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<document id="B9976F94124E8D3B56D9D88000176A9C" ID-CLB-Dataset="29444" ID-DOI="10.11646/zootaxa.4472.3.5" ID-GBIF-Dataset="fdbc45bb-a0df-4472-8cb7-ca718f01b8d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440287" ID-ZooBank="0BAB122B-66C0-4C30-B1FD-BC9E61B8F115" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538372442823" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2018" docId="0D5C096CFFA70D73B58AC4B2368CFB52" docLanguage="en" docName="zootaxa.4472.3.5.pdf" docOrigin="Zootaxa 4472 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scydmaenus (Corbulifer) eungellanus Jałoszyński 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="525" masterDocId="F1657114FFB50D67B51DC5053524FFBA" masterDocTitle="' Monster Scydmaenus' of Australia: revision of subgenus Corbulifer Franz (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="531" masterPageNumber="505" pageNumber="523" updateTime="1698559125286" updateUser="ExternalLinkService">
<mods:mods id="39BBF09099FCF3608D7A52F750D85EA2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7740A0FC293D4B1653F054D4E4F93FA2">
<mods:title id="91A03FE7AC1B6BCD207222E6B1ECB8C6">' Monster Scydmaenus' of Australia: revision of subgenus Corbulifer Franz (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="369F7D1B74F673B1CDED3425D8C94B27" type="personal">
<mods:role id="0037F84DB2789929DCA29A07462597C0">
<mods:roleTerm id="CABAD5334AA306155B4F3C85089B4D9D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBAAE5A9227E87CD4DFD9B0923240455">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="94E2AD348F078B0C3EA0E9DF69917B1C">text</mods:typeOfResource>
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<mods:title id="6805C369BA7DAE24DC31E16836A9B6FB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9F35C8E0F2C3B94645E6A832B52761CB">
<mods:date id="07924164594EB6CA62610C65E21A9AD7">2018</mods:date>
<mods:detail id="92F4851030243F11C6114AE0A09EE596" type="pubDate">
<mods:number id="5D5D629810A613F35F8BC6DB446FA748">2018-09-11</mods:number>
</mods:detail>
<mods:detail id="1CB9C5EE374ECD1807AD972B2A42E486" type="volume">
<mods:number id="D03263F57F76B032688164C443BF0181">4472</mods:number>
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<mods:number id="93A163D22AF4C819506B54B989A33D1F">3</mods:number>
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<mods:start id="9C24CED7023EB06D39D29858D2BBE447">505</mods:start>
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</mods:extent>
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<mods:classification id="22B219484084D9643A5E6B57720B890C">journal article</mods:classification>
<mods:identifier id="65E55A3FDCCA25DA836DE6515E81F1D7" type="CLB-Dataset">29444</mods:identifier>
<mods:identifier id="7CCFA5B67ACD57E4A6440A35A52E3917" type="DOI">10.11646/zootaxa.4472.3.5</mods:identifier>
<mods:identifier id="2DBA6567672F4C6D4DEB2E108BB33ACF" type="GBIF-Dataset">fdbc45bb-a0df-4472-8cb7-ca718f01b8d6</mods:identifier>
<mods:identifier id="4E09FA1AE6527DC6586BC98E96596C95" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A28F42D75D545E01F6E0E4FA727EE535" type="Zenodo-Dep">1440287</mods:identifier>
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</mods:mods>
<treatment id="0D5C096CFFA70D73B58AC4B2368CFB52" ID-DOI="http://doi.org/10.5281/zenodo.5975912" ID-GBIF-Taxon="148421405" ID-Zenodo-Dep="5975912" LSID="urn:lsid:plazi:treatment:0D5C096CFFA70D73B58AC4B2368CFB52" httpUri="http://treatment.plazi.org/id/0D5C096CFFA70D73B58AC4B2368CFB52" lastPageId="20" lastPageNumber="525" pageId="18" pageNumber="523">
<subSubSection id="CDEFEBF1FFA70D75B58AC4B23481FE49" pageId="18" pageNumber="523" type="nomenclature">
<paragraph id="854AB87AFFA70D75B58AC4B23789FE68" blockId="18.[151,685,439,499]" box="[151,685,439,466]" pageId="18" pageNumber="523">
<heading id="DE020F16FFA70D75B58AC4B23789FE68" bold="true" box="[151,685,439,466]" fontSize="11" level="1" pageId="18" pageNumber="523" reason="1">
<emphasis id="B7816468FFA70D75B58AC4B23789FE68" bold="true" box="[151,685,439,466]" pageId="18" pageNumber="523">
<taxonomicName id="42F5C3F9FFA70D75B58AC4B23747FE6B" ID-CoL="79ZTV" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[151,611,439,466]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="523" phylum="Arthropoda" rank="species" species="eungellanus" status="sp. nov." subGenus="Corbulifer">
<emphasis id="B7816468FFA70D75B58AC4B2340AFE6B" bold="true" box="[151,302,439,465]" italics="true" pageId="18" pageNumber="523">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFA70D75B422C4B2349AFE6B" bold="true" box="[319,446,439,465]" italics="true" pageId="18" pageNumber="523">Corbulifer</emphasis>
)
<emphasis id="B7816468FFA70D75B4D2C4B23747FE6B" bold="true" box="[463,611,439,465]" italics="true" pageId="18" pageNumber="523">eungellanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFA70D75B776C4BD3789FE68" box="[619,685,440,466]" pageId="18" pageNumber="523" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="854AB87AFFA70D75B58AC4DF3481FE49" blockId="18.[151,685,439,499]" box="[151,421,474,499]" pageId="18" pageNumber="523">
(
<figureCitation id="1DCEA4FFFFA70D75B582C4DF3404FE49" box="[159,288,474,499]" captionStart-0="FIGURES 42–44" captionStart-1="FIGURES 45–47" captionStartId-0="18.[151,264,1933,1956]" captionStartId-1="19.[151,264,1423,1446]" captionTargetBox-0="[161,1423,810,1917]" captionTargetBox-1="[151,1412,210,1401]" captionTargetId-0="figure@18.[151,1436,798,1924]" captionTargetId-1="figure@19.[151,1436,193,1403]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURES 42–44. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Mount William. Habitus of male (42), aedeagus in dorsal (43) and lateral (44) views." captionText-1="FIGURES 45–47. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Peases Lookout. Habitus of male (45), aedeagus in dorsal (46) and lateral (47) views." httpUri-0="https://zenodo.org/record/1440309/files/figure.png" httpUri-1="https://zenodo.org/record/1440311/files/figure.png" pageId="18" pageNumber="523">Figs 42–47</figureCitation>
,
<figureCitation id="1DCEA4FFFFA70D75B436C4DF3450FE49" box="[299,372,474,499]" captionStart="FIGURES 55–65" captionStartId="24.[151,264,1652,1675]" captionTargetBox="[151,1421,248,1572]" captionTargetId="figure@24.[151,1436,248,1586]" captionTargetPageId="24" captionText="FIGURES 55–65. Male metatibia in dorsal view. Scydmaenus tamborinensis Franz (55), S. monstrosetibialis sp. n. (56), S. bellenderanus sp. n. (57), S. pseudorobustus Franz (58), S. kiwarrakianus sp. n. (59), S. gushi sp. n. (60), S. gibraltar sp. n. (61), S. eungellanus sp. n., specimen from Mount William (62) and from Peases Lookout (63), S. fishermontanus sp. n. (64), S. super sp. n. (65)." httpUri="https://zenodo.org/record/1440317/files/figure.png" pageId="18" pageNumber="523">62–63</figureCitation>
,
<figureCitation id="1DCEA4FFFFA70D75B462C4DE34B8FE49" box="[383,412,475,499]" captionStart="FIGURE 70" captionStartId="25.[151,250,1999,2021]" captionTargetBox="[166,1423,886,1948]" captionTargetId="figure@25.[151,1436,867,1960]" captionTargetPageId="25" captionText="FIGURE 70. Distribution of Corbulifer species." httpUri="https://zenodo.org/record/1440323/files/figure.png" pageId="18" pageNumber="523">70</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFA70D75B58AC72734ECFD55" pageId="18" pageNumber="523" type="materials_examined">
<paragraph id="854AB87AFFA70D75B58AC72734ECFD55" blockId="18.[151,1436,546,751]" pageId="18" pageNumber="523">
<materialsCitation id="359DB227FFA70D75B58AC727317AFD39" ID-GBIF-Occurrence="1913889488" collectingDate="1979-04-19" collectorName="Eungella N. P. &amp; Mount &amp; William &amp; G. B. Monteith &amp; Rainforest &amp; Sieved" country="Australia" elevation="1100" latitude="-21.02" location="QUEENSLAND" longLatPrecision="759" longitude="148.36" pageId="18" pageNumber="523" specimenCount="1" specimenCount-male="1" stateProvince="QUEENSLAND" typeStatus="holotype">
<emphasis id="B7816468FFA70D75B58AC7273686FD81" bold="true" box="[151,930,546,571]" pageId="18" pageNumber="523">
<typeStatus id="5A4E06D8FFA70D75B58AC72735F6FD81" box="[151,210,546,571]" pageId="18" pageNumber="523">Type</typeStatus>
material.
<typeStatus id="5A4E06D8FFA70D75B44BC72734DDFD81" box="[342,505,546,571]" pageId="18" pageNumber="523" type="holotype">HOLOTYPE</typeStatus>
: (
<collectingCountry id="FDE2F8EAFFA70D75B70FC727379EFD81" box="[530,698,546,571]" name="Australia" pageId="18" pageNumber="523">AUSTRALIA</collectingCountry>
,
<collectingRegion id="47317698FFA70D75B7D0C72736B2FD81" box="[717,918,546,571]" country="Australia" name="Queensland" pageId="18" pageNumber="523">QUEENSLAND</collectingRegion>
):
</emphasis>
<specimenCount id="93F373F3FFA70D75B6B3C72636E1FD80" box="[942,965,547,570]" pageId="18" pageNumber="523" type="male">♂</specimenCount>
, two labels: &quot;
<collectingRegion id="47317698FFA70D75B176C726300EFD81" box="[1131,1322,547,571]" country="Australia" name="Queensland" pageId="18" pageNumber="523">QUEENSLAND</collectingRegion>
(MEQ) /
<collectorName id="2800DDACFFA70D75B58AC7433411FDE4" box="[151,309,582,607]" pageId="18" pageNumber="523">Eungella N.P.</collectorName>
,
<collectorName id="2800DDACFFA70D75B459C74234ABFDE4" box="[324,399,583,606]" pageId="18" pageNumber="523">Mount</collectorName>
/
<collectorName id="2800DDACFFA70D75B4B7C743372CFDE5" box="[426,520,582,607]" pageId="18" pageNumber="523">William</collectorName>
,
<collectingDate id="E10F6752FFA70D75B70BC742378DFDE5" box="[534,681,583,607]" pageId="18" pageNumber="523" value="1979-04-19">
<date id="F14B9EBAFFA70D75B70BC742378DFDE5" box="[534,681,583,607]" pageId="18" pageNumber="523" value="1979-04-19">19 Apr 1979</date>
</collectingDate>
/
<collectorName id="2800DDACFFA70D75B7D8C742364FFDE5" box="[709,875,582,607]" pageId="18" pageNumber="523">G.B. Monteith</collectorName>
&quot; [white, printed], &quot;Q.M. BERLESATE NO. 42 /
<geoCoordinate id="E0C1DEBDFFA70D75B58AC76E35C1FD39" box="[151,229,619,643]" direction="south" orientation="latitude" pageId="18" pageNumber="523" precision="555" value="-21.02">21.02S</geoCoordinate>
<geoCoordinate id="E0C1DEBDFFA70D75B5E7C76F3473FD39" box="[250,343,618,643]" direction="east" orientation="longitude" pageId="18" pageNumber="523" precision="555" value="148.36">148.36E</geoCoordinate>
/
<collectorName id="2800DDACFFA70D75B495C76F3724FD39" box="[392,512,618,643]" pageId="18" pageNumber="523">Rainforest</collectorName>
,
<elevation id="0ED85F49FFA70D75B705C76E3741FD39" box="[536,613,619,643]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="523" unit="m" value="1100.0">
<quantity id="420D159FFFA70D75B705C76E3741FD39" box="[536,613,619,643]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="523" unit="m" value="1100.0">1100m</quantity>
</elevation>
/
<collectorName id="2800DDACFFA70D75B78BC76F37C6FD39" box="[662,738,618,643]" pageId="18" pageNumber="523">Sieved</collectorName>
litter&quot; [white, printed] (QM)
</materialsCitation>
.
<materialsCitation id="359DB227FFA70D75B16BC76F37CFFD1D" ID-GBIF-Occurrence="1913889487" collectingDate="1979-04-19" collectorName="Eungella N. P. &amp; Mount &amp; William &amp; G. B. Monteith &amp; Rainforest &amp; Sieved" country="Australia" elevation="1100" latitude="-21.02" location="QUEENSLAND" longLatPrecision="759" longitude="148.36" pageId="18" pageNumber="523" specimenCount="2" specimenCount-female="2" stateProvince="QUEENSLAND" typeStatus="paratype">
<emphasis id="B7816468FFA70D75B16BC76F3447FD1D" bold="true" pageId="18" pageNumber="523">
<typeStatus id="5A4E06D8FFA70D75B16BC76F303EFD39" box="[1142,1306,618,643]" pageId="18" pageNumber="523" type="paratype">PARATYPES</typeStatus>
(4 exx.):
<collectingRegion id="47317698FFA70D75B58AC78B347AFD1D" box="[151,350,654,679]" country="Australia" name="Queensland" pageId="18" pageNumber="523">QUEENSLAND</collectingRegion>
:
</emphasis>
<specimenCount id="93F373F3FFA70D75B471C78A348BFD1C" box="[364,431,655,679]" pageId="18" pageNumber="523" type="female">2 ♀♀</specimenCount>
, same data as for
<typeStatus id="5A4E06D8FFA70D75B79BC78B37CFFD1D" box="[646,747,654,679]" pageId="18" pageNumber="523" type="holotype">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation id="359DB227FFA70D75B7E7C78A34ECFD55" ID-GBIF-Occurrence="1913889477" collectingDate="1992-11-17" collectingDateMax="1993-04" collectingDateMin="1992-11-17" collectorName="C. Qld &amp; Sx &amp; Pease's Lkt &amp; Lookout &amp; Eungella &amp; D. Cook &amp; G. B. Monteith" country="Russia" elevation="900" location="RF" longitude="148.51666" pageId="18" pageNumber="523" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="93F373F3FFA70D75B7E7C78A3619FD1C" box="[762,829,655,679]" pageId="18" pageNumber="523" type="male">2 ♂♂</specimenCount>
, two labels: &quot;
<collectorName id="2800DDACFFA70D75B6C0C78B3102FD1D" box="[989,1062,654,679]" pageId="18" pageNumber="523">C.Qld</collectorName>
: 21°07'
<collectorName id="2800DDACFFA70D75B167C78A31BEFD1D" box="[1146,1178,655,679]" pageId="18" pageNumber="523">Sx</collectorName>
<geoCoordinate id="E0C1DEBDFFA70D75B187C78A3024FD1D" box="[1178,1280,655,679]" direction="east" orientation="longitude" pageId="18" pageNumber="523" precision="925" value="148.51666">148°31'E</geoCoordinate>
/
<collectorName id="2800DDACFFA70D75B001C78A30B8FD1D" box="[1308,1436,654,679]" pageId="18" pageNumber="523">Pease's Lkt</collectorName>
{i.e.,
<collectorName id="2800DDACFFA70D75B5C5C7B7341BFD71" box="[216,319,690,715]" pageId="18" pageNumber="523">Lookout</collectorName>
},
<collectorName id="2800DDACFFA70D75B44BC7B7349EFD71" box="[342,442,690,715]" pageId="18" pageNumber="523">Eungella</collectorName>
/
<collectingDate id="E10F6752FFA70D75B4CBC7B63777FD71" box="[470,595,690,715]" pageId="18" pageNumber="523" value="1992-11-17">
<date id="F14B9EBAFFA70D75B4CBC7B63777FD71" box="[470,595,690,715]" pageId="18" pageNumber="523" value="1992-11-17">17 Nov 92</date>
</collectingDate>
-mid
<collectingDate id="E10F6752FFA70D75B78FC7B63624FD71" box="[658,768,691,715]" pageId="18" pageNumber="523" value="1993-04">
<date id="F14B9EBAFFA70D75B78FC7B63624FD71" box="[658,768,691,715]" pageId="18" pageNumber="523" value="1993-04">Apr 1993</date>
</collectingDate>
/
<collectorName id="2800DDACFFA70D75B600C7B73650FD71" box="[797,884,690,715]" pageId="18" pageNumber="523">D.Cook</collectorName>
&amp;
<collectorName id="2800DDACFFA70D75B697C7B73107FD71" box="[906,1059,690,715]" pageId="18" pageNumber="523">G.B.Monteith</collectorName>
,
<elevation id="0ED85F49FFA70D75B13AC7B7314EFD71" box="[1063,1130,690,715]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="523" unit="m" value="900.0">
<quantity id="420D159FFFA70D75B13AC7B7314EFD71" box="[1063,1130,690,715]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="523" unit="m" value="900.0">900m</quantity>
</elevation>
/
<collectingCountry id="FDE2F8EAFFA70D75B19AC7B6318DFD70" box="[1159,1193,691,714]" name="Russia" pageId="18" pageNumber="523">RF</collectingCountry>
Intercept &amp; Pitfalls&quot; [white, printed] (QM, cPJ).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="D18AE8F2FFA70D75B58AC2883748F878" httpUri="https://zenodo.org/record/1440309/files/figure.png" pageId="18" pageNumber="523" startId="18.[151,264,1933,1956]" targetBox="[161,1423,810,1917]" targetPageId="18">
<paragraph id="854AB87AFFA70D75B58AC2883748F878" blockId="18.[151,1436,1933,1986]" pageId="18" pageNumber="523">
<emphasis id="B7816468FFA70D75B58AC288347EF819" bold="true" box="[151,346,1933,1956]" pageId="18" pageNumber="523">FIGURES 42–44.</emphasis>
<taxonomicName id="42F5C3F9FFA70D75B47BC28B37D6F819" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[358,754,1933,1955]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="523" phylum="Arthropoda" rank="species" species="eungellanus" status="sp. nov." subGenus="Corbulifer">
<emphasis id="B7816468FFA70D75B47BC28B34C7F819" box="[358,483,1934,1955]" italics="true" pageId="18" pageNumber="523">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFA70D75B4E5C28B3747F819" box="[504,611,1934,1955]" italics="true" pageId="18" pageNumber="523">Corbulifer</emphasis>
)
<emphasis id="B7816468FFA70D75B765C28B37D6F819" box="[632,754,1934,1955]" italics="true" pageId="18" pageNumber="523">eungellanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFA70D75B7E2C28B361AF819" box="[767,830,1934,1955]" pageId="18" pageNumber="523" rank="species">
<emphasis id="B7816468FFA70D75B7E2C28B361AF819" bold="true" box="[767,830,1934,1955]" pageId="18" pageNumber="523">sp. n.</emphasis>
</taxonomicNameLabel>
, specimen from Mount William. Habitus of male (42), aedeagus in dorsal (43) and lateral (44) views.
</paragraph>
</caption>
<caption id="D18AE8F2FFA60D74B58AC08A3748FA7F" httpUri="https://zenodo.org/record/1440311/files/figure.png" pageId="19" pageNumber="524" startId="19.[151,264,1423,1446]" targetBox="[151,1412,210,1401]" targetPageId="19">
<paragraph id="854AB87AFFA60D74B58AC08A3748FA7F" blockId="19.[151,1435,1423,1477]" pageId="19" pageNumber="524">
<emphasis id="B7816468FFA60D74B58AC08A347EFA1F" bold="true" box="[151,346,1423,1446]" pageId="19" pageNumber="524">FIGURES 45–47.</emphasis>
<taxonomicName id="42F5C3F9FFA60D74B478C09537D6FA1F" authority="Jałoszyński, 2018" authorityName="Jałoszyński" authorityYear="2018" box="[357,754,1424,1446]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="524" phylum="Arthropoda" rank="species" species="eungellanus" status="sp. nov." subGenus="Corbulifer">
<emphasis id="B7816468FFA60D74B478C09534C6FA1F" box="[357,482,1424,1445]" italics="true" pageId="19" pageNumber="524">Scydmaenus</emphasis>
(
<emphasis id="B7816468FFA60D74B4EAC0953746FA1F" box="[503,610,1424,1445]" italics="true" pageId="19" pageNumber="524">Corbulifer</emphasis>
)
<emphasis id="B7816468FFA60D74B76AC09537D6FA1F" box="[631,754,1424,1445]" italics="true" pageId="19" pageNumber="524">eungellanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACB2D913FFA60D74B7E3C0953619FA1F" box="[766,829,1424,1445]" pageId="19" pageNumber="524" rank="species">
<emphasis id="B7816468FFA60D74B7E3C0953619FA1F" bold="true" box="[766,829,1424,1445]" pageId="19" pageNumber="524">sp. n.</emphasis>
</taxonomicNameLabel>
, specimen from Peases Lookout. Habitus of male (45), aedeagus in dorsal (46) and lateral (47) views.
</paragraph>
</caption>
<subSubSection id="CDEFEBF1FFA60D74B5DAC0F434DAF8BC" pageId="19" pageNumber="524" type="diagnosis">
<paragraph id="854AB87AFFA60D74B5DAC0F434DAF8BC" blockId="19.[151,1437,1521,2014]" pageId="19" pageNumber="524">
<emphasis id="B7816468FFA60D74B5DAC0F43465F9B0" bold="true" box="[199,321,1521,1546]" pageId="19" pageNumber="524">Diagnosis.</emphasis>
Male metatibia gradually and moderately broadened, slightly broader than metafemur, broadest near distal third, mesal tibial margin slightly concave in proximal third, convex at middle and nearly straight in distal third, lateral (external) tibial margin convex both in proximal and distal halves but with a short and indistinctly concave portion near distal 1/4–1/5, with a distinct submedian concavity, dorsolateral glandular 'pocket' present, with its mesal margin bearing long but relatively sparse setae, dorsomedian portion of the 'pocket' developed as a large and nearly circular pit; aedeagus in dorsal view nearly parallel-sided in proximal 3/4, with indistinct sub-basal constriction, apical portion nearly parallel-sided; apical projections very long and bent dorsoproximad at a sharp angle.
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFA60D73B5DAC214367BFC1F" lastPageId="20" lastPageNumber="525" pageId="19" pageNumber="524" type="description">
<paragraph id="854AB87AFFA60D74B5DAC214362AF8F4" blockId="19.[151,1437,1521,2014]" pageId="19" pageNumber="524">
<emphasis id="B7816468FFA60D74B5DAC214347DF890" bold="true" box="[199,345,1809,1834]" pageId="19" pageNumber="524">Description.</emphasis>
Body of male (
<figureCitation id="1DCEA4FFFFA60D74B715C214377BF890" box="[520,607,1809,1834]" captionStart="FIGURES 42–44" captionStartId="18.[151,264,1933,1956]" captionTargetBox="[161,1423,810,1917]" captionTargetId="figure@18.[151,1436,798,1924]" captionTargetPageId="18" captionText="FIGURES 42–44. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Mount William. Habitus of male (42), aedeagus in dorsal (43) and lateral (44) views." httpUri="https://zenodo.org/record/1440309/files/figure.png" pageId="19" pageNumber="524">Figs 42</figureCitation>
,
<figureCitation id="1DCEA4FFFFA60D74B776C21437ACF890" box="[619,648,1809,1834]" captionStart="FIGURES 45–47" captionStartId="19.[151,264,1423,1446]" captionTargetBox="[151,1412,210,1401]" captionTargetId="figure@19.[151,1436,193,1403]" captionTargetPageId="19" captionText="FIGURES 45–47. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Peases Lookout. Habitus of male (45), aedeagus in dorsal (46) and lateral (47) views." httpUri="https://zenodo.org/record/1440311/files/figure.png" pageId="19" pageNumber="524">45</figureCitation>
) strongly convex, moderately dark brown, covered with setae slightly lighter than cuticle; BL
<quantity id="420D159FFFA60D74B4BFC2303761F8F7" box="[418,581,1845,1870]" metricMagnitude="-3" metricUnit="m" metricValue="2.0149999999999997" metricValueMax="2.05" metricValueMin="1.98" pageId="19" pageNumber="524" unit="mm" value="2.0149999999999997" valueMax="2.05" valueMin="1.98">1.98–2.05 mm</quantity>
(mean
<quantity id="420D159FFFA60D74B784C2333626F8F4" box="[665,770,1846,1870]" metricMagnitude="-3" metricUnit="m" metricValue="2.02" pageId="19" pageNumber="524" unit="mm" value="2.02">2.02 mm</quantity>
).
</paragraph>
<paragraph id="854AB87AFFA60D73B5DAC25C3176FF6F" blockId="19.[151,1437,1521,2014]" lastBlockId="20.[151,1437,151,1256]" lastPageId="20" lastPageNumber="525" pageId="19" pageNumber="524">
Head broadest at small eyes, HL
<quantity id="420D159FFFA60D74B745C25F3624F8CB" box="[600,768,1882,1906]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="19" pageNumber="524" unit="mm" value="0.415" valueMax="0.43" valueMin="0.4">0.40–0.43 mm</quantity>
(mean
<quantity id="420D159FFFA60D74B640C25F36EFF8C8" box="[861,971,1882,1906]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="19" pageNumber="524" unit="mm" value="0.41">0.41 mm</quantity>
), HW
<quantity id="420D159FFFA60D74B100C25F31E1F8CB" box="[1053,1221,1882,1906]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="19" pageNumber="524" unit="mm" value="0.405" valueMax="0.41" valueMin="0.4">0.40–0.41 mm</quantity>
(mean
<quantity id="420D159FFFA60D74B03EC25F30B4F8C8" box="[1315,1424,1882,1906]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="524" unit="mm" value="0.4">0.40 mm</quantity>
); tempora in lateral view about twice as long as eyes, only slightly convergent posterad and posteriorly bent mesad at a slightly obtuse angle. Punctures on frons and vertex very fine and shallow, inconspicuous; setae short, sparse and suberect. Antennae slender, AnL
<quantity id="420D159FFFA60D74B710C2C03794F867" box="[525,688,1989,2014]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" metricValueMax="1.05" metricValueMin="1.03" pageId="19" pageNumber="524" unit="mm" value="1.04" valueMax="1.05" valueMin="1.03">1.03–1.05 mm</quantity>
(mean
<quantity id="420D159FFFA60D74B619C2C33649F864" box="[772,877,1990,2014]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" pageId="19" pageNumber="524" unit="mm" value="1.04">1.04 mm</quantity>
), antennomeres I–VI each distinctly elongate, VII and VIII each about as long as broad, IX and X much larger than VIII, IX about as long as broad, X slightly transverse, XI indistinctly shorter than IX–X combined, about 1.8 × as long as broad.
</paragraph>
<paragraph id="854AB87AFFA10D73B5DAC5E535D8FEFA" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="525">
Pronotum rounded, broadest slightly behind anterior third; PL
<quantity id="420D159FFFA10D73B697C5DA36D4FF4D" box="[906,1008,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="20" pageNumber="525" unit="mm" value="0.58">0.58 mm</quantity>
, PW
<quantity id="420D159FFFA10D73B132C5DA31F6FF4D" box="[1071,1234,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="20" pageNumber="525" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.50–0.55 mm</quantity>
(mean
<quantity id="420D159FFFA10D73B035C5DA30B4FF42" box="[1320,1424,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="20" pageNumber="525" unit="mm" value="0.53">0.53 mm</quantity>
). Antebasal pits indistinct. Punctures on pronotal disc very fine and shallow, inconspicuous; setae short, sparse and suberect.
</paragraph>
<paragraph id="854AB87AFFA10D73B5DAC4493799FE16" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="525">
Elytra oval, broadest in front of middle; EL 1.00–
<quantity id="420D159FFFA10D73B61FC4493642FEDE" box="[770,870,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="525" unit="mm" value="1.05">1.05 mm</quantity>
(mean
<quantity id="420D159FFFA10D73B6A0C4493103FEDE" box="[957,1063,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="20" pageNumber="525" unit="mm" value="1.03">1.03 mm</quantity>
), EW
<quantity id="420D159FFFA10D73B16DC4493030FEDE" box="[1136,1300,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="9.0" metricValueMin="8.299999999999999" pageId="20" pageNumber="525" unit="mm" value="0.865" valueMax="0.9" valueMin="0.83">0.83–0.90 mm</quantity>
(mean
<quantity id="420D159FFFA10D73B076C44935E3FE32" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="20" pageNumber="525" unit="mm" value="0.84">0.84 mm</quantity>
), EI 1.17–1.27; apices separately rounded. Punctures small and superficial, barely noticeable; setae longer and denser than those on pronotum, suberect to erect.
</paragraph>
<paragraph id="854AB87AFFA10D73B5DAC4B23060FDDA" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="525">
Metatibia (
<figureCitation id="1DCEA4FFFFA10D73B45EC4B234E0FE6A" box="[323,452,439,464]" captionStart="FIGURES 55–65" captionStartId="24.[151,264,1652,1675]" captionTargetBox="[151,1421,248,1572]" captionTargetId="figure@24.[151,1436,248,1586]" captionTargetPageId="24" captionText="FIGURES 55–65. Male metatibia in dorsal view. Scydmaenus tamborinensis Franz (55), S. monstrosetibialis sp. n. (56), S. bellenderanus sp. n. (57), S. pseudorobustus Franz (58), S. kiwarrakianus sp. n. (59), S. gushi sp. n. (60), S. gibraltar sp. n. (61), S. eungellanus sp. n., specimen from Mount William (62) and from Peases Lookout (63), S. fishermontanus sp. n. (64), S. super sp. n. (65)." httpUri="https://zenodo.org/record/1440317/files/figure.png" pageId="20" pageNumber="525">Figs 62–63</figureCitation>
) in dorsal view gradually and moderately broadened, broadest near distal third, mesal margin concave in proximal third, convex at middle and nearly straight in distal third, lateral (external) margin convex both in proximal and distal halves but with a short indistinctly concave portion near distal 1/4–1/5, with a deep concavity near middle; dorsolateral 'pocket' present, with its mesal margin bearing long but sparse and relatively inconspicuous setae, dorsomedian portion of 'pocket' developed as a deep and nearly circular pit.
</paragraph>
<paragraph id="854AB87AFFA10D73B5DAC769371BFC82" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="525">
Aedeagus (
<figureCitation id="1DCEA4FFFFA10D73B45BC76934EDFD3E" box="[326,457,620,645]" captionStart="FIGURES 42–44" captionStartId="18.[151,264,1933,1956]" captionTargetBox="[161,1423,810,1917]" captionTargetId="figure@18.[151,1436,798,1924]" captionTargetPageId="18" captionText="FIGURES 42–44. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Mount William. Habitus of male (42), aedeagus in dorsal (43) and lateral (44) views." httpUri="https://zenodo.org/record/1440309/files/figure.png" pageId="20" pageNumber="525">Figs 43–44</figureCitation>
,
<figureCitation id="1DCEA4FFFFA10D73B4CBC7693739FD3F" box="[470,541,620,645]" captionStart="FIGURES 45–47" captionStartId="19.[151,264,1423,1446]" captionTargetBox="[151,1412,210,1401]" captionTargetId="figure@19.[151,1436,193,1403]" captionTargetPageId="19" captionText="FIGURES 45–47. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Peases Lookout. Habitus of male (45), aedeagus in dorsal (46) and lateral (47) views." httpUri="https://zenodo.org/record/1440311/files/figure.png" pageId="20" pageNumber="525">46–47</figureCitation>
) relatively slender; AeL
<quantity id="420D159FFFA10D73B621C7693681FD3E" box="[828,933,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="20" pageNumber="525" unit="mm" value="0.4">0.40 mm</quantity>
; in dorsal view median lobe similarly broad near base and near middle, with lateral margins only indistinctly concave in between, gradually and weakly narrowing distad to form an elongate subrectangular apical portion, with apical projections long and bent dorsoproximad at a sharp angle. A slight variability in aedeagal structures was found between specimens from Mount William (
<figureCitation id="1DCEA4FFFFA10D73B449C7F934F1FCAE" box="[340,469,764,789]" captionStart="FIGURES 42–44" captionStartId="18.[151,264,1933,1956]" captionTargetBox="[161,1423,810,1917]" captionTargetId="figure@18.[151,1436,798,1924]" captionTargetPageId="18" captionText="FIGURES 42–44. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Mount William. Habitus of male (42), aedeagus in dorsal (43) and lateral (44) views." httpUri="https://zenodo.org/record/1440309/files/figure.png" pageId="20" pageNumber="525">Figs 43–44</figureCitation>
) and those from the Peases Lookout (
<figureCitation id="1DCEA4FFFFA10D73B69FC7F93122FCAF" box="[898,1030,764,789]" captionStart="FIGURES 45–47" captionStartId="19.[151,264,1423,1446]" captionTargetBox="[151,1412,210,1401]" captionTargetId="figure@19.[151,1436,193,1403]" captionTargetPageId="19" captionText="FIGURES 45–47. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Peases Lookout. Habitus of male (45), aedeagus in dorsal (46) and lateral (47) views." httpUri="https://zenodo.org/record/1440311/files/figure.png" pageId="20" pageNumber="525">Figs 46–47</figureCitation>
); males from the latter site have the apical projections more strongly bent.
</paragraph>
<paragraph id="854AB87AFFA10D73B5DAC641367BFC1F" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="525">
Female. Similar to male, but with unmodified metatibiae and much smaller eyes; length of temple equals 5–5.5 lengths of eye in lateral view. BL
<quantity id="420D159FFFA10D73B703C66237A3FCC5" box="[542,647,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="20" pageNumber="525" unit="mm" value="2.05">2.05 mm</quantity>
; HL
<quantity id="420D159FFFA10D73B7DFC66D360DFCC5" box="[706,809,872,896]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="20" pageNumber="525" unit="mm" value="0.4">0.40 mm</quantity>
, HW
<quantity id="420D159FFFA10D73B673C66D36F2FCC5" box="[878,982,872,896]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="20" pageNumber="525" unit="mm" value="0.43">0.43 mm</quantity>
, AnL
<quantity id="420D159FFFA10D73B102C66D31ACFCC5" box="[1055,1160,872,896]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="20" pageNumber="525" unit="mm" value="0.98">0.98 mm</quantity>
; PL
<quantity id="420D159FFFA10D73B1A3C6623041FCC5" box="[1214,1381,871,896]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="20" pageNumber="525" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
, PW
<quantity id="420D159FFFA10D73B58AC6893419FC1E" box="[151,317,908,933]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="20" pageNumber="525" unit="mm" value="0.54" valueMax="0.55" valueMin="0.53">0.53–0.55 mm</quantity>
; EL
<quantity id="420D159FFFA10D73B46CC6893732FC1E" box="[369,534,908,932]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" metricValueMax="1.1" metricValueMin="1.08" pageId="20" pageNumber="525" unit="mm" value="1.09" valueMax="1.1" valueMin="1.08">1.08–1.10 mm</quantity>
, EW
<quantity id="420D159FFFA10D73B748C68937DEFC1E" box="[597,762,908,933]" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="8.8" metricValueMin="8.5" pageId="20" pageNumber="525" unit="mm" value="0.865" valueMax="0.88" valueMin="0.85">0.85–0.88 mm</quantity>
, EI 1.26.
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFA10D73B5DAC6AA36EFFC72" box="[199,971,943,968]" pageId="20" pageNumber="525" type="distribution">
<paragraph id="854AB87AFFA10D73B5DAC6AA36EFFC72" blockId="20.[151,1437,151,1256]" box="[199,971,943,968]" pageId="20" pageNumber="525">
<emphasis id="B7816468FFA10D73B5DAC6AA3444FC72" bold="true" box="[199,352,943,968]" pageId="20" pageNumber="525">Distribution.</emphasis>
<collectingRegion id="47317698FFA10D73B47AC6AA34CBFC72" box="[359,495,943,968]" country="Australia" name="Queensland" pageId="20" pageNumber="525">Queensland</collectingRegion>
; north-eastern coast of
<collectingCountry id="FDE2F8EAFFA10D73B7E9C6AA367FFC72" box="[756,859,943,968]" name="Australia" pageId="20" pageNumber="525">Australia</collectingCountry>
(
<figureCitation id="1DCEA4FFFFA10D73B677C6AA369AFC72" box="[874,958,943,968]" captionStart="FIGURE 70" captionStartId="25.[151,250,1999,2021]" captionTargetBox="[166,1423,886,1948]" captionTargetId="figure@25.[151,1436,867,1960]" captionTargetPageId="25" captionText="FIGURE 70. Distribution of Corbulifer species." httpUri="https://zenodo.org/record/1440323/files/figure.png" pageId="20" pageNumber="525">Fig. 70</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFA10D73B5DAC6D13614FC57" box="[199,816,980,1005]" pageId="20" pageNumber="525" type="etymology">
<paragraph id="854AB87AFFA10D73B5DAC6D13614FC57" blockId="20.[151,1437,151,1256]" box="[199,816,980,1005]" pageId="20" pageNumber="525">
<emphasis id="B7816468FFA10D73B5DAC6D1346AFC57" bold="true" box="[199,334,980,1005]" pageId="20" pageNumber="525">Etymology.</emphasis>
Locotypical, after Eungella National Park.
</paragraph>
</subSubSection>
<subSubSection id="CDEFEBF1FFA10D73B5DAC6F2368CFB52" pageId="20" pageNumber="525" type="discussion">
<paragraph id="854AB87AFFA10D73B5DAC6F234E2FBE2" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="525">
<emphasis id="B7816468FFA10D73B5DAC6F2341FFBAA" bold="true" box="[199,315,1015,1040]" pageId="20" pageNumber="525">Remarks.</emphasis>
Males of
<taxonomicName id="42F5C3F9FFA10D73B4DEC6FD37C7FBAA" authorityName="Jałoszyński" authorityYear="2018" box="[451,739,1016,1040]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="525" phylum="Arthropoda" rank="species" species="eungellanus">
<emphasis id="B7816468FFA10D73B4DEC6FD37C7FBAA" box="[451,739,1016,1040]" italics="true" pageId="20" pageNumber="525">Scydmaenus eungellanus</emphasis>
</taxonomicName>
can be easily distinguished from congeners with weakly broadened metatibiae on the basis of a nearly circular impression at base of the dorsolateral metatibial 'pocket', well-visible in dorsal view.
</paragraph>
<paragraph id="854AB87AFFA10D73B5DAC161368CFB52" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="525">
Differences in the aedeagi were found between specimens collected in two localities within the Eungella National Park (
<figureCitation id="1DCEA4FFFFA10D73B45AC18234EDFB1A" box="[327,457,1159,1184]" captionStart="FIGURES 42–44" captionStartId="18.[151,264,1933,1956]" captionTargetBox="[161,1423,810,1917]" captionTargetId="figure@18.[151,1436,798,1924]" captionTargetPageId="18" captionText="FIGURES 42–44. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Mount William. Habitus of male (42), aedeagus in dorsal (43) and lateral (44) views." httpUri="https://zenodo.org/record/1440309/files/figure.png" pageId="20" pageNumber="525">Figs 43–44</figureCitation>
vs. 46–47); some males have the apical projections bent more strongly than others. However, proportions of body parts (
<figureCitation id="1DCEA4FFFFA10D73B727C1A937B5FB7E" box="[570,657,1196,1221]" captionStart="FIGURES 42–44" captionStartId="18.[151,264,1933,1956]" captionTargetBox="[161,1423,810,1917]" captionTargetId="figure@18.[151,1436,798,1924]" captionTargetPageId="18" captionText="FIGURES 42–44. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Mount William. Habitus of male (42), aedeagus in dorsal (43) and lateral (44) views." httpUri="https://zenodo.org/record/1440309/files/figure.png" pageId="20" pageNumber="525">Figs 42</figureCitation>
,
<figureCitation id="1DCEA4FFFFA10D73B783C1A9379FFB7F" box="[670,699,1196,1221]" captionStart="FIGURES 45–47" captionStartId="19.[151,264,1423,1446]" captionTargetBox="[151,1412,210,1401]" captionTargetId="figure@19.[151,1436,193,1403]" captionTargetPageId="19" captionText="FIGURES 45–47. Scydmaenus (Corbulifer) eungellanus sp. n., specimen from Peases Lookout. Habitus of male (45), aedeagus in dorsal (46) and lateral (47) views." httpUri="https://zenodo.org/record/1440311/files/figure.png" pageId="20" pageNumber="525">45</figureCitation>
) and the metatibial modifications (
<figureCitation id="1DCEA4FFFFA10D73B155C1A931ECFB7F" box="[1096,1224,1196,1221]" captionStart="FIGURES 55–65" captionStartId="24.[151,264,1652,1675]" captionTargetBox="[151,1421,248,1572]" captionTargetId="figure@24.[151,1436,248,1586]" captionTargetPageId="24" captionText="FIGURES 55–65. Male metatibia in dorsal view. Scydmaenus tamborinensis Franz (55), S. monstrosetibialis sp. n. (56), S. bellenderanus sp. n. (57), S. pseudorobustus Franz (58), S. kiwarrakianus sp. n. (59), S. gushi sp. n. (60), S. gibraltar sp. n. (61), S. eungellanus sp. n., specimen from Mount William (62) and from Peases Lookout (63), S. fishermontanus sp. n. (64), S. super sp. n. (65)." httpUri="https://zenodo.org/record/1440317/files/figure.png" pageId="20" pageNumber="525">Figs 62–63</figureCitation>
) do not differ, and these specimens are interpreted here as belonging in the same species.
</paragraph>
</subSubSection>
</treatment>
</document>
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