo.org/record/181460/files/figure.png" pageId="15" pageNumber="36">Fig. 8</figureCitation>
|
||
d) (relatively shorter, with straight proximal margin in
|
||
<taxonomicName id="D3DB4D259948FFA3C5F3D1C6FAE4FB2A" box="[1265,1373,1192,1217]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="ensifer">
|
||
<emphasis id="26AFEAB49948FFA3C5F3D1C6FAE4FB2A" box="[1265,1373,1192,1217]" italics="true" pageId="15" pageNumber="36">S. ensifer</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="8CE02A239948FFA3C46FD1C6FF1FFB01" captionStart="FIGURE 5" captionStartId="8.[151,255,1927,1951]" captionTargetBox="[179,1354,202,1883]" captionTargetId="figure@8.[151,1436,194,2013]" captionTargetPageId="8" captionText="FIGURE 5. Sulaplax ensifer spec. nov.: a, left P 3; b, male left chela; c, female left chela; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view. a, c, female paratype, ZRC 2007.0119, 5.0 × 6.5 mm; b, d – f, male holotype," httpUri="https://zenodo.org/record/181457/files/figure.png" pageId="15" pageNumber="36">Fig. 5</figureCitation>
|
||
d), and the straight G1 (
|
||
<figureCitation id="8CE02A239948FFA3C0BED1BEFDBBFB01" box="[444,514,1232,1258]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[167,1372,191,1865]" captionTargetId="figure@13.[164,1400,146,1866]" captionTargetPageId="13" captionText="FIGURE 8. Guaplax denticulata spec. nov.: a, right third maxilliped; b, male left chela; c, left P 3; d, male abdomen-pleotelson; e, left G 1, ventral view; f, left G 1, dorsal view; g, left G 2; h, G 2 on abdomen, inner view. a, female paratype, MNHN-B 3061, 3.4 × 4.1 mm; b – h, male, ZRC 2007.0120, 2.9 × 3.4 mm. Scale bars, a, e – g, 0.5 mm; b – d, h, 1 mm." httpUri="https://zenodo.org/record/181460/files/figure.png" pageId="15" pageNumber="36">Fig. 8</figureCitation>
|
||
e, f) (curving outwards at half in
|
||
<taxonomicName id="D3DB4D259948FFA3C287D1BEFC48FB02" box="[901,1009,1232,1257]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="ensifer">
|
||
<emphasis id="26AFEAB49948FFA3C287D1BEFC48FB02" box="[901,1009,1232,1257]" italics="true" pageId="15" pageNumber="36">S. ensifer</emphasis>
|
||
</taxonomicName>
|
||
, fig. 5e, f).
|
||
</paragraph>
|
||
<paragraph id="146436A69948FFA3C1C7D196FCD8F921" blockId="15.[151,1437,152,1738]" pageId="15" pageNumber="36">
|
||
The only female specimen of
|
||
<taxonomicName id="D3DB4D259948FFA3C324D196FD73FAFA" box="[550,714,1272,1297]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="denticulata">
|
||
<emphasis id="26AFEAB49948FFA3C324D196FD73FAFA" box="[550,714,1272,1297]" italics="true" pageId="15" pageNumber="36">G. denticulata</emphasis>
|
||
</taxonomicName>
|
||
(MNHN-B30397, 3.4 ×
|
||
<quantity id="D3239B439948FFA3C2F3D196FBF4FAF9" box="[1009,1101,1272,1298]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="15" pageNumber="36" unit="mm" value="4.1">4.1 mm</quantity>
|
||
) has a torn thoracic sternum (
|
||
<figureCitation id="8CE02A239948FFA3C19DD04EFF51FAD1" box="[159,232,1312,1338]" captionStart="FIGURE 9" captionStartId="14.[151,255,1571,1595]" captionTargetBox="[160,1401,194,1531]" captionTargetId="figure@14.[151,1436,194,1547]" captionTargetPageId="14" captionText="FIGURE 9. Guaplax denticulata spec. nov.: a, thoracic sternum of female; b – d, female abdomen-pleotelson, outer side (b and d) and inner surface (c); e – h, second to fifth left pleopods, anterior view. Female paratype, MNHN-B 3061, 3.4 × 4.1 mm. Numbers and “ V ” refer to thoracic sternites and vulvae, respectively. Scale bars, 1 mm." httpUri="https://zenodo.org/record/181461/files/figure.png" pageId="15" pageNumber="36">Fig. 9</figureCitation>
|
||
a).
|
||
<bibRefCitation id="704A4B579948FFA3C00CD04EFD6AFAD1" author="Ng" box="[270,723,1312,1338]" pageId="15" pageNumber="40" refString="Ng, P. K. L. & Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement 3, 1 - 82." type="book chapter" year="1996">Ng & Chuang (1996: 60, figs. 25, 26)</bibRefCitation>
|
||
observed that the zoeae of
|
||
<taxonomicName id="D3DB4D259948FFA3C51FD04EFA8EFAD1" authority="Ng, 1988" authorityName="Ng" authorityYear="1988" box="[1053,1335,1312,1338]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="mangalis">
|
||
<emphasis id="26AFEAB49948FFA3C51FD04EFB15FAD2" box="[1053,1196,1312,1337]" italics="true" pageId="15" pageNumber="36">N. mangalis</emphasis>
|
||
(Ng, 1988)
|
||
</taxonomicName>
|
||
hatched inside the internal brood cavity and left their mother from a pair of “pseudovulvae” at thinly chitinised posterior part of the thoracic sternal cuticle. The true vulvae are too small for zoeae to come out.
|
||
<bibRefCitation id="704A4B579948FFA3C403D01EFF5CFA59" pageId="15" pageNumber="40" refString="Naruse, T., Shokita, S. & Kawahara, T. (2005) Neorhynchoplax yaeyamaensis, a new false spider crab (Decapoda: Brachyura: Hymenosomatidae) from the Yaeyama Group, the Ryukyu Islands, Japan. Zootaxa, 877, 1 - 7." type="journal article">
|
||
Naruse
|
||
<emphasis id="26AFEAB49948FFA3C45CD01EFA22FA62" box="[1374,1435,1392,1417]" italics="true" pageId="15" pageNumber="36">et al.</emphasis>
|
||
(2005)
|
||
</bibRefCitation>
|
||
also reported that the zoeae of
|
||
<taxonomicName id="D3DB4D259948FFA3C36AD0F6FB53FA59" authority="Naruse, Shokita & Kawahara, 2005" authorityName="Naruse, Shokita & Kawahara" authorityYear="2005" box="[616,1258,1432,1458]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="yaeyamaensis">
|
||
<emphasis id="26AFEAB49948FFA3C36AD0F6FC88FA5A" box="[616,817,1432,1457]" italics="true" pageId="15" pageNumber="36">N. yaeyamaensis</emphasis>
|
||
<bibRefCitation id="704A4B579948FFA3C239D0F6FB53FA59" author="Naruse" box="[827,1258,1432,1458]" pageId="15" pageNumber="40" refString="Naruse, T., Shokita, S. & Kawahara, T. (2005) Neorhynchoplax yaeyamaensis, a new false spider crab (Decapoda: Brachyura: Hymenosomatidae) from the Yaeyama Group, the Ryukyu Islands, Japan. Zootaxa, 877, 1 - 7." type="journal article" year="2005">Naruse, Shokita & Kawahara, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
, were released from the pseudovulvae when the abdomen of the preserved specimen was opened. As such, the torn thoracic sternum of the female specimen of
|
||
<taxonomicName id="D3DB4D259948FFA3C33AD086FD64F9EA" box="[568,733,1512,1537]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="species" species="denticulata">
|
||
<emphasis id="26AFEAB49948FFA3C33AD086FD64F9EA" box="[568,733,1512,1537]" italics="true" pageId="15" pageNumber="36">G. denticulata</emphasis>
|
||
</taxonomicName>
|
||
might have been caused by a recent release of its eggs, larvae or juveniles. Such an internal brood cavity is possible in the
|
||
<taxonomicName id="D3DB4D259948FFA3C28AD37EFBDDF9C1" box="[904,1124,1552,1578]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="36" phylum="Arthropoda" rank="family">Hymenosomatidae</taxonomicName>
|
||
because there is a vacuum in the central part of the body, that is devoid of endosternites and endopleurites in the whole median part (except the sella turcica, posteriorly), the phragmae being only located on lateral sides of the body and leaving an empty place. The sternal wall is thus strongly depressed, even reaching the dorsal wall of the carapace (Guinot & Richer de
|
||
<bibRefCitation id="704A4B579948FFA3C092D3DEFD9CF921" author="Forges" box="[400,549,1712,1738]" pageId="15" pageNumber="39" refString="Guinot, D. & Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay, 1838 et les Inachoididae Dana, 1851 (Crustacea, Decapoda, Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges 1997</bibRefCitation>
|
||
: 474, 477, figs. 2C, E, 6F).
|
||
</paragraph>
|
||
<paragraph id="146436A69957FFBCC195D5F0FB3EFF53" blockId="16.[151,1159,158,184]" box="[151,1159,158,184]" pageId="16" pageNumber="37">
|
||
<emphasis id="26AFEAB49957FFBCC195D5F0FEACFF53" bold="true" box="[151,277,158,184]" pageId="16" pageNumber="37">TABLE 1.</emphasis>
|
||
Key characters of cave-dwelling hymenosomatid genera and allied genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5CC1652D9957FFBDC07ED586FC09FBB2" lastPageId="17" lastPageNumber="38" pageId="16" pageNumber="37" type="description">
|
||
<paragraph id="146436A69957FFBCC07ED586FAF8FF16" blockId="16.[156,1421,231,1890]" box="[380,1345,231,253]" pageId="16" pageNumber="37">
|
||
<taxonomicName id="D3DB4D259957FFBCC07ED586FE69FF16" box="[380,464,232,253]" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="26AFEAB49957FFBCC07ED586FE69FF16" box="[380,464,232,253]" italics="true" pageId="16" pageNumber="37">Sulaplax</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="26AFEAB49957FFBCC0D4D589FD89FF17" bold="true" box="[470,560,231,252]" pageId="16" pageNumber="37">
|
||
<taxonomicNameLabel id="3D9C57CF9957FFBCC0D4D589FD89FF17" box="[470,560,231,252]" pageId="16" pageNumber="37" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<taxonomicName id="D3DB4D259957FFBCC35AD586FD12FF16" box="[600,683,232,253]" class="Malacostraca" family="Hymenosomatidae" genus="Guaplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="26AFEAB49957FFBCC35AD586FD12FF16" box="[600,683,232,253]" italics="true" pageId="16" pageNumber="37">Guaplax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3D9C57CF9957FFBCC3B3D589FCB3FF16" box="[689,778,231,253]" pageId="16" pageNumber="37" rank="genus">
|
||
<emphasis id="26AFEAB49957FFBCC3B3D589FCBDFF17" bold="true" box="[689,772,231,252]" pageId="16" pageNumber="37">gen. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="26AFEAB49957FFBCC22CD586FAF8FF16" box="[814,1345,232,253]" italics="true" pageId="16" pageNumber="37">
|
||
<taxonomicName id="D3DB4D259957FFBCC22CD586FC75FF16" box="[814,972,232,253]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="37" phylum="Arthropoda" rank="genus">Neorhynchoplax</taxonomicName>
|
||
<taxonomicName id="D3DB4D259957FFBCC501D586FB3BFF16" box="[1027,1154,232,253]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="37" phylum="Arthropoda" rank="genus">Cancrocaeca</taxonomicName>
|
||
<taxonomicName id="D3DB4D259957FFBCC5D7D586FAF8FF16" box="[1237,1345,232,253]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="37" phylum="Arthropoda" rank="genus">Limnopilos</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED463FA3FFEC9" blockId="16.[156,1421,231,1890]" box="[156,1414,269,290]" pageId="16" pageNumber="37">Rostrum Unilobed, deflexed, Trilobed, developed Unilobed or trilobed, Absent Vestigial or absent</paragraph>
|
||
<paragraph id="146436A69957FFBCC07ED45CFC29FEAC" blockId="16.[156,1421,231,1890]" box="[380,912,306,327]" pageId="16" pageNumber="37">vestigial developed</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED439FA3FFE87" blockId="16.[156,1421,231,1890]" box="[156,1414,343,364]" pageId="16" pageNumber="37">Anterior surface of Vertical Relatively vertical Strongly projecting Relatively vertical Relatively vertical</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED413FC4BFE79" blockId="16.[156,1421,231,1890]" box="[156,1010,381,402]" pageId="16" pageNumber="37">cephalothorax (in lat- beyond rest of ceph-</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED4CCFC31FE5C" blockId="16.[156,1421,231,1890]" box="[156,904,418,439]" pageId="16" pageNumber="37">eral view) alothorax</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED4A9FADAFE37" blockId="16.[156,1421,231,1890]" box="[156,1379,455,476]" pageId="16" pageNumber="37">Configuration of the Located at outer Located at lower Located at just inner Located at just inner Located at just</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED482FAC2FDEA" blockId="16.[156,1421,231,1890]" box="[156,1403,492,513]" pageId="16" pageNumber="37">base of basal anten- lower region with region with respect lower region with lower region with inner region with</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED77FFAC4FDCD" blockId="16.[156,1421,231,1890]" box="[156,1405,529,550]" pageId="16" pageNumber="37">nal article against respect to base of to base of ocular respect to base of respect to base of respect to base of</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED759FAD4FDA7" blockId="16.[156,1421,231,1890]" box="[156,1389,567,588]" pageId="16" pageNumber="37">base of ocular pedun- ocular peduncle peduncle ocular peduncle ocular peduncle ocular peduncle</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED732FF01FD9A" blockId="16.[156,1421,231,1890]" box="[156,184,604,625]" pageId="16" pageNumber="37">cle</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED7EFFAE9FD7D" blockId="16.[156,1421,231,1890]" box="[156,1360,641,662]" pageId="16" pageNumber="37">Dorsal surface Well defined Well defined Well defined Poorly defined Well defined</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED7C8FF5AFD50" blockId="16.[156,1421,231,1890]" box="[156,227,678,699]" pageId="16" pageNumber="37">regions</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED7A2FBC9FCED" blockId="16.[156,1421,231,1890]" pageId="16" pageNumber="37">Eyes Fused to orbit, Fused to orbit, Complete Fused to orbit, Complete no pigment no pigment no pigment</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED678FAA2FCC0" blockId="16.[156,1421,231,1890]" box="[156,1307,790,811]" pageId="16" pageNumber="37">Longitudinal groove Absent Absent Absent Present Present</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED655FED9FCBB" blockId="16.[156,1421,231,1890]" box="[156,352,827,848]" pageId="16" pageNumber="37">on side wall of cara-</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED60EFF71FC9E" blockId="16.[156,1421,231,1890]" box="[156,200,864,885]" pageId="16" pageNumber="37">pace</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED6E8FAB0FB02" blockId="16.[156,1421,231,1890]" pageId="16" pageNumber="37">
|
||
Female vulvae On imaginary line On imaginary line On various parts on On imaginary line On imaginary line joining anterior mar- joining posterior medial fused plate of joining bases of P3 joining sutures gins of P2 coxae on margins of P2 coxae thoracic sternum on medial fused between sternites 5 medial fused plate of on medial fused (from level of poste- plate of the thoracic and 6 on medial thoracic sternum; plate of thoracic ster- rior margins of P1 sternum; with basal fused plate of the without basal mount num; without basal coxae to anterior mount thoracic sternum; mount margins of P3 coxae) with basal mount (confirmed except for
|
||
<taxonomicName id="D3DB4D259957FFBCC5F5D1C1FAB8FB01" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="37" phylum="Arthropoda" rank="species" species="microrhynchus">
|
||
<emphasis id="26AFEAB49957FFBCC5F5D1C1FAB8FB01" italics="true" pageId="16" pageNumber="37">L. microrhynchus</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED197FAA5FAE5" blockId="16.[156,1421,231,1890]" box="[156,1308,1273,1294]" pageId="16" pageNumber="37">Ambulatory dactyli: Absent Absent Present or absent Absent Present</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED071FEFDFADF" blockId="16.[156,1421,231,1890]" box="[156,324,1311,1332]" pageId="16" pageNumber="37">subterminal tooth</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED02AFACDFAB2" blockId="16.[156,1421,231,1890]" box="[156,1396,1348,1369]" pageId="16" pageNumber="37">Male abdomen-pleo- 4 segmented (3rd–5th 4 segmented (3rd– 4–5 segmented (3rd– 6 segmented (no 6 segmented (no</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED007FC5FF9D3" blockId="16.[156,1421,231,1890]" pageId="16" pageNumber="37">telson fused); pleotelson 5th fused); pleotel- 5th or 4th–5th fused) fusion), barreled; fusion), barreled; narrow, distinctly less son narrow, dis- [see Ng & Chuang pleotelson wide, pleotelson wide, than half of maxi- tinctly less than half 1996]; pleotelson more than half of more than half of mum width of abdo- of maximum width narrow, distinctly maximum width of maximum width of men; fused segment of abdomen; fused less than half of abdomen abdomen with proximal part segment with proxi- maximum width of</paragraph>
|
||
<paragraph id="146436A69957FFBCC07ED326FC3FF9B6" blockId="16.[156,1421,231,1890]" box="[380,902,1608,1629]" pageId="16" pageNumber="37">not constricted mal part constricted abdomen</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED300FA32F968" blockId="16.[156,1421,231,1890]" box="[156,1419,1646,1667]" pageId="16" pageNumber="37">Female abdomen-ple- 1st–2nd and 2nd–3rd 1st–2nd movable, 3rd–5th, 2nd–5th, or 6 demarcated seg- 6 distinctly demar-</paragraph>
|
||
<paragraph id="146436A69957FFBCC19ED3FDFB44F889" blockId="16.[156,1421,231,1890]" pageId="16" pageNumber="37">otelson segmented, 3rd–pleo- 2nd–3rd and 5th– 2nd-pleotelson fused ments, boundary cated segments, telson fused, 5th–ple- pleotelson only with between 1st and 2nd boundary between otelson with suture suture on outer side. movable 1st and 2nd or on outer side only between 2nd and 3rd segments movable</paragraph>
|
||
<paragraph id="146436A69957FFBDC5D7D2D2FEEAFF5C" blockId="16.[1237,1403,1980,2001]" lastBlockId="17.[156,1425,162,1113]" lastPageId="17" lastPageNumber="38" pageId="16" pageNumber="37">to be continued... table 1 continued...</paragraph>
|
||
<paragraph id="146436A69956FFBDC19ED5A9FAD5FDED" blockId="17.[156,1425,162,1113]" pageId="17" pageNumber="38">
|
||
Female pleopods 2nd–5th present, 2nd–5th present, Present in some spe- Biramous, setose, Biramous, setose, biramous, short, biramous, short, cies (e.g.
|
||
<emphasis id="26AFEAB49956FFBDC288D583FC57FEE9" box="[906,1006,237,258]" italics="true" pageId="17" pageNumber="38">N. manga-</emphasis>
|
||
long plumose pleo- plumose long pleowithin pouch within pouch
|
||
<emphasis id="26AFEAB49956FFBDC22CD47CFCFDFECC" box="[814,836,274,295]" italics="true" pageId="17" pageNumber="38">lis</emphasis>
|
||
, unpubl. data) pods developing pods developing from distal outer from distal outer angle of inner sur- angle of inner surface of 2nd–5th face of 2nd– 5th abdominal segment abdominal segment (not observed in
|
||
<taxonomicName id="D3DB4D259956FFBDC475D4A2FADAFDED" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="38" phylum="Arthropoda" rank="species" species="microrhynchus">
|
||
<emphasis id="26AFEAB49956FFBDC475D4A2FADAFDED" italics="true" pageId="17" pageNumber="38">L. microrhynchus</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="146436A69956FFBDC19ED778FA98FD70" blockId="17.[156,1425,162,1113]" pageId="17" pageNumber="38">G1 Bent at half, tapering, Straight, tapering, tip Distal portion Stout, bent and Stout, bent and tip simple simple curved, tapers to twisted on proximal twisted on proximal simple tip 1/4, with processes 3/5, with processes on apex on apex</paragraph>
|
||
<paragraph id="146436A69956FFBDC19ED7C5FB02FC2F" blockId="17.[156,1425,162,1113]" pageId="17" pageNumber="38">
|
||
Brooding of eggs Internal brood cavity Internal brood cavity Internal brood cavity Eggs brooded within Eggs attached to inner surface of endopods; exopods dome-shaped abdo- extending inwards, men, not attached to partially covering endopods; 3rd–5th “dome” (not exopods lie along observed in
|
||
<emphasis id="26AFEAB49956FFBDC449D60BFAE6FC91" box="[1355,1375,869,890]" italics="true" pageId="17" pageNumber="38">L.</emphasis>
|
||
lateral margin of
|
||
<taxonomicName id="D3DB4D259956FFBDC5D7D6E4FADAFC74" box="[1237,1379,906,927]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="38" phylum="Arthropoda" rank="species" species="microrhynchus">
|
||
<emphasis id="26AFEAB49956FFBDC5D7D6E4FADAFC74" box="[1237,1379,906,927]" italics="true" pageId="17" pageNumber="38">microrhynchus</emphasis>
|
||
</taxonomicName>
|
||
) abdomen, covering
|
||
</paragraph>
|
||
<paragraph id="146436A69956FFBDC501D6BBFB08FC01" blockId="17.[156,1425,162,1113]" box="[1027,1201,981,1002]" pageId="17" pageNumber="38">margin of “dome”</paragraph>
|
||
<paragraph id="146436A69956FFBDC19ED694FC09FBB2" blockId="17.[156,1425,162,1113]" pageId="17" pageNumber="38">Habitat Freshwater in caves Freshwater in caves Intertidal flat to Freshwater in caves Brackish-water freshwater environments environments</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |