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<document id="57575D147EC7FDA6ED32C72A8A0D4DAC" ID-CLB-Dataset="29444" ID-DOI="10.11646/zootaxa.4472.3.5" ID-GBIF-Dataset="fdbc45bb-a0df-4472-8cb7-ca718f01b8d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440287" ID-ZooBank="0BAB122B-66C0-4C30-B1FD-BC9E61B8F115" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538372442823" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2018" docId="0D5C096CFFBE0D6AB58AC10B30BEFE3D" docLanguage="en" docName="zootaxa.4472.3.5.pdf" docOrigin="Zootaxa 4472 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scydmaenus (Corbulifer) pseudorobustus Franz" docType="treatment" docVersion="7" lastPageNumber="517" masterDocId="F1657114FFB50D67B51DC5053524FFBA" masterDocTitle="' Monster Scydmaenus' of Australia: revision of subgenus Corbulifer Franz (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="531" masterPageNumber="505" pageNumber="516" updateTime="1698559125286" updateUser="ExternalLinkService">
<mods:mods id="25C31ABECE3EAEB870307390D76CE36B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A95F283D44B94F6A4FA516A200F03F21">
<mods:title id="C816716EBFE8CAECD379EDFFB1B68E39">' Monster Scydmaenus' of Australia: revision of subgenus Corbulifer Franz (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="F1B9C41161F0CA93AEDAB8057FB9C9A3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E7E6E585F1E9E0FA7350DAC6BE3ACF92">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="14A9797E25BC0B5A307D5F89D5186F75">text</mods:typeOfResource>
<mods:relatedItem id="F8F511852390DF7FF7B7FDC90B3C60D6" type="host">
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<mods:title id="789A344CBEBCFE10327EADE52A473920">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BE2AED1B2F3B9D78E519B8F8020D32FA">
<mods:date id="6E0C6A3349A00FACA29BF8F2BF10B899">2018</mods:date>
<mods:detail id="CB33D1719A6CDFB6F4CBD3CBEBEA7786" type="pubDate">
<mods:number id="038EC00152FF846CFAB5F029169EC379">2018-09-11</mods:number>
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<mods:number id="78D7C79EFC818F380EC9373C72EA6385">4472</mods:number>
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<mods:number id="C27008ADFD1863FECA16C990BBD00656">3</mods:number>
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<mods:classification id="0BAD1140C8AAE95B3C3756870357C4A6">journal article</mods:classification>
<mods:identifier id="4BB9720E5FE1F98CAA53722DD05D8BF3" type="CLB-Dataset">29444</mods:identifier>
<mods:identifier id="301D84A8D6A882393984F795564BA57B" type="DOI">10.11646/zootaxa.4472.3.5</mods:identifier>
<mods:identifier id="2B2E81D95B3F8973A203708DA09F7415" type="GBIF-Dataset">fdbc45bb-a0df-4472-8cb7-ca718f01b8d6</mods:identifier>
<mods:identifier id="3BDCFDBB5F15EDB0602DC62EB5B0E9A0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="790F494525C8F977D3923565FD624193" type="Zenodo-Dep">1440287</mods:identifier>
<mods:identifier id="C561400EE0099A4694FF8FB5AD929C77" type="ZooBank">0BAB122B-66C0-4C30-B1FD-BC9E61B8F115</mods:identifier>
</mods:mods>
<treatment id="0D5C096CFFBE0D6AB58AC10B30BEFE3D" ID-DOI="http://doi.org/10.5281/zenodo.5975904" ID-GBIF-Taxon="148421399" ID-Zenodo-Dep="5975904" LSID="urn:lsid:plazi:treatment:0D5C096CFFBE0D6AB58AC10B30BEFE3D" httpUri="http://treatment.plazi.org/id/0D5C096CFFBE0D6AB58AC10B30BEFE3D" lastPageId="13" lastPageNumber="517" pageId="11" pageNumber="516">
<subSubSection id="CDEFEBF1FFBE0D6CB58AC10B3481FBF0" pageId="11" pageNumber="516" type="nomenclature">
<paragraph id="854AB87AFFBE0D6CB58AC10B37FBFB92" blockId="11.[151,735,1038,1098]" box="[151,735,1038,1064]" pageId="11" pageNumber="516">
<heading id="DE020F16FFBE0D6CB58AC10B37FBFB92" bold="true" box="[151,735,1038,1064]" fontSize="11" level="1" pageId="11" pageNumber="516" reason="1">
<taxonomicName id="42F5C3F9FFBE0D6CB58AC10B37FBFB92" ID-CoL="4W3RM" authority="Franz" authorityName="Franz" box="[151,735,1038,1064]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="516" phylum="Arthropoda" rank="species" species="pseudorobustus" subGenus="Corbulifer">
<emphasis id="B7816468FFBE0D6CB58AC10B37FBFB92" bold="true" box="[151,735,1038,1064]" pageId="11" pageNumber="516">
