114 lines
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114 lines
13 KiB
XML
<document id="EBB95ADCA950AF87591E81F86071D073" ID-CLB-Dataset="33603" ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1487" ID-GBIF-Dataset="c6e677be-f8a3-4973-98bd-ddb1d954ed06" ID-PMC="PMC3260754" ID-Pensoft-Pub="1313-2970-130-131" ID-PubMed="22259271" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-131" ModsDocOrigin="ZooKeys 130" ModsDocTitle="A new, enigmatic family for new genus and species of Polyneoptera from the Upper Permian of Russia" checkinTime="1451249886334" checkinUser="pensoft" docAuthor="Gorochov, Andrej V." docDate="2011" docId="A8FE1F9871658210991E9755ED40668A" docLanguage="en" docName="ZooKeys 130: 131-136" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1487" docTitle="Alexarasniidae Gorochov, fam. n." docType="treatment" docVersion="6" lastPageNumber="133" masterDocId="FFE7FFBEFFE0846C4B5EFFDC4664FFEA" masterDocTitle="A new, enigmatic family for new genus and species of Polyneoptera from the Upper Permian of Russia" masterLastPageNumber="136" masterPageNumber="131" pageNumber="132" updateTime="1732836818524" updateUser="ExternalLinkService">
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<mods:mods id="D6F1385CA0E47E35782148A46ECCF263" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FD230A94CCD5613795AE4C4E132AEF55">
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<mods:title id="27F388114F62045E911D7D0D21376D17">A new, enigmatic family for new genus and species of Polyneoptera from the Upper Permian of Russia</mods:title>
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<mods:name id="6A03ACD0CFACA146FD4328CD7B83971A" type="personal">
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<mods:roleTerm id="D68BBE22F30E2C6D015BC2F9A9641B2B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E109516CD84C99935E59EEB5DCF8070B">Gorochov, Andrej V.</mods:namePart>
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<mods:typeOfResource id="E430E4BCAE15D7A65BADC573A8D2E0C7">text</mods:typeOfResource>
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<mods:title id="BFFFDEE73042F9805BD22492891C03EB">ZooKeys</mods:title>
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<mods:part id="0FCB73975C52E2E72A372298EEE10BB7">
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<mods:date id="5BAF8F2DAD9517966ED167A0EA1EBA6F">2011</mods:date>
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<mods:detail id="B67786EDA64A346B20FDCA681CC6C78C" type="volume">
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<mods:number id="6958447F23A95BEE8DC886F648695A71">130</mods:number>
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<mods:extent id="5455E79BE53547A4871248A3D7308D91" unit="page">
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<mods:start id="C9EB301AB9E67AE9913EAC0D10CFEF48">131</mods:start>
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<mods:end id="30986A3CEA1DAF1013378547C0A97BF1">136</mods:end>
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<mods:location id="9945462193A7F1FA74E49A11D478C1EE">
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<mods:url id="A5B25AC75354396DC8530AE345247D9B">http://dx.doi.org/10.3897/zookeys.130.1487</mods:url>
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</mods:location>
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<mods:classification id="8E6D1415C5E85ED3123B59066DA50069">journal article</mods:classification>
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<mods:identifier id="D3ED72B920569031869AD320CE142398" type="DOI">http://dx.doi.org/10.3897/zookeys.130.1487</mods:identifier>
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<mods:identifier id="2ADA53F15DA24471168534E11B545C4C" type="Pensoft-Pub">1313-2970-130-131</mods:identifier>
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<treatment id="A8FE1F9871658210991E9755ED40668A" ID-GBIF-Taxon="152030798" LSID="urn:lsid:zoobank.org:act:D6A2D4C4-6E69-4450-B12F-2488A6D9B6B7" httpUri="http://treatment.plazi.org/id/A8FE1F9871658210991E9755ED40668A" lastPageId="2" lastPageNumber="133" pageId="1" pageNumber="132">
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<subSubSection id="1EF363D9D0F07E07439D3A7B05D69C56" pageId="1" pageNumber="132" type="nomenclature">
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<paragraph id="2BC9BBC86057167ABBDDDDAC24018C53" pageId="1" pageNumber="132">
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Family
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<taxonomicName id="972FB0AD73820F7A233E70BD60A7DD15" ID-CoL="8MPCF" LSID="urn:lsid:zoobank.org:act:D6A2D4C4-6E69-4450-B12F-2488A6D9B6B7" authority="Gorochov" class="Insecta" family="Alexarasniidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Embioptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="family">Alexarasniidae Gorochov</taxonomicName>
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<taxonomicNameLabel id="48B54177BA86C9ED5773045F479FF343" pageId="1" pageNumber="132">fam. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="4DB7F3F8049958C399AC417523F25752" pageId="1" pageNumber="132" type="type genus">
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<paragraph id="D2D0EC7220BF231FE8322C9B0BB4F10E" pageId="1" pageNumber="132">Type genus.</paragraph>
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<paragraph id="68ABF1A20D7C6AFB63667700D94A89FB" pageId="1" pageNumber="132">
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<taxonomicName id="02755E2D62037AFCCC579F666389A34E" class="Insecta" family="Alexarasniidae" genus="Alexarasnia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alexarasnia" order="Embioptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Alexarasnia</taxonomicName>
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gen. n.
