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<document id="134B0A6B154CBD7042F57733371F2E26" ID-CLB-Dataset="29444" ID-DOI="10.11646/zootaxa.4472.3.5" ID-GBIF-Dataset="fdbc45bb-a0df-4472-8cb7-ca718f01b8d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440287" ID-ZooBank="0BAB122B-66C0-4C30-B1FD-BC9E61B8F115" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538372442823" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2018" docId="0D5C096CFFB40D62B58AC0E835C5F84E" docLanguage="en" docName="zootaxa.4472.3.5.pdf" docOrigin="Zootaxa 4472 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scydmaenus (Corbulifer) Franz" docType="treatment" docVersion="7" lastPageNumber="510" masterDocId="F1657114FFB50D67B51DC5053524FFBA" masterDocTitle="' Monster Scydmaenus' of Australia: revision of subgenus Corbulifer Franz (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="531" masterPageNumber="505" pageNumber="506" updateTime="1698559125286" updateUser="ExternalLinkService">
<mods:mods id="4655BBDD08C9130D02EE66E95AB36E1C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="50925C46ED1FFBF32D462FA4FBE11D20">' Monster Scydmaenus' of Australia: revision of subgenus Corbulifer Franz (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="45C50826E592431395CF1FCA97226E6A">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="89CE63D1F9D4843918D6F70FD56D2288">Zootaxa</mods:title>
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<mods:part id="36082249892B8D0D035A01FC7C4E0147">
<mods:date id="367D052813CDAED20AC0762C79938A0F">2018</mods:date>
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<mods:number id="E795173993806AC242031CE67C67C5C2">2018-09-11</mods:number>
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<mods:number id="F0D93A1933DC7303D3C9C419EFBEDCE1">4472</mods:number>
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<mods:identifier id="8C0AC384B50875343CF57602710007CA" type="DOI">10.11646/zootaxa.4472.3.5</mods:identifier>
<mods:identifier id="CA90645B32D9C505F7D4943FD7470AE2" type="GBIF-Dataset">fdbc45bb-a0df-4472-8cb7-ca718f01b8d6</mods:identifier>
<mods:identifier id="ACF767648322DBA0B56474740EBE7E6C" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0D5C096CFFB40D62B58AC0E835C5F84E" ID-DOI="http://doi.org/10.5281/zenodo.5975890" ID-GBIF-Taxon="148421403" ID-Zenodo-Dep="5975890" LSID="urn:lsid:plazi:treatment:0D5C096CFFB40D62B58AC0E835C5F84E" httpUri="http://treatment.plazi.org/id/0D5C096CFFB40D62B58AC0E835C5F84E" lastPageId="5" lastPageNumber="510" pageId="1" pageNumber="506">
<subSubSection id="CDEFEBF1FFB40D66B58AC0E83451F9DD" pageId="1" pageNumber="506" type="nomenclature">
<paragraph id="854AB87AFFB40D66B58AC0E8373EF9BD" blockId="1.[151,538,1517,1543]" box="[151,538,1517,1543]" pageId="1" pageNumber="506">
<heading id="DE020F16FFB40D66B58AC0E8373EF9BD" bold="true" box="[151,538,1517,1543]" fontSize="11" level="1" pageId="1" pageNumber="506" reason="1">
<taxonomicName id="42F5C3F9FFB40D66B58AC0E8373EF9BD" ID-CoL="8XSCS" authority="Franz" authorityName="Franz" box="[151,538,1517,1543]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="subGenus" subGenus="Corbulifer">
<emphasis id="B7816468FFB40D66B58AC0E8373EF9BD" bold="true" box="[151,538,1517,1543]" pageId="1" pageNumber="506">
<emphasis id="B7816468FFB40D66B58AC0E8340AF9BD" bold="true" box="[151,302,1517,1543]" italics="true" pageId="1" pageNumber="506">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFB40D66B422C0E8349AF9BD" bold="true" box="[319,446,1517,1543]" italics="true" pageId="1" pageNumber="506">Corbulifer</emphasis>
) Franz
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="854AB87AFFB40D66B58AC3363451F9DD" blockId="1.[151,1436,1586,1639]" pageId="1" pageNumber="506">
<taxonomicName id="42F5C3F9FFB40D66B58AC3363700F9F2" authority="(Corbulifer Franz, 1975)" authorityName="Corbulifer Franz" authorityYear="1975" baseAuthorityName="Corbulifer Franz" baseAuthorityYear="1975" box="[151,548,1586,1608]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B7816468FFB40D66B58AC3363430F9F2" box="[151,276,1587,1608]" italics="true" pageId="1" pageNumber="506">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFB40D66B43BC33634B5F9F2" box="[294,401,1587,1608]" italics="true" pageId="1" pageNumber="506">Corbulifer</emphasis>
Franz, 1975)
</taxonomicName>
: 286 (as subgenus of
<taxonomicName id="42F5C3F9FFB40D66B612C33636A8F9F2" baseAuthorityName="Corbulifer Franz" baseAuthorityYear="1975" box="[783,908,1587,1608]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B7816468FFB40D66B612C33636A8F9F2" box="[783,908,1587,1608]" italics="true" pageId="1" pageNumber="506">Scydmaenus</emphasis>
</taxonomicName>
). Type species:
<taxonomicName id="42F5C3F9FFB40D66B121C33635DEF9DD" authority="Franz, 1975" authorityName="Franz" authorityYear="1975" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="tamborinensis">
<emphasis id="B7816468FFB40D66B121C3363076F9F2" box="[1084,1362,1587,1608]" italics="true" pageId="1" pageNumber="506">Scydmaenus tamborinensis</emphasis>
Franz, 1975
</taxonomicName>
(des. orig.).
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFB40D66B58AC3913656F8A2" pageId="1" pageNumber="506" type="diagnosis">
<paragraph id="854AB87AFFB40D66B58AC3913656F8A2" blockId="1.[151,1436,1684,2033]" pageId="1" pageNumber="506">
<emphasis id="B7816468FFB40D66B58AC3913453F917" bold="true" box="[151,375,1684,1709]" pageId="1" pageNumber="506">Revised diagnosis.</emphasis>
<taxonomicName id="42F5C3F9FFB40D66B499C391372BF916" baseAuthorityName="Corbulifer Franz" baseAuthorityYear="1975" box="[388,527,1684,1708]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B7816468FFB40D66B499C391372BF916" box="[388,527,1684,1708]" italics="true" pageId="1" pageNumber="506">Scydmaenus</emphasis>
</taxonomicName>
with posteroventral region of head capsule (bearing posterior tentorial pits) laterally confluent with genae and postgenae; entirely obliterated notosternal sutures; present hypomeral grooves; lacking basal elytral foveae; apices of parameres free, forming a pair of apical projections; protarsi not broadened in males; and metatibiae strongly modified (broadened) in males.
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFB40D63B5DAC2213136FEA6" lastPageId="4" lastPageNumber="509" pageId="1" pageNumber="506" type="description">
<paragraph id="854AB87AFFB40D66B5DAC22135F2F876" blockId="1.[151,1436,1684,2033]" pageId="1" pageNumber="506">
<emphasis id="B7816468FFB40D66B5DAC2213450F887" bold="true" box="[199,372,1828,1853]" pageId="1" pageNumber="506">Redescription.</emphasis>
Body ~
<quantity id="420D159FFFB40D66B4C5C221373EF886" box="[472,538,1828,1852]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="506" unit="mm" value="2.0">2 mm</quantity>
in length, body shape (
<figureCitation id="1DCEA4FFFFB40D66B638C2213659F886" box="[805,893,1828,1853]" captionStart="FIGURES 17–23" captionStartId="7.[151,264,1798,1821]" captionTargetBox="[163,1423,274,1764]" captionTargetId="figure@7.[151,1436,263,1777]" captionTargetPageId="7" captionText="FIGURES 17–23. Scydmaenus (Corbulifer) tamborinensis Franz. Holotype of S. tamborinensis tamborinensis, habitus of male (17), aedeagus in dorsal (18) and lateral (19) views. Aedeagi of holotype of S. tamborinensis cunninghamensis (20–21) and S. tamborinensis maipotonensis (22–23) in dorsal (20, 22) and lateral (21, 23) views." httpUri="https://zenodo.org/record/1440295/files/figure.png" pageId="1" pageNumber="506">Figs 17</figureCitation>
,
<figureCitation id="1DCEA4FFFFB40D66B697C221368CF886" box="[906,936,1828,1852]" captionStart="FIGURES 24–26" captionStartId="9.[151,264,1802,1825]" captionTargetBox="[163,1422,843,1774]" captionTargetId="figure@9.[151,1436,831,1781]" captionTargetPageId="9" captionText="FIGURES 24–26. Scydmaenus (Corbulifer) monstrosetibialis sp. n. Habitus of male (24), aedeagus in dorsal (25) and lateral (26) views." httpUri="https://zenodo.org/record/1440297/files/figure.png" pageId="1" pageNumber="506">24</figureCitation>
,
<figureCitation id="1DCEA4FFFFB40D66B6A8C22136F0F886" box="[949,980,1828,1852]" captionStart="FIGURES 27–29" captionStartId="10.[151,264,1935,1958]" captionTargetBox="[163,1424,849,1912]" captionTargetId="figure@10.[151,1436,837,1924]" captionTargetPageId="10" captionText="FIGURES 27–29. Scydmaenus (Corbulifer) bellenderanus sp. n. Habitus of male (27), aedeagus in dorsal (28) and lateral (29) views." httpUri="https://zenodo.org/record/1440299/files/figure.png" pageId="1" pageNumber="506">27</figureCitation>
,
<figureCitation id="1DCEA4FFFFB40D66B6FDC22136DAF886" box="[992,1022,1828,1852]" captionStart="FIGURES 30–32" captionStartId="12.[151,264,1350,1373]" captionTargetBox="[161,1412,206,1316]" captionTargetId="figure@12.[151,1436,193,1329]" captionTargetPageId="12" captionText="FIGURES 30–32. Scydmaenus (Corbulifer) pseudorobustus Franz. Habitus of male (30), aedeagus in dorsal (31) and lateral (32) views." httpUri="https://zenodo.org/record/1440301/files/figure.png" pageId="1" pageNumber="506">30</figureCitation>
,
<figureCitation id="1DCEA4FFFFB40D66B116C221310DF886" box="[1035,1065,1828,1852]" captionStart="FIGURES 33–35" captionStartId="14.[151,264,1966,1989]" captionTargetBox="[163,1418,825,1920]" captionTargetId="figure@14.[151,1436,812,1932]" captionTargetPageId="14" captionText="FIGURES 33–35. Scydmaenus (Corbulifer) kiwarrakianus sp. n. Habitus of male (33), aedeagus in dorsal (34) and lateral (35) views." httpUri="https://zenodo.org/record/1440303/files/figure.png" pageId="1" pageNumber="506">33</figureCitation>
,
<figureCitation id="1DCEA4FFFFB40D66B12BC2213171F887" box="[1078,1109,1828,1853]" captionStart="FIGURES 36–38" captionStartId="15.[151,264,1844,1867]" captionTargetBox="[164,1421,704,1816]" captionTargetId="figure@15.[151,1436,695,1823]" captionTargetPageId="15" captionText="FIGURES 36–38. Scydmaenus (Corbulifer) gushi sp. n. Habitus of male (36), aedeagus in dorsal (37) and lateral (38) views." httpUri="https://zenodo.org/record/1440305/files/figure.png" pageId="1" pageNumber="506">36</figureCitation>
,
<figureCitation id="1DCEA4FFFFB40D66B17CC221315BF886" box="[1121,1151,1828,1852]" captionStart="FIGURES 39–41" captionStartId="17.[151,264,1314,1337]" captionTargetBox="[155,1410,205,1280]" captionTargetId="figure@17.[151,1436,193,1293]" captionTargetPageId="17" captionText="FIGURES 39–41. Scydmaenus (Corbulifer) gibraltar sp. n. Habitus of male (39), aedeagus in dorsal (40) and lateral (41) views." httpUri="https://zenodo.org/record/1440307/files/figure.png" pageId="1" pageNumber="506">39</figureCitation>
,
<figureCitation id="1DCEA4FFFFB40D66B191C221318EF886" box="[1164,1194,1828,1852]" captionStart="FIGURES 42–44" captionStartId="18.[151,264,1933,1956]" captionTargetBox="[161,1423,810,1917]" captionTargetId="figure@18.[151,1436,798,1924]" captionTargetPageId="18" captionText="FIGURES 42–44. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Mount William. Habitus of male (42), aedeagus in dorsal (43) and lateral (44) views." httpUri="https://zenodo.org/record/1440309/files/figure.png" pageId="1" pageNumber="506">42</figureCitation>
,
<figureCitation id="1DCEA4FFFFB40D66B1AAC22131F2F887" box="[1207,1238,1828,1853]" captionStart="FIGURES 45–47" captionStartId="19.[151,264,1423,1446]" captionTargetBox="[151,1412,210,1401]" captionTargetId="figure@19.[151,1436,193,1403]" captionTargetPageId="19" captionText="FIGURES 45–47. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Peases Lookout. Habitus of male (45), aedeagus in dorsal (46) and lateral (47) views." httpUri="https://zenodo.org/record/1440311/files/figure.png" pageId="1" pageNumber="506">45</figureCitation>
,
<figureCitation id="1DCEA4FFFFB40D66B1FFC2213024F886" box="[1250,1280,1828,1852]" captionStart="FIGURES 48–50" captionStartId="21.[151,264,1270,1293]" captionTargetBox="[158,1428,203,1240]" captionTargetId="figure@21.[151,1436,193,1249]" captionTargetPageId="21" captionText="FIGURES 48–50. Scydmaenus (Corbulifer) fishermontanus sp. n. Habitus of male (48), aedeagus in dorsal (49) and lateral (50) views." httpUri="https://zenodo.org/record/1440313/files/figure.png" pageId="1" pageNumber="506">48</figureCitation>
,
<figureCitation id="1DCEA4FFFFB40D66B010C221300EF887" box="[1293,1322,1828,1853]" captionStart="FIGURES 51–54" captionStartId="23.[151,264,1954,1977]" captionTargetBox="[166,1427,900,1906]" captionTargetId="figure@23.[151,1436,886,1916]" captionTargetPageId="23" captionText="FIGURES 51–54. Scydmaenus (Corbulifer) super sp. n. Habitus of male (51), elytral apex of male (52), aedeagus in dorsal (53) and lateral (54) views." httpUri="https://zenodo.org/record/1440315/files/figure.png" pageId="1" pageNumber="506">51</figureCitation>
) uniform, 'ant-like', i.e., constricted between head and pronotum and between pronotum and elytra, with flattened head, moderately convex pronotum and moderately to strongly convex elytra; pigmentation uniformly brown, typically with slightly reddish or chestnut hue; vestiture of setae moderately dense and composed of thin, suberect to erect setae.
</paragraph>
<paragraph id="854AB87AFFB40D65B5DAC2DD3424FCE7" blockId="1.[151,1436,1684,2033]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="507" pageId="1" pageNumber="506">
Head (
<figureCitation id="1DCEA4FFFFB40D66B40EC2DD3471F84A" box="[275,341,2008,2033]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="1" pageNumber="506">Fig. 1</figureCitation>
) divided into a large exposed anterior part and a much smaller posterior 'neck' region, the latter retracted into prothorax. Anterior part short, subquadrate or indistinctly subtrapezoidal, flattened; eyes small, 2–5 times shorter than tempora, weakly convex, finely faceted, narrowly but clearly separated from mandibular bases; tempora weakly convergent posterad, anteriorly weakly rounded, posteriorly strongly bent mesad at an obtuse angle; vertex and frons confluent, convex, except for a shallow posteromedian impression on vertex; occipital constriction broader than half width of head. Labrum (
<figureCitation id="1DCEA4FFFFB70D65B7E4C4223619FEFA" box="[761,829,295,320]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 2</figureCitation>
) strongly transverse, anterior and lateral margins with nine shallow and broad emarginations, so that labral margin appears dentate; dorsal surface with paired, sparsely and symmetrically distributed setae of various lengths; ventral surface (epistome) with several (typically seven) long and thick sensilla inserted near labral base and one pair of much shorter peg-like sensilla inserted slightly behind the long ones. Mandibles (
<figureCitation id="1DCEA4FFFFB70D65B70AC4B2377FFE6A" box="[535,603,439,464]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 3</figureCitation>
) large and robust, subtriangular and slightly asymmetrical, each with one mesal tooth (
<figureCitation id="1DCEA4FFFFB70D65B43AC4D9344FFE4E" box="[295,363,476,501]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 3</figureCitation>
;
<emphasis id="B7816468FFB70D65B46AC4D834B7FE4E" box="[375,403,477,500]" italics="true" pageId="2" pageNumber="507">mt</emphasis>
) with bifurcate apex, larger and more deeply divided on the right mandible, setose prostheca (
<figureCitation id="1DCEA4FFFFB70D65B582C4FA35CCFDA2" box="[159,232,511,536]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 3</figureCitation>
;
<emphasis id="B7816468FFB70D65B5E7C704343FFDA2" box="[250,283,513,536]" italics="true" pageId="2" pageNumber="507">pst</emphasis>
) present and long, with its proximal half composed of particularly dense setae. Maxillae (
<figureCitation id="1DCEA4FFFFB70D65B051C4FA30B5FDA2" box="[1356,1425,511,536]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
) generalized, as in most
<taxonomicName id="42F5C3F9FFB70D65B481C7213766FD87" authorityName="Leach" authorityYear="1815" box="[412,578,548,573]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
, with short and broad cardo (
<figureCitation id="1DCEA4FFFFB70D65B698C72136E3FD86" box="[901,967,548,573]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
;
<emphasis id="B7816468FFB70D65B6C9C72136CAFD86" box="[980,1006,548,572]" italics="true" pageId="2" pageNumber="507">cd</emphasis>
), subtriangular basistipes (
<figureCitation id="1DCEA4FFFFB70D65B006C721307AFD86" box="[1307,1374,548,573]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
;
<emphasis id="B7816468FFB70D65B076C72130A8FD86" box="[1387,1420,548,572]" italics="true" pageId="2" pageNumber="507">bst</emphasis>
), elongate mediostipes (
<figureCitation id="1DCEA4FFFFB70D65B48EC74234F3FDDA" box="[403,471,583,608]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
;
<emphasis id="B7816468FFB70D65B4FBC74C3729FDDA" box="[486,525,585,608]" italics="true" pageId="2" pageNumber="507">mst</emphasis>
), elongate palpifer (
<figureCitation id="1DCEA4FFFFB70D65B7EEC7423613FDDA" box="[755,823,583,608]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
;
<emphasis id="B7816468FFB70D65B658C74D364CFDDA" box="[837,872,584,608]" italics="true" pageId="2" pageNumber="507">ppf</emphasis>
) and elongate galea (
<figureCitation id="1DCEA4FFFFB70D65B141C7423184FDDA" box="[1116,1184,583,608]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
;
<emphasis id="B7816468FFB70D65B1B2C74D31F7FDDA" box="[1199,1235,584,608]" italics="true" pageId="2" pageNumber="507">gal</emphasis>
) and lacinia (
<figureCitation id="1DCEA4FFFFB70D65B073C7423582FD3E" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
;
<emphasis id="B7816468FFB70D65B5A9C76935F2FD3E" box="[180,214,620,644]" italics="true" pageId="2" pageNumber="507">lac</emphasis>
). Maxillary palps (
<figureCitation id="1DCEA4FFFFB70D65B4ADC76934D5FD3E" box="[432,497,620,645]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
) moderately large, not different from those in
<taxonomicName id="42F5C3F9FFB70D65B11FC76931A9FD3E" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[1026,1165,620,644]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB70D65B11FC76931A9FD3E" box="[1026,1165,620,644]" italics="true" pageId="2" pageNumber="507">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB70D65B189C76831F4FD3E" box="[1172,1232,621,644]" pageId="2" pageNumber="507" sensu="stricto">s. str.</taxonomicNameLabel>
Submentum (
<figureCitation id="1DCEA4FFFFB70D65B073C7693582FD12" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
;
<emphasis id="B7816468FFB70D65B5A8C79435C6FD12" box="[181,226,657,680]" italics="true" pageId="2" pageNumber="507">smn</emphasis>
) short and broad, anteriorly deeply emarginate; mentum (
<figureCitation id="1DCEA4FFFFB70D65B66BC78A369FFD12" box="[886,955,655,680]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
;
<emphasis id="B7816468FFB70D65B6D7C79436C8FD12" box="[970,1004,657,680]" italics="true" pageId="2" pageNumber="507">mn</emphasis>
) subtrapezoidal, slightly broader than long, anteriorly emarginate; prementum (
<figureCitation id="1DCEA4FFFFB70D65B76BC7B1379BFD76" box="[630,703,692,717]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 4</figureCitation>
;
<emphasis id="B7816468FFB70D65B7D2C7B037DBFD76" box="[719,767,693,716]" italics="true" pageId="2" pageNumber="507">pmn</emphasis>
) large, with anterior margin subtriangular and projecting anterad, insertions of labial palps broadly separated. Labial palpomere II (
<figureCitation id="1DCEA4FFFFB70D65B6CDC7D23131FD4A" box="[976,1045,727,752]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 4</figureCitation>
;
<emphasis id="B7816468FFB70D65B13DC7DD3160FD4A" box="[1056,1092,728,752]" italics="true" pageId="2" pageNumber="507">lp2</emphasis>
) much longer than I and much broader than III. Gular plate (
<figureCitation id="1DCEA4FFFFB70D65B4ECC7F93713FCAE" box="[497,567,764,789]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
;
<emphasis id="B7816468FFB70D65B755C7F83740FCAE" box="[584,612,765,788]" italics="true" pageId="2" pageNumber="507">gp</emphasis>
) broad and distinctly microreticulate, gular sutures indistinct; posterior tentorial pits (
<figureCitation id="1DCEA4FFFFB70D65B42BC61A345FFC82" box="[310,379,799,824]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 1</figureCitation>
;
<emphasis id="B7816468FFB70D65B497C624348AFC82" box="[394,430,801,824]" italics="true" pageId="2" pageNumber="507">ptp</emphasis>
) small, elongate, located in front of transverse impression demarcating the 'neck' region ventrally.
</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC66D300AFC1F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="507">
Antennae (
<figureCitation id="1DCEA4FFFFB70D65B459C66234ADFC3A" box="[324,393,871,896]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1418,1441]" captionTargetBox="[151,1436,193,1397]" captionTargetId="figure@3.[151,1436,193,1397]" captionTargetPageId="3" captionText="FIGURES 1–5. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="2" pageNumber="507">Fig. 5</figureCitation>
) shorter than body, slender, with antennomeres VII and VIII distinctly asymmetrical, with their mesal margins shorter than external lateral margins; antennomeres IX–XI forming a trimerous club.
</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC6B534B6FB8F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="507">Pronotum in dorsal view oval, broadest slightly behind anterior third, weakly elongate; all margins rounded, anterior and posterior pronotal corners strongly obtuse and rounded; pronotal base with two pairs of very small, shallow and often diffuse antebasal pits, distance between median pits several times larger than that between each median and lateral pit.</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC14534DFFB52" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="507">
Prosternum (
<figureCitation id="1DCEA4FFFFB70D65B441C13A3487FBE2" box="[348,419,1087,1112]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 6</figureCitation>
) with its basisternal part (
<figureCitation id="1DCEA4FFFFB70D65B7C0C13A3602FBE2" box="[733,806,1087,1112]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 6</figureCitation>
;
<emphasis id="B7816468FFB70D65B625C145367DFBE2" box="[824,857,1088,1112]" italics="true" pageId="2" pageNumber="507">bst</emphasis>
) very short, less then half as long as coxal part; prosternal process (
<figureCitation id="1DCEA4FFFFB70D65B46CC1613492FBC7" box="[369,438,1124,1149]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 6</figureCitation>
;
<emphasis id="B7816468FFB70D65B4DFC16034CDFBC6" box="[450,489,1125,1148]" italics="true" pageId="2" pageNumber="507">psp</emphasis>
) developed as a short longitudinal ridge, which is very weakly elevated and in intact specimens hidden between contiguous procoxae; notosternal sutures entirely absent, prosternum laterally completely fused with hypomera; hypomera (
<figureCitation id="1DCEA4FFFFB70D65B7B1C1A937D1FB7F" box="[684,757,1196,1221]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 6</figureCitation>
;
<emphasis id="B7816468FFB70D65B61BC1A93605FB7E" box="[774,801,1196,1220]" italics="true" pageId="2" pageNumber="507">hy</emphasis>
) lacking hypomeral ridges, but with distinct posterior hypomeral grooves (
<figureCitation id="1DCEA4FFFFB70D65B463C1CA34E6FB52" box="[382,450,1231,1256]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 6</figureCitation>
;
<emphasis id="B7816468FFB70D65B4D3C1D534CEFB52" box="[462,490,1232,1256]" italics="true" pageId="2" pageNumber="507">hg</emphasis>
).
</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC1F035FFFAEF" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="507">
Mesonotum (
<figureCitation id="1DCEA4FFFFB70D65B441C1F13484FAB6" box="[348,416,1268,1293]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 7</figureCitation>
) with strongly elongate mesoscutum (
<figureCitation id="1DCEA4FFFFB70D65B650C1F136B5FAB6" box="[845,913,1268,1293]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 7</figureCitation>
;
<emphasis id="B7816468FFB70D65B683C1F136E0FAB6" box="[926,964,1268,1292]" italics="true" pageId="2" pageNumber="507">sc2</emphasis>
) indistinctly delimited from subtriangular mesocutellum (
<figureCitation id="1DCEA4FFFFB70D65B459C01234ADFA8A" box="[324,393,1303,1328]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 7</figureCitation>
;
<emphasis id="B7816468FFB70D65B48BC01D34E7FA8A" box="[406,451,1304,1328]" italics="true" pageId="2" pageNumber="507">scl2</emphasis>
), scutoscutellar suture absent; in intact specimens scutellum not visible between elytral bases.
</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC05A36D0FA5F" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="507">
Mesoventrite (
<figureCitation id="1DCEA4FFFFB70D65B472C05A3493FAC2" box="[367,439,1375,1400]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 8</figureCitation>
) with narrow and sharply demarcated anterior ridge (
<figureCitation id="1DCEA4FFFFB70D65B121C05A31A2FAC2" box="[1084,1158,1375,1400]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 8</figureCitation>
;
<emphasis id="B7816468FFB70D65B185C0643195FAC2" box="[1176,1201,1377,1400]" italics="true" pageId="2" pageNumber="507">ar</emphasis>
); area behind ridge functioning as a procoxal rest (
<figureCitation id="1DCEA4FFFFB70D65B4F2C0813717FA26" box="[495,563,1412,1437]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 8</figureCitation>
;
<emphasis id="B7816468FFB70D65B722C0803740FA26" box="[575,612,1413,1436]" italics="true" pageId="2" pageNumber="507">pcr</emphasis>
) flattened and asetose, not divided at middle, posteriorly demarcated by a sharp stepwise transverse edge, region behind the edge impressed and densely setose; mesoventral intercoxal process (
<figureCitation id="1DCEA4FFFFB70D65B5E7C0C9341AFA5E" box="[250,318,1484,1509]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 8</figureCitation>
;
<emphasis id="B7816468FFB70D65B456C0C834A1FA5E" box="[331,389,1485,1508]" italics="true" pageId="2" pageNumber="507">msvp</emphasis>
) carinate, weakly elevated, with a distinct posterior tip.
</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC0F4314BF9B2" blockId="2.[151,1437,151,2013]" box="[199,1135,1519,1544]" pageId="2" pageNumber="507">Metanotum with alacristae reaching half length of metascutum; hind wings present.</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC311306BF9CF" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="507">
Metaventrite (
<figureCitation id="1DCEA4FFFFB70D65B47BC311348CF996" box="[358,424,1556,1581]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 8</figureCitation>
) slightly broader than long, with rounded sides; metanepisterna (
<figureCitation id="1DCEA4FFFFB70D65B19BC31131EFF996" box="[1158,1227,1556,1581]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 8</figureCitation>
;
<emphasis id="B7816468FFB70D65B1CAC3113035F996" box="[1239,1297,1556,1580]" italics="true" pageId="2" pageNumber="507">aest3</emphasis>
) very broad and shifted ventrad, entirely exposed in ventral view in intact specimens; metaventral intercoxal process (
<figureCitation id="1DCEA4FFFFB70D65B04DC33230B2F9EA" box="[1360,1430,1591,1616]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 8</figureCitation>
;
<emphasis id="B7816468FFB70D65B58AC35835E9F9CE" box="[151,205,1629,1652]" italics="true" pageId="2" pageNumber="507">mtvp</emphasis>
) short and broad, with small median lobe projecting caudad and lateral corners also projecting caudad.
</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC37A378CF906" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="507">
Elytra oval, lacking basal foveae and humeral denticles, anterior margin of elytron (adjacent to elytral articulating lobe (
<figureCitation id="1DCEA4FFFFB70D65B440C3A13485F906" box="[349,417,1700,1725]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 7</figureCitation>
;
<emphasis id="B7816468FFB70D65B4B3C3A134F4F906" box="[430,464,1700,1724]" italics="true" pageId="2" pageNumber="507">eal</emphasis>
)) slightly concave.
</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC3CC3127F8BF" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="507">
Legs moderately long and robust; metatrochanters (
<figureCitation id="1DCEA4FFFFB70D65B627C3C236A3F95A" box="[826,903,1735,1760]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 8</figureCitation>
;
<emphasis id="B7816468FFB70D65B681C3CD3698F95A" box="[924,956,1736,1760]" italics="true" pageId="2" pageNumber="507">tr3</emphasis>
) strongly elongate; other podites as in
<taxonomicName id="42F5C3F9FFB70D65B58AC3E93406F8BE" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[151,290,1772,1796]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB70D65B58AC3E93406F8BE" box="[151,290,1772,1796]" italics="true" pageId="2" pageNumber="507">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB70D65B435C3E83441F8BE" box="[296,357,1773,1796]" pageId="2" pageNumber="507" sensu="stricto">s. str.</taxonomicNameLabel>
, except for different male dimorphic structures (see below).
</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC20A3446F8CA" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="507">
Abdomen short, with first visible sternite clearly longer than second; suture between V and VI distinct; pygidium and propygidium (
<figureCitation id="1DCEA4FFFFB70D65B4C6C2313705F8F6" box="[475,545,1844,1869]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 9</figureCitation>
;
<emphasis id="B7816468FFB70D65B733C230376EF8F6" box="[558,586,1845,1868]" italics="true" pageId="2" pageNumber="507">pg</emphasis>
,
<emphasis id="B7816468FFB70D65B744C23037AAF8F6" box="[601,654,1845,1868]" italics="true" pageId="2" pageNumber="507">prpg</emphasis>
) exposed, propygidium with its posterodorsal margin forming lateral denticles or lobes.
</paragraph>
<paragraph id="854AB87AFFB70D65B5DAC2793021F867" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="507">
Spermatheca (
<figureCitation id="1DCEA4FFFFB70D65B476C27934E4F82E" box="[363,448,1916,1941]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 10</figureCitation>
) located deeply in abdomen, asymmetrical, nearly hemispherical with 'basal' surface impressed and bearing a broad subconical projection with distally located insertion of ductus spermathecae and a small weakly sclerotized accessory gland (
<figureCitation id="1DCEA4FFFFB70D65B769C2C137E3F866" box="[628,711,1988,2013]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1833,1856]" captionTargetBox="[151,1436,409,1813]" captionTargetId="figure@4.[151,1436,409,1813]" captionTargetPageId="4" captionText="FIGURES 6–10. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter." httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="2" pageNumber="507">Fig. 10</figureCitation>
;
<emphasis id="B7816468FFB70D65B7CEC2C037DFF866" box="[723,763,1989,2012]" italics="true" pageId="2" pageNumber="507">acg</emphasis>
) connected with spermatheca by a broad duct.
</paragraph>
<caption id="D18AE8F2FFB60D64B58AC08F37EEF9A1" httpUri="https://zenodo.org/record/1440289/files/figure.png" pageId="3" pageNumber="508" startId="3.[151,264,1418,1441]" targetBox="[151,1436,193,1397]" targetPageId="3">
<paragraph id="854AB87AFFB60D64B58AC08F37EEF9A1" blockId="3.[151,1436,1418,1563]" pageId="3" pageNumber="508">
<emphasis id="B7816468FFB60D64B58AC08F341DFA25" bold="true" box="[151,313,1418,1441]" pageId="3" pageNumber="508">FIGURES 1–5.</emphasis>
Morphological structures of
<taxonomicName id="42F5C3F9FFB60D64B743C08E36D2FA1A" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[606,1014,1418,1440]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="kiwarrakianus" status="sp. nov." subGenus="Corbulifer">
<emphasis id="B7816468FFB60D64B743C08E37FFFA1A" box="[606,731,1419,1440]" italics="true" pageId="3" pageNumber="508">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFB60D64B7F7C08E3671FA1A" box="[746,853,1419,1440]" italics="true" pageId="3" pageNumber="508">Corbulifer</emphasis>
)
<emphasis id="B7816468FFB60D64B67EC08E36D2FA1A" box="[867,1014,1419,1440]" italics="true" pageId="3" pageNumber="508">kiwarrakianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB60D64B6E6C08E3117FA1A" box="[1019,1075,1419,1440]" pageId="3" pageNumber="508" rank="species">
<emphasis id="B7816468FFB60D64B6E6C08E3117FA1A" bold="true" box="[1019,1075,1419,1440]" pageId="3" pageNumber="508">sp. n.</emphasis>
</taxonomicNameLabel>
Head in ventral view (1); labrum in dorsal view (2); mandibles in dorsal view (3); prementum in ventral view (4); distal portion of right antenna in dorsal view (5). Abbreviations: a6–11, antennomere VI–XI; bst, basistipes; cd, cardo; gal, galea; gp, gular plate; lac, lacinia; lp1–3, labial palp I–III; mn, mentum; mst, mediostipes; mt, mesal tooth; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum.