<emphasis id="B7816468FFBE0D6CB58AC10B340AFB92" bold="true" box="[151,302,1038,1064]" italics="true" pageId="11" pageNumber="516">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFBE0D6CB422C10B349AFB92" bold="true" box="[319,446,1038,1064]" italics="true" pageId="11" pageNumber="516">Corbulifer</emphasis>
)
<emphasis id="B7816468FFBE0D6CB4D2C10B37A8FB92" bold="true" box="[463,652,1038,1064]" italics="true" pageId="11" pageNumber="516">pseudorobustus</emphasis>
Franz
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="854AB87AFFBE0D6CB58AC1343481FBF0" blockId="11.[151,735,1038,1098]" box="[151,421,1073,1098]" pageId="11" pageNumber="516">
(
<figureCitation id="1DCEA4FFFFBE0D6CB582C1343405FBF0" box="[159,289,1073,1098]" captionStart="FIGURES 30–32" captionStartId="12.[151,264,1350,1373]" captionTargetBox="[161,1412,206,1316]" captionTargetId="figure@12.[151,1436,193,1329]" captionTargetPageId="12" captionText="FIGURES 30–32. Scydmaenus (Corbulifer) pseudorobustus Franz. Habitus of male (30), aedeagus in dorsal (31) and lateral (32) views." httpUri="https://zenodo.org/record/1440301/files/figure.png" pageId="11" pageNumber="516">Figs 30–32</figureCitation>
,
<figureCitation id="1DCEA4FFFFBE0D6CB436C134346EFBF0" box="[299,330,1073,1098]" captionStart="FIGURES 55–65" captionStartId="24.[151,264,1652,1675]" captionTargetBox="[151,1421,248,1572]" captionTargetId="figure@24.[151,1436,248,1586]" captionTargetPageId="24" captionText="FIGURES 55–65. Male metatibia in dorsal view. Scydmaenus tamborinensis Franz (55), S. monstrosetibialis sp. n. (56), S. bellenderanus sp. n. (57), S. pseudorobustus Franz (58), S. kiwarrakianus sp. n. (59), S. gushi sp. n. (60), S. gibraltar sp. n. (61), S. eungellanus sp. n., specimen from Mount William (62) and from Peases Lookout (63), S. fishermontanus sp. n. (64), S. super sp. n. (65)." httpUri="https://zenodo.org/record/1440317/files/figure.png" pageId="11" pageNumber="516">58</figureCitation>
,
<figureCitation id="1DCEA4FFFFBE0D6CB448C1343457FBF0" box="[341,371,1073,1098]" captionStart="FIGURES 66–69" captionStartId="25.[151,264,721,744]" captionTargetBox="[162,1433,227,683]" captionTargetId="figure@25.[151,1436,205,690]" captionTargetPageId="25" captionText="FIGURES 66–69. Original labels of primary type specimens of previously described Corbulifer species. Scydmaenus tamborinensis tamborinensis Franz (66); Scydmaenus tamborinensis cunninghamensis Franz (67); Scydmaenus tamborinensis maipotonensis Franz (68); Scydmaenus pseudorobustus Franz (69)." httpUri="https://zenodo.org/record/1440321/files/figure.png" pageId="11" pageNumber="516">69</figureCitation>
,
<figureCitation id="1DCEA4FFFFBE0D6CB462C13734B8FBF0" box="[383,412,1074,1098]" captionStart="FIGURE 70" captionStartId="25.[151,250,1999,2021]" captionTargetBox="[166,1423,886,1948]" captionTargetId="figure@25.[151,1436,867,1960]" captionTargetPageId="25" captionText="FIGURE 70. Distribution of Corbulifer species." httpUri="https://zenodo.org/record/1440323/files/figure.png" pageId="11" pageNumber="516">70</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFBE0D6CB58AC17C36F1FB28" box="[151,981,1145,1170]" pageId="11" pageNumber="516" type="reference_group">
<paragraph id="854AB87AFFBE0D6CB58AC17C36F1FB28" blockId="11.[151,981,1145,1170]" box="[151,981,1145,1170]" pageId="11" pageNumber="516">
<taxonomicName id="42F5C3F9FFBE0D6CB58AC17C360BFB28" ID-CoL="4W3RM" authority="Franz, 1975: 288" authorityName="Franz" authorityPageNumber="288" authorityYear="1975" box="[151,815,1145,1170]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="516" phylum="Arthropoda" rank="species" species="pseudorobustus" subGenus="Corbulifer">
<emphasis id="B7816468FFBE0D6CB58AC17C3406FB2B" box="[151,290,1145,1169]" italics="true" pageId="11" pageNumber="516">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFBE0D6CB42CC17C3483FB2B" box="[305,423,1145,1169]" italics="true" pageId="11" pageNumber="516">Corbulifer</emphasis>
)
<emphasis id="B7816468FFBE0D6CB4A5C17C3743FB2B" box="[440,615,1145,1169]" italics="true" pageId="11" pageNumber="516">pseudorobustus</emphasis>
Franz, 1975: 288
</taxonomicName>
, figs 266–267.