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</paragraph>
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</subSubSection>
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<subSubSection id="6C8FE61938E833FBFECD9B8A0770FC06" pageId="1" pageNumber="132" type="composition">
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<paragraph id="1B36F7627E1652E34DCFEB1D478A05D2" pageId="1" pageNumber="132">Composition.</paragraph>
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<paragraph id="3623A29ED4ED2BE44799A155560FA5D2" pageId="1" pageNumber="132">
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Only the genus
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<taxonomicName id="061B38BA8B9BD7FCA60532139EAFDE24" class="Insecta" family="Alexarasniidae" genus="Alexarasnia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alexarasnia" order="Embioptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Alexarasnia</taxonomicName>
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gen. n.
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</paragraph>
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</subSubSection>
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<subSubSection id="29750AC350A411CF70710F97A51E3D4B" lastPageId="2" lastPageNumber="133" pageId="1" pageNumber="132" type="diagnosis">
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<paragraph id="3491F8CC9FA13ED6940A65CFCD443064" pageId="1" pageNumber="132">Diagnosis.</paragraph>
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<paragraph id="A34ABCE31A3461E777998036E231953A" pageId="1" pageNumber="132">
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Tegmen (Figs 1-4) differs from that of
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<taxonomicName id="40531F18422CF4024FBDA74770B6291F" genus="Titanoptera" lsidName="Titanoptera" pageId="1" pageNumber="132" rank="genus">Titanoptera</taxonomicName>
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, majority representatives of
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<taxonomicName id="678E7F9E49E0E086389A82854ABFB1A6" genus="Orthoptera" lsidName="Orthoptera" pageId="1" pageNumber="132" rank="genus">Orthoptera</taxonomicName>
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, and Paleozoic and Triassic
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<taxonomicName id="C85463FBEFBE21A859FB4C17C85AFC2A" genus="Phasmatoptera" lsidName="Phasmatoptera" pageId="1" pageNumber="132" rank="genus">Phasmatoptera</taxonomicName>
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in the absence of precostal area (presence of this area is a synapomorphy of all these orders). From the other representatives of two latter orders, this family differs in the reduction of Sc branches in the tegmen and/or clearly more numerous longitudinal tegminal veins (CuA has six or more branches in
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<taxonomicName id="8CB5F4509A0DBA4C9AEB62F99B71112C" class="Insecta" family="Alexarasniidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Embioptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="family">Alexarasniidae</taxonomicName>
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and four or less branches in all the other representatives of
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<taxonomicName id="0AA795B027E30A27A9C208BD4B3B3880" genus="Phasmatoptera" lsidName="Phasmatoptera" pageId="1" pageNumber="132" rank="genus">Phasmatoptera</taxonomicName>
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). Tegminal venation of the new family is distinguished from that of all the other orders of
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<taxonomicName id="642C0E902C4FB93B5D9E57E6AC63AB56" genus="Polyneoptera" lsidName="Polyneoptera" pageId="1" pageNumber="132" rank="genus">Polyneoptera</taxonomicName>
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by the partly parallel longitudinal veins in combination with the following characters: straight CuP, very narrow radial and interradial areas, and reduction of branches of RA and RS (from
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<taxonomicName id="BB6CE24688B9B7A9B65EBFD2EADE1114" class="Insecta" family="Lycidae" genus="Dictyoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dictyoptera" order="Coleoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Dictyoptera</taxonomicName>
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); reduction of branches of Sc and/or RS, partial fusion of distal parts of some longitudinal veins with formation of long loop-like cells along anal edge of tegmen, and presence of intercalary veins between majority of longitudinal veins (from Grylloblattida as well as from the families
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<taxonomicName id="DFC0D0A49B720ECF00069339FA7823FB" class="Insecta" family="Lemmatophoridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Grylloblattodea" pageId="1" pageNumber="132" phylum="Arthropoda" rank="family">Lemmatophoridae</taxonomicName>
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Sellards, 1909 and
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<taxonomicName id="98B8A69A777C80C3DE97764383121AB2" class="Insecta" family="Atactophlebiidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Grylloblattodea" pageId="1" pageNumber="132" phylum="Arthropoda" rank="family">Atactophlebiidae</taxonomicName>
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Martynov, 1930 possibly belonging to the extinct order Eoblattida;
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<bibRefCitation id="F40BCCFEDA83331A4764B9309D9CB393" author="Gorochov, AV" journalOrPublisher="Cholodkovsky" pageId="4" pageNumber="135" pagination="1 - 54" title="Primitive Titanoptera and early evolution of Polyneoptera. Meetings in memory of N. A." volume="57" year="2004">Gorochov 2004</bibRefCitation>
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); the latter characters as well as distal part of Sc not fused with RA (from
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<taxonomicName id="3B824253D60DE20AACD3EC90F6B7FED8" class="Insecta" family="Erebidae" genus="Plecoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plecoptera" order="Lepidoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="genus">Plecoptera</taxonomicName>
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and the other taxa of Eoblattida); tegmen not leathery with well-developed venation (from
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<taxonomicName id="77D8ECB760DB58DBE1923C162C154495" genus="Dermaptera" lsidName="Dermaptera" pageId="1" pageNumber="132" rank="genus">Dermaptera</taxonomicName>
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), and distinctly more numerous longitudinal veins (from
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<taxonomicName id="A58870D945812CB078465CFE7988A38B" genus="Embioptera" lsidName="Embioptera" pageId="1" pageNumber="132" rank="genus">Embioptera</taxonomicName>
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). Parallel venation of the tegmen is also present in the enigmatic family
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<taxonomicName id="0093FB08232CF69EA4A91CFD0AC745CE" family="Chresmodidae" lsidName="" pageId="1" pageNumber="132" rank="family">Chresmodidae</taxonomicName>
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Handlirsch, 1906 belonging to an unknown order of
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<taxonomicName id="0C3CA75375A58A1DE31BDED786E54838" genus="Polyneoptera" lsidName="Polyneoptera" pageId="1" pageNumber="132" rank="genus">Polyneoptera</taxonomicName>
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(
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<bibRefCitation id="3BB075C3D983E4CEEBAE9177AA81B1BE" author="Delclos, X" journalOrPublisher="Neues Jahrbuch fuer Geologie und Palaeontologie - Abhandlungen" pageId="4" pageNumber="135" pagination="353 - 381" title="The enigmatic Mesozoic insect taxon Chresmodidae (Polyneoptera): new palaeobiological and phylogenetic data, with the description of a new species from the Lower Cretaceous of Brazil." volume="247" year="2008">
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<normalizedToken id="CBD01B40C53FDAC23288220B2700FC9C" originalValue="Delclòs">Delclos</normalizedToken>
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et al 2008
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</bibRefCitation>
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;
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<bibRefCitation id="60AEF79E018DD48AB59CEEA5059F1789" author="Zhang, X-W" journalOrPublisher="Zootaxa" pageId="4" pageNumber="135" pagination="26 - 40" title="A new genus and species of Chresmodidae (Insecta: Gryllones) from Upper Jurassic-Lower Cretaceous of Yixian Formation, Inner Mongolia, China." volume="1702" year="2008 a">Zhang et al 2008a</bibRefCitation>
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,
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<bibRefCitation id="451722CD125A90D5903B0E4F3245CAF0" author="Zhang, X-W" journalOrPublisher="Zootaxa" pageId="4" pageNumber="135" pagination="53 - 62" title="A water-skiing chresmodid from the Middle Jurassic in Daohugou, Inner Mongolia, China (Polyneoptera: Orthopterida)." volume="1762" year="2008 b">b</bibRefCitation>
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);
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<taxonomicName id="1478F847591BC497965D9B6418BEAE40" class="Insecta" family="Alexarasniidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Embioptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="family">Alexarasniidae</taxonomicName>
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is distinguished from it by the distal half of tegminal CuP situated not parallel to the anal tegminal edge and more numerous branches of CuA in tegmen (6-9 instead of 3-4).
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</paragraph>
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<caption id="7A8AEA9A929FC5340129374516980D56" pageId="2" pageNumber="133">
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<paragraph id="9BA23186378AF3ED1F67D8C71D511713" pageId="2" pageNumber="133">
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<pageBreakToken id="4C1577A782BE869EF63C33CA1DC1D49E" pageId="2" pageNumber="133" start="start">Figures</pageBreakToken>
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1-2.
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<taxonomicName id="2B928F3A26F934CDD06176685530BB82" class="Insecta" family="Alexarasniidae" genus="Alexarasnia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alexarasnia rossica" order="Embioptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="rossica">Alexarasnia rossica</taxonomicName>
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gen. et sp. n., tegmen, holotype PIN 3840/63 (part) 1 scheme of venation 2 photograph of fossil. Scale bar 5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |