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<document id="8D2B2055AA21490DBC245E0DB39D5BEF" ID-CLB-Dataset="153839" ID-DOI="10.11646/zootaxa.5190.2.1" ID-GBIF-Dataset="60faa853-726d-4b71-a3cb-c21a0087515f" ID-ISSN="1175-5326" ID-Zenodo-Dep="7120060" ID-ZooBank="92BFDA7A-28C0-4816-8A91-CFB842C95E96" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664372331544" checkinUser="plazi" docAuthor="Zarei, Fatah, Esmaeili, Hamid Reza, Abbasi, Keyvan, Kovačić, Marcelo, Schliewen, Ulrich K. &amp; Stepien, Carol A." docDate="2022" docId="4154F53A8A0FFF89FF72AA5CFE554BB7" docLanguage="en" docName="zootaxa.5190.2.1.pdf" docOrigin="Zootaxa 5190 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hyrcanogobius bergi Iljin, CE 1928" docType="treatment" docVersion="6" lastPageNumber="170" masterDocId="BD6D8D428A1CFF9AFFE5AC4EFF994C6A" masterDocTitle="Gobies (Teleostei: Gobiidae) of the oldest and deepest Caspian Sea sub-basin: an evidence-based annotated checklist and a key for species identification" masterLastPageNumber="193" masterPageNumber="151" pageNumber="170" updateTime="1699361546833" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="09ADF1EDCF69FB5EF088DE26A488C2A0">Gobies (Teleostei: Gobiidae) of the oldest and deepest Caspian Sea sub-basin: an evidence-based annotated checklist and a key for species identification</mods:title>
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<mods:namePart id="A3ACF89406A42EE3C8E4686E169524D3">Zarei, Fatah</mods:namePart>
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<mods:affiliation id="FB367A119D712CE0393ED69CBD63944C">Ichthyology and Molecular Systematics Research Laboratory, Zoology Section, Department of Biology, College of Sciences, Shiraz University, Shiraz, Iran. &amp; fataahzarei @ gmail. com; https: // orcid. org / 0000 - 0001 - 5552 - 4301</mods:affiliation>
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<mods:namePart id="7F0CB16FE7D8ECEB669A35524789AFFD">Esmaeili, Hamid Reza</mods:namePart>
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<mods:affiliation id="9E9B53C17FD15D559178F65B21A2FC6B">Ichthyology and Molecular Systematics Research Laboratory, Zoology Section, Department of Biology, College of Sciences, Shiraz University, Shiraz, Iran. &amp; hresmaeili @ shirazu. ac. ir; https: // orcid. org / 0000 - 0002 - 9736 - 397 X</mods:affiliation>
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<mods:namePart id="C8945B384A36F76F28CC26ADAC4971D2">Abbasi, Keyvan</mods:namePart>
<mods:affiliation id="A73EA24ED44C57B4C29850B7652780D1">Inland Waters Aquaculture Research Center, Iranian Fisheries Sciences Research Institute, Agricultural Research, Education and Extension Organization, Bandar Anzali, Iran.</mods:affiliation>
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<mods:namePart id="98132E4477AC56049539379F247B20C6">Kovačić, Marcelo</mods:namePart>
<mods:affiliation id="A0B44E923FB5C6CC84D374701CB00667">Natural History Museum Rijeka, Lorenzov prolaz 1, HR- 51000 Rijeka, Croatia.</mods:affiliation>
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<mods:namePart id="D0D140EC301656553A67F917E107BFA5">Schliewen, Ulrich K.</mods:namePart>
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<mods:affiliation id="4D6C8E85A6AD0498F80C91B0D01F52A8">SNSB-Bavarian State Collection of Zoology, Department of Ichthyology, Münchhausenstr. 21, 81247 München, Germany. schliewen @ snsb. de; https: // orcid. org / 0000 - 0003 - 0187 - 4319</mods:affiliation>
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<mods:namePart id="E31B8C51582C4332B98FDD58E21950A2">Stepien, Carol A.</mods:namePart>
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<mods:affiliation id="2C7B871CDC6A545991D3AEF0457DCDE3">National Museum of Natural History, Department of Vertebrate Zoology, Smithsonian Institution, Washington D. C, USA. StepienCA @ si. edu; https: // orcid. org / 0000 - 0002 - 5544 - 4333</mods:affiliation>
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<mods:title id="01CE9608729253C3724A951BE841AD1C">Zootaxa</mods:title>
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<mods:date id="54959FE1BC3438610A53C811F24F7AB6">2022</mods:date>
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<treatment id="4154F53A8A0FFF89FF72AA5CFE554BB7" ID-DOI="http://doi.org/10.5281/zenodo.7126322" ID-GBIF-Taxon="201137467" ID-Zenodo-Dep="7126322" LSID="urn:lsid:plazi:treatment:4154F53A8A0FFF89FF72AA5CFE554BB7" httpUri="http://treatment.plazi.org/id/4154F53A8A0FFF89FF72AA5CFE554BB7" lastPageNumber="170" pageId="19" pageNumber="170">
<subSubSection id="81E717A78A0FFF89FF72AA5CFCA34A47" box="[151,826,1554,1581]" pageId="19" pageNumber="170" type="nomenclature">
<paragraph id="C942442C8A0FFF89FF72AA5CFCA34A47" blockId="19.[151,1437,1554,2013]" box="[151,826,1554,1581]" pageId="19" pageNumber="170">
<emphasis id="FB89983E8A0FFF89FF72AA5CFDAC4A46" bold="true" box="[151,565,1554,1581]" pageId="19" pageNumber="170">
21.
<taxonomicName id="0EFD3FAF8A0FFF89FF24AA5CFDAC4A46" ID-CoL="3P5GM" authority="Iljin, 1928" authorityName="Iljin, CE" authorityYear="1928" box="[193,565,1554,1581]" class="Actinopterygii" family="Gobiidae" genus="Hyrcanogobius" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="170" phylum="Chordata" rank="species" species="bergi">
<emphasis id="FB89983E8A0FFF89FF24AA5CFE2C4A47" bold="true" box="[193,437,1554,1581]" italics="true" pageId="19" pageNumber="170">Hyrcanogobius bergi</emphasis>
<bibRefCitation id="AD6C39DD8A0FFF89FE59AA5CFDAC4A46" author="Iljin, B. S." box="[444,565,1554,1580]" pageId="19" pageNumber="170" pagination="1 - 13" refId="ref27372" refString="Iljin, B. S. (1928) Two new genera and a new species of Gobiidae from the Caspian sea. Reports of the Astrakhan Scientific Fishery Station, 3, 1 - 13." type="journal article" year="1928">Iljin, 1928</bibRefCitation>
</taxonomicName>
</emphasis>
(
<figureCitation id="51C658A98A0FFF89FDA1AA5DFD0E4A47" box="[580,663,1554,1581]" captionStart="FIGURE 15" captionStartId="20.[151,250,694,718]" captionTargetBox="[172,1416,181,671]" captionTargetId="figure-20@20.[172,1416,181,671]" captionTargetPageId="20" captionText="FIGURE 15. Hyrcanogobius bergi. From top: ZM-CBSU S017-1, 29.5 mm SL; ZM-CBSU S017-12, 23.6 mm SL, off Astara, southern Caspian Sea, Gilan Province, Iran." figureDoi="http://doi.org/10.5281/zenodo.7120106" httpUri="https://zenodo.org/record/7120106/files/figure.png" pageId="19" pageNumber="170">Fig. 15</figureCitation>
), Berg’s Goby
</paragraph>
</subSubSection>
<subSubSection id="81E717A78A0FFF89FF72AA79FBD14A1F" pageId="19" pageNumber="170" type="reference_group">
<paragraph id="C942442C8A0FFF89FF72AA79FBD14A1F" blockId="19.[151,1437,1554,2013]" pageId="19" pageNumber="170">
<treatmentCitationGroup id="E9ED63028A0FFF89FF72AA79FBD14A1F" pageId="19" pageNumber="170">
<treatmentCitation id="485C623D8A0FFF89FF72AA79FDAF4A3B" author="Iljin, B. S." box="[151,566,1591,1617]" page="44" pageId="19" pageNumber="170" year="1928">
<taxonomicName id="0EFD3FAF8A0FFF89FF72AA79FDAF4A3B" ID-CoL="3P5GM" authority="Iljin, 1928: 44" authorityName="Iljin, CE" authorityPageNumber="44" authorityYear="1928" box="[151,566,1591,1617]" class="Actinopterygii" family="Gobiidae" genus="Hyrcanogobius" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="170" phylum="Chordata" rank="species" species="bergi">
<emphasis id="FB89983E8A0FFF89FF72AA79FE1F4A3B" box="[151,390,1591,1617]" italics="true" pageId="19" pageNumber="170">Hyrcanogobius bergi</emphasis>
<materialsCitation id="79954E718A0FFF89FE6AAA79FDAF4A3B" ID-GBIF-Occurrence="3914046396" accessionNumber="ZIN 25417" box="[399,566,1591,1617]" collectionCode="ZIN" country="Russia" county="Emba" location="North Caspian Sea" municipality="Ural" pageId="19" pageNumber="170" specimenCount="1" typeStatus="syntype">
<bibRefCitation id="AD6C39DD8A0FFF89FE6AAA79FDAF4A3B" author="Iljin, B. S." box="[399,566,1591,1617]" pageId="19" pageNumber="170" pagination="1 - 13" refId="ref27372" refString="Iljin, B. S. (1928) Two new genera and a new species of Gobiidae from the Caspian sea. Reports of the Astrakhan Scientific Fishery Station, 3, 1 - 13." type="journal article" year="1928">Iljin, 1928: 44</bibRefCitation>
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, figs. 7–11; type locality:
<location id="CC2212F78A0FFF89FC86AA79FBA54A3B" LSID="urn:lsid:plazi:treatment:4154F53A8A0FFF89FF72AA5CFE554BB7:CC2212F78A0FFF89FC86AA79FBA54A3B" box="[867,1084,1591,1617]" country="Russia" county="Emba" municipality="Ural" name="North Caspian Sea" pageId="19" pageNumber="170">North Caspian Sea</location>
, near the mouths of the
<location id="CC2212F78A0FFF89FAB1AA79FA014A3B" LSID="urn:lsid:plazi:treatment:4154F53A8A0FFF89FF72AA5CFE554BB7:CC2212F78A0FFF89FAB1AA79FA014A3B" box="[1364,1432,1591,1617]" country="Russia" county="Emba" municipality="Ural" name="Volga" pageId="19" pageNumber="170">Volga</location>
,
<collectingMunicipality id="2926DE568A0FFF89FF23AA15FF624A1F" box="[198,251,1627,1653]" pageId="19" pageNumber="170">Ural</collectingMunicipality>
, and
<collectingCounty id="20233CA08A0FFF89FED0AA15FEEF4A1F" box="[309,374,1627,1653]" pageId="19" pageNumber="170">Emba</collectingCounty>
rivers,
<collectingCountry id="B1EA04BC8A0FFF89FE2EAA15FD8F4A1F" box="[459,534,1627,1653]" name="Russia" pageId="19" pageNumber="170">Russia</collectingCountry>
and
<collectingCountry id="B1EA04BC8A0FFF89FDA9AA15FD4B4A1F" box="[588,722,1627,1653]" name="Kazakhstan" pageId="19" pageNumber="170">Kazakhstan</collectingCountry>
;
<typeStatus id="1646FA8E8A0FFF89FD3BAA15FCDD4A1F" box="[734,836,1627,1653]" pageId="19" pageNumber="170" type="syntype">syntypes</typeStatus>
: ZIN 25417 (5), Volga.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="81E717A78A0FFF89FF72AA31FCAC4AF3" box="[151,821,1663,1689]" pageId="19" pageNumber="170" type="etymology">
<paragraph id="C942442C8A0FFF89FF72AA31FCAC4AF3" blockId="19.[151,1437,1554,2013]" box="[151,821,1663,1689]" pageId="19" pageNumber="170">
<emphasis id="FB89983E8A0FFF89FF72AA31FEBB4AF3" bold="true" box="[151,290,1663,1689]" pageId="19" pageNumber="170">Etymology:</emphasis>
Named after L. S. Berg, Russian ichthyologist.
</paragraph>
</subSubSection>
<subSubSection id="81E717A78A0FFF89FF72AAECFC9C4BFF" pageId="19" pageNumber="170" type="distribution">
<paragraph id="C942442C8A0FFF89FF72AAECFC9C4BFF" blockId="19.[151,1437,1554,2013]" pageId="19" pageNumber="170">
<emphasis id="FB89983E8A0FFF89FF72AAECFE594AD6" bold="true" box="[151,448,1698,1725]" pageId="19" pageNumber="170">Distribution and habitat:</emphasis>
The coastal waters of all three areas of the Caspian Sea (
<figureCitation id="51C658A98A0FFF89FBCFAAEDFB084AD7" box="[1066,1169,1699,1725]" captionStart="FIGURE 16" captionStartId="20.[151,250,1195,1219]" captionTargetBox="[168,1419,779,1172]" captionTargetId="figure-48@20.[168,1419,779,1172]" captionTargetPageId="20" captionText="FIGURE 16. Distribution of (A) Hyrcanogobius bergi, (B) Knipowitschia caucasica, (C) Knipowitschia iljini, and (D) Knipowitschia longecaudata." figureDoi="http://doi.org/10.5281/zenodo.7120110" httpUri="https://zenodo.org/record/7120110/files/figure.png" pageId="19" pageNumber="170">Fig. 16A</figureCitation>
).