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFBE0D6CB58AC1C43415F9A4" pageId="11" pageNumber="516" type="materials_examined">
<paragraph id="854AB87AFFBE0D6CB58AC1C4378DFAD0" blockId="11.[151,1437,1217,2034]" pageId="11" pageNumber="516">
<emphasis id="B7816468FFBE0D6CB58AC1C4368EFB60" bold="true" box="[151,938,1217,1242]" pageId="11" pageNumber="516">
<typeStatus id="5A4E06D8FFBE0D6CB58AC1C435F6FB60" box="[151,210,1217,1242]" pageId="11" pageNumber="516">Type</typeStatus>
material.
<typeStatus id="5A4E06D8FFBE0D6CB446C1C434DAFB60" box="[347,510,1217,1242]" pageId="11" pageNumber="516" type="holotype">HOLOTYPE</typeStatus>
: (
<collectionCode id="E3E420BFFFBE0D6CB705C1C437E4FB60" box="[536,704,1217,1242]" pageId="11" pageNumber="516">
<collectingCountry id="FDE2F8EAFFBE0D6CB705C1C437E4FB60" box="[536,704,1217,1242]" name="Australia" pageId="11" pageNumber="516">AUSTRALIA</collectingCountry>
</collectionCode>
,
<collectionCode id="E3E420BFFFBE0D6CB7C8C1C436BAFB60" box="[725,926,1217,1242]" pageId="11" pageNumber="516">
<collectingRegion id="47317698FFBE0D6CB7C8C1C436BAFB60" box="[725,926,1217,1242]" country="Australia" name="Queensland" pageId="11" pageNumber="516">QUEENSLAND</collectingRegion>
</collectionCode>
):
</emphasis>
<specimenCount id="93F373F3FFBE0D6CB6A5C1C736EBFB63" box="[952,975,1218,1241]" pageId="11" pageNumber="516" type="male">♂</specimenCount>
, six labels: &quot;Deviding {sic!} Range / Highway to Warwick / Queensld., Austr.&quot; [white, printed, with handwritten &quot;Pa82&quot; in blue ink on the reverse side], &quot;
<taxonomicName id="42F5C3F9FFBE0D6CB582C00C3414FA98" authorityName="Jałoszyński" authorityYear="2018" box="[159,304,1289,1314]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="531" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (
<taxonomicName id="42F5C3F9FFBE0D6CB451C00C34E0FA98" authorityName="Franz" authorityYear="1975" box="[332,452,1289,1314]" class="Insecta" family="Staphylinidae" genus="Corbulifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="530" phylum="Arthropoda" rank="genus">Corbulifer</taxonomicName>
) /
<taxonomicName id="42F5C3F9FFBE0D6CB4FCC00C37ABFA98" box="[481,655,1289,1314]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="516" phylum="Arthropoda" rank="species" species="pseudorobustus">pseudorobustus</taxonomicName>
/ det.H.Franz&quot; [white, handwritten and printed], &quot;
<specimenCount id="93F373F3FFBE0D6CB1A6C00F31F6FA98" box="[1211,1234,1290,1314]" pageId="11" pageNumber="516" type="male">♂</specimenCount>
&quot; {white, printed], &quot;
<typeStatus id="5A4E06D8FFBE0D6CB5BDC02B35CDFAFC" box="[160,233,1326,1350]" pageId="11" pageNumber="516">Typus</typeStatus>
&quot; [red, handwritten], &quot;
<collectionCode id="E3E420BFFFBE0D6CB4C1C02B3714FAFC" box="[476,560,1326,1350]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="11" pageNumber="516">SAMA</collectionCode>
Database / No. 25-037032&quot; [white, printed], &quot;
<collectionCode id="E3E420BFFFBE0D6CB127C02B31AAFAFC" box="[1082,1166,1326,1350]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="11" pageNumber="516">SAMA</collectionCode>
Digital Image /
<date id="F14B9EBAFFBE0D6CB054C02B35F6FAD0" pageId="11" pageNumber="516" value="2013-08-21">21.Aug 2013</date>
&quot; [green, printed and handwritten] (
<collectionCode id="E3E420BFFFBE0D6CB746C05737BAFAD0" box="[603,670,1362,1386]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="11" pageNumber="516">SAM</collectionCode>
).
