582 lines
90 KiB
XML
582 lines
90 KiB
XML
<document id="D82FEA14A70A8B021502B18864B5ACBF" ID-CLB-Dataset="21611" ID-DOI="10.5852/ejt.2020.670" ID-GBIF-Dataset="c16b18d9-fad7-4700-8f8e-a0e59272069c" ID-ISSN="2118-9773" ID-Zenodo-Dep="3906721" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593019764452" checkinUser="valdenar" docAuthor="Zaova, Dušica, Cvetkoska, Aleksandra, Mitic-Kopanja, Danijela, Jovanovska, Elena, Ognjanova-Rumenova, Nadja & Levkov, Zlatko" docDate="2020" docId="A4376C2BFFBEFFFE8319F75AFC534D80" docLanguage="en" docName="ejt-670_zavoa_cvetkoska_mitic-kopanja.pdf.imf" docOrigin="European Journal of Taxonomy 670" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tertiarius minutulus Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="7" masterDocId="580E1453FFBCFFF78175F333FFEC4C5A" masterDocTitle="Tertiarius minutulus sp. nov. (Stephanodiscaceae, Bacillariophyta) - a new fossil diatom species from Lake Ohrid" masterLastPageNumber="14" masterPageNumber="1" pageNumber="3" updateTime="1698842861002" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="BFD0639E47845F10EE67BBDAA6A4FB9A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7DA6A651A23274E1FE7636C8C178F8FE">
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<mods:title id="93879A52ADF06343E61AD24E2FF22BAD">Tertiarius minutulus sp. nov. (Stephanodiscaceae, Bacillariophyta) - a new fossil diatom species from Lake Ohrid</mods:title>
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</mods:titleInfo>
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<mods:name id="952753948F1A90B847DADDA18F7E3CAC" type="personal">
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<mods:role id="3207A928BEE853DCEA1F36623FAD2154">
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<mods:roleTerm id="EB74B7A4EABDF364B89026A7856AE5DE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EF8038BDFA3A949A9FF27A0D745E0269">Zaova, Dušica</mods:namePart>
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||
<mods:affiliation id="20D41844EC406F5283383422F637B098">Institute of Biology, Faculty of Natural Sciences, Arhimedova 3, 1000 Skopje, R. North Macedonia. & Corresponding author: zaovadusica @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="8AD5B50C6427AEB7F3AA7CDCC4E38C52" type="email">zaovadusica@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="607BCA81D1D83E85E94C89EE11D8E642" type="personal">
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<mods:role id="2BC115E91894646D73F48624E2019315">
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<mods:roleTerm id="02AA689B79896DD940B2BA35DAF7AEA5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6E2BFEA35094EA3ADA7D848D5B86BE7C">Cvetkoska, Aleksandra</mods:namePart>
|
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<mods:affiliation id="2E7D9F9DE2B8E28EC04300173EBFDB70">Department of Aquatic Ecology, Netherlands Institute of Ecology (NIOO-KNAW), Wageningen, Netherlands. & Email: acvetkoska @ yahoo. com</mods:affiliation>
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<mods:nameIdentifier id="2FAF45C717413EE9C5D3940B96000E39" type="email">acvetkoska@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="D06FC58C758C130EB841092FFBC0D73E" type="personal">
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<mods:role id="146236072AEBDB68E9B52B34DE0AEE0A">
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<mods:roleTerm id="7A93A9472D2ACC041BC76EFC44E6A11A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F7BE194AC79B33513ADE9102262CE2EA">Mitic-Kopanja, Danijela</mods:namePart>
|
||
<mods:affiliation id="03824D0201C09AB0802BA1344D7EFD92">Institute of Biology, Faculty of Natural Sciences, Arhimedova 3, 1000 Skopje, R. North Macedonia. & Email: mitickopanja @ yahoo. com</mods:affiliation>
|
||
<mods:nameIdentifier id="BECBDED757E9BF47DB3E7BEF3C501A3B" type="email">mitickopanja@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="F8F67C6B2F66B57CF74F41F42B594647" type="personal">
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<mods:role id="407AEDAC49278A10CBAAFB7D83C563FC">
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<mods:roleTerm id="3B266F6A36D67BD969EE34D7BBFA8101">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="393473F67FE17970A7FF0D1D80E06102">Jovanovska, Elena</mods:namePart>
|
||
<mods:affiliation id="55C8EA92B4EFE66556DBE1F579E0A74B">Institute for Ecology, Evolution and Diversity, Goethe University, Frankfurt, Germany. & Department of Animal Ecology and Systematics, Justus Liebig University Giessen, Giessen, Germany. & Email: jovanovska. eci @ gmail. com</mods:affiliation>
|
||
<mods:nameIdentifier id="7E97C4AD29B0CBD0F2BB7D7EC32AEE31" type="email">jovanovska.eci@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="6BCF8E2C6E1780491C2C6DBD6EDC4790" type="personal">
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<mods:role id="E8749451E95E83A0B504F73DFB2415BE">
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<mods:roleTerm id="CADD44E82A69C2E32F64E6BAB07EE8B9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A3F425D49A64B986782D5FFB489C0F6E">Ognjanova-Rumenova, Nadja</mods:namePart>
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<mods:affiliation id="7B6603343FBBECB8E98EAA3D139DD958">Department of Palaeontology, Stratigraphy and Sedimentology, Geological Institute, Bulgarian Academy of Sciences, Sofia, Bulgaria. & Email: ognjanovanadja @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="E5F7B89D28B80E49DDA5E4D50EA018E5" type="email">ognjanovanadja@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="62894838E675AC9CCC4580E303E55350" type="personal">
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<mods:roleTerm id="CA9A1171B6423A2D64B83406AC881CFE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A843F21DA4ADD227DAB5BB4E2730F0BD">Levkov, Zlatko</mods:namePart>
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<mods:affiliation id="676531F4E78CB48931A19D13DEF410F2">Institute of Biology, Faculty of Natural Sciences, Arhimedova 3, 1000 Skopje, R. North Macedonia. & Email: zlevkov @ pmf. ukim. mk</mods:affiliation>
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<mods:nameIdentifier id="170D8834F3851D9A53A4284F155D3665" type="email">zlevkov@pmf.ukim.mk</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="2B233E5636D2E04C312A6FBE64AE351F">text</mods:typeOfResource>
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<mods:titleInfo id="811F7E3386D59DCEFBD0401BA92F4A95">
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<mods:title id="DDC91EBD731C16097CDBA745CD0DBC52">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="C313945E9E8EB0A9913085A3E8E4FEC3">
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<mods:date id="F5219C05F810524A7EB9035DFCF84FC0">2020</mods:date>
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<mods:detail id="FE3CCEB5A003B1843BCB9C12523A2733" type="pubDate">
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<mods:number id="A2D72B90AE6645B58D2B8EB7C8022F13">2020-06-19</mods:number>
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</mods:detail>
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<mods:detail id="5D194E9014EE431A30C70330C5BEB0D3" type="issue">
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<mods:number id="E12D5C7D086F3592A123B438579A8CB5">670</mods:number>
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</mods:detail>
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<mods:extent id="CAE97AE4AE2FE391D0925E7D22552BBD" unit="page">
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<mods:start id="5C93010102FD032336A5B01967B98122">1</mods:start>
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<mods:end id="A5D16CFA131FF309AD8CF5BE4A831C86">14</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="810A8ED0A3811A3BE83C4D45DD968256">journal article</mods:classification>
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<mods:identifier id="94C248D65DAEC878BD97B7724FD07C0E" type="CLB-Dataset">21611</mods:identifier>
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<mods:identifier id="EDC2BBB8697572961FF0FD1FB2EBD648" type="DOI">10.5852/ejt.2020.670</mods:identifier>
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<mods:identifier id="D60AB23B2058EDC55BA3FE26712E0ED7" type="GBIF-Dataset">c16b18d9-fad7-4700-8f8e-a0e59272069c</mods:identifier>
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<mods:identifier id="43E6A7563016DCFB22A4AAAB82D6C956" type="ISSN">2118-9773</mods:identifier>
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<mods:identifier id="9B9DCF5212EFDC2BFE2A99B4D98B7CDD" type="Zenodo-Dep">3906721</mods:identifier>
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</mods:mods>
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<treatment id="A4376C2BFFBEFFFE8319F75AFC534D80" ID-GBIF-Taxon="164723333" LSID="urn:lsid:plazi:treatment:A4376C2BFFBEFFFE8319F75AFC534D80" httpUri="http://treatment.plazi.org/id/A4376C2BFFBEFFFE8319F75AFC534D80" lastPageId="9" lastPageNumber="7" pageId="2" pageNumber="3">