</paragraph>
</caption>
<paragraph id="854AB87AFFB60D64B5DAC34237A9F83A" blockId="3.[151,1436,1607,2029]" pageId="3" pageNumber="508">
Aedeagus (
<figureCitation id="1DCEA4FFFFB60D64B458C34234E3F9DA" box="[325,455,1607,1632]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1455,1478]" captionTargetBox="[151,1436,213,1417]" captionTargetId="figure@5.[151,1436,213,1417]" captionTargetPageId="5" captionText="FIGURES 11–16. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Right metatibia of male in dorsal (11) and ventral (12) views; left protarsus of male in dorsal view (13); enlarged spatulate setae on protarsus (14); aedeagus in laterodorsal view (15), details of aedeagal apex in laterodorsal view (16). Abbreviations: app, apical projection; bf, basal foramen; cp, copulatory piece; lf, lateral flange; ost, ostium; pm, paramere." httpUri="https://zenodo.org/record/1440293/files/figure.png" pageId="3" pageNumber="508">Figs 15–16</figureCitation>
) symmetrical, strongly elongate and curved or recurved in lateral view, basal foramen (
<figureCitation id="1DCEA4FFFFB60D64B582C36E35D7F93E" box="[159,243,1643,1668]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1455,1478]" captionTargetBox="[151,1436,213,1417]" captionTargetId="figure@5.[151,1436,213,1417]" captionTargetPageId="5" captionText="FIGURES 11–16. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Right metatibia of male in dorsal (11) and ventral (12) views; left protarsus of male in dorsal view (13); enlarged spatulate setae on protarsus (14); aedeagus in laterodorsal view (15), details of aedeagal apex in laterodorsal view (16). Abbreviations: app, apical projection; bf, basal foramen; cp, copulatory piece; lf, lateral flange; ost, ostium; pm, paramere." httpUri="https://zenodo.org/record/1440293/files/figure.png" pageId="3" pageNumber="508">Fig. 15</figureCitation>
;
<emphasis id="B7816468FFB60D64B41DC3693432F93E" box="[256,278,1644,1668]" italics="true" pageId="3" pageNumber="508">bf</emphasis>
) close to base of median lobe on its dorsal surface; ostium (
<figureCitation id="1DCEA4FFFFB60D64B6A3C36E3136F93E" box="[958,1042,1643,1668]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1455,1478]" captionTargetBox="[151,1436,213,1417]" captionTargetId="figure@5.[151,1436,213,1417]" captionTargetPageId="5" captionText="FIGURES 11–16. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Right metatibia of male in dorsal (11) and ventral (12) views; left protarsus of male in dorsal view (13); enlarged spatulate setae on protarsus (14); aedeagus in laterodorsal view (15), details of aedeagal apex in laterodorsal view (16). Abbreviations: app, apical projection; bf, basal foramen; cp, copulatory piece; lf, lateral flange; ost, ostium; pm, paramere." httpUri="https://zenodo.org/record/1440293/files/figure.png" pageId="3" pageNumber="508">Fig. 16</figureCitation>
;
<emphasis id="B7816468FFB60D64B103C369311BF939" box="[1054,1087,1644,1667]" italics="true" pageId="3" pageNumber="508">ost</emphasis>
) subapically on dorsal wall, in all species flanked by variously shaped but always strongly elevated and typically asymmetrical lateral flanges (
<figureCitation id="1DCEA4FFFFB60D64B582C3B635D0F976" box="[159,244,1715,1740]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1455,1478]" captionTargetBox="[151,1436,213,1417]" captionTargetId="figure@5.[151,1436,213,1417]" captionTargetPageId="5" captionText="FIGURES 11–16. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Right metatibia of male in dorsal (11) and ventral (12) views; left protarsus of male in dorsal view (13); enlarged spatulate setae on protarsus (14); aedeagus in laterodorsal view (15), details of aedeagal apex in laterodorsal view (16). Abbreviations: app, apical projection; bf, basal foramen; cp, copulatory piece; lf, lateral flange; ost, ostium; pm, paramere." httpUri="https://zenodo.org/record/1440293/files/figure.png" pageId="3" pageNumber="508">Fig. 16</figureCitation>
;
<emphasis id="B7816468FFB60D64B41CC3B13434F976" box="[257,272,1716,1740]" italics="true" pageId="3" pageNumber="508">lf</emphasis>
); parameres (
<figureCitation id="1DCEA4FFFFB60D64B4B3C3B63727F976" box="[430,515,1715,1740]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1455,1478]" captionTargetBox="[151,1436,213,1417]" captionTargetId="figure@5.[151,1436,213,1417]" captionTargetPageId="5" captionText="FIGURES 11–16. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Right metatibia of male in dorsal (11) and ventral (12) views; left protarsus of male in dorsal view (13); enlarged spatulate setae on protarsus (14); aedeagus in laterodorsal view (15), details of aedeagal apex in laterodorsal view (16). Abbreviations: app, apical projection; bf, basal foramen; cp, copulatory piece; lf, lateral flange; ost, ostium; pm, paramere." httpUri="https://zenodo.org/record/1440293/files/figure.png" pageId="3" pageNumber="508">Fig. 15</figureCitation>
;
<emphasis id="B7816468FFB60D64B70DC3B13716F971" box="[528,562,1716,1739]" italics="true" pageId="3" pageNumber="508">pm</emphasis>
) fused with aedeagus, but fusion sites marked by shallow furrows, apices of parameres free and forming apical projections (
<figureCitation id="1DCEA4FFFFB60D64B7AFC3D23622F94A" box="[690,774,1751,1776]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1455,1478]" captionTargetBox="[151,1436,213,1417]" captionTargetId="figure@5.[151,1436,213,1417]" captionTargetPageId="5" captionText="FIGURES 11–16. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Right metatibia of male in dorsal (11) and ventral (12) views; left protarsus of male in dorsal view (13); enlarged spatulate setae on protarsus (14); aedeagus in laterodorsal view (15), details of aedeagal apex in laterodorsal view (16). Abbreviations: app, apical projection; bf, basal foramen; cp, copulatory piece; lf, lateral flange; ost, ostium; pm, paramere." httpUri="https://zenodo.org/record/1440293/files/figure.png" pageId="3" pageNumber="508">Fig. 16</figureCitation>
;
<emphasis id="B7816468FFB60D64B60EC3DC3619F94A" box="[787,829,1753,1776]" italics="true" pageId="3" pageNumber="508">app</emphasis>
) of aedeagus, parameral subapical setae forming two longitudinal dorsal rows on aedeagal apex, additionally dorsal wall of median lobe slightly in front of ostium with oblique or transversely arcuate rows of setae; endophallic structures asymmetrical, some of the sclerites protruding from ostium, among them a subtriangular copulatory piece (
<figureCitation id="1DCEA4FFFFB60D64B622C24636B7F8E6" box="[831,915,1859,1884]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1455,1478]" captionTargetBox="[151,1436,213,1417]" captionTargetId="figure@5.[151,1436,213,1417]" captionTargetPageId="5" captionText="FIGURES 11–16. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Right metatibia of male in dorsal (11) and ventral (12) views; left protarsus of male in dorsal view (13); enlarged spatulate setae on protarsus (14); aedeagus in laterodorsal view (15), details of aedeagal apex in laterodorsal view (16). Abbreviations: app, apical projection; bf, basal foramen; cp, copulatory piece; lf, lateral flange; ost, ostium; pm, paramere." httpUri="https://zenodo.org/record/1440293/files/figure.png" pageId="3" pageNumber="508">Fig. 16</figureCitation>
;
<emphasis id="B7816468FFB60D64B682C241369EF8E1" box="[927,954,1860,1883]" italics="true" pageId="3" pageNumber="508">cp</emphasis>
), which is movable and can flip so that its tip can be directed dorsad or dorsoproximad.
</paragraph>
<paragraph id="854AB87AFFB60D63B5DAC28E3136FEA6" blockId="3.[151,1436,1607,2029]" lastBlockId="4.[151,1436,151,357]" lastPageId="4" lastPageNumber="509" pageId="3" pageNumber="508">
Sexual dimorphism profound; males of all species have strongly, often monstrously modified metatibiae (
<figureCitation id="1DCEA4FFFFB60D64B077C28E35FAF872" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1455,1478]" captionTargetBox="[151,1436,213,1417]" captionTargetId="figure@5.[151,1436,213,1417]" captionTargetPageId="5" captionText="FIGURES 11–16. Morphological structures of Scydmaenus (Corbulifer) kiwarrakianus sp. n. Right metatibia of male in dorsal (11) and ventral (12) views; left protarsus of male in dorsal view (13); enlarged spatulate setae on protarsus (14); aedeagus in laterodorsal view (15), details of aedeagal apex in laterodorsal view (16). Abbreviations: app, apical projection; bf, basal foramen; cp, copulatory piece; lf, lateral flange; ost, ostium; pm, paramere." httpUri="https://zenodo.org/record/1440293/files/figure.png" pageId="3" pageNumber="508">Figs 11–12</figureCitation>
,
<figureCitation id="1DCEA4FFFFB60D64B5F6C2AA3411F872" box="[235,309,1967,1992]" captionStart="FIGURES 55–65" captionStartId="24.[151,264,1652,1675]" captionTargetBox="[151,1421,248,1572]" captionTargetId="figure@24.[151,1436,248,1586]" captionTargetPageId="24" captionText="FIGURES 55–65. Male metatibia in dorsal view. Scydmaenus tamborinensis Franz (55), S. monstrosetibialis sp. n. (56), S. bellenderanus sp. n. (57), S. pseudorobustus Franz (58), S. kiwarrakianus sp. n. (59), S. gushi sp. n. (60), S. gibraltar sp. n. (61), S. eungellanus sp. n., specimen from Mount William (62) and from Peases Lookout (63), S. fishermontanus sp. n. (64), S. super sp. n. (65)." httpUri="https://zenodo.org/record/1440317/files/figure.png" pageId="3" pageNumber="508">55–65</figureCitation>
), which are abruptly or gradually broadened, and typically with a glandular dorsolateral, lateral or/ and ventral 'pocket'; in species lacking the 'pocket' a group of tiny pores can be seen in a submedian dorsal or dorsolateral region; 'pocket' is often accompanied by a group of conspicuously long setae. In one species also elytra are modified in males, with glandular subapical structures (a flattened area with a convexity, both bearing modified setae). Eyes in males are typically larger than those in females (except for one species). Protarsomeres in males are not broadened, and in both sexes tarsomeres I–III bear spatulate adhesive setae.
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFB10D63B5DAC4223632FEDF" pageId="4" pageNumber="509" type="distribution">
<paragraph id="854AB87AFFB10D63B5DAC4223632FEDF" blockId="4.[151,1436,151,357]" pageId="4" pageNumber="509">
<emphasis id="B7816468FFB10D63B5DAC422370FFEFA" bold="true" box="[199,555,295,320]" pageId="4" pageNumber="509">Distribution and composition.</emphasis>
<taxonomicName id="42F5C3F9FFB10D63B72FC42D378CFEFA" authorityName="Franz" authorityYear="1975" box="[562,680,296,320]" class="Insecta" family="Staphylinidae" genus="Corbulifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="genus">
<emphasis id="B7816468FFB10D63B72FC42D378CFEFA" box="[562,680,296,320]" italics="true" pageId="4" pageNumber="509">Corbulifer</emphasis>
</taxonomicName>
comprises 10 species (2 known previously and 8 described as new) distributed in forests along the eastern coast of
<collectingCountry id="FDE2F8EAFFB10D63B7BAC4493635FEDF" box="[679,785,332,357]" name="Australia" pageId="4" pageNumber="509">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="D18AE8F2FFB10D63B58AC22C317BF863" httpUri="https://zenodo.org/record/1440291/files/figure.png" pageId="4" pageNumber="509" startId="4.[151,264,1833,1856]" targetBox="[151,1436,409,1813]" targetPageId="4">
<paragraph id="854AB87AFFB10D63B58AC22C317BF863" blockId="4.[151,1436,1833,2009]" pageId="4" pageNumber="509">
<emphasis id="B7816468FFB10D63B58AC22C346EF885" bold="true" box="[151,330,1833,1856]" pageId="4" pageNumber="509">FIGURES 6–10.</emphasis>
Morphological structures of
<taxonomicName id="42F5C3F9FFB10D63B766C22F313CF885" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[635,1048,1834,1856]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="kiwarrakianus" status="sp. nov." subGenus="Corbulifer">
<emphasis id="B7816468FFB10D63B766C22F37DCF885" box="[635,760,1834,1855]" italics="true" pageId="4" pageNumber="509">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFB10D63B614C22F3650F885" box="[777,884,1834,1855]" italics="true" pageId="4" pageNumber="509">Corbulifer</emphasis>
)
<emphasis id="B7816468FFB10D63B698C22F313CF885" box="[901,1048,1834,1855]" italics="true" pageId="4" pageNumber="509">kiwarrakianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB10D63B13DC22F3178F885" box="[1056,1116,1834,1855]" pageId="4" pageNumber="509" rank="species">
<emphasis id="B7816468FFB10D63B13DC22F3178F885" bold="true" box="[1056,1116,1834,1855]" pageId="4" pageNumber="509">sp. n.</emphasis>
</taxonomicNameLabel>
Prothorax in ventral view (6); mesonotum and elytral base in dorsal view (7); pterothorax in ventral view (8); distal portion of abdomen in dorsal view (9); spermatheca (10). Abbreviations: acg, accessory gland; aest3, metanepisternum; ar, anterior ridge; bst, basisternum; eal, elytral articulating lobe; el, elytron; hg, hypomeral groove; hy, hypomeron; mscc, mesocoxal cavity; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; pcxc, procoxal cavity; pg, pygidium; pre, prepectus; prpg, propygidium; psp, prosternal process; sc2, mesoscutum; scl2, mesoscutellum; sp, spermatheca; tr3, metarochanter.
</paragraph>
</caption>
<caption id="D18AE8F2FFB00D62B58AC0AA36B4F998" httpUri="https://zenodo.org/record/1440293/files/figure.png" pageId="5" pageNumber="510" startId="5.[151,264,1455,1478]" targetBox="[151,1436,213,1417]" targetPageId="5">
<paragraph id="854AB87AFFB00D62B58AC0AA36B4F998" blockId="5.[151,1436,1455,1570]" pageId="5" pageNumber="510">
<emphasis id="B7816468FFB00D62B58AC0AA3476FA7F" bold="true" box="[151,338,1455,1478]" pageId="5" pageNumber="510">FIGURES 11–16.</emphasis>
Morphological structures of
<taxonomicName id="42F5C3F9FFB00D62B76BC0B53129FA7F" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[630,1037,1456,1478]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="kiwarrakianus" status="sp. nov." subGenus="Corbulifer">
<emphasis id="B7816468FFB00D62B76BC0B537D7FA7F" box="[630,755,1456,1477]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFB00D62B61FC0B53649FA7F" box="[770,877,1456,1477]" italics="true" pageId="5" pageNumber="510">Corbulifer</emphasis>
)
<emphasis id="B7816468FFB00D62B666C0B53129FA7F" box="[891,1037,1456,1477]" italics="true" pageId="5" pageNumber="510">kiwarrakianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB00D62B10EC0B5316FFA7F" box="[1043,1099,1456,1477]" pageId="5" pageNumber="510" rank="species">
<emphasis id="B7816468FFB00D62B10EC0B5316FFA7F" bold="true" box="[1043,1099,1456,1477]" pageId="5" pageNumber="510">sp. n.</emphasis>
</taxonomicNameLabel>
Right metatibia of male in dorsal (11) and ventral (12) views; left protarsus of male in dorsal view (13); enlarged spatulate setae on protarsus (14); aedeagus in laterodorsal view (15), details of aedeagal apex in laterodorsal view (16). Abbreviations: app, apical projection; bf, basal foramen; cp, copulatory piece; lf, lateral flange; ost, ostium; pm, paramere.
</paragraph>
</caption>
<subSubSection id="CDEFEBF1FFB00D62B5DAC34B35C5F84E" pageId="5" pageNumber="510" type="discussion">
<paragraph id="854AB87AFFB00D62B5DAC34B35C5F84E" blockId="5.[151,1436,1614,2036]" pageId="5" pageNumber="510">
<emphasis id="B7816468FFB00D62B5DAC34B341FF9DD" bold="true" box="[199,315,1614,1639]" pageId="5" pageNumber="510">Remarks.</emphasis>
Characters of the
<typeStatus id="5A4E06D8FFB00D62B70BC3553762F9D2" box="[534,582,1616,1640]" pageId="5" pageNumber="510">type</typeStatus>
species of
<taxonomicName id="42F5C3F9FFB00D62B7D0C34A367CF9DD" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[717,856,1615,1639]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB00D62B7D0C34A367CF9DD" box="[717,856,1615,1639]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB00D62B67EC3553685F9DD" box="[867,929,1616,1639]" pageId="5" pageNumber="510" sensu="stricto">s. str.</taxonomicNameLabel>
were studied and illustrated by Jałoszyński (2016).
<taxonomicName id="42F5C3F9FFB00D62B5E8C3763448F931" authorityName="Franz" authorityYear="1975" box="[245,364,1651,1675]" class="Insecta" family="Staphylinidae" genus="Corbulifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B7816468FFB00D62B5E8C3763448F931" box="[245,364,1651,1675]" italics="true" pageId="5" pageNumber="510">Corbulifer</emphasis>
</taxonomicName>
differs from
<taxonomicName id="42F5C3F9FFB00D62B712C37637BEF931" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[527,666,1651,1675]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB00D62B712C37637BEF931" box="[527,666,1651,1675]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB00D62B7BBC37137C3F931" box="[678,743,1652,1675]" pageId="5" pageNumber="510" sensu="stricto">s. str.</taxonomicNameLabel>
in the following characters: posteroventral region of head capsule, on which the posterior tentorial pits are located, laterally confluent with genae and postgenae (in
<taxonomicName id="42F5C3F9FFB00D62B58AC3BE3406F969" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[151,290,1723,1747]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB00D62B58AC3BE3406F969" box="[151,290,1723,1747]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB00D62B435C3B93447F969" box="[296,355,1724,1747]" pageId="5" pageNumber="510" sensu="stricto">s. str.</taxonomicNameLabel>
developed as an elevated 'platform' laterally demarcated by stepwise ridges); notosternal sutures entirely obliterated (in
<taxonomicName id="42F5C3F9FFB00D62B481C3DA3702F94D" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[412,550,1759,1783]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB00D62B481C3DA3702F94D" box="[412,550,1759,1783]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB00D62B733C3E5374EF94D" box="[558,618,1760,1783]" pageId="5" pageNumber="510" sensu="stricto">s. str.</taxonomicNameLabel>
vestiges of sutures can be seen at the anterior prosternal margin); elytra lacking basal foveae (in
<taxonomicName id="42F5C3F9FFB00D62B4AFC2063718F8A1" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[434,572,1795,1819]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB00D62B4AFC2063718F8A1" box="[434,572,1795,1819]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB00D62B75BC20137A7F8A1" box="[582,643,1796,1819]" pageId="5" pageNumber="510" sensu="stricto">s. str.</taxonomicNameLabel>
each elytron with a pair of small and in some species poorly visible foveae); apices of parameres free, forming a pair of apical projections (parameral apices fused with the median lobe in
<taxonomicName id="42F5C3F9FFB00D62B5A9C24E341AF8D9" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[180,318,1867,1891]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB00D62B5A9C24E341AF8D9" box="[180,318,1867,1891]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB00D62B458C24934A7F8D9" box="[325,387,1867,1891]" pageId="5" pageNumber="510" sensu="stricto">s. str.</taxonomicNameLabel>
); protarsi not broadened in males (in
<taxonomicName id="42F5C3F9FFB00D62B635C24E3697F8D9" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[808,947,1867,1891]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB00D62B635C24E3697F8D9" box="[808,947,1867,1891]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB00D62B6A4C24936D0F8D9" box="[953,1012,1868,1891]" pageId="5" pageNumber="510" sensu="stricto">s. str.</taxonomicNameLabel>
distinctly broadened); and metatibiae strongly modified in males (in
<taxonomicName id="42F5C3F9FFB00D62B715C26A37B6F83D" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[520,658,1903,1927]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB00D62B715C26A37B6F83D" box="[520,658,1903,1927]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB00D62B783C27537F9F83D" box="[670,733,1904,1927]" pageId="5" pageNumber="510" sensu="stricto">s. str.</taxonomicNameLabel>
unmodified). However, character variability remains poorly studied among species of
<taxonomicName id="42F5C3F9FFB00D62B4CFC2963779F811" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[466,605,1939,1963]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B7816468FFB00D62B4CFC2963779F811" box="[466,605,1939,1963]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFB00D62B777C2913788F811" box="[618,684,1940,1963]" pageId="5" pageNumber="510" sensu="stricto">s. str.</taxonomicNameLabel>
and all other subgenera of
<taxonomicName id="42F5C3F9FFB00D62B11EC29631A9F811" authorityName="Jałoszyński" authorityYear="2018" box="[1027,1165,1939,1963]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B7816468FFB00D62B11EC29631A9F811" box="[1027,1165,1939,1963]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
, and consequently the diagnosis of
<taxonomicName id="42F5C3F9FFB00D62B43AC2B234B9F875" authorityName="Franz" authorityYear="1975" box="[295,413,1975,1999]" class="Insecta" family="Staphylinidae" genus="Corbulifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B7816468FFB00D62B43AC2B234B9F875" box="[295,413,1975,1999]" italics="true" pageId="5" pageNumber="510">Corbulifer</emphasis>
</taxonomicName>
(and all other subgenera) as well as the subgeneric division of
<taxonomicName id="42F5C3F9FFB00D62B178C2B231CBF875" authorityName="Jałoszyński" authorityYear="2018" box="[1125,1263,1975,1999]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B7816468FFB00D62B178C2B231CBF875" box="[1125,1263,1975,1999]" italics="true" pageId="5" pageNumber="510">Scydmaenus</emphasis>
</taxonomicName>
may change in future.
</paragraph>
</subSubSection>
</treatment>
</document>
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