</paragraph>
<paragraph id="854AB87AFFBE0D6CB5DAC0703415F9A4" blockId="11.[151,1437,1217,2034]" pageId="11" pageNumber="516">
<materialsCitation id="359DB227FFBE0D6CB5DAC07031F8FA08" ID-GBIF-Occurrence="1913889500" collectingDate="1982-12-17" collectorName="Monteith &amp; Yeates &amp; Thompson &amp; QM Berlesate" elevation="720" latitude="-28.316668" location="Rathdowney Mt Chinighee" longLatPrecision="1232" longitude="152.96666" pageId="11" pageNumber="516" specimenCount="1" stateProvince="QUEENSLAND">
<emphasis id="B7816468FFBE0D6CB5DAC07037D4FA34" bold="true" box="[199,752,1397,1422]" pageId="11" pageNumber="516">
Additional material (2 exx.):
<collectingRegion id="47317698FFBE0D6CB739C07037CFFA34" box="[548,747,1397,1422]" country="Australia" name="Queensland" pageId="11" pageNumber="516">QUEENSLAND</collectingRegion>
:
</emphasis>
<quantity id="420D159FFFBE0D6CB7E5C070361CFA34" box="[760,824,1397,1422]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="516" unit="km" value="12.0">12km</quantity>
SE of
<location id="802AEEA1FFBE0D6CB69AC070319FFA34" LSID="urn:lsid:plazi:treatment:0D5C096CFFBE0D6AB58AC10B30BEFE3D:802AEEA1FFBE0D6CB69AC070319FFA34" box="[903,1211,1397,1422]" latitude="-28.316668" longLatPrecision="1232" longitude="152.96666" name="Rathdowney Mt Chinighee" pageId="11" pageNumber="516" stateProvince="QUEENSLAND">Rathdowney Mt Chinighee</location>
,
<geoCoordinate id="E0C1DEBDFFBE0D6CB1DAC0733005FA34" box="[1223,1313,1398,1422]" direction="south" orientation="latitude" pageId="11" pageNumber="516" precision="925" value="-28.316668">28°19'S</geoCoordinate>
,
<geoCoordinate id="E0C1DEBDFFBE0D6CB033C07030B3FA34" box="[1326,1431,1397,1422]" direction="east" orientation="longitude" pageId="11" pageNumber="516" precision="925" value="152.96666">152°58'E</geoCoordinate>
,
<elevation id="0ED85F49FFBE0D6CB58AC09F35C0FA0B" box="[151,228,1434,1458]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="11" pageNumber="516" unit="m" value="720.0">
<quantity id="420D159FFFBE0D6CB58AC09F35C0FA0B" box="[151,228,1434,1458]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="11" pageNumber="516" unit="m" value="720.0">720 m</quantity>
</elevation>
,
<collectingDate id="E10F6752FFBE0D6CB5E9C09F3451FA08" box="[244,373,1434,1458]" pageId="11" pageNumber="516" value="1982-12-17">
<date id="F14B9EBAFFBE0D6CB5E9C09F3451FA08" box="[244,373,1434,1458]" pageId="11" pageNumber="516" value="1982-12-17">17.12.1982</date>
</collectingDate>
,
<collectorName id="2800DDACFFBE0D6CB499C09C34C9FA08" box="[388,493,1433,1458]" pageId="11" pageNumber="516">Monteith</collectorName>
,
<collectorName id="2800DDACFFBE0D6CB4E0C09F3762FA0B" box="[509,582,1434,1457]" pageId="11" pageNumber="516">Yeates</collectorName>
&amp;
<collectorName id="2800DDACFFBE0D6CB76FC09C37C9FA08" box="[626,749,1433,1458]" pageId="11" pageNumber="516">Thompson</collectorName>
,
<collectorName id="2800DDACFFBE0D6CB7E1C09F36B8FA08" box="[764,924,1433,1458]" pageId="11" pageNumber="516">QM Berlesate</collectorName>
496, rainforest litter (QM)
</materialsCitation>
;
<materialsCitation id="359DB227FFBE0D6CB1F1C09C3409F9A4" ID-GBIF-Occurrence="1913889501" collectingDate="1993-06-16" collectionCode="ANIC" collectorName="Beaury SF &amp; D. S. Chandler &amp; Berlesate" elevation="755" latitude="-28.29" location="Urbenville" longLatPrecision="740" longitude="152.23" pageId="11" pageNumber="516" specimenCode="ANIC 1648" specimenCount="1" specimenCount-male="1" stateProvince="NEW SOUTH WALES">
<collectingRegion id="47317698FFBE0D6CB1F1C09C35D2FA6C" country="Australia" name="New South Wales" pageId="11" pageNumber="516">
<emphasis id="B7816468FFBE0D6CB1F1C09C35D2FA6C" bold="true" pageId="11" pageNumber="516">NEW SOUTH WALES</emphasis>
</collectingRegion>
:
<specimenCount id="93F373F3FFBE0D6CB41AC0BB3417FA6F" box="[263,307,1470,1494]" pageId="11" pageNumber="516" type="male">1 ♂</specimenCount>
,
<geoCoordinate id="E0C1DEBDFFBE0D6CB423C0BB34A8FA6C" box="[318,396,1470,1494]" direction="south" orientation="latitude" pageId="11" pageNumber="516" precision="555" value="-28.29">28.29S</geoCoordinate>
<geoCoordinate id="E0C1DEBDFFBE0D6CB48EC0B834D0FA6C" box="[403,500,1469,1494]" direction="east" orientation="longitude" pageId="11" pageNumber="516" precision="555" value="152.23">152.23E</geoCoordinate>
, Tooloom Scrub Floral Res., rest area,
<elevation id="0ED85F49FFBE0D6CB6BBC0B836CCFA6C" box="[934,1000,1469,1494]" metricMagnitude="2" metricUnit="m" metricValue="7.55" pageId="11" pageNumber="516" unit="m" value="755.0">
<quantity id="420D159FFFBE0D6CB6BBC0B836CCFA6C" box="[934,1000,1469,1494]" metricMagnitude="2" metricUnit="m" metricValue="7.55" pageId="11" pageNumber="516" unit="m" value="755.0">755m</quantity>
</elevation>
,
<collectorName id="2800DDACFFBE0D6CB6EEC0BB314EFA6C" box="[1011,1130,1470,1494]" pageId="11" pageNumber="516">Beaury SF</collectorName>
,
<quantity id="420D159FFFBE0D6CB168C0B83191FA6C" box="[1141,1205,1469,1494]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="11" pageNumber="516" unit="km" value="23.0">23km</quantity>
W of
<location id="802AEEA1FFBE0D6CB1E7C0B83051FA6C" LSID="urn:lsid:plazi:treatment:0D5C096CFFBE0D6AB58AC10B30BEFE3D:802AEEA1FFBE0D6CB1E7C0B83051FA6C" box="[1274,1397,1469,1494]" latitude="-28.29" longLatPrecision="740" longitude="152.23" name="Urbenville" pageId="11" pageNumber="516" stateProvince="NEW SOUTH WALES">Urbenville</location>
,
<collectingDate id="E10F6752FFBE0D6CB09DC0B8342EFA40" pageId="11" pageNumber="516" value="1993-06-16">
<date id="F14B9EBAFFBE0D6CB09DC0B8342EFA40" pageId="11" pageNumber="516" value="1993-06-16">16 Jun. 1993</date>
</collectingDate>
,
<collectorName id="2800DDACFFBE0D6CB405C0E734E3FA40" box="[280,455,1505,1530]" pageId="11" pageNumber="516">D. S. Chandler</collectorName>
,
<collectorName id="2800DDACFFBE0D6CB4C8C0E43719FA40" box="[469,573,1505,1530]" pageId="11" pageNumber="516">Berlesate</collectorName>
<specimenCode id="D5531001FFBE0D6CB755C0E737F6FA40" box="[584,722,1505,1530]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="516">ANIC 1648</specimenCode>
, subtropical closed forest
<taxonomicName id="42F5C3F9FFBE0D6CB11EC0E431D5FA43" box="[1027,1265,1505,1529]" class="Magnoliopsida" family="Urticaceae" genus="Dendrocnide" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="11" pageNumber="516" phylum="Tracheophyta" rank="species" species="excelsa">
<emphasis id="B7816468FFBE0D6CB11EC0E431D5FA43" box="[1027,1265,1505,1529]" italics="true" pageId="11" pageNumber="516">Dendrocnide excelsa</emphasis>
</taxonomicName>
(giant stinger) litter (
<collectionCode id="E3E420BFFFBE0D6CB5C6C3033402F9A4" box="[219,294,1542,1566]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="516">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFBE0D6CB5DAC32C3005F968" pageId="11" pageNumber="516" type="diagnosis">
<paragraph id="854AB87AFFBE0D6CB5DAC32C3005F968" blockId="11.[151,1437,1217,2034]" pageId="11" pageNumber="516">
<emphasis id="B7816468FFBE0D6CB5DAC32C3485F9F8" bold="true" box="[199,417,1577,1602]" pageId="11" pageNumber="516">Revised diagnosis.</emphasis>
Male metatibia gradually and moderately broadened, indistinctly broader than metafemur, broadest distinctly distad middle, lateral (external) tibial margin bisinuate with a distinct submedian concavity, mesal tibial margin concave in proximal 1/5 and nearly evenly convex in distal 4/5, dorsolateral glandular 'pocket' present, with mesal margin bearing long setae; aedeagus in dorsal view broadest near base, with apical portion not delimited, apex divided into two very long and pointed lateral projections strongly bent dorsoproximad.
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFBE0D6BB5DAC3D834BAF8CB" lastPageId="12" lastPageNumber="517" pageId="11" pageNumber="516" type="description">
<paragraph id="854AB87AFFBE0D6CB5DAC3D83799F8A0" blockId="11.[151,1437,1217,2034]" pageId="11" pageNumber="516">
<emphasis id="B7816468FFBE0D6CB5DAC3D83450F94C" bold="true" box="[199,372,1757,1782]" pageId="11" pageNumber="516">Redescription.</emphasis>
Body of male (
<figureCitation id="1DCEA4FFFFBE0D6CB72DC3D837A0F94C" box="[560,644,1757,1782]" captionStart="FIGURES 30–32" captionStartId="12.[151,264,1350,1373]" captionTargetBox="[161,1412,206,1316]" captionTargetId="figure@12.[151,1436,193,1329]" captionTargetPageId="12" captionText="FIGURES 30–32. Scydmaenus (Corbulifer) pseudorobustus Franz. Habitus of male (30), aedeagus in dorsal (31) and lateral (32) views." httpUri="https://zenodo.org/record/1440301/files/figure.png" pageId="11" pageNumber="516">Fig. 30</figureCitation>
) strongly convex, reddish brown, covered with setae slightly lighter than cuticle; BL
<quantity id="420D159FFFBE0D6CB44FC20734D1F8A3" box="[338,501,1794,1818]" metricMagnitude="-3" metricUnit="m" metricValue="1.855" metricValueMax="1.88" metricValueMin="1.83" pageId="11" pageNumber="516" unit="mm" value="1.855" valueMax="1.88" valueMin="1.83">1.83–1.88 mm</quantity>
(mean
<quantity id="420D159FFFBE0D6CB754C2043795F8A0" box="[585,689,1793,1818]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="11" pageNumber="516" unit="mm" value="1.85">1.85 mm</quantity>
).
</paragraph>
<paragraph id="854AB87AFFBE0D6CB5DAC2203745F848" blockId="11.[151,1437,1217,2034]" pageId="11" pageNumber="516">
Head broadest at very small eyes, HL
<quantity id="420D159FFFBE0D6CB762C2203600F887" box="[639,804,1829,1854]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="11" pageNumber="516" unit="mm" value="0.365" valueMax="0.38" valueMin="0.35">0.35–0.38 mm</quantity>
(mean
<quantity id="420D159FFFBE0D6CB661C22336C2F884" box="[892,998,1830,1854]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="11" pageNumber="516" unit="mm" value="0.37">0.37 mm</quantity>
), HW 0.40; tempora in lateral view as long as 5 × length of eye, only slightly convergent posterad and posteriorly bent mesad at a slightly obtuse angle. Punctures on frons and vertex very fine and shallow, inconspicuous; setae short, sparse and suberect. Antennae slender, AnL
<quantity id="420D159FFFBE0D6CB433C29434F5F813" box="[302,465,1937,1962]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" metricValueMax="10.3" metricValueMin="9.5" pageId="11" pageNumber="516" unit="mm" value="0.99" valueMax="1.03" valueMin="0.95">0.95–1.03 mm</quantity>
(mean
<quantity id="420D159FFFBE0D6CB739C29737AFF810" box="[548,651,1938,1962]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" pageId="11" pageNumber="516" unit="mm" value="0.99">0.99 mm</quantity>
), antennomeres I–VI each distinctly elongate, VII and VIII each about as long as broad, IX and X much larger than VIII, each slightly transverse, XI indistinctly longer than IX–X combined, nearly twice as long as broad.
</paragraph>
<caption id="D18AE8F2FFB90D6BB58AC043342DFAC1" httpUri="https://zenodo.org/record/1440301/files/figure.png" pageId="12" pageNumber="517" startId="12.[151,264,1350,1373]" targetBox="[161,1412,206,1316]" targetPageId="12">
<paragraph id="854AB87AFFB90D6BB58AC043342DFAC1" blockId="12.[151,1435,1350,1403]" pageId="12" pageNumber="517">
<emphasis id="B7816468FFB90D6BB58AC0433471FAE6" bold="true" box="[151,341,1350,1373]" pageId="12" pageNumber="517">FIGURES 30–32.</emphasis>
<taxonomicName id="42F5C3F9FFB90D6BB441C0423686FAE7" authority="Franz. Habitus" authorityName="Franz. Habitus" box="[348,930,1351,1373]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="pseudorobustus" subGenus="Corbulifer">
<emphasis id="B7816468FFB90D6BB441C04234FDFAE6" box="[348,473,1351,1372]" italics="true" pageId="12" pageNumber="517">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFB90D6BB4F7C0423771FAE6" box="[490,597,1351,1372]" italics="true" pageId="12" pageNumber="517">Corbulifer</emphasis>
)
<emphasis id="B7816468FFB90D6BB778C0423627FAE6" box="[613,771,1351,1372]" italics="true" pageId="12" pageNumber="517">pseudorobustus</emphasis>
Franz. Habitus
</taxonomicName>
of male (30), aedeagus in dorsal (31) and lateral (32) views.
</paragraph>
</caption>
<paragraph id="854AB87AFFB90D6BB5DAC0AC34D9F9B3" blockId="12.[151,1437,1448,2013]" pageId="12" pageNumber="517">
Pronotum rounded, broadest slightly behind anterior third; PL
<quantity id="420D159FFFB90D6BB69AC0AD310FFA7A" box="[903,1067,1448,1473]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="12" pageNumber="517" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50–0.53 mm</quantity>
(mean
<quantity id="420D159FFFB90D6BB162C0AD31C3FA7A" box="[1151,1255,1448,1473]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="12" pageNumber="517" unit="mm" value="0.51">0.51 mm</quantity>
), PW
<quantity id="420D159FFFB90D6BB031C0AD35E7FA5E" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="12" pageNumber="517" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48 mm</quantity>
(mean
<quantity id="420D159FFFB90D6BB404C0C934A7FA5F" box="[281,387,1484,1509]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="12" pageNumber="517" unit="mm" value="0.46">0.46 mm</quantity>
). Antebasal pits indistinct. Punctures on pronotal disc very fine and shallow, inconspicuous; setae short, sparse and suberect.
</paragraph>
<paragraph id="854AB87AFFB90D6BB5DAC31134F5F9CE" blockId="12.[151,1437,1448,2013]" pageId="12" pageNumber="517">
Elytra oval, broadest in front of middle; EL
<quantity id="420D159FFFB90D6BB7CAC3103665F996" box="[727,833,1557,1581]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="12" pageNumber="517" unit="mm" value="0.98">0.98 mm</quantity>
, EW
<quantity id="420D159FFFB90D6BB69AC31036D6F996" box="[903,1010,1557,1581]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="12" pageNumber="517" unit="mm" value="0.73">0.73 mm</quantity>
, EI 1.34; apices separately rounded. Punctures relatively large and dense but very shallow, inconspicuous; setae longer and denser than those on pronotum, suberect to erect.
</paragraph>
<paragraph id="854AB87AFFB90D6BB5DAC38530A5F95A" blockId="12.[151,1437,1448,2013]" pageId="12" pageNumber="517">
Metatibia (
<figureCitation id="1DCEA4FFFFB90D6BB459C38534BCF923" box="[324,408,1664,1689]" captionStart="FIGURES 55–65" captionStartId="24.[151,264,1652,1675]" captionTargetBox="[151,1421,248,1572]" captionTargetId="figure@24.[151,1436,248,1586]" captionTargetPageId="24" captionText="FIGURES 55–65. Male metatibia in dorsal view. Scydmaenus tamborinensis Franz (55), S. monstrosetibialis sp. n. (56), S. bellenderanus sp. n. (57), S. pseudorobustus Franz (58), S. kiwarrakianus sp. n. (59), S. gushi sp. n. (60), S. gibraltar sp. n. (61), S. eungellanus sp. n., specimen from Mount William (62) and from Peases Lookout (63), S. fishermontanus sp. n. (64), S. super sp. n. (65)." httpUri="https://zenodo.org/record/1440317/files/figure.png" pageId="12" pageNumber="517">Fig. 58</figureCitation>
) in dorsal view gradually and moderately broadened, broadest between middle and distal third, mesal margin concave in proximal 1/5, remaining portion nearly evenly convex, lateral (external) margin bisinuate with distinct submedian concavity; dorsolateral 'pocket' present, with mesal margin bearing long setae.
</paragraph>
<paragraph id="854AB87AFFB90D6BB5DAC3E936BBF8F7" blockId="12.[151,1437,1448,2013]" pageId="12" pageNumber="517">
Aedeagus (
<figureCitation id="1DCEA4FFFFB90D6BB45AC3E934EEF8BF" box="[327,458,1772,1797]" captionStart="FIGURES 30–32" captionStartId="12.[151,264,1350,1373]" captionTargetBox="[161,1412,206,1316]" captionTargetId="figure@12.[151,1436,193,1329]" captionTargetPageId="12" captionText="FIGURES 30–32. Scydmaenus (Corbulifer) pseudorobustus Franz. Habitus of male (30), aedeagus in dorsal (31) and lateral (32) views." httpUri="https://zenodo.org/record/1440301/files/figure.png" pageId="12" pageNumber="517">Figs 31–32</figureCitation>
) moderately slender; AeL
<quantity id="420D159FFFB90D6BB61EC3E83649F8BE" box="[771,877,1773,1797]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="12" pageNumber="517" unit="mm" value="0.43">0.43 mm</quantity>
; in dorsal view median lobe broadest near base, relatively strongly narrowing distad with bisinuate lateral margins and broad, subtrapezoidal apical portion; apical projections conspicuously long, bent dorsoproximad at a sharp angle.
</paragraph>
<paragraph id="854AB87AFFB90D6BB5DAC25D34BAF8CB" blockId="12.[151,1437,1448,2013]" box="[199,414,1880,1905]" pageId="12" pageNumber="517">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFB90D6BB5DAC27931FEF82F" box="[199,1242,1916,1941]" pageId="12" pageNumber="517" type="distribution">
<paragraph id="854AB87AFFB90D6BB5DAC27931FEF82F" blockId="12.[151,1437,1448,2013]" box="[199,1242,1916,1941]" pageId="12" pageNumber="517">
<emphasis id="B7816468FFB90D6BB5DAC2793444F82F" bold="true" box="[199,352,1916,1941]" pageId="12" pageNumber="517">Distribution.</emphasis>
<collectingRegion id="47317698FFB90D6BB47AC27934CFF82F" box="[359,491,1916,1941]" country="Australia" name="Queensland" pageId="12" pageNumber="517">Queensland</collectingRegion>
and
<collectingRegion id="47317698FFB90D6BB73FC27837C9F82F" box="[546,749,1916,1941]" country="Australia" name="New South Wales" pageId="12" pageNumber="517">
New South
<collectingCountry id="FDE2F8EAFFB90D6BB7BBC27937C9F82F" box="[678,749,1916,1941]" name="United Kingdom" pageId="12" pageNumber="517">Wales</collectingCountry>
</collectingRegion>
; central-eastern coast of
<collectingCountry id="FDE2F8EAFFB90D6BB11EC279314EF82F" box="[1027,1130,1916,1941]" name="Australia" pageId="12" pageNumber="517">Australia</collectingCountry>
(
<figureCitation id="1DCEA4FFFFB90D6BB164C27931E9F82F" box="[1145,1229,1916,1941]" captionStart="FIGURE 70" captionStartId="25.[151,250,1999,2021]" captionTargetBox="[166,1423,886,1948]" captionTargetId="figure@25.[151,1436,867,1960]" captionTargetPageId="25" captionText="FIGURE 70. Distribution of Corbulifer species." httpUri="https://zenodo.org/record/1440323/files/figure.png" pageId="12" pageNumber="517">Fig. 70</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFB90D6AB5DAC2A530BEFE3D" lastPageId="13" lastPageNumber="518" pageId="12" pageNumber="517" type="discussion">
<paragraph id="854AB87AFFB90D6AB5DAC2A530BEFE3D" blockId="12.[151,1437,1448,2013]" lastBlockId="13.[151,1436,151,392]" lastPageId="13" lastPageNumber="518" pageId="12" pageNumber="517">
<emphasis id="B7816468FFB90D6BB5DAC2A5341FF803" bold="true" box="[199,315,1952,1977]" pageId="12" pageNumber="517">Remarks.</emphasis>
Males of several species have the metatibiae modified in a similar way as those of
<taxonomicName id="42F5C3F9FFB90D6BB09AC2A53461F866" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="pseudorobustus">
<emphasis id="B7816468FFB90D6BB09AC2A53461F866" italics="true" pageId="12" pageNumber="517">S. pseudorobustus</emphasis>
</taxonomicName>
:
<taxonomicName id="42F5C3F9FFB90D6BB449C2C034D2F866" authorityName="Jałoszyński" authorityYear="2018" box="[340,502,1988,2012]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="eungellanus">
<emphasis id="B7816468FFB90D6BB449C2C034D2F866" box="[340,502,1988,2012]" italics="true" pageId="12" pageNumber="517">S. eungellanus</emphasis>
</taxonomicName>
,
<taxonomicName id="42F5C3F9FFB90D6BB719C2C037A7F866" authorityName="Jałoszyński" authorityYear="2018" box="[516,643,1988,2012]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="gibraltar">
<emphasis id="B7816468FFB90D6BB719C2C037A7F866" box="[516,643,1988,2012]" italics="true" pageId="12" pageNumber="517">S. gibraltar</emphasis>
</taxonomicName>
,
<taxonomicName id="42F5C3F9FFB90D6BB78DC2C0366AF866" authorityName="Jałoszyński" authorityYear="2018" box="[656,846,1988,2012]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="kiwarrakianus">
<emphasis id="B7816468FFB90D6BB78DC2C0366AF866" box="[656,846,1988,2012]" italics="true" pageId="12" pageNumber="517">S. kiwarrakianus</emphasis>
</taxonomicName>
and
<taxonomicName id="42F5C3F9FFB90D6BB69FC2C036FDF866" authorityName="Jałoszyński" authorityYear="2018" box="[898,985,1988,2012]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="gushi">
<emphasis id="B7816468FFB90D6BB69FC2C036FDF866" box="[898,985,1988,2012]" italics="true" pageId="12" pageNumber="517">S. gushi</emphasis>
</taxonomicName>
(all described as new). All of them have the metatibiae gradually swollen and with a dorsolateral 'pocket' which divides the tibia into proximal and distal portions. In
<taxonomicName id="42F5C3F9FFB80D6AB43DC5B934CFFF6E" box="[288,491,188,212]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="pseudorobustus">
<emphasis id="B7816468FFB80D6AB43DC5B934CFFF6E" box="[288,491,188,212]" italics="true" pageId="13" pageNumber="518">S. pseudorobustus</emphasis>
</taxonomicName>
the lateral (external) metatibial margin is very weekly concave at the 'pocket', in all remaining similar species the concavity is much deeper, so that the tibia appears recurved. Moreover, in
<taxonomicName id="42F5C3F9FFB80D6AB09AC5E53462FEA6" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="pseudorobustus">
<emphasis id="B7816468FFB80D6AB09AC5E53462FEA6" italics="true" pageId="13" pageNumber="518">S. pseudorobustus</emphasis>
</taxonomicName>
the proximal portion of metatibia is distinctly longer than the distal one, and consequently the tibia is broadest in its distal half. Only in males of
<taxonomicName id="42F5C3F9FFB80D6AB78FC42D3611FEFA" authorityName="Jałoszyński" authorityYear="2018" box="[658,821,296,320]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="eungellanus">
<emphasis id="B7816468FFB80D6AB78FC42D3611FEFA" box="[658,821,296,320]" italics="true" pageId="13" pageNumber="518">S. eungellanus</emphasis>
</taxonomicName>
the proximal portion of metatibia is indistinctly longer than the distal portion, but adults of this species have the metatibiae of a different shape than those in
<taxonomicName id="42F5C3F9FFB80D6AB09AC4493461FE32" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="518" phylum="Arthropoda" rank="species" species="pseudorobustus">
<emphasis id="B7816468FFB80D6AB09AC4493461FE32" italics="true" pageId="13" pageNumber="518">S. pseudorobustus</emphasis>
</taxonomicName>
. Aedeagi of all these species differ markedly in the shape of the median lobe and apical structures.
</paragraph>
</subSubSection>
</treatment>
</document>
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