<taxonomicName id="0EFD3FAF8A0FFF89FB46AAEDFA054AD7" authorityName="Kessler" authorityYear="1877" box="[1187,1436,1699,1725]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="170" phylum="Chordata" rank="species" species="longecaudatus">
<emphasis id="FB89983E8A0FFF89FB46AAEDFA054AD7" box="[1187,1436,1699,1725]" italics="true" pageId="19" pageNumber="170">Gobius longecaudatus</emphasis>
</taxonomicName>
var. “c”
<bibRefCitation id="AD6C39DD8A0FFF89FECCAA89FE5F4A8B" author="Kessler, K. T." box="[297,454,1735,1761]" pageId="19" pageNumber="170" refId="ref27724" refString="Kessler, K. T. (1877) The Aralo - Caspian Expedition. IV. Fishes of the Aralo - Caspio - Pontine Ichthyological Region. Naturgeschichte der Fische Islands, St. Petersburg, 360 pp." type="book" year="1877">Kessler, 1877</bibRefCitation>
described from
<collectingRegion id="0B398ACE8A0FFF89FD62AA89FD5B4A8B" box="[647,706,1735,1761]" country="Azerbaijan" name="Baku" pageId="19" pageNumber="170">Baku</collectingRegion>
Bay (
<collectingCountry id="B1EA04BC8A0FFF89FCEEAA89FC134A8B" box="[779,906,1735,1761]" name="Azerbaijan" pageId="19" pageNumber="170">Azerbaijan</collectingCountry>
) is a synonym (
<bibRefCitation id="AD6C39DD8A0FFF89FBADAA89FAF24A8B" author="Pinchuk, V. I. &amp; Miller, P. J." box="[1096,1387,1735,1761]" pageId="19" pageNumber="170" pagination="274 - 278" refId="ref29298" refString="Pinchuk, V. I. &amp; Miller, P. J. (2004 a) Anatirostrum profundorum (Berg, 1927). In: Miller, P. J. (Ed.), The Freshwater Fishes of Europe Vol. 8 / II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 274 - 278." type="book chapter" year="2004">Miller &amp; Pinchuk 2004a</bibRefCitation>
). Its records are from the western coasts of the North and western Middle Caspian Sea (in
<collectingCountry id="B1EA04BC8A0FFF89FB73AAA5FB7B4B6F" box="[1174,1250,1771,1797]" name="Russia" pageId="19" pageNumber="170">Russia</collectingCountry>
) and the eastern South Caspian Sea (
<bibRefCitation id="AD6C39DD8A0FFF89FE43AB41FD254B43" author="Pinchuk, V. I. &amp; Miller, P. J." box="[422,700,1807,1833]" pageId="19" pageNumber="170" pagination="274 - 278" refId="ref29298" refString="Pinchuk, V. I. &amp; Miller, P. J. (2004 a) Anatirostrum profundorum (Berg, 1927). In: Miller, P. J. (Ed.), The Freshwater Fishes of Europe Vol. 8 / II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 274 - 278." type="book chapter" year="2004">Miller &amp; Pinchuk 2004a</bibRefCitation>
), also from the western South Caspian Sea (this study).
<bibRefCitation id="AD6C39DD8A0FFF89FACEAB41FE2E4B27" author="Shakirova, F. M. &amp; Sukhanova, A. I." pageId="19" pageNumber="170" pagination="35 - 45" refId="ref30507" refString="Shakirova, F. M. &amp; Sukhanova, A. I. (1994) Iktiofauna Turkmenistana (sostav i rasprostranenie). Izvestiya Akademii Nauk Turkmenistana, Seriya Biologicheskikh Nauk, 3, 35 - 45. [in Russian]" type="journal article" year="1994">Shakirova &amp; Sukhanova (1994)</bibRefCitation>
also record this species from the Atrek lakes of
<collectingCountry id="B1EA04BC8A0FFF89FC16AB7DFB084B27" box="[1011,1169,1843,1869]" name="Turkmenistan" pageId="19" pageNumber="170">Turkmenistan</collectingCountry>
. Semi-pelagic and also benthic in inshore waters, inhabiting euryhaline to freshwaters (
<bibRefCitation id="AD6C39DD8A0FFF89FC71AB19FB374B1B" author="Pinchuk, V. I. &amp; Miller, P. J." box="[916,1198,1879,1905]" pageId="19" pageNumber="170" pagination="274 - 278" refId="ref29298" refString="Pinchuk, V. I. &amp; Miller, P. J. (2004 a) Anatirostrum profundorum (Berg, 1927). In: Miller, P. J. (Ed.), The Freshwater Fishes of Europe Vol. 8 / II Gobiidae 2. AULA-Verlag GmbH, Verlag fur Wissenschaft und Forschung, Wiebelsheim, pp. 274 - 278." type="book chapter" year="2004">Miller &amp; Pinchuk 2004a</bibRefCitation>
). In
<collectingCountry id="B1EA04BC8A0FFF89FB05AB19FA894B1B" box="[1248,1296,1879,1905]" name="Iran" pageId="19" pageNumber="170">Iran</collectingCountry>
, this species was recorded by us in the coastal waters off Astara.
</paragraph>
</subSubSection>
<subSubSection id="81E717A78A0FFF89FF72ABD0FE554BB7" pageId="19" pageNumber="170" type="materials_examined">
<paragraph id="C942442C8A0FFF89FF72ABD0FA9B4BD3" blockId="19.[151,1437,1554,2013]" box="[151,1282,1950,1977]" pageId="19" pageNumber="170">
<emphasis id="FB89983E8A0FFF89FF72ABD0FE1A4BD2" bold="true" box="[151,387,1950,1976]" pageId="19" pageNumber="170">Material examined:</emphasis>
<materialsCitation id="79954E718A0FFF89FE6FABD1FD134BD3" ID-GBIF-Occurrence="3914046423" box="[394,650,1951,1977]" collectionCode="ZM-CBSU" pageId="19" pageNumber="170" specimenCode="S017-3, 34" specimenCount="1">
<collectionCode id="AFECDCE98A0FFF89FE6FABD1FD9F4BD3" box="[394,518,1951,1977]" pageId="19" pageNumber="170">ZM-CBSU</collectionCode>
<specimenCode id="995BEC578A0FFF89FDE8ABD1FDF84BD3" box="[525,609,1951,1977]" collectionCode="ZM-CBSU" pageId="19" pageNumber="170">S017-3</specimenCode>
,
<specimenCode id="995BEC578A0FFF89FD89ABD1FD134BD3" box="[620,650,1951,1977]" collectionCode="ZM-CBSU" pageId="19" pageNumber="170">34</specimenCode>
</materialsCitation>
, &amp;
<materialsCitation id="79954E718A0FFF89FD56ABD1FB674BD3" ID-GBIF-Occurrence="3914046350" box="[691,1278,1950,1977]" collectionCode="ZM-CBSU" country="Iran" latitude="-85.5" location="Astara" pageId="19" pageNumber="170" specimenCount="1" stateProvince="Gilan">
<collectionCode id="AFECDCE98A0FFF89FD56ABD1FCB64BD3" box="[691,815,1951,1977]" pageId="19" pageNumber="170">ZM-CBSU</collectionCode>
S085, 5, off
<location id="CC2212F78A0FFF89FC25ABD1FB954BD3" LSID="urn:lsid:plazi:treatment:4154F53A8A0FFF89FF72AA5CFE554BB7:CC2212F78A0FFF89FC25ABD1FB954BD3" box="[960,1036,1951,1977]" country="Iran" latitude="-85.5" name="Astara" pageId="19" pageNumber="170" stateProvince="Gilan">Astara</location>
,
<collectingRegion id="0B398ACE8A0FFF89FBF2ABD1FB5B4BD3" box="[1047,1218,1951,1977]" country="Iran" name="Gilan" pageId="19" pageNumber="170">Gilan Province</collectingRegion>
,
<collectingCountry id="B1EA04BC8A0FFF89FB28ABD1FB674BD3" box="[1229,1278,1951,1977]" name="Iran" pageId="19" pageNumber="170">Iran</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="C942442C8A0FFF89FF72AB8CFE554BB7" blockId="19.[151,1437,1554,2013]" box="[151,460,1986,2013]" pageId="19" pageNumber="170">
<emphasis id="FB89983E8A0FFF89FF72AB8CFF714BB6" bold="true" box="[151,232,1986,2012]" pageId="19" pageNumber="170">IUCN:</emphasis>
LC (
<bibRefCitation id="AD6C39DD8A0FFF89FEC4AB8DFE594BB7" author="Freyhof, J." box="[289,448,1987,2013]" pageId="19" pageNumber="170" pagination="135608" refId="ref26535" refString="Freyhof, J. (2020) Knipowitschia bergi. The IUCN Red List of Threatened Species, 2020, e. T 135608 A 137270791. https: // doi. org / 10.2305 / IUCN. UK. 2020 - 3. RLTS. T 135608 A 137270791. en" type="journal article" year="2020">Freyhof 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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