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<document id="6B533EE7A1817274ABCDA10B96317FFB" ID-CLB-Dataset="29756" ID-DOI="10.11646/zootaxa.4442.3.12" ID-GBIF-Dataset="beac85c9-35b0-4510-b8f1-489649182af7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1303703" ID-ZooBank="9534D594-8AF3-4D43-ADF6-DB6A6F44DE17" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530598479361" checkinUser="plazi" docAuthor="Frolov, Andrey V. &amp; Montreuil, Olivier" docDate="2018" docId="0D063C199A53FFDA31F2EB50DFEDFC89" docLanguage="en" docName="zootaxa.4442.3.12.pdf" docOrigin="Zootaxa 4442 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Adorodocia hirsuta Frolov &amp; Montreuil 2018, new species" docType="treatment" docVersion="7" lastPageNumber="500" masterDocId="F13F44619A53FFD83165EF23DB18FF9E" masterDocTitle="A contribution to the revision of Adorodocia Brenske (Coleoptera: Scarabaeidae: Rutelinae: Adoretini): a new peculiar species from Ankaratra, Madagascar" masterLastPageNumber="500" masterPageNumber="498" pageNumber="498" updateTime="1698552308232" updateUser="ExternalLinkService">
<mods:mods id="EDAEE08FD5A1BD313070EE70C22F359F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="179A8690533B94293ABE8B6AA27B6B33">
<mods:title id="E675384150FDF4E45129BBBF84897869">A contribution to the revision of Adorodocia Brenske (Coleoptera: Scarabaeidae: Rutelinae: Adoretini): a new peculiar species from Ankaratra, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="78A33747352EFFEE5CE5BBFAC6661D0D" type="personal">
<mods:role id="FCD0EFA82CE6CAAE318CB02DF2983C0C">
<mods:roleTerm id="50BB9D0F635E8D20DB8193F8BF90C938">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A3CB36F9029AD53CE1DDB9D835853CA0">Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:name id="0CB1D01253DC2DDEB7E965FFFED7A68B" type="personal">
<mods:role id="C4ED3709EBADB3845177972F90520F61">
<mods:roleTerm id="A7FC518BCA5C65362C971E7167E42727">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A7715C46BF4A9C5C2A8BBC8819AE6849">Montreuil, Olivier</mods:namePart>
</mods:name>
<mods:typeOfResource id="2053F5102A63FA6A40E11A874481B9D9">text</mods:typeOfResource>
<mods:relatedItem id="617996996C90479C631D18E7D01B4D80" type="host">
<mods:titleInfo id="3DBAF748F512322B046B1F0208D0EDC6">
<mods:title id="38338F26DF0199BA3827CADB4567317D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B3FC62FB7F7616DCC74B9F81C54E1155">
<mods:date id="93F414F519FFBAF487FE168D5E7E2CFF">2018</mods:date>
<mods:detail id="E2A3258E2E7998DBE3E2C77D63AAA561" type="pubDate">
<mods:number id="6D41417D786782FAC0DD2DE8E48BD1B6">2018-07-03</mods:number>
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<mods:detail id="B85E73E2C5618FCFEF68F5E10B743DAA" type="volume">
<mods:number id="57D416103D376215D8B39BE9E4704EBF">4442</mods:number>
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<mods:number id="90909333E24F30E103196415198DDFDD">3</mods:number>
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<mods:classification id="FE4FAE5B7A30BD4AF196DD53718D1B62">journal article</mods:classification>
<mods:identifier id="C66E32D6F4470A78904743B8852CD5EA" type="CLB-Dataset">29756</mods:identifier>
<mods:identifier id="45DF99BFE0DE121E01D9F91F5771B055" type="DOI">10.11646/zootaxa.4442.3.12</mods:identifier>
<mods:identifier id="A92CD2392FEE88171D0B17390ECCEAD6" type="GBIF-Dataset">beac85c9-35b0-4510-b8f1-489649182af7</mods:identifier>
<mods:identifier id="0149DC522FA482122C6400696D742848" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="26CEB51F415EE5CC42F396B74EC30489" type="Zenodo-Dep">1303703</mods:identifier>
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</mods:mods>
<treatment id="0D063C199A53FFDA31F2EB50DFEDFC89" ID-DOI="http://doi.org/10.5281/zenodo.5975774" ID-GBIF-Taxon="150794985" ID-Zenodo-Dep="5975774" LSID="urn:lsid:plazi:treatment:0D063C199A53FFDA31F2EB50DFEDFC89" httpUri="http://treatment.plazi.org/id/0D063C199A53FFDA31F2EB50DFEDFC89" lastPageId="2" lastPageNumber="500" pageId="0" pageNumber="498">
<subSubSection id="CDB5DE849A53FFD831F2EB50D8B0FB13" box="[151,936,1139,1165]" pageId="0" pageNumber="498" type="nomenclature">
<paragraph id="85108D0F9A53FFD831F2EB50D8B0FB13" blockId="0.[151,1437,1139,2030]" box="[151,936,1139,1165]" pageId="0" pageNumber="498">
<heading id="DE583A639A53FFD831F2EB50D8B0FB13" bold="true" box="[151,936,1139,1165]" fontSize="11" level="1" pageId="0" pageNumber="498" reason="1">
<emphasis id="B7DB511D9A53FFD831F2EB50D8B0FB13" bold="true" box="[151,936,1139,1165]" pageId="0" pageNumber="498">
<taxonomicName id="42AFF68C9A53FFD831F2EB50D999FB13" ID-CoL="655VL" authority="Frolov &amp; Montreuil" authorityName="Frolov &amp; Montreuil" authorityYear="2018" box="[151,641,1139,1165]" class="Insecta" family="Scarabaeidae" genus="Adorodocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="498" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. nov.">
<emphasis id="B7DB511D9A53FFD831F2EB50DA98FB13" bold="true" box="[151,384,1139,1165]" italics="true" pageId="0" pageNumber="498">Adorodocia hirsuta</emphasis>
Frolov &amp; Montreuil
</taxonomicName>
,
<taxonomicNameLabel id="ACE8EC669A53FFD833EBEB50D806FB13" box="[654,798,1139,1165]" pageId="0" pageNumber="498" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="1D94918A9A53FFD83248EB50D887FB13" box="[813,927,1139,1165]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1899,1922]" captionTargetBox="[214,1392,231,1820]" captionTargetId="figure@1.[151,1436,217,1849]" captionTargetPageId="1" captionText="FIGURES 1–4. Adorodocia hirsuta Frolov &amp; Montreuil, new species, holotype. 1, habitus; 2, ventral side of metasternum; 3, aedeagus in left lateral view and parameres in dorsal view; 4, locality records of A. hirsuta Frolov &amp; Montreuil, new species (star) and other species of Adorodocia (circle) in Madagascar (from Montreuil &amp; Frolov 2014, modified). Scale bars = 1 mm for Figs. 1–3." httpUri="https://zenodo.org/record/1303705/files/figure.png" pageId="0" pageNumber="498">
Figs. 1–
<docIssue id="C1CCF2E09A53FFD832F5EB50D887FB13" box="[912,927,1139,1165]" pageId="0" pageNumber="498">3</docIssue>
</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CDB5DE849A53FFD831F2EBB5D82EFAA1" pageId="0" pageNumber="498" type="materials_examined">
<paragraph id="85108D0F9A53FFD831F2EBB5D82EFAA1" blockId="0.[151,1437,1139,2030]" pageId="0" pageNumber="498">
<materialsCitation id="35C787529A53FFD831F2EBB5DE95FB4D" ID-GBIF-Occurrence="1950907665" collectingDate="1951-09" collectingDateMax="1951-11-28" collectingDateMin="1951-09" collectorName="P. Viette &amp; Chasse &amp; Museum Paris &amp; Mission P." country="Madagascar" county="Massif de l'Ankaratra" elevation="1850" location="Foret d'Ambahona" municipality="Manjakatompo" pageId="0" pageNumber="498" specimenCount="2" stateProvince="Centre" typeStatus="holotype">
<emphasis id="B7DB511D9A53FFD831F2EBB5DA5BFB31" bold="true" box="[151,323,1174,1199]" pageId="0" pageNumber="498">
<typeStatus id="5A1433AD9A53FFD831F2EBB5DBCAFB31" box="[151,210,1174,1199]" pageId="0" pageNumber="498">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="5A1433AD9A53FFD83032EBB5DADAFB31" box="[343,450,1174,1199]" pageId="0" pageNumber="498" type="holotype">Holotype</typeStatus>
, male, labeled: “
<collectingCountry id="FDB8CD9F9A53FFD833E9EBB5D80EFB31" box="[652,790,1174,1199]" name="Madagascar" pageId="0" pageNumber="498">Madagascar</collectingCountry>
<collectingRegion id="476B43ED9A53FFD83246EBB4D868FB31" box="[803,880,1175,1199]" country="France" name="Centre" pageId="0" pageNumber="498">Centre</collectingRegion>
,
<collectingCounty id="6C71F5839A53FFD832E4EBB5DF9DFB31" box="[897,1157,1174,1199]" pageId="0" pageNumber="498">Massif de l’Ankaratra</collectingCounty>
,
<collectingMunicipality id="657417759A53FFD835F3EBB5DE58FB31" box="[1174,1344,1174,1199]" pageId="0" pageNumber="498">Manjakatompo</collectingMunicipality>
/
<location id="8070DBD49A53FFD83404EBB4DA34FB4D" LSID="urn:lsid:plazi:treatment:0D063C199A53FFDA31F2EB50DFEDFC89:8070DBD49A53FFD83404EBB4DA34FB4D" country="Madagascar" county="Massif de l'Ankaratra" municipality="Manjakatompo" name="Foret d'Ambahona" pageId="0" pageNumber="498" stateProvince="Centre">Forêt d’Ambahona</location>
, Alt.
<elevation id="0E826A3C9A53FFD8300BEB99DAA6FB4D" box="[366,446,1210,1235]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="0" pageNumber="498" unit="m" value="1850.0">
<quantity id="425720EA9A53FFD8300BEB99DAA6FB4D" box="[366,446,1210,1235]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="0" pageNumber="498" unit="m" value="1850.0">1850m</quantity>
</elevation>
,
<collectorName id="285AE8D99A53FFD830A8EB98D937FB4D" box="[461,559,1210,1235]" pageId="0" pageNumber="498">P. Viette</collectorName>
,
<collectorName id="285AE8D99A53FFD83358EB99D994FB4D" box="[573,652,1210,1235]" pageId="0" pageNumber="498">Chasse</collectorName>
n°62 du
<collectingDate id="E15552279A53FFD8339FEB99D860FB4D" box="[762,888,1210,1235]" pageId="0" pageNumber="498" value="1951-11-28">
<date id="F111ABCF9A53FFD8339FEB99D860FB4D" box="[762,888,1210,1235]" pageId="0" pageNumber="498" value="1951-11-28">28.11.1951</date>
</collectingDate>
/
<collectorName id="285AE8D99A53FFD832F7EB99DF21FB4D" box="[914,1081,1210,1235]" pageId="0" pageNumber="498">Muséum Paris</collectorName>
,
<collectorName id="285AE8D99A53FFD83523EB99DFA6FB4C" box="[1094,1214,1210,1235]" pageId="0" pageNumber="498">Mission P.</collectorName>
Viette
<collectingDate id="E15552279A53FFD83471EB99DE95FB4D" box="[1300,1421,1210,1235]" pageId="0" pageNumber="498" value="1951-09">
<date id="F111ABCF9A53FFD83471EB99DE95FB4D" box="[1300,1421,1210,1235]" pageId="0" pageNumber="498" value="1951-09">Sept. 1951</date>
</collectingDate>
</materialsCitation>
–
<materialsCitation id="35C787529A53FFD831F2EBFCD82EFAA1" ID-GBIF-Occurrence="1950907565" collectingDate="1952-03" country="Madagascar" county="Ankaratra" determinerName="Montreuil" location="Rakotovao" municipality="Manjakatompo" pageId="0" pageNumber="498" specimenCount="2" typeStatus="holotype">
<collectingDate id="E15552279A53FFD831F2EBFCDA0BFB69" box="[151,275,1246,1271]" pageId="0" pageNumber="498" value="1952-03">
<date id="F111ABCF9A53FFD831F2EBFCDA0BFB69" box="[151,275,1246,1271]" pageId="0" pageNumber="498" value="1952-03">Mars 1952</date>
</collectingDate>
/
<taxonomicName id="42AFF68C9A53FFD8304BEBFDD911FB69" authority="Frolov &amp; Montreuil, 2018" authorityName="Frolov &amp; Montreuil" authorityYear="2018" box="[302,521,1246,1271]" class="Insecta" family="Scarabaeidae" genus="Adorodocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="498" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. nov.">Adorodocia hirsuta</taxonomicName>
<taxonomicNameLabel id="ACE8EC669A53FFD83376EBFCD94AFB68" box="[531,594,1247,1270]" pageId="0" pageNumber="498" rank="species">n. sp.</taxonomicNameLabel>
Frolov &amp;
<determinerName id="70CC61CB9A53FFD833B4EBFDD859FB69" box="[721,833,1246,1271]" pageId="0" pageNumber="498">Montreuil</determinerName>
det.
<docYear id="9DC961F49A53FFD8321BEBFCD8AEFB69" box="[894,950,1247,1271]" pageId="0" pageNumber="498">2018</docYear>
/
<typeStatus id="5A1433AD9A53FFD832B7EBFDDF25FB69" box="[978,1085,1246,1271]" pageId="0" pageNumber="498" type="holotype">Holotype</typeStatus>
” (Muséum national d’Histoire naturelle, Paris).
<typeStatus id="5A1433AD9A53FFD83032EA20DAA1FA85" box="[343,441,1283,1307]" pageId="0" pageNumber="498" type="paratype">Paratype</typeStatus>
no. 1, male, labeled: “
<collectingCounty id="6C71F5839A53FFD833D0EA21D832FA85" box="[693,810,1282,1307]" pageId="0" pageNumber="498">Ankaratra</collectingCounty>
,
<collectingMunicipality id="657417759A53FFD83253EA21D8C7FA85" box="[822,991,1282,1307]" pageId="0" pageNumber="498">Manjakatompo</collectingMunicipality>
/ 1.52
<location id="8070DBD49A53FFD8354BEA21DFBFFA85" LSID="urn:lsid:plazi:treatment:0D063C199A53FFDA31F2EB50DFEDFC89:8070DBD49A53FFD8354BEA21DFBFFA85" box="[1070,1191,1282,1307]" country="Madagascar" county="Ankaratra" municipality="Manjakatompo" name="Rakotovao" pageId="0" pageNumber="498">Rakotovao</location>
/ Institut Scientifique
<collectingCountry id="FDB8CD9F9A53FFD831F2EA05DA39FAA1" box="[151,289,1318,1343]" name="Madagascar" pageId="0" pageNumber="498">Madagascar</collectingCountry>
” (
<location id="8070DBD49A53FFD8305CEA05DA85FAA0" LSID="urn:lsid:plazi:treatment:0D063C199A53FFDA31F2EB50DFEDFC89:8070DBD49A53FFD8305CEA05DA85FAA0" box="[313,413,1318,1342]" country="Madagascar" county="Ankaratra" municipality="Manjakatompo" name="Museum" pageId="0" pageNumber="498">Muséum</location>
national
<location id="8070DBD49A53FFD83360EA05D96EFAA1" LSID="urn:lsid:plazi:treatment:0D063C199A53FFDA31F2EB50DFEDFC89:8070DBD49A53FFD83360EA05D96EFAA1" box="[517,630,1318,1343]" country="Madagascar" county="Ankaratra" municipality="Manjakatompo" name="d'Histoire" pageId="0" pageNumber="498">d’Histoire</location>
naturelle,
<location id="8070DBD49A53FFD8338BEA05D833FAA1" LSID="urn:lsid:plazi:treatment:0D063C199A53FFDA31F2EB50DFEDFC89:8070DBD49A53FFD8338BEA05D833FAA1" box="[750,811,1318,1343]" country="Madagascar" county="Ankaratra" municipality="Manjakatompo" name="Paris" pageId="0" pageNumber="498">Paris</location>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CDB5DE849A53FFDA31A2EA69DAEEFEFB" lastPageId="2" lastPageNumber="500" pageId="0" pageNumber="498" type="description">
<paragraph id="85108D0F9A53FFD831A2EA69DE0DFA19" blockId="0.[151,1437,1139,2030]" pageId="0" pageNumber="498">
<emphasis id="B7DB511D9A53FFD831A2EA69DA49FAFD" bold="true" box="[199,337,1354,1379]" pageId="0" pageNumber="498">Description</emphasis>
. Male,
<typeStatus id="5A1433AD9A53FFD830CFEA69D914FAFD" box="[426,524,1354,1379]" pageId="0" pageNumber="498" type="holotype">holotype</typeStatus>
(
<figureCitation id="1D94918A9A53FFD83378EA69D979FAFD" box="[541,609,1354,1379]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1899,1922]" captionTargetBox="[214,1392,231,1820]" captionTargetId="figure@1.[151,1436,217,1849]" captionTargetPageId="1" captionText="FIGURES 1–4. Adorodocia hirsuta Frolov &amp; Montreuil, new species, holotype. 1, habitus; 2, ventral side of metasternum; 3, aedeagus in left lateral view and parameres in dorsal view; 4, locality records of A. hirsuta Frolov &amp; Montreuil, new species (star) and other species of Adorodocia (circle) in Madagascar (from Montreuil &amp; Frolov 2014, modified). Scale bars = 1 mm for Figs. 1–3." httpUri="https://zenodo.org/record/1303705/files/figure.png" pageId="0" pageNumber="498">Fig. 1</figureCitation>
). Body length
<quantity id="425720EA9A53FFD8326AEA69D860FAFC" box="[783,888,1354,1379]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="0" pageNumber="498" unit="mm" value="16.5">16.5 mm</quantity>
. Yellowish brown, elytra lighter, pronotum with complete, longitudinal light brown strip on each side and basal, short light brown mark more medially.
</paragraph>
<paragraph id="85108D0F9A53FFD831A2EAB1DE7CF989" blockId="0.[151,1437,1139,2030]" pageId="0" pageNumber="498">
<emphasis id="B7DB511D9A53FFD831A2EAB1DA1CFA35" box="[199,260,1426,1451]" italics="true" pageId="0" pageNumber="498">Head</emphasis>
. Clypeus flat, its regularly raised front margin semicircular. Punctures large and rather sparse, bearing short, adpressed setae. Frons flat, without any relief. Punctures large and sparse, bearing short and adpressed setae. Only one yellowish, long and raised seta near inner edge of each eye. Labrum with medial prominent truncated process. Antennal club elongated, much longer than funiculus but shorter than funiculus and scape combined.
</paragraph>
<paragraph id="85108D0F9A53FFD831A2E901DE5FF939" blockId="0.[151,1437,1139,2030]" pageId="0" pageNumber="498">
<emphasis id="B7DB511D9A53FFD831A2E901DA00F9A5" box="[199,280,1570,1595]" italics="true" pageId="0" pageNumber="498">Thorax</emphasis>
. Pronotal margins thick, especially lateral ones bearing a few long setae in their anterior part. Lateral borders regularly curved, anterolateral angles almost right, posterolateral angles obtuse. Basal margin prominent in middle. Punctation large and sparse, bearing short and adpressed setae similar to those on head. Prosternal apophyse strong between posterior edge of procoxae, its anterior edge forming a spine-shaped prominence.
</paragraph>
<paragraph id="85108D0F9A53FFD831A2E991D994F970" blockId="0.[151,1437,1139,2030]" pageId="0" pageNumber="498">
<taxonomicName id="42AFF68C9A53FFD831A2E991DA2DF955" box="[199,309,1714,1739]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="0" pageNumber="498" phylum="Arthropoda" rank="genus">
<emphasis id="B7DB511D9A53FFD831A2E991DA2DF955" box="[199,309,1714,1739]" italics="true" pageId="0" pageNumber="498">Scutellum</emphasis>
</taxonomicName>
. Smooth at base, with dense large punctures at apex and along sides, bearing short and adpressed setae similar to those on head and pronotum.
</paragraph>
<paragraph id="85108D0F9A53FFD831A2E9D9DFB5F8E0" blockId="0.[151,1437,1139,2030]" pageId="0" pageNumber="498">
<emphasis id="B7DB511D9A53FFD831A2E9D9DA15F88D" box="[199,269,1786,1811]" italics="true" pageId="0" pageNumber="498">Elytra</emphasis>
. Striae indistinguishable. Elytra with slightly raised, narrow stripes (“odd interstriae”), much narrower than flat “even interstriae”, however this sculpture is poorly defined. Punctation much denser than that on pronotum and head, punctures evenly distributed, bearing short, adpressed setae, a little longer and thick, scale-like setae near apex. Sides with a continuous row of numerous long setae being shorter near apex.
</paragraph>
<paragraph id="85108D0F9A53FFD831A2E8AEDE9CF870" blockId="0.[151,1437,1139,2030]" pageId="0" pageNumber="498">
<emphasis id="B7DB511D9A53FFD831A2E8AEDA28F83B" box="[199,304,1929,1957]" italics="true" pageId="0" pageNumber="498">8th tergite</emphasis>
. Strongly convex. Surface with rough aspect due to microreticulate tegument and superficial, dense, more or less coalescent punctures. Short adpressed setae present only basally, in apical part and on sides; disc glabrous. A few long, flexuous, raised setae in middle of apical part and a row along middle of posterior margin.
</paragraph>
<caption id="D1D0DD879A52FFD931F2E848DA38F843" httpUri="https://zenodo.org/record/1303705/files/figure.png" pageId="1" pageNumber="499" startId="1.[151,264,1899,1922]" targetBox="[214,1392,231,1820]" targetPageId="1">
<paragraph id="85108D0F9A52FFD931F2E848DA38F843" blockId="1.[151,1436,1899,2014]" pageId="1" pageNumber="499">
<emphasis id="B7DB511D9A52FFD931F2E848DA23F81F" bold="true" box="[151,315,1899,1922]" pageId="1" pageNumber="499">FIGURES 1–4.</emphasis>
<taxonomicName id="42AFF68C9A52FFD93027E84FD9C3F81C" authority="Frolov &amp; Montreuil" authorityName="Frolov &amp; Montreuil" authorityYear="2018" box="[322,731,1900,1922]" class="Insecta" family="Scarabaeidae" genus="Adorodocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="499" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. nov.">
<emphasis id="B7DB511D9A52FFD93027E84FD91FF81F" box="[322,519,1900,1921]" italics="true" pageId="1" pageNumber="499">Adorodocia hirsuta</emphasis>
Frolov &amp; Montreuil
</taxonomicName>
,
<taxonomicNameLabel id="ACE8EC669A52FFD93383E84FD87BF81C" box="[742,867,1900,1922]" pageId="1" pageNumber="499" rank="species">new species</taxonomicNameLabel>
, holotype. 1, habitus; 2, ventral side of metasternum; 3, aedeagus in left lateral view and parameres in dorsal view; 4, locality records of
<taxonomicName id="42AFF68C9A52FFD932BFE8A8DE0EF83F" authority="Frolov &amp; Montreuil" authorityName="Frolov &amp; Montreuil" authorityYear="2018" box="[986,1302,1931,1953]" class="Insecta" family="Scarabaeidae" genus="Adorodocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="499" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. nov.">
<emphasis id="B7DB511D9A52FFD932BFE8A8DF27F83E" box="[986,1087,1931,1952]" italics="true" pageId="1" pageNumber="499">A. hirsuta</emphasis>
Frolov &amp; Montreuil
</taxonomicName>
,
<taxonomicNameLabel id="ACE8EC669A52FFD93444E8A8DE84F83F" box="[1313,1436,1931,1953]" pageId="1" pageNumber="499" rank="species">new species</taxonomicNameLabel>
(star) and other species of
<taxonomicName id="42AFF68C9A52FFD930CDE88AD907F820" authorityName="Brenske" authorityYear="1893" box="[424,543,1961,1982]" class="Insecta" family="Scarabaeidae" genus="Adorodocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="499" phylum="Arthropoda" rank="genus">
<emphasis id="B7DB511D9A52FFD930CDE88AD907F820" box="[424,543,1961,1982]" italics="true" pageId="1" pageNumber="499">Adorodocia</emphasis>
</taxonomicName>
(circle) in Madagascar (from Montreuil &amp; Frolov 2014, modified). Scale bars = 1 mm for Figs. 1–3.
</paragraph>
</caption>
<paragraph id="85108D0F9A51FFDA31A2EFB4DE7FFF4A" blockId="2.[151,1437,151,791]" pageId="2" pageNumber="500">
<emphasis id="B7DB511D9A51FFDA31A2EFB4DA4BFF2E" box="[199,339,151,176]" italics="true" pageId="2" pageNumber="500">Ventral side.</emphasis>
Metasternum completely covered by yellowish, long, flexuous setae (
<figureCitation id="1D94918A9A51FFDA3506EFB4DFB0FF2E" box="[1123,1192,151,176]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1899,1922]" captionTargetBox="[214,1392,231,1820]" captionTargetId="figure@1.[151,1436,217,1849]" captionTargetPageId="1" captionText="FIGURES 1–4. Adorodocia hirsuta Frolov &amp; Montreuil, new species, holotype. 1, habitus; 2, ventral side of metasternum; 3, aedeagus in left lateral view and parameres in dorsal view; 4, locality records of A. hirsuta Frolov &amp; Montreuil, new species (star) and other species of Adorodocia (circle) in Madagascar (from Montreuil &amp; Frolov 2014, modified). Scale bars = 1 mm for Figs. 1–3." httpUri="https://zenodo.org/record/1303705/files/figure.png" pageId="2" pageNumber="500">Fig. 2</figureCitation>
). Sternites with a mix of regularly distributed white, short and adpressed setae and a transversal row of yellowish, long, raised setae.
</paragraph>
<paragraph id="85108D0F9A51FFDA31A2EFC2D857FF66" blockId="2.[151,1437,151,791]" box="[199,847,223,248]" pageId="2" pageNumber="500">
<emphasis id="B7DB511D9A51FFDA31A2EFC2DBE4FF66" box="[199,252,225,248]" italics="true" pageId="2" pageNumber="500">Legs</emphasis>
. Protibiae tridentate, the basal tooth slightly marked.
</paragraph>
<paragraph id="85108D0F9A51FFDA31A2EE27DFA2FE82" blockId="2.[151,1437,151,791]" box="[199,1210,260,285]" pageId="2" pageNumber="500">
<emphasis id="B7DB511D9A51FFDA31A2EE27DA2CFE83" box="[199,308,260,285]" italics="true" pageId="2" pageNumber="500">Aedeagus</emphasis>
. Aedeagus of typical shape for
<taxonomicName id="42AFF68C9A51FFDA33F2EE27D802FE83" authorityName="Brenske" authorityYear="1893" box="[663,794,260,285]" class="Insecta" family="Scarabaeidae" genus="Adorodocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="B7DB511D9A51FFDA33F2EE27D802FE83" box="[663,794,260,285]" italics="true" pageId="2" pageNumber="500">Adorodocia</emphasis>
</taxonomicName>
with partly fused parameres (
<figureCitation id="1D94918A9A51FFDA350DEE27DFB4FE82" box="[1128,1196,260,285]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1899,1922]" captionTargetBox="[214,1392,231,1820]" captionTargetId="figure@1.[151,1436,217,1849]" captionTargetPageId="1" captionText="FIGURES 1–4. Adorodocia hirsuta Frolov &amp; Montreuil, new species, holotype. 1, habitus; 2, ventral side of metasternum; 3, aedeagus in left lateral view and parameres in dorsal view; 4, locality records of A. hirsuta Frolov &amp; Montreuil, new species (star) and other species of Adorodocia (circle) in Madagascar (from Montreuil &amp; Frolov 2014, modified). Scale bars = 1 mm for Figs. 1–3." httpUri="https://zenodo.org/record/1303705/files/figure.png" pageId="2" pageNumber="500">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="85108D0F9A51FFDA31A2EE04DE09FEDE" blockId="2.[151,1437,151,791]" box="[199,1297,295,320]" pageId="2" pageNumber="500">
<emphasis id="B7DB511D9A51FFDA31A2EE04DA55FEDE" bold="true" box="[199,333,295,320]" pageId="2" pageNumber="500">Variability.</emphasis>
<typeStatus id="5A1433AD9A51FFDA3036EE0BDAADFEDE" box="[339,437,296,320]" pageId="2" pageNumber="500" type="paratype">Paratype</typeStatus>
differs from the
<typeStatus id="5A1433AD9A51FFDA3317EE04D9CCFEDE" box="[626,724,295,320]" pageId="2" pageNumber="500" type="holotype">holotype</typeStatus>
by its slightly smaller size (body length 16.0 mm).
</paragraph>
<paragraph id="85108D0F9A51FFDA31A2EE6FDAEEFEFB" blockId="2.[151,1437,151,791]" box="[199,502,332,357]" pageId="2" pageNumber="500">
<emphasis id="B7DB511D9A51FFDA31A2EE6FDA3DFEFB" bold="true" box="[199,293,332,357]" pageId="2" pageNumber="500">Female.</emphasis>
Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="CDB5DE849A51FFDA31A2EE4CDF18FE19" box="[199,1024,367,392]" pageId="2" pageNumber="500" type="etymology">
<paragraph id="85108D0F9A51FFDA31A2EE4CDF18FE19" blockId="2.[151,1437,151,791]" box="[199,1024,367,392]" pageId="2" pageNumber="500">
<emphasis id="B7DB511D9A51FFDA31A2EE4CDA56FE16" bold="true" box="[199,334,367,392]" pageId="2" pageNumber="500">Etymology.</emphasis>
Species named in reference to its long setae on metasternum.
</paragraph>
</subSubSection>
<subSubSection id="CDB5DE849A51FFDA31A2EEB7D961FD6D" pageId="2" pageNumber="500" type="diagnosis">
<paragraph id="85108D0F9A51FFDA31A2EEB7D961FD6D" blockId="2.[151,1437,151,791]" pageId="2" pageNumber="500">
<emphasis id="B7DB511D9A51FFDA31A2EEB7DAC9FE33" bold="true" box="[199,465,404,429]" pageId="2" pageNumber="500">Differential diagnosis.</emphasis>
<taxonomicName id="42AFF68C9A51FFDA30B9EEB7D9A1FE33" authority="Frolov &amp; Montreuil, 2018" authorityName="Frolov &amp; Montreuil" authorityYear="2018" box="[476,697,404,429]" class="Insecta" family="Scarabaeidae" genus="Adorodocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. nov.">
<emphasis id="B7DB511D9A51FFDA30B9EEB7D9A1FE33" box="[476,697,404,429]" italics="true" pageId="2" pageNumber="500">Adorodocia hirsuta</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ACE8EC669A51FFDA33A9EEB6D84EFE33" box="[716,854,404,429]" pageId="2" pageNumber="500" rank="species">new species</taxonomicNameLabel>
presents several characters which make it unique within the genus
<taxonomicName id="42AFF68C9A51FFDA3032EE94DAC2FE4E" authorityName="Brenske" authorityYear="1893" box="[343,474,439,464]" class="Insecta" family="Scarabaeidae" genus="Adorodocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="B7DB511D9A51FFDA3032EE94DAC2FE4E" box="[343,474,439,464]" italics="true" pageId="2" pageNumber="500">Adorodocia</emphasis>
</taxonomicName>
: metasternum completely covered by long and raised setae, while other species of the genus bear shorter and adpressed setae only on the sides and have the central part appearing almost glabrous; body obviously more slender (rate body length/body width 2.2), while other species are wider (rate body length/body width approximately 1.8); elytra only covered by sparse, evenly distributed short adpressed setae and apical hump of elytra without any particular setae, while the other species bear obvious scale-like setae and a few long and strong, yellowish setae both on apical elytral hump and on elytral disc; 8th tergite with reduced setae, while other species bear a mix of very long and thick, yellowish raised setae and white, scale-like adpressed setae. The parameres of
<emphasis id="B7DB511D9A51FFDA3054ED94DAB0FD51" box="[305,424,694,719]" italics="true" pageId="2" pageNumber="500">
<taxonomicName id="42AFF68C9A51FFDA3054ED94DABCFD51" authority="Frolov &amp; Montreuil, 2018" authorityName="Frolov &amp; Montreuil" authorityYear="2018" box="[305,420,694,719]" class="Insecta" family="Scarabaeidae" genus="Adorodocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. nov.">A. hirsuta</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="ACE8EC669A51FFDA30D5ED9BD920FD51" box="[432,568,694,719]" pageId="2" pageNumber="500" rank="species">new species</taxonomicNameLabel>
are most similar to those of
<taxonomicName id="42AFF68C9A51FFDA321AED94DE10FD51" authority="Montreuil &amp; Frolov, 2014" authorityName="Montreuil &amp; Frolov" authorityYear="2014" box="[895,1288,694,719]" class="Insecta" family="Scarabaeidae" genus="Adorodocia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="recta">
<emphasis id="B7DB511D9A51FFDA321AED94D8CFFD51" box="[895,983,695,719]" italics="true" pageId="2" pageNumber="500">A. recta</emphasis>
<bibRefCitation id="E13EF0FE9A51FFDA32BAED95DE10FD51" box="[991,1288,694,719]" pageId="2" pageNumber="500" refId="ref1298" refString="Montreuil, O. &amp; Frolov, A. (2014) Revision of the genus Adorodocia Brenske, 1893, (Coleoptera, Rutelidae, Adoretinae) endemic to Madagascar. Annales de la Societe Entomologique de France, 49 (4), 460 - 475. [2013] https: // doi. org / 10.1080 / 00379271.2014.895124" type="journal article">Montreuil &amp; Frolov, 2014</bibRefCitation>
</taxonomicName>
, but differ in more curved apices in lateral view (
<figureCitation id="1D94918A9A51FFDA3342EDF9D973FD6D" box="[551,619,730,755]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1899,1922]" captionTargetBox="[214,1392,231,1820]" captionTargetId="figure@1.[151,1436,217,1849]" captionTargetPageId="1" captionText="FIGURES 1–4. Adorodocia hirsuta Frolov &amp; Montreuil, new species, holotype. 1, habitus; 2, ventral side of metasternum; 3, aedeagus in left lateral view and parameres in dorsal view; 4, locality records of A. hirsuta Frolov &amp; Montreuil, new species (star) and other species of Adorodocia (circle) in Madagascar (from Montreuil &amp; Frolov 2014, modified). Scale bars = 1 mm for Figs. 1–3." httpUri="https://zenodo.org/record/1303705/files/figure.png" pageId="2" pageNumber="500">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CDB5DE849A51FFDA31A2EDDDDFEDFC89" box="[199,1269,766,791]" pageId="2" pageNumber="500" type="distribution">
<paragraph id="85108D0F9A51FFDA31A2EDDDDFEDFC89" blockId="2.[151,1437,151,791]" box="[199,1269,766,791]" pageId="2" pageNumber="500">
<emphasis id="B7DB511D9A51FFDA31A2EDDDDA41FC89" bold="true" box="[199,345,766,791]" pageId="2" pageNumber="500">Distribution</emphasis>
. The new species is only known from an isolated forest in Ankaratra area (
<figureCitation id="1D94918A9A51FFDA35C7EDDDDFF0FC89" box="[1186,1256,766,791]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1899,1922]" captionTargetBox="[214,1392,231,1820]" captionTargetId="figure@1.[151,1436,217,1849]" captionTargetPageId="1" captionText="FIGURES 1–4. Adorodocia hirsuta Frolov &amp; Montreuil, new species, holotype. 1, habitus; 2, ventral side of metasternum; 3, aedeagus in left lateral view and parameres in dorsal view; 4, locality records of A. hirsuta Frolov &amp; Montreuil, new species (star) and other species of Adorodocia (circle) in Madagascar (from Montreuil &amp; Frolov 2014, modified). Scale bars = 1 mm for Figs. 1–3." httpUri="https://zenodo.org/record/1303705/files/figure.png" pageId="2" pageNumber="500">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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