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<mods:title id="EA67EFCECB2AE6F811B02396C1D92C63">A taxonomic guide to the brittle-stars (Echinodermata, Ophiuroidea) from the State of Paraiba continental shelf, Northeastern Brazil</mods:title>
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<mods:namePart id="BC9D1A2B381DC29A35BA8CB03709E440">Gondim, Anne I.</mods:namePart>
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<mods:namePart id="B774F31827D2F027A22F42BDB6165B2D">Alonso, Carmen</mods:namePart>
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<mods:namePart id="E938BA82A01D7FD7FC24AA46615A82B1">Dias, Thelma L. P.</mods:namePart>
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<mods:namePart id="628354A903C9B319445206ABDA437EA6">Manso, Cynthia L. C.</mods:namePart>
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<mods:namePart id="782FFD94E6260A56FEB1B2B2837D8D1A">Christoffersen, Martin L.</mods:namePart>
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<treatment id="790F0102B26C399EFCDD9B45F62AB4E4" ID-GBIF-Taxon="152044877" LSID="urn:lsid:plazi:treatment:790F0102B26C399EFCDD9B45F62AB4E4" httpUri="http://treatment.plazi.org/id/790F0102B26C399EFCDD9B45F62AB4E4" lastPageNumber="58" pageId="13" pageNumber="58" scope_class="Ophiuroidea" scope_family="Amphiuridae" scope_phylum="Echinodermata">
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<paragraph id="27A221674C207ED0F218413247ED05D9" pageId="13" pageNumber="58">
<taxonomicName id="A0B9026E8ABA664CF579FD4A1682CDCE" ID-CoL="6SQ7G" LSID="http://species-id.net/wiki/Ophiophragmus_brachyactis" authority="H. L. Clark, 1915" authorityName="H. L. Clark" authorityYear="1915" class="Ophiuroidea" family="Amphiuridae" genus="Ophiophragmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiophragmus brachyactis" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="brachyactis">
<pageBreakToken id="DF2E748B24A5620B920EED16307184EA" pageId="13" pageNumber="58" start="start">Ophiophragmus</pageBreakToken>
brachyactis H.L. Clark, 1915
</taxonomicName>
Figure 7
<normalizedToken id="4BCDF059D259481EC5D8375520482C4A" originalValue="ae">a-e</normalizedToken>
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<subSubSection id="2F8FDE12F38265B356A40D3CFA930431" pageId="13" pageNumber="58" type="description">
<paragraph id="8346C8C06BEEFD999941CDCF6365BDE0" pageId="13" pageNumber="58">Description.</paragraph>
<paragraph id="65284081626F1F8DE3E68DD6C55FA3AD" pageId="13" pageNumber="58">Disk circular (dd = 4.45 mm). Covered by imbricating scales of irregular shape (Fig. 7a). Margin of interradial field with 12-13 small, enlarged, blunt spines that decrease in size in the direction of the radial shields (Fig. 7a). Radial shields slightly longer than wide, united, except at proximal end, where a small triangular scale occurs between the pair of radial shields. Ventral interradius covered by scales similar to dorsal scales (Fig. 7b). Oral shields arrow-shaped (Fig. 7c). Adoral shields enlarged laterally, contiguous in proximal region. Two papillae on each side of jaw angle, the distal one longer and wider (Fig. 7c). Infradental papillae robust, and rectangular. Dorsal arm plate wider than long, rectangular, with rounded borders (Fig. 7d). Ventral arm plate pentagonal, with small notch on distal margin (Fig. 7e). Two small, perpendicular, tentacle scales, inner scale slightly longer than outer scale. Three small, compressed, blunt, arm spines, the ventral one largest (Fig. 7e).</paragraph>
<caption id="660BCE58A158E0275B9B8C189A611ADA" pageId="13" pageNumber="58">
<paragraph id="A473903C04499D47F88D05698C4449C1" pageId="13" pageNumber="58">
Figure 7. Species of the family
<taxonomicName id="5A99CDA1BC69605A8ED9637EE5772F06" class="Ophiuroidea" family="Amphiuridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="family">Amphiuridae</taxonomicName>
.
<taxonomicName id="3F27FFD35E9B9DB6CC80061496082457" class="Ophiuroidea" family="Amphiuridae" genus="Ophiophragmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiophragmus brachyactis" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="brachyactis">Ophiophragmus brachyactis</taxonomicName>
A dorsal view, detail of the fence of papillae B ventral view C jaw D dorsal view of the arms E ventral view of the arms.
<taxonomicName id="CDA8EB2720A3BE03E18FE367D220D9C5" class="Ophiuroidea" family="Amphiuridae" genus="Ophiostigma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiostigma isocanthum" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="isocanthum">Ophiostigma isocanthum</taxonomicName>
F dorsal view G ventral view H detail of the radial shields I jaw J dorsal view of the arms L ventral view of the arms. Scale bar = 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="CBAFA603EA3C1EC2C77F7D068EFD16B1" pageId="13" pageNumber="58" type="distribution">
<paragraph id="9FACAAB11F2976E15CC7E5F16354CD4C" pageId="13" pageNumber="58">Distribution.</paragraph>
<paragraph id="4E7B51144736CC64C9A2D71590304DA8" pageId="13" pageNumber="58">
Florida and Dry Tortugas, the Antilles, Gulf of Mexico, and Brazil (
<bibRefCitation id="34A34D1FF92C6824E30B0F11373DA565" author="Thomas, LP" journalOrPublisher="Bulletin of Marine Science of the Gulf and Caribbean" pageId="36" pageNumber="81" pagination="623 - 694" title="The shallow water Amphiurid brittle stars (Echinodermata, Ophiuroidea) of Florida." volume="12" year="1962">Thomas 1962</bibRefCitation>
,
<bibRefCitation id="D0490F31ADB7749D80EA7A59039078E5" author="Hendler, G" journalOrPublisher="Smithsonian Institution Press, Washington" pageId="33" pageNumber="78" title="Sea stars, sea urchins and allies: echinoderms of Florida and the Caribbean." year="1995">Hendler et al. 1995</bibRefCitation>
). In Brazil from
<normalizedToken id="2FA12692CDBFF42CAE53A34FDDF006F0" originalValue="Paraíba">Paraiba</normalizedToken>
(
<bibRefCitation id="30DB233D5F4D34DCFC76E0A9C9DAE325" author="Gondim, AI" journalOrPublisher="Memoirs of Museum Victoria" pageId="32" pageNumber="77" title="Echinodermata da Praia do Cabo Branco, Joao Pessoa, Paraiba, Brasil. Biota Neotropica 8: 151 - 159. http: // www. biotaneotropica. org. br / v 8 n 2 / pt / abstract? inventory + bn 02408022008," url="10.1590/S1676-06032008000200016" year="2008">Gondim et al. 2008</bibRefCitation>
),
<normalizedToken id="D9E31EE8BA2FDD68D42C2B5151239120" originalValue="Espírito">Espirito</normalizedToken>
Santo, and Rio de Janeiro (
<bibRefCitation id="E16AEBC41BF400714BCA14C7E5162919" author="Manso, CLC" journalOrPublisher="Revista Brasileira de Biologia" pageId="34" pageNumber="79" pagination="965 - 968" title="Sobre a ocorrencia de Ophionema intricata Luetken, 1869 e Ophiophragmus brachyactis H. L. Clark, 1915 no Brasil (Echinodermata, Ophiuroidea, Amphiurudae)." volume="48" year="1988">Manso 1988</bibRefCitation>
,
<bibRefCitation id="62BD55A7EE0F717F6FB4E3A554A94DA9" author="Manso, CLC" journalOrPublisher="Revista Brasileira de Biologia" pageId="34" pageNumber="79" pagination="189 - 195" title="Ofiuroides da Plataforma Brasileira. Parte II: Norte do Estado do Rio de Janeiro, Estado do Espirito Santo, Sul do Estado da Bahia e Bancos Royal Charlotte, Hostpur e Davis (Echinodermata: Ophiuroidea)." volume="35" year="1993">1993</bibRefCitation>
). Depth 22 to 87 m. Recorded from 30 m in this study.
</paragraph>
</subSubSection>
<subSubSection id="8A1261CDC11571DF28AD3FC9E0F4D339" pageId="13" pageNumber="58" type="remarks">
<paragraph id="5999A19A273D145728136615CB5837CC" pageId="13" pageNumber="58">Remarks.</paragraph>
<paragraph id="279E033E252166E03287F8DACEB9F8B6" pageId="13" pageNumber="58">
Species known from bottoms with sand, in which bryozoans predominate (
<bibRefCitation id="296C8AD36854AB06E88C4E4575672AE5" author="Manso, CLC" journalOrPublisher="Revista Brasileira de Biologia" pageId="34" pageNumber="79" pagination="965 - 968" title="Sobre a ocorrencia de Ophionema intricata Luetken, 1869 e Ophiophragmus brachyactis H. L. Clark, 1915 no Brasil (Echinodermata, Ophiuroidea, Amphiurudae)." volume="48" year="1988">Manso 1988</bibRefCitation>
). We consider
<taxonomicName id="B6E0D5548DE8A15B2A576F1E0982F4BA" class="Ophiuroidea" family="Amphiuridae" genus="Amphiodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphiodia riisei" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="riisei">Amphiodia riisei</taxonomicName>
distinct from
<taxonomicName id="DBB082665DEED4FA4BCB82D8A4D5767F" class="Ophiuroidea" family="Amphiuridae" genus="Ophiophragmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiophragmus brachyactis" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="brachyactis">Ophiophragmus brachyactis</taxonomicName>
and use the name
<taxonomicName id="0B69E1F320E814238DB7A5FD251DEE61" class="Ophiuroidea" family="Amphiuridae" genus="Amphiodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphiodia riisei" order="Amphilepidida" pageId="13" pageNumber="58" phylum="Echinodermata" rank="species" species="riisei">Amphiodia riisei</taxonomicName>
for the former taxon (see discussion under this species).
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