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<subSubSection id="64848EB6FFBEFFF58319F75AFC2448DE" box="[620,968,1129,1156]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="2C21DD3DFFBEFFF58319F75AFC2448DE" blockId="2.[620,968,1129,1193]" box="[620,968,1129,1156]" pageId="2" pageNumber="3">
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<heading id="77696A51FFBEFFF58319F75AFC2448DE" box="[620,968,1129,1156]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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||
<taxonomicName id="EB9EA6BEFFBEFFF58319F75AFC8A48DE" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[620,870,1129,1156]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="2" pageNumber="3" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBEFFF58319F75AFC8A48DE" bold="true" box="[620,870,1129,1156]" italics="true" pageId="2" pageNumber="3">Tertiarius minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBEFFF58218F759FC2448DE" box="[877,968,1130,1156]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="64848EB6FFBEFFF58392F7BCFCA148F3" box="[743,845,1167,1193]" pageId="2" pageNumber="3" type="description">
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<paragraph id="2C21DD3DFFBEFFF58392F7BCFCA148F3" blockId="2.[620,968,1129,1193]" box="[743,845,1167,1193]" pageId="2" pageNumber="3">
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<figureCitation id="B4A5C1B8FFBEFFF58392F7BCFCA148F3" box="[743,845,1167,1193]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="3.[189,232,1965,1991]" captionStartId-1="4.[189,232,1894,1920]" captionStartId-2="5.[189,232,1791,1817]" captionTargetBox-0="[189,1398,1405,1923]" captionTargetBox-1="[238,1350,265,1854]" captionTargetBox-2="[274,1313,265,1749]" captionTargetId-0="figure@3.[189,1398,1405,1923]" captionTargetId-1="figure@4.[238,1350,265,1854]" captionTargetId-2="figure@5.[274,1313,265,1749]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Fig. 1. Tertiarius minutulus sp. nov., LM valve views, Lake Ohrid, North Macedonia. The asterisk indicates the holotype. Scale bar = 10 µm." captionText-1="Fig. 2. Tertiarius minutulus sp. nov., SEM external valve views. A. Valve view showing the marginal openings of the fultoportulae (see white arrow). B–D. View of the whole frustule. E. Close view of marginal openings of the fultoportula (see white arrow). F. Close view of the external openings of the valve face fultoportulae (see white arrows). Scale bars: A–D = 2.0 µm; E–F = 1.0 µm." captionText-2="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi-0="http://doi.org/10.5281/zenodo.3906723" figureDoi-1="http://doi.org/10.5281/zenodo.3906725" figureDoi-2="http://doi.org/10.5281/zenodo.3906727" httpUri-0="https://zenodo.org/record/3906723/files/figure.png" httpUri-1="https://zenodo.org/record/3906725/files/figure.png" httpUri-2="https://zenodo.org/record/3906727/files/figure.png" pageId="2" pageNumber="3">Figs 1–3</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="64848EB6FFBEFFF581C8F7E0FCAC4947" box="[189,832,1235,1310]" pageId="2" pageNumber="3" type="etymology">
|
||
<paragraph id="2C21DD3DFFBEFFF581C8F7E0FEA848B7" blockId="2.[189,324,1235,1261]" box="[189,324,1235,1261]" pageId="2" pageNumber="3">
|
||
<heading id="77696A51FFBEFFF581C8F7E0FEA848B7" bold="true" box="[189,324,1235,1261]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="1EEA012FFFBEFFF581C8F7E0FEA848B7" bold="true" box="[189,324,1235,1261]" pageId="2" pageNumber="3">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBEFFF581C8F630FCAC4947" blockId="2.[189,832,1283,1310]" box="[189,832,1283,1310]" pageId="2" pageNumber="3">
|
||
The specific epithet ʻ
|
||
<taxonomicName id="EB9EA6BEFFBEFFF580C1F630FDC44947" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[436,552,1283,1309]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="2" pageNumber="3" phylum="Ochrophyta" rank="species" species="minutulus">
|
||
<emphasis id="1EEA012FFFBEFFF580C1F630FDC44947" box="[436,552,1283,1309]" italics="true" pageId="2" pageNumber="3">minutulus</emphasis>
|
||
</taxonomicName>
|
||
ʼ refers to its small size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="64848EB6FFBEFFF581C8F67BFEE34A37" pageId="2" pageNumber="3" type="materials_examined">
|
||
<paragraph id="2C21DD3DFFBEFFF581C8F67BFE80493B" blockId="2.[189,364,1351,1378]" box="[189,364,1351,1378]" pageId="2" pageNumber="3">
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||
<heading id="77696A51FFBEFFF581C8F67BFE80493B" bold="true" box="[189,364,1351,1378]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="1EEA012FFFBEFFF581C8F67BFE80493B" bold="true" box="[189,364,1351,1378]" pageId="2" pageNumber="3">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBEFFF5819FF64BFEB049C8" blockId="2.[189,1398,1400,1502]" box="[234,348,1400,1426]" pageId="2" pageNumber="3">
|
||
<emphasis id="1EEA012FFFBEFFF5819FF64BFEB049C8" bold="true" box="[234,348,1400,1426]" pageId="2" pageNumber="3">
|
||
<typeStatus id="F325639FFFBEFFF5819FF64BFEB049C8" box="[234,348,1400,1426]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBEFFF581C8F6ADFADC4987" blockId="2.[189,1398,1400,1502]" pageId="2" pageNumber="3">
|
||
<materialsCitation id="9CF6D760FFBEFFF581C8F6ADFADC4987" ID-GBIF-Occurrence="2642705302" collectingDate="2013-03" collectingDateMax="2013-06-31" collectingDateMin="2013-03" collectionCode="REPUBLIC, OF, NORTH" country="REPUBLIC OF NORTH MACEDONIA" elevation="452" latitude="41.049168" location="Lake Ohrid" longLatPrecision="1" longitude="20.715" pageId="2" pageNumber="3" specimenCode="MKNDC 010093" specimenCount="1" stateProvince="Ohrid" typeStatus="holotype">
|
||
<collectingCountry id="54899DADFFBEFFF581C8F6ADFD7849E2" box="[189,660,1438,1464]" pageId="2" pageNumber="3">REPUBLIC OF NORTH MACEDONIA</collectingCountry>
|
||
•
|
||
<location id="29418BE6FFBEFFF583D2F6AEFCC149ED" LSID="urn:lsid:plazi:treatment:A4376C2BFFBEFFFE8319F75AFC534D80:29418BE6FFBEFFF583D2F6AEFCC149ED" box="[679,813,1437,1463]" country="REPUBLIC OF NORTH MACEDONIA" latitude="41.049168" longLatPrecision="1" longitude="20.715" name="Lake Ohrid" pageId="2" pageNumber="3" stateProvince="Ohrid">Lake Ohrid</location>
|
||
, core DEEP-5045-1;
|
||
<geoCoordinate id="49AABBFAFFBEFFF58554F6AEFB2A49E2" box="[1057,1222,1437,1464]" degrees="41.049167" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="41.049168">41.049167° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="49AABBFAFFBEFFF585BAF6AEFA9D49E2" box="[1231,1393,1437,1464]" degrees="20.715000" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="20.715">20.715000° E</geoCoordinate>
|
||
; sediment sample from
|
||
<elevation id="A7B33A0EFFBEFFF580BCF6F0FD6C4987" box="[457,640,1475,1502]" metricMagnitude="2" metricUnit="m" metricValue="4.5192" pageId="2" pageNumber="3" unit="m" value="451.92">
|
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<quantity id="EB6670D8FFBEFFF580BCF6F0FDD44984" box="[457,568,1475,1502]" metricMagnitude="2" metricUnit="m" metricValue="4.5192" pageId="2" pageNumber="3" unit="m" value="451.92">451.92 m</quantity>
|
||
depth
|
||
</elevation>
|
||
of the composite core;
|
||
<date id="5820FBFDFFBEFFF582E0F6F7FBA34987" box="[917,1103,1475,1502]" pageId="2" pageNumber="3" value="2013-03" valueMax="2013-06-31" valueMin="2013-03-00">
|
||
<collectingDate id="48640215FFBEFFF582E0F6F7FBA34987" box="[917,1103,1475,1502]" pageId="2" pageNumber="3" value="2013-03" valueMax="2013-06-31" valueMin="2013-03-00">Mar.–Jun. 2013</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="7C387546FFBEFFF5852EF6F0FAC04987" box="[1115,1324,1475,1501]" collectionCode="MKNDC" country="Macedonia" httpUri="http://grbio.org/cool/40e3-v9id" name="Institute of Biology" pageId="2" pageNumber="3" type="Herbarium">MKNDC 010093</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBEFFF5819FF534FEA94A7B" blockId="2.[189,1398,1543,1645]" box="[234,325,1543,1569]" pageId="2" pageNumber="3">
|
||
<emphasis id="1EEA012FFFBEFFF5819FF534FEA94A7B" bold="true" box="[234,325,1543,1569]" pageId="2" pageNumber="3">Isotype</emphasis>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBEFFF581C8F51FFEE34A37" blockId="2.[189,1398,1543,1645]" pageId="2" pageNumber="3">
|
||
<materialsCitation id="9CF6D760FFBEFFF581C8F51FFEE04A37" ID-GBIF-Occurrence="2642705301" collectingDate="2013-03" collectingDateMax="2013-06-31" collectingDateMin="2013-03" collectionCode="REPUBLIC, OF, NORTH" country="REPUBLIC OF NORTH MACEDONIA" elevation="452" latitude="41.049168" location="REPUBLIC OF NORTH MACEDONIA" longLatPrecision="1" longitude="20.715" pageId="2" pageNumber="3" specimenCode="BM 81918" specimenCount="1" stateProvince="Ohrid">
|
||
<collectingCountry id="54899DADFFBEFFF581C8F51FFD494A1C" box="[189,677,1580,1606]" pageId="2" pageNumber="3">REPUBLIC OF NORTH MACEDONIA</collectingCountry>
|
||
•
|
||
<specimenCount id="3A9816B4FFBEFFF583B1F51FFCF04A1D" box="[708,796,1580,1607]" pageId="2" pageNumber="3" type="generic">1 spec.</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="7C387546FFBEFFF58580F51EFA9A4A1C" box="[1269,1398,1580,1607]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="2" pageNumber="3">BM 81918</specimenCode>
|
||
(slide)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="64848EB6FFBEFFF581C8F5A5FB554ABB" pageId="2" pageNumber="3" type="distribution">
|
||
<paragraph id="2C21DD3DFFBEFFF581C8F5A5FEB14AF5" blockId="2.[189,349,1685,1712]" box="[189,349,1685,1712]" pageId="2" pageNumber="3">
|
||
<heading id="77696A51FFBEFFF581C8F5A5FEB14AF5" bold="true" box="[189,349,1685,1712]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="1EEA012FFFBEFFF581C8F5A5FEB14AF5" bold="true" box="[189,349,1685,1712]" pageId="2" pageNumber="3">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBEFFF581C8F5F5FB554ABB" blockId="2.[189,1209,1734,1761]" box="[189,1209,1734,1761]" pageId="2" pageNumber="3">
|
||
Lake
|
||
<collectingRegion id="EE5A13DFFFBEFFF5818AF5F5FEAF4ABA" box="[255,323,1734,1760]" country="Macedonia" name="Ohrid" pageId="2" pageNumber="3">Ohrid</collectingRegion>
|
||
core 5045-1, site DEEP, at a depth of
|
||
<quantity id="EB6670D8FFBEFFF58271F5F5FC9F4ABB" box="[772,883,1734,1761]" metricMagnitude="2" metricUnit="m" metricValue="4.5192" pageId="2" pageNumber="3" unit="m" value="451.92">451.92 m</quantity>
|
||
of the composite sequence.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="64848EB6FFBEFFF481C8F439FD264FC6" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="2C21DD3DFFBEFFF581C8F439FEA24B7E" blockId="2.[189,334,1802,1828]" box="[189,334,1802,1828]" pageId="2" pageNumber="3">
|
||
<heading id="77696A51FFBEFFF581C8F439FEA24B7E" bold="true" box="[189,334,1802,1828]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="1EEA012FFFBEFFF581C8F439FEA24B7E" bold="true" box="[189,334,1802,1828]" pageId="2" pageNumber="3">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBEFFF5819FF408FDE14B0F" blockId="2.[189,1399,1851,2027]" box="[234,525,1851,1877]" pageId="2" pageNumber="3">
|
||
<emphasis id="1EEA012FFFBEFFF5819FF408FDE14B0F" bold="true" box="[234,525,1851,1877]" pageId="2" pageNumber="3">Light microscope (LM)</emphasis>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBEFFF581C8F453FB5F4BB1" blockId="2.[189,1399,1851,2027]" pageId="2" pageNumber="3">
|
||
Valves circular, 3.5–8.0 µm in diameter, central area 3.0–
|
||
<quantity id="EB6670D8FFBEFFF5822DF453FC474B21" box="[856,939,1888,1915]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="3" unit="mm" value="6.5">6.5 µm</quantity>
|
||
in diameter (
|
||
<figureCitation id="B4A5C1B8FFBEFFF58530F453FB654B20" box="[1093,1161,1888,1914]" captionStart="Fig" captionStartId="3.[189,232,1965,1991]" captionTargetBox="[189,1398,1405,1923]" captionTargetId="figure@3.[189,1398,1405,1923]" captionTargetPageId="3" captionText="Fig. 1. Tertiarius minutulus sp. nov., LM valve views, Lake Ohrid, North Macedonia. The asterisk indicates the holotype. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3906723" httpUri="https://zenodo.org/record/3906723/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
). Valve face uneven with two parts of distinctly different morphology. Marginal area with short, radiating striae exceeding ¹/8–¼ of valve diameter, with ca 24–30 striae in
|
||
<quantity id="EB6670D8FFBEFFF583AAF498FCC14B9C" box="[735,813,1963,1990]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="mm" value="10.0">10 µm</quantity>
|
||
. Central area uneven and colliculate with scattered central areolae that are not radially arranged. Frustules rectangular in connective view.
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBFFFF4819FF238FDD44D7C" blockId="3.[189,1399,267,541]" box="[234,568,267,294]" pageId="3" pageNumber="4">
|
||
<emphasis id="1EEA012FFFBFFFF4819FF238FE344D7F" bold="true" box="[234,472,267,294]" pageId="3" pageNumber="4">SEM external view</emphasis>
|
||
(
|
||
<figureCitation id="B4A5C1B8FFBFFFF4809DF238FDC34D7C" box="[488,559,267,294]" captionStart="Fig" captionStartId="4.[189,232,1894,1920]" captionTargetBox="[238,1350,265,1854]" captionTargetId="figure@4.[238,1350,265,1854]" captionTargetPageId="4" captionText="Fig. 2. Tertiarius minutulus sp. nov., SEM external valve views. A. Valve view showing the marginal openings of the fultoportulae (see white arrow). B–D. View of the whole frustule. E. Close view of marginal openings of the fultoportula (see white arrow). F. Close view of the external openings of the valve face fultoportulae (see white arrows). Scale bars: A–D = 2.0 µm; E–F = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906725" httpUri="https://zenodo.org/record/3906725/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBFFFF481C8F21CFE624E47" blockId="3.[189,1399,267,541]" pageId="3" pageNumber="4">
|
||
Valve face uneven and colliculate (
|
||
<figureCitation id="B4A5C1B8FFBFFFF48321F21CFD754D13" box="[596,665,303,329]" captionStart="Fig" captionStartId="4.[189,232,1894,1920]" captionTargetBox="[238,1350,265,1854]" captionTargetId="figure@4.[238,1350,265,1854]" captionTargetPageId="4" captionText="Fig. 2. Tertiarius minutulus sp. nov., SEM external valve views. A. Valve view showing the marginal openings of the fultoportulae (see white arrow). B–D. View of the whole frustule. E. Close view of marginal openings of the fultoportula (see white arrow). F. Close view of the external openings of the valve face fultoportulae (see white arrows). Scale bars: A–D = 2.0 µm; E–F = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906725" httpUri="https://zenodo.org/record/3906725/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
A–C), with small granules in central area (
|
||
<figureCitation id="B4A5C1B8FFBFFFF485FCF21CFB224D13" box="[1161,1230,303,329]" captionStart="Fig" captionStartId="4.[189,232,1894,1920]" captionTargetBox="[238,1350,265,1854]" captionTargetId="figure@4.[238,1350,265,1854]" captionTargetPageId="4" captionText="Fig. 2. Tertiarius minutulus sp. nov., SEM external valve views. A. Valve view showing the marginal openings of the fultoportulae (see white arrow). B–D. View of the whole frustule. E. Close view of marginal openings of the fultoportula (see white arrow). F. Close view of the external openings of the valve face fultoportulae (see white arrows). Scale bars: A–D = 2.0 µm; E–F = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906725" httpUri="https://zenodo.org/record/3906725/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
D–E). Areolae present in central area with simple round openings. Marginal striae short, occluded by cribra, perforated with pores of variable size. Line of bigger pores bordering each costa (
|
||
<figureCitation id="B4A5C1B8FFBFFFF48295F246FBCF4DCA" box="[992,1059,373,400]" captionStart="Fig" captionStartId="4.[189,232,1894,1920]" captionTargetBox="[238,1350,265,1854]" captionTargetId="figure@4.[238,1350,265,1854]" captionTargetPageId="4" captionText="Fig. 2. Tertiarius minutulus sp. nov., SEM external valve views. A. Valve view showing the marginal openings of the fultoportulae (see white arrow). B–D. View of the whole frustule. E. Close view of marginal openings of the fultoportula (see white arrow). F. Close view of the external openings of the valve face fultoportulae (see white arrows). Scale bars: A–D = 2.0 µm; E–F = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906725" httpUri="https://zenodo.org/record/3906725/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
D–E). Marginal fultoportulae situated on costae, close to margin edge (
|
||
<figureCitation id="B4A5C1B8FFBFFFF483EDF2AAFD314DE9" box="[664,733,409,435]" captionStart="Fig" captionStartId="4.[189,232,1894,1920]" captionTargetBox="[238,1350,265,1854]" captionTargetId="figure@4.[238,1350,265,1854]" captionTargetPageId="4" captionText="Fig. 2. Tertiarius minutulus sp. nov., SEM external valve views. A. Valve view showing the marginal openings of the fultoportulae (see white arrow). B–D. View of the whole frustule. E. Close view of marginal openings of the fultoportula (see white arrow). F. Close view of the external openings of the valve face fultoportulae (see white arrows). Scale bars: A–D = 2.0 µm; E–F = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906725" httpUri="https://zenodo.org/record/3906725/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
A–B), situated on every 6
|
||
<superScript id="DBEB7075FFBFFFF48573F2A4FBCF4DE9" attach="none" box="[1030,1059,407,435]" fontSize="6" pageId="3" pageNumber="4">th–</superScript>
|
||
7
|
||
<superScript id="DBEB7075FFBFFFF48544F2A4FBD24DFF" attach="left" box="[1073,1086,407,421]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
costa. Externally marginal fultoportulae with simple, slightly elevated and round opening (
|
||
<figureCitation id="B4A5C1B8FFBFFFF482B5F28FFBAF4D8D" box="[960,1091,444,471]" captionStart="Fig" captionStartId="4.[189,232,1894,1920]" captionTargetBox="[238,1350,265,1854]" captionTargetId="figure@4.[238,1350,265,1854]" captionTargetPageId="4" captionText="Fig. 2. Tertiarius minutulus sp. nov., SEM external valve views. A. Valve view showing the marginal openings of the fultoportulae (see white arrow). B–D. View of the whole frustule. E. Close view of marginal openings of the fultoportula (see white arrow). F. Close view of the external openings of the valve face fultoportulae (see white arrows). Scale bars: A–D = 2.0 µm; E–F = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906725" httpUri="https://zenodo.org/record/3906725/files/figure.png" pageId="3" pageNumber="4">Fig. 2C, E</figureCitation>
|
||
). One to three valve face fultoportulae present, with external small openings, round (
|
||
<figureCitation id="B4A5C1B8FFBFFFF4821AF2ECFC584DA0" box="[879,948,479,506]" captionStart="Fig" captionStartId="4.[189,232,1894,1920]" captionTargetBox="[238,1350,265,1854]" captionTargetId="figure@4.[238,1350,265,1854]" captionTargetPageId="4" captionText="Fig. 2. Tertiarius minutulus sp. nov., SEM external valve views. A. Valve view showing the marginal openings of the fultoportulae (see white arrow). B–D. View of the whole frustule. E. Close view of marginal openings of the fultoportula (see white arrow). F. Close view of the external openings of the valve face fultoportulae (see white arrows). Scale bars: A–D = 2.0 µm; E–F = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906725" httpUri="https://zenodo.org/record/3906725/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
A–B). Single solid ligulate girdle band present (
|
||
<figureCitation id="B4A5C1B8FFBFFFF48057F130FE6C4E47" box="[290,384,515,541]" captionStart="Fig" captionStartId="4.[189,232,1894,1920]" captionTargetBox="[238,1350,265,1854]" captionTargetId="figure@4.[238,1350,265,1854]" captionTargetPageId="4" captionText="Fig. 2. Tertiarius minutulus sp. nov., SEM external valve views. A. Valve view showing the marginal openings of the fultoportulae (see white arrow). B–D. View of the whole frustule. E. Close view of marginal openings of the fultoportula (see white arrow). F. Close view of the external openings of the valve face fultoportulae (see white arrows). Scale bars: A–D = 2.0 µm; E–F = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906725" httpUri="https://zenodo.org/record/3906725/files/figure.png" pageId="3" pageNumber="4">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBFFFF4819FF177FDD94E04" blockId="3.[189,1399,580,925]" box="[234,565,580,606]" pageId="3" pageNumber="4">
|
||
<emphasis id="1EEA012FFFBFFFF4819FF177FE394E04" bold="true" box="[234,469,580,606]" pageId="3" pageNumber="4">SEM internal view</emphasis>
|
||
(
|
||
<figureCitation id="B4A5C1B8FFBFFFF48090F177FDC74E04" box="[485,555,580,606]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBFFFF481C8F15BFD264FC6" blockId="3.[189,1399,580,925]" pageId="3" pageNumber="4">
|
||
Internal valve face flat with several central areolae occluded with domed cribra (arrow in
|
||
<figureCitation id="B4A5C1B8FFBFFFF4847FF15BFA844ED8" box="[1290,1384,616,642]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig. 3B</figureCitation>
|
||
). Marginal areolae loculate, crossed with inwardly raised circumferential silica trabeculae (arrowhead in
|
||
<figureCitation id="B4A5C1B8FFBFFFF481C8F19DFEF44E92" box="[189,280,686,712]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig. 3B</figureCitation>
|
||
), while central areolae occluded with domed cribra (
|
||
<figureCitation id="B4A5C1B8FFBFFFF482F3F19DFC274E92" box="[902,971,686,712]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
|
||
B–C). Costae strongly thickened, ca
|
||
<quantity id="EB6670D8FFBFFFF481C8F1E1FF154EB6" box="[189,249,722,748]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="4" unit="in" value="30.0">30 in</quantity>
|
||
<quantity id="EB6670D8FFBFFFF48074F1E1FEA24EB6" box="[257,334,722,748]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="mm" value="10.0">10 µm</quantity>
|
||
(
|
||
<figureCitation id="B4A5C1B8FFBFFFF4802BF1E1FE574EB6" box="[350,443,722,748]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig. 3A</figureCitation>
|
||
, E–F). Marginal fultoportulae with short tube located on short and narrow costae (
|
||
<figureCitation id="B4A5C1B8FFBFFFF481B3F1C6FECF4F55" box="[198,291,757,783]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig. 3A</figureCitation>
|
||
). Each marginal fultoportula with two satellite pores in lateral position (arrow in
|
||
<figureCitation id="B4A5C1B8FFBFFFF485A1F1C6FADD4F55" box="[1236,1329,757,783]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig. 3A</figureCitation>
|
||
). One to three valve face fultoportulae present. Valve face fultoportulae with short central tube surrounded with 2–3 satellite pores (arrow in
|
||
<figureCitation id="B4A5C1B8FFBFFFF4837EF00FFD894F0C" box="[523,613,828,854]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig. 3E</figureCitation>
|
||
). One rimoportula present, located at valve face/mantle junction in the middle of costa (arrow in
|
||
<figureCitation id="B4A5C1B8FFBFFFF48366F06CFD844F23" box="[531,616,863,889]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig. 3F</figureCitation>
|
||
) or on its side (arrow in
|
||
<figureCitation id="B4A5C1B8FFBFFFF482F0F06CFC364F23" box="[901,986,863,889]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig 3D</figureCitation>
|
||
). Rimoportula consists of short and narrow labium with oblique slit (
|
||
<figureCitation id="B4A5C1B8FFBFFFF4834AF0B1FD6F4FC6" box="[575,643,898,924]" captionStart="Fig" captionStartId="5.[189,232,1791,1817]" captionTargetBox="[274,1313,265,1749]" captionTargetId="figure@5.[274,1313,265,1749]" captionTargetPageId="5" captionText="Fig. 3. Tertiarius minutulus sp. nov., SEM internal valve views. A. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow). B. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow). C. View of the whole frustule. D. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle. E. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows). F. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm." figureDoi="http://doi.org/10.5281/zenodo.3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
|
||
D–F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="64848EB6FFBFFFF481C8F0F7FBEF4856" box="[189,1027,964,1037]" pageId="3" pageNumber="4" type="distribution">
|
||
<paragraph id="2C21DD3DFFBFFFF481C8F0F7FEBA4F84" blockId="3.[189,342,964,990]" box="[189,342,964,990]" pageId="3" pageNumber="4">
|
||
<heading id="77696A51FFBFFFF481C8F0F7FEBA4F84" bold="true" box="[189,342,964,990]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="1EEA012FFFBFFFF481C8F0F7FEBA4F84" bold="true" box="[189,342,964,990]" pageId="3" pageNumber="4">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBFFFF481C8F0C1FBEF4856" blockId="3.[189,1027,1010,1037]" box="[189,1027,1010,1037]" pageId="3" pageNumber="4">
|
||
Freshwater fossil species observed only at its
|
||
<typeStatus id="F325639FFFBFFFF483A3F0C0FCE54857" box="[726,777,1011,1037]" pageId="3" pageNumber="4">type</typeStatus>
|
||
locality, Lake
|
||
<collectingRegion id="EE5A13DFFFBFFFF482C3F0C1FC124856" box="[950,1022,1010,1036]" country="Macedonia" name="Ohrid" pageId="3" pageNumber="4">Ohrid</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="64848EB6FFBFFFF181C8F707FA9A4DEF" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="diagnosis">
|
||
<paragraph id="2C21DD3DFFBFFFF481C8F707FE2A4814" blockId="3.[189,454,1076,1102]" box="[189,454,1076,1102]" pageId="3" pageNumber="4">
|
||
<heading id="77696A51FFBFFFF481C8F707FE2A4814" bold="true" box="[189,454,1076,1102]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="1EEA012FFFBFFFF481C8F707FE2A4814" bold="true" box="[189,454,1076,1102]" pageId="3" pageNumber="4">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBFFFF181C8F750FA9A4DEF" blockId="3.[189,1399,1123,1361]" lastBlockId="6.[189,1399,267,437]" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
|
||
The main differential features of
|
||
<taxonomicName id="EB9EA6BEFFBFFFF4833DF757FD374827" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[584,731,1123,1149]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="3" pageNumber="4" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBFFFF4833DF757FD374827" box="[584,731,1123,1149]" italics="true" pageId="3" pageNumber="4">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBFFFF48391F750FCAC4827" box="[740,832,1123,1149]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
that can be observed in LM are its small valve size (diameter = 3.5–8.0 µm) and the presence of central scattered areolae that are not radially arranged.
|
||
<taxonomicName id="EB9EA6BEFFBFFFF481C8F799FBF0489E" authority="(Pantocsek) Hakansson & Khursevich (1997: 22)" authorityName="Hakansson & Khursevich" authorityPageNumber="22" authorityYear="1997" baseAuthorityName="Pantocsek" box="[189,1052,1193,1220]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="3" pageNumber="4" phylum="Ochrophyta" rank="species" species="pygmaeus">
|
||
<emphasis id="1EEA012FFFBFFFF481C8F799FE5E4899" box="[189,434,1194,1219]" italics="true" pageId="3" pageNumber="4">Tertiarius pygmaeus</emphasis>
|
||
(Pantocsek)
|
||
<bibRefCitation id="480FA0CCFFBFFFF48323F79AFBF0489E" author="Hakansson H. & Khursevich G." box="[598,1052,1193,1220]" pageId="3" pageNumber="4" pagination="19 - 33" refId="ref6776" refString="Hakansson H. & Khursevich G. 1997. Tertiarius gen. nov., a new genus in the Bacillariophyceae, the transfer of some Cyclotelloid species and a comparison to closely related genera. Diatom Research 12: 19 - 33. https: // doi. org / 10.1080 / 0269249 x. 1997.9705399" type="journal article" year="1997">Håkansson & Khursevich (1997: 22)</bibRefCitation>
|
||
</taxonomicName>
|
||
is characterized by a valve diameter ranging from 8.0–16.0 µm, and occasionally the areolae are not arranged in radial striae (e.g.,
|
||
<bibRefCitation id="480FA0CCFFBFFFF481C8F7C3FE924950" author="Pantocsek J." box="[189,382,1264,1290]" pageId="3" pageNumber="4" refId="ref7664" refString="Pantocsek J. 1892. Beitrage zur Kenntniss der fossilen Bacillarien Ungarns. III. Susswasser Bacillarien. Anhang: Analysen 15 neuer Depots von Bulgarien, Japan, Mahren, Rusland und Ungarn. Julius Platzko, Nagytapolcsany." type="book" year="1892">Pantocsek 1892</bibRefCitation>
|
||
: figs 2: 22, 4: 59;
|
||
<bibRefCitation id="480FA0CCFFBFFFF4831DF7C3FCD14950" author="Houk V. & Klee R. & Tanaka H." box="[616,829,1264,1290]" pageId="3" pageNumber="4" pagination="1 - 498" refId="ref6833" refString="Houk V., Klee R. & Tanaka H. 2010. Atlas of freshwater centric diatoms with a brief key and descriptions, Part III. Stephanodiscaceae A, Cyclotella, Tertiarius, Discostella. Fottea 10 (Supplement): 1 - 498." type="journal article" year="2010">
|
||
Houk
|
||
<emphasis id="1EEA012FFFBFFFF483C1F7C2FD1F4950" box="[692,755,1264,1290]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
: fig. 296: 1, 4). Important differences between
|
||
<taxonomicName id="EB9EA6BEFFBFFFF481C8F627FEA14977" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[189,333,1299,1325]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="3" pageNumber="4" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBFFFF481C8F627FEA14977" box="[189,333,1299,1325]" italics="true" pageId="3" pageNumber="4">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBFFFF48027F627FE404974" box="[338,428,1300,1326]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="EB9EA6BEFFBFFFF48097F627FD9F4977" authorityName="Hakansson & Khursevich" authorityYear="1997" baseAuthorityName="Pantocsek" box="[482,627,1300,1325]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="3" pageNumber="4" phylum="Ochrophyta" rank="species" species="pygmaeus">
|
||
<emphasis id="1EEA012FFFBFFFF48097F627FD9F4977" box="[482,627,1300,1325]" italics="true" pageId="3" pageNumber="4">T. pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
can be observed with SEM. The marginal striae in
|
||
<taxonomicName id="EB9EA6BEFFBFFFF485B7F627FABF4977" authorityName="Hakansson & Khursevich" authorityYear="1997" baseAuthorityName="Pantocsek" box="[1218,1363,1300,1325]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="3" pageNumber="4" phylum="Ochrophyta" rank="species" species="pygmaeus">
|
||
<emphasis id="1EEA012FFFBFFFF485B7F627FABF4977" box="[1218,1363,1300,1325]" italics="true" pageId="3" pageNumber="4">T. pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
do not have a complex alveolar structure and are composed of regular rows of fine pori. Externally, distinct spines are present on every costa. Internally, the alveolus is round and small in
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1853AF23FFB334D7F" authorityName="Hakansson & Khursevich" authorityYear="1997" baseAuthorityName="Pantocsek" box="[1103,1247,268,293]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="pygmaeus">
|
||
<emphasis id="1EEA012FFFBAFFF1853AF23FFB334D7F" box="[1103,1247,268,293]" italics="true" pageId="6" pageNumber="7">T. pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
whereas it is larger and more elongated in
|
||
<taxonomicName id="EB9EA6BEFFBAFFF18365F203FD734D13" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[528,671,303,329]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBAFFF18365F203FD734D13" box="[528,671,303,329]" italics="true" pageId="6" pageNumber="7">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBAFFF183D0F203FD134D10" box="[677,767,304,330]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
In
|
||
<taxonomicName id="EB9EA6BEFFBAFFF18256F203FC584D13" authorityName="Hakansson & Khursevich" authorityYear="1997" baseAuthorityName="Pantocsek" box="[803,948,304,329]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="pygmaeus">
|
||
<emphasis id="1EEA012FFFBAFFF18256F203FC584D13" box="[803,948,304,329]" italics="true" pageId="6" pageNumber="7">T. pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
, the marginal fultoportulae are present on each costa, the number of valve face fultoportulae is one to seven scattered in the central area, and the rimoportula is situated inside of the alvelolar opening on a side of costa. Additionally, the costae are much thicker and with a lower density (
|
||
<quantity id="EB6670D8FFBAFFF183CDF2A8FCFD4DEF" box="[696,785,411,437]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="6" pageNumber="7" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">8–10 in</quantity>
|
||
<quantity id="EB6670D8FFBAFFF1826DF2A8FC8A4DEF" box="[792,870,411,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="mm" value="10.0">10 µm</quantity>
|
||
) in
|
||
<taxonomicName id="EB9EA6BEFFBAFFF182E6F2AFFBC94DEF" authorityName="Hakansson & Khursevich" authorityYear="1997" baseAuthorityName="Pantocsek" box="[915,1061,412,437]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="pygmaeus">
|
||
<emphasis id="1EEA012FFFBAFFF182E6F2AFFBC94DEF" box="[915,1061,412,437]" italics="true" pageId="6" pageNumber="7">T. pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="EB9EA6BEFFBAFFF185F6F2AFFAF84DEF" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[1155,1300,411,437]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBAFFF185F6F2AFFAF84DEF" box="[1155,1300,411,437]" italics="true" pageId="6" pageNumber="7">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBAFFF1846EF2A8FA9A4DEF" box="[1307,1398,411,437]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="78E18DB5FFBFFFF481C8F49EFD404BB1" ID-DOI="http://doi.org/10.5281/zenodo.3906723" ID-Zenodo-Dep="3906723" httpUri="https://zenodo.org/record/3906723/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1965,1991]" targetBox="[189,1398,1405,1923]" targetPageId="3">
|
||
<paragraph id="2C21DD3DFFBFFFF481C8F49EFD404BB1" blockId="3.[189,1399,1965,2027]" pageId="3" pageNumber="4">
|
||
<emphasis id="1EEA012FFFBFFFF481C8F49EFEE34B92" bold="true" box="[189,271,1965,1992]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
|
||
<taxonomicName id="EB9EA6BEFFBFFFF4806FF49DFDE04B9D" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[282,524,1965,1991]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="3" pageNumber="4" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBFFFF4806FF49DFDE04B9D" box="[282,524,1965,1991]" italics="true" pageId="3" pageNumber="4">Tertiarius minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBFFFF48362F49DFD944B92" box="[535,632,1966,1992]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, LM valve views, Lake Ohrid, North Macedonia. The asterisk indicates the holotype. Scale bar = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="78E18DB5FFB8FFF381C8F455FB424BB0" ID-DOI="http://doi.org/10.5281/zenodo.3906725" ID-Zenodo-Dep="3906725" httpUri="https://zenodo.org/record/3906725/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1894,1920]" targetBox="[238,1350,265,1854]" targetPageId="4">
|
||
<paragraph id="2C21DD3DFFB8FFF381C8F455FB424BB0" blockId="4.[189,1399,1894,2026]" pageId="4" pageNumber="5">
|
||
<emphasis id="1EEA012FFFB8FFF381C8F455FEE04BDA" bold="true" box="[189,268,1894,1920]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
|
||
<taxonomicName id="EB9EA6BEFFB8FFF38060F454FDE94BDA" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[277,517,1894,1920]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="4" pageNumber="5" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFB8FFF38060F454FDE94BDA" box="[277,517,1894,1920]" italics="true" pageId="4" pageNumber="5">Tertiarius minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFB8FFF38378F455FD804BDA" box="[525,620,1894,1920]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, SEM external valve views.
|
||
<emphasis id="1EEA012FFFB8FFF382B6F455FC344BDA" bold="true" box="[963,984,1894,1920]" pageId="4" pageNumber="5">A</emphasis>
|
||
. Valve view showing the marginal openings of the fultoportulae (see white arrow).
|
||
<emphasis id="1EEA012FFFB8FFF3827CF4BAFCAC4BF9" bold="true" box="[777,832,1929,1956]" pageId="4" pageNumber="5">B–D</emphasis>
|
||
. View of the whole frustule.
|
||
<emphasis id="1EEA012FFFB8FFF385DDF4BAFB504BF9" bold="true" box="[1192,1212,1929,1955]" pageId="4" pageNumber="5">E</emphasis>
|
||
. Close view of marginal openings of the fultoportula (see white arrow).
|
||
<emphasis id="1EEA012FFFB8FFF3822BF49EFC9C4B9D" bold="true" box="[862,880,1965,1991]" pageId="4" pageNumber="5">F</emphasis>
|
||
. Close view of the external openings of the valve face fultoportulae (see white arrows). Scale bars: A–D = 2.0 µm; E–F = 1.0 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="78E18DB5FFB9FFF281C8F5CCFCD84BB6" ID-DOI="http://doi.org/10.5281/zenodo.3906727" ID-Zenodo-Dep="3906727" httpUri="https://zenodo.org/record/3906727/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1791,1817]" targetBox="[274,1313,265,1749]" targetPageId="5">
|
||
<paragraph id="2C21DD3DFFB9FFF281C8F5CCFCD84BB6" blockId="5.[189,1399,1790,2028]" pageId="5" pageNumber="6">
|
||
<emphasis id="1EEA012FFFB9FFF281C8F5CCFEE74B43" bold="true" box="[189,267,1791,1817]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
|
||
<taxonomicName id="EB9EA6BEFFB9FFF28061F5CCFDEE4B42" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[276,514,1790,1816]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="5" pageNumber="6" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFB9FFF28061F5CCFDEE4B42" box="[276,514,1790,1816]" italics="true" pageId="5" pageNumber="6">Tertiarius minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFB9FFF2837FF5CCFD844B43" box="[522,616,1791,1817]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, SEM internal valve views.
|
||
<emphasis id="1EEA012FFFB9FFF282C2F5CCFC204B43" bold="true" box="[951,972,1791,1817]" pageId="5" pageNumber="6">A</emphasis>
|
||
. View of the whole frustule and the marginal fultoportulae surrounded by two satellite pores, cowlings absent (see white arrow).
|
||
<emphasis id="1EEA012FFFB9FFF28465F411FAC84B66" bold="true" box="[1296,1316,1826,1852]" pageId="5" pageNumber="6">B</emphasis>
|
||
. Close view of marginal areolae with circumferential silica trabeculae and central areolae occluded by domed cribra (see white arrow).
|
||
<emphasis id="1EEA012FFFB9FFF28096F45BFE144BD8" bold="true" box="[483,504,1896,1922]" pageId="5" pageNumber="6">C</emphasis>
|
||
. View of the whole frustule.
|
||
<emphasis id="1EEA012FFFB9FFF28239F45AFC8D4BD9" bold="true" box="[844,865,1897,1923]" pageId="5" pageNumber="6">D</emphasis>
|
||
. Close view of rimoportula (see white arrow) located on costa at junction between valve face/ mantle.
|
||
<emphasis id="1EEA012FFFB9FFF28205F4BFFC684BFC" bold="true" box="[880,900,1932,1958]" pageId="5" pageNumber="6">E</emphasis>
|
||
. Internal valve view showing the central fultoportulae with short central tube surrounded with 2–3 satellite pores (see white arrows).
|
||
<emphasis id="1EEA012FFFB9FFF28464F49CFACF4B93" bold="true" box="[1297,1315,1967,1993]" pageId="5" pageNumber="6">F</emphasis>
|
||
. Valve view. Scale bars: A, C, E–F = 2.0 µm; B, D = 1.0 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="64848EB6FFBAFFFE81C8F2EEFC534D80" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7" type="discussion">
|
||
<paragraph id="2C21DD3DFFBAFFF181C8F2EEFC2B48BD" blockId="6.[189,1399,477,1256]" pageId="6" pageNumber="7">
|
||
Another species with small-sized valves is
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1838FF2EDFE2C4E41" authority="Ognjanova-Rumenova, Jovanovska, Cvetkoska & Levkov" authorityName="Ognjanova-Rumenova, Jovanovska, Cvetkoska & Levkov" authorityYear="2015" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="mariovensis">
|
||
<emphasis id="1EEA012FFFBAFFF1838FF2EDFC5C4DAD" box="[762,944,478,503]" italics="true" pageId="6" pageNumber="7">T. mariovensis</emphasis>
|
||
Ognjanova-Rumenova, Jovanovska, Cvetkoska & Levkov
|
||
</taxonomicName>
|
||
(diameter 4.0–19.0 µm), recently described from a diatomite near the village
|
||
<collectingRegion id="EE5A13DFFFBAFFF181C8F117FECB4E64" box="[189,295,548,574]" country="Macedonia" name="Bitola" pageId="6" pageNumber="7">Manastir</collectingRegion>
|
||
, Mariovo Neogene Basin, Republic of North
|
||
<collectingCountry id="54899DADFFBAFFF18232F117FC254E64" box="[839,969,548,574]" name="Macedonia" pageId="6" pageNumber="7">Macedonia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="480FA0CCFFBAFFF182AFF117FF314E39" author="Ognjanova-Rumenova N. & Jovanovska E. & Cvetkoska A. & Levkov Z." pageId="6" pageNumber="7" pagination="51 - 62" refId="ref7609" refString="Ognjanova-Rumenova N., Jovanovska E., Cvetkoska A. & Levkov Z. 2015. Two new Tertiarius (Bacillariophyta, Coscinodiscophyceae) species from Mariovo Neogene Basin, Macedonia. Fottea 15 (1): 51 - 62. https: // doi. org / 10.5507 / fot. 2015.005" type="journal article" year="2015">
|
||
Ognjanova-Rumenova
|
||
<emphasis id="1EEA012FFFBAFFF1859AF116FAC74E64" box="[1263,1323,548,574]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
2015: 56
|
||
</bibRefCitation>
|
||
, figs 36–105). This species was also observed in several sediment samples from Lake
|
||
<collectingRegion id="EE5A13DFFFBAFFF185A2F17BFAF74E38" box="[1239,1307,584,610]" country="Macedonia" name="Ohrid" pageId="6" pageNumber="7">Ohrid</collectingRegion>
|
||
DEEP- 5045-1 core. However, both species can easily be differentiated by the structure of the central area. In
|
||
<taxonomicName id="EB9EA6BEFFBAFFF18195F1A2FE614EF0" authorityName="Ognjanova-Rumenova, Jovanovska, Cvetkoska & Levkov" authorityYear="2015" box="[224,397,657,682]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="mariovensis">
|
||
<emphasis id="1EEA012FFFBAFFF18195F1A2FE614EF0" box="[224,397,657,682]" italics="true" pageId="6" pageNumber="7">T. mariovensis</emphasis>
|
||
</taxonomicName>
|
||
, the areolae in the central area form radially arranged striae. The number of valve face fultoportulae in
|
||
<taxonomicName id="EB9EA6BEFFBAFFF180CFF186FD8B4E94" authorityName="Ognjanova-Rumenova, Jovanovska, Cvetkoska & Levkov" authorityYear="2015" box="[442,615,693,718]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="mariovensis">
|
||
<emphasis id="1EEA012FFFBAFFF180CFF186FD8B4E94" box="[442,615,693,718]" italics="true" pageId="6" pageNumber="7">T. mariovensis</emphasis>
|
||
</taxonomicName>
|
||
is higher (2–9) and the tube of the central fultoportula is always surrounded by three satellite pores. Additionally, the marginal fultoportulae are positioned on every costa, while the rimoportula is located inside and in the middle of the alveolus, but not connected to a costa as in
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1800EF013FDE24F63" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[379,526,799,825]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBAFFF1800EF013FDE24F63" box="[379,526,799,825]" italics="true" pageId="6" pageNumber="7">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBAFFF18362F013FD984F60" box="[535,628,800,826]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Another small-sized species,
|
||
<taxonomicName id="EB9EA6BEFFBAFFF182ACF013FE594F07" authority="Khursevich & Kociolek (2002: 333" authorityName="Khursevich & Kociolek" authorityYear="2002" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="distinctus">
|
||
<emphasis id="1EEA012FFFBAFFF182ACF013FB294F63" box="[985,1221,799,825]" italics="true" pageId="6" pageNumber="7">Tertiarius distinctus</emphasis>
|
||
<bibRefCitation id="480FA0CCFFBAFFF185BAF02CFE594F07" author="Khursevich G. & Kociolek J. P." pageId="6" pageNumber="7" pagination="331 - 349" refId="ref7021" refString="Khursevich G. & Kociolek J. P. 2002. New Tertiarius (Bacillariophyta: Stephanodiscaceae) species from Western North America. In: John J. (ed.) Proceedings of the 15 th International Diatom Symposium: 331 - 349. Biopress Limited, Bristol." type="book chapter" year="2002">Khursevich & Kociolek (2002: 333</bibRefCitation>
|
||
</taxonomicName>
|
||
, figs 1–5, 12–22), has a comparable valve size as
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1857DF077FB764F07" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[1032,1178,835,861]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBAFFF1857DF077FB764F07" box="[1032,1178,835,861]" italics="true" pageId="6" pageNumber="7">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBAFFF185D7F077FB124F04" box="[1186,1278,836,862]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(diameter 4.5–14.0 µm), but coarser areolae arranged in short or long radial striae composed of large areolae, while the marginal striae are composed of 4–6 rows of small pori. Marginal fultoportulae are present on every costa, while in the central area there are 1–5 valve face fultoportulae. The rimoportula is located near the base of a costa within the alveolus. Differences between
|
||
<taxonomicName id="EB9EA6BEFFBAFFF182C8F0E0FBA64FB6" authorityName="Khursevich & Kociolek" authorityYear="2002" box="[957,1098,978,1004]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="distinctus">
|
||
<emphasis id="1EEA012FFFBAFFF182C8F0E0FBA64FB6" box="[957,1098,978,1004]" italics="true" pageId="6" pageNumber="7">T. distinctus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EB9EA6BEFFBAFFF185F1F0E0FAF94FB6" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[1156,1301,978,1004]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBAFFF185F1F0E0FAF94FB6" box="[1156,1301,978,1004]" italics="true" pageId="6" pageNumber="7">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBAFFF18469F0E0FA9B4FB7" box="[1308,1399,979,1005]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be noticed in the structure of the marginal and central striae, the number of valve face and marginal fultoportulae, and the position of the rimoportula. Smaller specimens of
|
||
<taxonomicName id="EB9EA6BEFFBAFFF18545F728FE5E4802" authority="Khursevich & Kociolek (2002: 336" authorityName="Khursevich & Kociolek" authorityYear="2002" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="indigenus">
|
||
<emphasis id="1EEA012FFFBAFFF18545F728FB2F486E" box="[1072,1219,1050,1076]" italics="true" pageId="6" pageNumber="7">T. indigenus</emphasis>
|
||
<bibRefCitation id="480FA0CCFFBAFFF185BBF729FE5E4802" author="Khursevich G. & Kociolek J. P." pageId="6" pageNumber="7" pagination="331 - 349" refId="ref7021" refString="Khursevich G. & Kociolek J. P. 2002. New Tertiarius (Bacillariophyta: Stephanodiscaceae) species from Western North America. In: John J. (ed.) Proceedings of the 15 th International Diatom Symposium: 331 - 349. Biopress Limited, Bristol." type="book chapter" year="2002">Khursevich & Kociolek (2002: 336</bibRefCitation>
|
||
</taxonomicName>
|
||
, figs 6–11, 23–32) have a valve size comparable to
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1857DF70CFB754802" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[1032,1177,1086,1112]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBAFFF1857DF70CFB754802" box="[1032,1177,1086,1112]" italics="true" pageId="6" pageNumber="7">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBAFFF185D3F70CFAEC4803" box="[1190,1280,1087,1113]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(diameter 5.0–
|
||
<quantity id="EB6670D8FFBAFFF18184F751FEB64826" box="[241,346,1122,1149]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="6" pageNumber="7" unit="mm" value="27.5">27.5 µm</quantity>
|
||
). However, differences between these two species can be noticed in the striae structure (longer marginal striae composed of 4–5 regular rows or pori), central area (colliculate central area with 3–5 large areolae and several ʻbumpsʼ), presence of fultoportulae on each costa and one to several rimportulae located on thinner costae within the alveolar chamber.
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBAFFF181C8F623FC064A7F" blockId="6.[189,1399,1295,1573]" pageId="6" pageNumber="7">
|
||
<taxonomicName id="EB9EA6BEFFBAFFF181C8F623FD324973" authority="Khursevich & Fedenya" authorityName="Khursevich & Fedenya" box="[189,734,1295,1322]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="baicalensis">
|
||
<emphasis id="1EEA012FFFBAFFF181C8F623FE524973" box="[189,446,1295,1321]" italics="true" pageId="6" pageNumber="7">Tertiarius baicalensis</emphasis>
|
||
Khursevich & Fedenya
|
||
</taxonomicName>
|
||
(in
|
||
<bibRefCitation id="480FA0CCFFBAFFF18261F63CFB854973" author="Khursevich G. K. & Fedenya S. A. & Kuzmin M. I. & Karabanov E. B. & Williams D. F. & Prokopenko A. A." box="[788,1129,1295,1322]" pageId="6" pageNumber="7" pagination="305 - 321" refId="ref7254" refString="Khursevich G. K., Fedenya S. A., Kuzmin M. I., Karabanov E. B., Williams D. F. & Prokopenko A. A. 2003. Morphology of new taxa of the Class Centrophyceae (Bacillariophyta) from the Pliocene and Pleistocene deposits of Lake Baikal (Siberia). Algologia 13 (3): 305 - 321. https: // doi. org / 10.1615 / InterJAlgae. v 5. i 3.70" type="journal article" year="2003">
|
||
Khursevich
|
||
<emphasis id="1EEA012FFFBAFFF182D0F623FC0F4973" box="[933,995,1295,1321]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
2003: 306
|
||
</bibRefCitation>
|
||
, figs 1: 1, 2, 4, 5, 14) is characterized by circular valves with flat face and diameter of
|
||
<quantity id="EB6670D8FFBAFFF182BCF600FB8F4914" box="[969,1123,1331,1358]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.66" metricValueMin="0.64" pageId="6" pageNumber="7" unit="mm" value="11.5" valueMax="16.6" valueMin="6.4">6.4–16.6 µm</quantity>
|
||
, three to 17 valve face fultoportulae arranged in radial rows and one to four sessile rimoportulae located in the submarginal zone of the valve face. Differences between
|
||
<taxonomicName id="EB9EA6BEFFBAFFF183BBF64FFC8C49CF" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[718,864,1403,1429]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBAFFF183BBF64FFC8C49CF" box="[718,864,1403,1429]" italics="true" pageId="6" pageNumber="7">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBAFFF1821CF648FC2949CF" box="[873,965,1403,1429]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="EB9EA6BEFFBAFFF18575F64FFB4E49CF" box="[1024,1186,1403,1429]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="baicalensis">
|
||
<emphasis id="1EEA012FFFBAFFF18575F64FFB4E49CF" box="[1024,1186,1403,1429]" italics="true" pageId="6" pageNumber="7">T. baicalensis</emphasis>
|
||
</taxonomicName>
|
||
can be noticed in the ornamentation of the central area (areolae organized in radial striae in
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1855BF6ACFB3C49E3" box="[1070,1232,1439,1465]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="baicalensis">
|
||
<emphasis id="1EEA012FFFBAFFF1855BF6ACFB3C49E3" box="[1070,1232,1439,1465]" italics="true" pageId="6" pageNumber="7">T. baicalensis</emphasis>
|
||
</taxonomicName>
|
||
), number and position of valve face fultoportulae, number of satellite pores on marginal fultoportulae (three satellite pores in
|
||
<taxonomicName id="EB9EA6BEFFBAFFF18051F6D4FE2A4A5A" box="[292,454,1510,1536]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="baicalensis">
|
||
<emphasis id="1EEA012FFFBAFFF18051F6D4FE2A4A5A" box="[292,454,1510,1536]" italics="true" pageId="6" pageNumber="7">T. baicalensis</emphasis>
|
||
</taxonomicName>
|
||
), and position of rimoportula(e) that are located in the submarginal zone of the valve face (not connected with costa or within the alveolar chamber).
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBAFFF181C8F57FFCD14B64" blockId="6.[189,1399,1612,1854]" pageId="6" pageNumber="7">
|
||
Another small-celled species,
|
||
<taxonomicName id="EB9EA6BEFFBAFFF18357F57FFCC64A3C" authorityName="Jovanovska, Cvetkoska, Tofilovska, Ognjanova-Rumenova & Levkov" authorityYear="2016" box="[546,810,1612,1638]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Cribrionella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="ohridana">
|
||
<emphasis id="1EEA012FFFBAFFF18357F57FFCC64A3C" box="[546,810,1612,1638]" italics="true" pageId="6" pageNumber="7">Cribrionella ohridana</emphasis>
|
||
</taxonomicName>
|
||
, was recently described from the same core from Lake
|
||
<collectingRegion id="EE5A13DFFFBAFFF18074F543FEA94AD0" box="[257,325,1648,1674]" country="Macedonia" name="Ohrid" pageId="6" pageNumber="7">Ohrid</collectingRegion>
|
||
(
|
||
<bibRefCitation id="480FA0CCFFBAFFF18022F543FD934AD1" author="Jovanovska E. & Cvetkoska A. & Tofilovska S. & Ognjanova-Rumenova N. & Levkov Z." box="[343,639,1648,1675]" pageId="6" pageNumber="7" pagination="31 - 42" refId="ref6957" refString="Jovanovska E., Cvetkoska A., Tofilovska S., Ognjanova-Rumenova N. & Levkov Z. 2016 b. Description of a new fossil diatom genus, Cribrionella gen. nov. (Bacillariophyta) from Quaternary sediments of Lake Ohrid. Phytotaxa 252: 31 - 42. https: // doi. org / 10.11646 / phytotaxa. 252.1.3" type="journal article" year="2016">
|
||
Jovanovska
|
||
<emphasis id="1EEA012FFFBAFFF1809EF542FDCB4AD0" box="[491,551,1648,1674]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
2016b
|
||
</bibRefCitation>
|
||
). Both species share several characters such as small size (2.0–
|
||
<quantity id="EB6670D8FFBAFFF181C8F5A7FEFE4AF4" box="[189,274,1684,1711]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="7" unit="mm" value="7.5">7.5 µm</quantity>
|
||
in diameter), the presence of areolae in the central area and a single rimoportula situated on a costa, marginal fultoportulae located on each 4
|
||
<superScript id="DBEB7075FFBAFFF18383F585FCF84A89" attach="none" box="[758,788,1718,1747]" fontSize="6" pageId="6" pageNumber="7">th–</superScript>
|
||
5
|
||
<superScript id="DBEB7075FFBAFFF18254F585FCC24A9E" attach="left" box="[801,814,1718,1732]" fontSize="6" pageId="6" pageNumber="7">th</superScript>
|
||
costa surrounded by two satellite pores and the presence of inwardly raised circumferential silica trabeculae. However, differences between these two species can be noticed in the absence of valve face fultoportula in
|
||
<taxonomicName id="EB9EA6BEFFBAFFF182CEF433FBA44B43" authorityName="Jovanovska, Cvetkoska, Tofilovska, Ognjanova-Rumenova & Levkov" authorityYear="2016" box="[955,1096,1791,1818]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Cribrionella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="ohridana">
|
||
<emphasis id="1EEA012FFFBAFFF182CEF433FBA44B43" box="[955,1096,1791,1818]" italics="true" pageId="6" pageNumber="7">C. ohridana</emphasis>
|
||
</taxonomicName>
|
||
and areolae internally not occluded with domed cribra as in
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1833FF417FD374B67" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[586,731,1827,1853]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFBAFFF1833FF417FD374B67" box="[586,731,1827,1853]" italics="true" pageId="6" pageNumber="7">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFBAFFF18397F417FCD14B64" box="[738,829,1828,1854]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="2C21DD3DFFBAFFFE81C8F456FC534D80" blockId="6.[189,1399,1893,2028]" lastBlockId="9.[189,1399,267,474]" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7">
|
||
One of the dominant and very frequent species in fossil diatom assemblages of Lake
|
||
<collectingRegion id="EE5A13DFFFBAFFF185ECF456FB314B25" box="[1177,1245,1893,1919]" country="Macedonia" name="Ohrid" pageId="6" pageNumber="7">Ohrid</collectingRegion>
|
||
is
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1858BF456FDF24BF9" authority="(Jurilj) Cvetkoska." authorityName="Cvetkoska." baseAuthorityName="Jurilj" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Cyclotella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minuscula">
|
||
<emphasis id="1EEA012FFFBAFFF1858BF456FEDA4BF9" italics="true" pageId="6" pageNumber="7">Cyclotella minuscula</emphasis>
|
||
(Jurilj) Cvetkoska.
|
||
</taxonomicName>
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1835DF4BAFCCF4BF9" box="[552,803,1929,1955]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Cyclotella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minuscula">
|
||
<emphasis id="1EEA012FFFBAFFF1835DF4BAFCCF4BF9" box="[552,803,1929,1955]" italics="true" pageId="6" pageNumber="7">Cyclotella minuscula</emphasis>
|
||
</taxonomicName>
|
||
was recently transferred to
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1850CF4BAFEB84B9D" authority="(Schutt) De Toni & Forti" authorityName="De Toni & Forti" baseAuthorityName="Schutt" class="Bacillariophyceae" family="Coscinodiscaceae" genus="Lindavia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Coscinodiscales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="1EEA012FFFBAFFF1850CF4BAFB0D4BF9" box="[1145,1249,1929,1955]" italics="true" pageId="6" pageNumber="7">Lindavia</emphasis>
|
||
(Schütt) De Toni & Forti
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName id="EB9EA6BEFFBAFFF1800BF49DFB684B9D" authority="(Jurilj) Nakov, Guillory, Julius, Theriot & Alverson" authorityName="Nakov, Guillory, Julius, Theriot & Alverson" baseAuthorityName="Jurilj" box="[382,1156,1965,1992]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Cyclotella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minuscula">
|
||
<emphasis id="1EEA012FFFBAFFF1800BF49DFDFB4B9D" box="[382,535,1965,1991]" italics="true" pageId="6" pageNumber="7">L. minuscula</emphasis>
|
||
(Jurilj) Nakov, Guillory, Julius, Theriot & Alverson
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="480FA0CCFFBAFFF185E1F49EFA824B92" author="Nakov T. & Guillory W. X. & Julius M. L. & Theriot E. C. & Alverson A. J." box="[1172,1390,1965,1992]" pageId="6" pageNumber="7" pagination="249 - 264" refId="ref7524" refString="Nakov T., Guillory W. X., Julius M. L., Theriot E. C. & Alverson A. J. 2015. Towards a phylogenetic classification of species belonging to the diatom genus Cyclotella (Bacillariophyceae): Transfer of species formerly placed in Puncticulata, Handmannia, Pliocaenicus and Cyclotella to the genus Lindavia. Phytotaxa 217 (3): 249 - 264. https: // doi. org / 10.11646 / phytotaxa. 217.3.2" type="journal article" year="2015">
|
||
Nakov
|
||
<emphasis id="1EEA012FFFBAFFF18598F49DFAC54B9D" box="[1261,1321,1965,1991]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
) and later to the genus
|
||
<taxonomicName id="EB9EA6BEFFBAFFF180B4F4E2FD114BB1" authority="Kiss & Acs" authorityName="Kiss & Acs" box="[449,765,1999,2027]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Pantocsekiella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="1EEA012FFFBAFFF180B4F4E2FD824BB1" box="[449,622,2001,2027]" italics="true" pageId="6" pageNumber="7">Pantocsekiella</emphasis>
|
||
Kiss & Ács
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName id="EB9EA6BEFFBAFFF18250F4E1FB4E4BB1" authority="(Jurilj) Kiss & Acs" authorityName="Kiss & Acs" baseAuthorityName="Jurilj" box="[805,1186,1999,2028]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Cyclotella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="6" pageNumber="7" phylum="Ochrophyta" rank="species" species="minuscula">
|
||
<emphasis id="1EEA012FFFBAFFF18250F4E1FC574BB1" box="[805,955,2001,2027]" italics="true" pageId="6" pageNumber="7">P. minuscula</emphasis>
|
||
(Jurilj) Kiss & Ács
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="480FA0CCFFBAFFF185C4F4FCFA874BB6" author="Acs E. & Ari E. & Duleba M. & Dressler M. & Genkal S. I. & Jako E. & Rimet F. & Ector L. & Kiss K. T." box="[1201,1387,1999,2028]" pageId="6" pageNumber="7" pagination="56 - 78" refId="ref6250" refString="Acs E., Ari E., Duleba M., Dressler M., Genkal S. I., Jako E., Rimet F., Ector L. & Kiss K. T. 2016. Pantocsekiella, a new centric diatom genus based on morphological and genetic studies. Fottea 16 (1): 56 - 78. https: // doi. org / 10.5507 / fot. 2015.028" type="journal article" year="2016">
|
||
Ács
|
||
<emphasis id="1EEA012FFFBAFFF1859DF4E1FACF4BB1" box="[1256,1315,2001,2027]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
). Both species,
|
||
<taxonomicName id="EB9EA6BEFFB5FFFE802BF23FFDA74D7F" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[350,587,267,293]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFB5FFFE802BF23FFDA74D7F" box="[350,587,267,293]" italics="true" pageId="9" pageNumber="10">Tertiarius minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFB5FFFE8324F23FFD464D7C" box="[593,682,268,294]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="EB9EA6BEFFB5FFFE8395F238FC344D7F" box="[736,984,267,293]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Cyclotella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="minuscula">
|
||
<emphasis id="1EEA012FFFB5FFFE8395F238FC344D7F" box="[736,984,267,293]" italics="true" pageId="9" pageNumber="10">Cyclotella minuscula</emphasis>
|
||
</taxonomicName>
|
||
, have a similar valve size (diameter
|
||
<quantity id="EB6670D8FFB5FFFE81C8F203FEF64D10" box="[189,282,304,330]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="9" pageNumber="10" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 µm</quantity>
|
||
in
|
||
<taxonomicName id="EB9EA6BEFFB5FFFE8037F203FE324D13" box="[322,478,303,330]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Cyclotella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="minuscula">
|
||
<emphasis id="1EEA012FFFB5FFFE8037F203FE324D13" box="[322,478,303,330]" italics="true" pageId="9" pageNumber="10">C. minuscula</emphasis>
|
||
</taxonomicName>
|
||
), a colliculate central area, a small marginal zone, a single valve face central fultoportula, a marginal fultoportula with two satellite pores located on each 4
|
||
<superScript id="DBEB7075FFB5FFFE8523F261FB984D34" attach="none" box="[1110,1140,338,366]" fontSize="6" pageId="9" pageNumber="10">th–</superScript>
|
||
10
|
||
<superScript id="DBEB7075FFB5FFFE85E5F261FB714D3A" attach="left" box="[1168,1181,338,352]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
|
||
costa and a single rimoportula positioned at a rib, beneath the ring of marginal fultoportulae. Difference between these two species can be observed only in a presence/absence of areolae in the central area: in
|
||
<taxonomicName id="EB9EA6BEFFB5FFFE85AEF2AFFA9B4DEF" box="[1243,1399,411,438]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Cyclotella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="minuscula">
|
||
<emphasis id="1EEA012FFFB5FFFE85AEF2AFFA9B4DEF" box="[1243,1399,411,438]" italics="true" pageId="9" pageNumber="10">C. minuscula</emphasis>
|
||
</taxonomicName>
|
||
areolae are absent, while in
|
||
<taxonomicName id="EB9EA6BEFFB5FFFE8371F2F3FD794D83" authority="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov, 2020" authorityName="Zaova & Cvetkoska & Mitic-Kopanja & Jovanovska & Ognjanova-Rumenova & Levkov" authorityYear="2020" box="[516,661,447,473]" class="Bacillariophyceae" family="Stephanodiscaceae" genus="Tertiarius" higherTaxonomySource="GBIF" kingdom="Chromista" order="Thalassiosirales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="minutulus" status="sp. nov.">
|
||
<emphasis id="1EEA012FFFB5FFFE8371F2F3FD794D83" box="[516,661,447,473]" italics="true" pageId="9" pageNumber="10">T. minutulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="05D9BC54FFB5FFFE83E9F2F3FD1B4D80" box="[668,759,448,474]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
they are present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |