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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.186.75412" ID-GBIF-Dataset="ad4a5506-c7fa-4a6f-bf30-5fe30cff71af" ID-PMC="PMC8677714" ID-Pensoft-Pub="1314-2003-186-139" ID-Pensoft-UUID="70DC0EB03FF3548ABBF5113F5E1EB62B" ID-PubMed="35002361" ModsDocID="1314-2003-186-139" checkinTime="1639109976156" checkinUser="pensoft" docAuthor="Saez, Llorenc, Bogunic, Faruk, Cambria, Salvatore, Riera, Jesus &amp; Bogdanovic, Sandro" docDate="2021" docId="0D988EF069CF5B1896B2A0B427C1A8C1" docLanguage="en" docName="PhytoKeys 186: 139-158" docOrigin="PhytoKeys 186" docPubDate="2021-12-09" docSource="http://dx.doi.org/10.3897/phytokeys.186.75412" docTitle="Thymus richardii subsp. vigoi Riera, Güemes &amp; Rosselló, Fl. Montiber. 37: 78. 2007" docType="treatment" docVersion="5" id="70DC0EB03FF3548ABBF5113F5E1EB62B" lastPageNumber="139" masterDocId="70DC0EB03FF3548ABBF5113F5E1EB62B" masterDocTitle="On the identity of Thymus humifusus var. aureopunctatus (Lamiaceae) and taxonomic notes on the Th. richardii complex" masterLastPageNumber="158" masterPageNumber="139" pageNumber="139" updateTime="1732707109020" updateUser="ExternalLinkService">
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<mods:title id="41C279E92BF98BD2F4CF463EE6628B02">On the identity of Thymus humifusus var. aureopunctatus (Lamiaceae) and taxonomic notes on the Th. richardii complex</mods:title>
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<mods:name id="CC952DFD318860BE5D88CA1D8D9E1AAD" type="personal">
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<mods:namePart id="AAD675791D8DCA248864B9662439E0D7">Saez, Llorenc</mods:namePart>
<mods:affiliation id="B5059FBFEB15E2A3501DD0FB0BBCC1D5">Systematics and Evolution of Vascular Plants (UAB) - Associated Unit to CSIC, Departament de Biologia Animal, Biologia Vegetal i Ecologia, Facultat de Biociencies, Universitat Autonoma de Barcelona, 08193 Bellaterra, Spain</mods:affiliation>
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<mods:namePart id="972EEA0713744092890FD29F6D56E83A">Bogunic, Faruk</mods:namePart>
<mods:affiliation id="01E6175D053EF29EFE94B681FA2972B2">University of Sarajevo, Faculty of Forestry, Zagrebacka 20, 71 000 Sarajevo, Bosnia and Herzegovina</mods:affiliation>
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<mods:namePart id="B372531F1098C48109A7A865F7ADEADB">Cambria, Salvatore</mods:namePart>
<mods:affiliation id="1C09B057AC17DFA5A3FF22E3E6C6EFD9">Department of Biological, Geological and Environmental Sciences, University of Catania, Via A. Longo 19, 95125 Catania, Italy</mods:affiliation>
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<mods:namePart id="59A7704696032E7A5309BC2BB9442BB3">Riera, Jesus</mods:namePart>
<mods:affiliation id="F0A30624A1F8D9D438677B35DF81438D">Jardin Botanico. Universidad de Valencia. C / Quart, 80. 46008 Valencia, Spain</mods:affiliation>
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<mods:namePart id="A14F6875D5EA8363DC5845F3EB707E64">Bogdanovic, Sandro</mods:namePart>
<mods:nameIdentifier id="3132E35BBBA7C2D699C8D33A4DC3D2B0" type="ORCID">https://orcid.org/0000-0002-1952-6059</mods:nameIdentifier>
<mods:affiliation id="69DAA6B6E70AF01C16B65F6C364E8AC9">University of Zagreb, Faculty of Agriculture, Department of Agricultural Botany, Svetosimunska 25, 10000 Zagreb, Croatia &amp; Centre of Excellence for Biodiversity and Molecular Plant Breeding, Svetosimunska 25, 10000 Zagreb, Croatia</mods:affiliation>
<mods:nameIdentifier id="226D7B3397CA63943375C8712AD31885" type="email">sbogdanovic@agr.hr</mods:nameIdentifier>
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<mods:title id="AAA2B09FC137397BE18C72DD1B5D323A">PhytoKeys</mods:title>
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<mods:part id="4937AE4213BD5AA9696255EC69D6CFA8">
<mods:date id="0217A9398A0021C2A464551D7C1A3E71">2021</mods:date>
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<mods:number id="640114A9E003D917C146BECE0F0B5FA2">2021-12-09</mods:number>
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<mods:number id="D2D3A67892F22F0230C1149755BA25BA">186</mods:number>
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<mods:start id="B39CABE1AC13C3B5AE3A5047326603FA">139</mods:start>
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<mods:url id="C0D9776216648A9B73C89A3D028B2BCD">http://dx.doi.org/10.3897/phytokeys.186.75412</mods:url>
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<mods:identifier id="C827292DEDD77B1B403075A1216D323B" type="DOI">http://dx.doi.org/10.3897/phytokeys.186.75412</mods:identifier>
<mods:identifier id="5E5BF191E54F0F37DE52971E53167B73" type="Pensoft-Pub">1314-2003-186-139</mods:identifier>
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<treatment id="0D988EF069CF5B1896B2A0B427C1A8C1" ID-GBIF-Taxon="191261309" LSID="urn:lsid:plazi:treatment:0D988EF069CF5B1896B2A0B427C1A8C1" httpUri="http://treatment.plazi.org/id/0D988EF069CF5B1896B2A0B427C1A8C1" lastPageNumber="139" pageId="0" pageNumber="139" scope_family="Lamiaceae">
<subSubSection id="52317079B1FDE1BD7DE0396DD762A1BC" pageId="0" pageNumber="139" type="nomenclature">
<paragraph id="2A0415B54636CC4688B04B187DE5F8D1" pageId="0" pageNumber="139">
5.
<taxonomicName id="7164AAE61088D51E67B4618A4C2640A6" ID-CoL="5LJJS" LSID="0D988EF0-69CF-5B18-96B2-A0B427C1A8C1" authority="Riera, Gueemes &amp; Rossello, Fl. Montiber. 37: 78. 2007" authorityName="Riera, Güemes &amp; Rosselló, Fl. Montiber. 37: 78." authorityYear="2007" class="Magnoliopsida" family="Lamiaceae" genus="Thymus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thymus richardii subsp. vigoi" order="Lamiales" pageId="0" pageNumber="139" phylum="Tracheophyta" rank="subSpecies" species="richardii" subSpecies="vigoi">
Thymus richardii subsp. vigoi Riera,
<normalizedToken id="F5763FB5AC7904787A4EA9D8274E0EB5" originalValue="Güemes">Gueemes</normalizedToken>
&amp;
<normalizedToken id="281BBFAC3D7628CB0262B6C8862B1F3F" originalValue="Rosselló">Rossello</normalizedToken>
, Fl. Montiber. 37: 78. 2007
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="6C66E25240891017BC4938E4B6781687" pageId="0" pageNumber="139" type="reference_group">
<paragraph id="E7D88E8F04D018D3D6ACB5842E650B2B" pageId="0" pageNumber="139">
<taxonomicName id="55426310EAF4FA6FEB0A401A47621A6C" class="Magnoliopsida" family="Lamiaceae" genus="Thymus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thymus richardii subsp. vigoi" order="Lamiales" pageId="0" pageNumber="139" phylum="Tracheophyta" rank="subSpecies" species="richardii" subSpecies="vigoi">Thymus richardii vigoi</taxonomicName>
Type. Spain, Valencia, Villalonga, La Safor, ad
<normalizedToken id="D26179A3A95F477D6FD04FDA8D0048F3" originalValue="lOrxa">l'Orxa</normalizedToken>
, 30SYJ3706, 600 m, 4 July 2000,
<emphasis id="2299B4B68AF0CAB4D744978F5F89EA3C" italics="true" pageId="0" pageNumber="139">
J. Riera &amp; J.
<normalizedToken id="C7FDF9E2A1375EB898EF0C3CF02C700A" originalValue="Güemes">Gueemes</normalizedToken>
</emphasis>
(holotype: VAL 185406!; isotype: MA 757804!;
<bibRefCitation id="56E6978194BCB9625052F3CCB0C56BE8" author="Riera, J" journalOrPublisher="Flora Montiberica" pageId="0" pageNumber="139" pagination="77 - 80" refId="B21" refString="Riera, J, Guemes, J, Rossello, JA, 2007. Thymus richardii (Lamiaceae) in the Iberian Peninsula. Flora Montiberica 37: 77 - 80" title="Thymus richardii (Lamiaceae) in the Iberian Peninsula." volume="37" year="2007">Riera et al. 2007</bibRefCitation>
).
</paragraph>
<paragraph id="076C8D28BB2F55893D416F3BCA85DF07" pageId="0" pageNumber="139">
<taxonomicName id="2714F8807A5A7556D7474AAF81070DE1" authorityName="O.Bolos &amp; Vigo" authorityYear="1983" class="Magnoliopsida" family="Lamiaceae" genus="Thymus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thymus richardii var. valentinus" order="Lamiales" pageId="0" pageNumber="139" phylum="Tracheophyta" rank="variety" species="richardii" variety="valentinus">Thymus richardii valentinus</taxonomicName>
=
<taxonomicName id="CD5EC26BC031B20B9489106FA331B7CE" authorityName="O.Bolos &amp; Vigo" authorityYear="1983" class="Magnoliopsida" family="Lamiaceae" genus="Thymus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thymus richardii subsp. var. var. valentinus" order="Lamiales" pageId="0" pageNumber="139" phylum="Tracheophyta" rank="variety" species="richardii" subSpecies="var." variety="valentinus">Thymus richardii var. valentinus</taxonomicName>
O.
<normalizedToken id="1A41512161760A9694AC20EF9EBE3EC0" originalValue="Bolòs">Bolos</normalizedToken>
&amp; Vigo, Collect. Bot. (Barcelona) 14: 95. 1983
</paragraph>
</subSubSection>
<subSubSection id="F96C25C02754B6D916599C4AFFBA0D43" pageId="0" pageNumber="139" type="materials_examined">
<paragraph id="C342EC1A33F093E2129FD993BDD5D2C5" pageId="0" pageNumber="139">
<typeStatus id="058E3B611E870621AB534562394546B1">Type</typeStatus>
.
</paragraph>
<paragraph id="5F878101AAD719F5CE58B281AE596AF6" pageId="0" pageNumber="139">
<materialsCitation id="14FA8A357A7A4FC40830004D9770ED46" accessionNumber="BC 119858" collectingDate="1950-09-15" collectorName="P. Canigueral" country="Spain" location="Gandia" municipality="Valentia" specimenCount="1" stateProvince="Valencia province" typeStatus="holotype">
<collectingCountry id="1AB7E93D664267195E8D248C44C7DE2F" name="Spain">Spain</collectingCountry>
,
<collectingRegion id="AED1687549331A7E42BF850ECAE8C521" country="Spain" name="Comunitat Valenciana">Valencia province</collectingRegion>
,
<collectingMunicipality id="BC4BEC739E1C7C7F7A8E4CD50AE0E007">Valentia</collectingMunicipality>
, c.
<location id="C9B2623BED9DCF6B3AB9270197A2E658" LSID="urn:lsid:plazi:treatment:0D988EF069CF5B1896B2A0B427C1A8C1:C9B2623BED9DCF6B3AB9270197A2E658" country="Spain" municipality="Valentia" name="Gandia" stateProvince="Valencia province">Gandia</location>
,
<collectingDate id="435D54DEE7AE5B03FF2DE3A8CF26593C" value="1950-09-15">15 Sept 1950</collectingDate>
,
<emphasis id="6D70B4350D04CD02DFAC8D75CE7DF286" italics="true" pageId="0" pageNumber="139">
<collectorName id="C4DADF5A9D1A7AABA3EDC0436BAA833B">
P.
<normalizedToken id="C1DD0A8F0E862EFCEA52B2AA87C0571B" originalValue="Cañigueral">Canigueral</normalizedToken>
</collectorName>
</emphasis>
(
<typeStatus id="F3587FEF5508D3B0EC89904F3810ADF9">holotype</typeStatus>
:
<accessionNumber id="44518230CD4179F3DE07B3A079ADF0E6" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/BC119858">BC 119858</accessionNumber>
!;
<bibRefCitation id="C073A167AE3A7FF3976213B77197DC98" author="Bolos, O" journalOrPublisher="Collectanea Botanica (Barcelona)" pageId="0" pageNumber="139" pagination="89 - 103" refId="B6" refString="Bolos, O, Vigo, J, 1983. Notes sobre taxonomia i nomenclatura de les plantes, II. Collectanea Botanica (Barcelona) 14: 89 - 103" title="Notes sobre taxonomia i nomenclatura de les plantes, II." volume="14" year="1983">
<normalizedToken id="2D5E7C5E54F5E27C6970A5353CCC07A3" originalValue="Bolòs">Bolos</normalizedToken>
and Vigo 1983
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0D57F0D97919B90520C868F9B9CC52BF" pageId="0" pageNumber="139" type="description">
<paragraph id="B1B33B50F10595E4E8882BB146232A69" pageId="0" pageNumber="139">Description.</paragraph>
<paragraph id="9D638FAF9A29069A080C883631778B2C" pageId="0" pageNumber="139">
Stems up to 16 cm long, suberect to erect. Leaf blade up to 11
<normalizedToken id="77B1E1E9F79C2FCB1F34EED86B1F8948" originalValue="×">x</normalizedToken>
8.1 mm, ovate-triangular, denticulate. Inflorescence 15-34 mm long, usually oblong; bracts up to 8
<normalizedToken id="7024B7D965DA8EDD68E7D3EE58978C07" originalValue="×">x</normalizedToken>
7 mm, similar to leaves, denticulate, usually glabrous. Calyx 5.8-7 mm long, sparsely hairy, with eglandular hairs up to 0.7 mm long, sometimes with sparse stipitate glandular hairs; calyx tube 2.1-2.9 mm long, with eglandular hairs up to 0.5 mm long on the ventral surface; central tooth of upper lip 1.8-2.6 mm long, lower teeth 3-3.5 mm long, with pectinate hairs up to 0.7 mm long. Corolla 7-10 mm long, whitish to pale rose, sometimes cream.
</paragraph>
</subSubSection>
<subSubSection id="13ABC9D3E8246959C5462DF2B9FB88D0" pageId="0" pageNumber="139" type="chromosome number">
<paragraph id="301703869442EC9F9112E3036D3FC318" pageId="0" pageNumber="139">Chromosome number.</paragraph>
<paragraph id="D951E9ADC439A05A5085C82380EA9495" pageId="0" pageNumber="139">Unknown.</paragraph>
</subSubSection>
<subSubSection id="12702901BAD9C9F027CD1BC114997913" pageId="0" pageNumber="139" type="distribution">
<paragraph id="698976F4E42AE2586F8D708475961B58" pageId="0" pageNumber="139">Distribution.</paragraph>
<paragraph id="9875F499EED8A0B3F7C9D8BD215AE95F" pageId="0" pageNumber="139">Endemic to Alicante and Valencia provinces (Spain).</paragraph>
</subSubSection>
<subSubSection id="4B22CAA64CF797093DD17D76300B2532" pageId="0" pageNumber="139" type="habitat">
<paragraph id="64E2E6F4E6773212C4C7752104662812" pageId="0" pageNumber="139">Habitat.</paragraph>
<paragraph id="0C7BB509FFD2604152A0731A9ACEF495" pageId="0" pageNumber="139">Open scrub, on limestone soil, 130-600 m a.s.l.</paragraph>
</subSubSection>
<subSubSection id="3D04849357D4BF6A6DC7167E956643E7" pageId="0" pageNumber="139" type="remarks">
<paragraph id="97C611E2477AF3BB19C034052F74F156" pageId="0" pageNumber="139">Remarks.</paragraph>
<paragraph id="B9AA4DCC3F54B5EC7B6F73E7967B06CB" pageId="0" pageNumber="139">
Plants which were considered to be hybrids between
<taxonomicName id="64F2137B55E039D3BFE60200C3B7B3F1" class="Magnoliopsida" family="Lamiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Th. richardii subsp. vigoi" order="Lamiales" pageId="0" pageNumber="139" phylum="Tracheophyta" rank="subSpecies" species="richardii" subSpecies="vigoi">Th. richardii subsp. vigoi</taxonomicName>
and
<taxonomicName id="CF63786D2233CB43F0B146EAE36493A2" lsidName="T. piperella" pageId="0" pageNumber="139" rank="species" species="piperella">
<emphasis id="BFD7A928C7F39316191C77420F3A539E" italics="true" pageId="0" pageNumber="139">T. piperella</emphasis>
</taxonomicName>
L. have been called
<taxonomicName id="367E55BF344F505FAE1B2789A7A546AB" isHybrid="true" lsidName="T. bolosii" pageId="0" pageNumber="139" rank="species" species="bolosii">
<emphasis id="32CF68F40D571A10F165C9F1EEEC5BE4" italics="true" pageId="0" pageNumber="139">
T.
<normalizedToken id="8BABC3D1ACA069A7B56333CD0E553F12" originalValue="×">x</normalizedToken>
bolosii
</emphasis>
</taxonomicName>
. The hybrid has been reported from a small area of Serra de la Safor, eastern Spain (
<bibRefCitation id="1AB81112FF9E94DBA53C94E8D58A2D12" author="Riera, J" journalOrPublisher="Flora Montiberica" pageId="0" pageNumber="139" pagination="77 - 80" refId="B22" refString="Riera, J, Guemes, J, Rossello, JA, 2020. Thymus x bolosii (Lamiaceae) a new wild hybrid from the eastern Iberian Peninsula. Flora Montiberica 78: 77 - 80" title="Thymus x bolosii (Lamiaceae) a new wild hybrid from the eastern Iberian Peninsula." volume="78" year="2020">Riera et al. 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="88137CB6B8DC40DDE5878D9FB6C19552" pageId="0" pageNumber="139" type="materials_examined">
<paragraph id="C4D39733BEF8D08C11B10ECCF714C68C" pageId="0" pageNumber="139">Specimens examined.</paragraph>
<paragraph id="D8D26A527AF111DB16323B794C990DFD" pageId="0" pageNumber="139">
<materialsCitation id="CA923FBC623AFF95E10B1C58E41C74EE" accessionNumber="BC 674556, BC 119858" collectingDate="1950-09-15" collectingDateMax="2000-07-04" collectingDateMin="1950-09-15" collectorName="La Vilallonga, La Safor, J. B. Peris, G. Stuebing, P. Canigueral, J. Riera, J. Gueemes" country="Spain" county="Alicante province" elevation="136" location="La Safor" municipality="La Vilallonga" specimenCount="1" stateProvince="Valencia province">
<emphasis id="659E27DEA4B7F7AF81F3B3CD8E8AC8DE" bold="true" pageId="0" pageNumber="139">
<collectingCountry id="1666526DB6AF0EEA5D45DDF8C54DAB43" name="Spain">Spain</collectingCountry>
.
</emphasis>
<collectingCounty id="80149A79BB7D2E2D1335E4F1E0E22294">Alicante province</collectingCounty>
:
<collectorName id="35931D32DE1C0F965C53D8D43388A746">
<collectingMunicipality id="A02A85D2829E4ECB1980587D2D337F39">La Vilallonga</collectingMunicipality>
</collectorName>
,
<collectorName id="D30E372B43AD4FBCCBEE7DC27A9622F2">
<location id="17CE9B4B84A38DC26987E6452226008B" LSID="urn:lsid:plazi:treatment:0D988EF069CF5B1896B2A0B427C1A8C1:17CE9B4B84A38DC26987E6452226008B" country="Spain" county="Alicante province" municipality="La Vilallonga" name="La Safor" stateProvince="Valencia province">La Safor</location>
</collectorName>
,
<quantity id="E850567E1531C7EF22F2671A8A118DC6" metricMagnitude="2" metricUnit="m" metricValue="1.3599999999999999" unit="m" value="136.0">
<elevation id="181B3AE3B35FB45551C6794F8A3011E2" metricMagnitude="2" metricUnit="m" metricValue="1.3599999999999999" unit="m" value="136.0">136 m</elevation>
</quantity>
,
<collectingDate id="1C67357E321DA335D212407D03AE7AB2" value="1984-06-22">22 June 1984</collectingDate>
,
<emphasis id="A25FF5C1AFE9275E79458D53031BACED" italics="true" pageId="0" pageNumber="139">
<collectorName id="6DBFFFC9E30E81762502858CF85DAEB7">J.B. Peris</collectorName>
&amp;
<collectorName id="A55FA899DFE210F3CF82FF8B1E6CF8B9">
G.
<normalizedToken id="4D730A8D15049C0F21A73D7F9EEB1090" originalValue="Stübing">Stuebing</normalizedToken>
</collectorName>
</emphasis>
(
<accessionNumber id="914E116550880780C987B8BED3CB95DA" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/BC674556">BC 674556</accessionNumber>
);
<collectingRegion id="83EE0EDB8744CF305A38DE825A474ED2" country="Spain" name="Comunitat Valenciana">Valencia province</collectingRegion>
, Valentia, c. Gandia,
<collectingDate id="8963453A147105A14CDC2657819F4724" value="1950-09-15">15 Sept 1950</collectingDate>
,
<emphasis id="4BAE12AFD58FE05E5D79E891902151F7" italics="true" pageId="0" pageNumber="139">
<collectorName id="67161DEDA8C0E6615220250442983354">
P.
<normalizedToken id="B2F5B8858B7EA1AABEEF78030256182F" originalValue="Cañigueral">Canigueral</normalizedToken>
</collectorName>
</emphasis>
(
<accessionNumber id="2A5C4F77C8B97585D3C84BFB0CF23E0A" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/BC119858">BC 119858</accessionNumber>
); Villalonga,
<collectorName id="340582B73BAD2CCDF636717E436ADC9F">La Safor</collectorName>
, ad
<normalizedToken id="B61BAA6C7B140F2EFAF4071556277F24" originalValue="lOrxa">l'Orxa</normalizedToken>
, 30SYJ
<quantity id="2F6CF71C947C2D7D1FE040352CFE347F" metricMagnitude="6" metricUnit="m" metricValue="3.7065999999999995" unit="m" value="3706600.0">
<elevation id="01B18410D929730EBECD923284138A9E" metricMagnitude="6" metricUnit="m" metricValue="3.7065999999999995" unit="m" value="3706600.0">3706, 600 m</elevation>
</quantity>
,
<collectingDate id="1A3349CCB3028734A3D10A477AB26E1B" value="2000-07-04">4 July 2000</collectingDate>
,
<emphasis id="BB66442223878B7F5B58085BB2777651" italics="true" pageId="0" pageNumber="139">
<collectorName id="F40C58A9CA04769069F5A46005231C96">J. Riera</collectorName>
&amp;
<collectorName id="BF6779F77F011527B1738CDA2DAAC460">
J.
<normalizedToken id="0627D42C200E08C555237AB0EC973F4F" originalValue="Güemes">Gueemes</normalizedToken>
</collectorName>
</emphasis>
(VAL 185406!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>