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<document ID-CLB-Dataset="122795" ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-GBIF-Dataset="5f938945-980c-433c-942a-f05d39529a25" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende &amp; Pinto-da-Rocha, Ricardo" docDate="2022" docId="1B054189204659C99211985EBCC44301" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Ayacucho inermis Benedetti &amp; Pinto-da-Rocha 2022, comb. nov." docType="treatment" docVersion="5" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1732651490011" updateUser="ExternalLinkService">
<mods:mods id="78114EC0B67ADECC944B91FA7DB9E246" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D28267B891AAB43480295FA56AD05816">
<mods:title id="68F1019059FD455807E1023157B3515A">Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:role id="7797A4F9FDAFC46F60D3D16760FA7E65">
<mods:roleTerm id="C0A77EF5CF571306677EFCA43C30B880">Author</mods:roleTerm>
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<mods:namePart id="BB25C3018AB1623DC34580714DB982B2">Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation id="E5AA2FAF255DAF8AC5CDCE840CCF1CCA">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
<mods:nameIdentifier id="0057FFFF6902A64BECF08D2C3691B628" type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A0EEBAEBE17C087F4CEE4C33185DE041">Pinto-da-Rocha, Ricardo</mods:namePart>
<mods:nameIdentifier id="AAADCA8D86D000417104A05F9BA53E9E" type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
<mods:affiliation id="CA48CB567DBA8D67D6399B98122EEE76">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:title id="ADA922D60151F2844AE8BE756E1C8A91">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:part id="4E3C988D75487D174D7EFCB5610D68EC">
<mods:date id="C6E58C94C19F9CC2A770BA1EBC09B715">2022</mods:date>
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<mods:number id="BEB287F72D388FE5DC02231409A5ABF7">2022-08-16</mods:number>
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<mods:number id="077DAFD630CB24EF9844DB44975A23F3">80</mods:number>
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<mods:classification id="A3537962A4A0DC4A1B965590A3F4797F">journal article</mods:classification>
<mods:identifier id="0F79AE6C693398AC29E0CE9131CB92C5" type="DOI">http://dx.doi.org/10.3897/asp.80.e73829</mods:identifier>
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<treatment id="1B054189204659C99211985EBCC44301" ID-GBIF-Taxon="199468715" LSID="urn:lsid:plazi:treatment:1B054189204659C99211985EBCC44301" httpUri="http://treatment.plazi.org/id/1B054189204659C99211985EBCC44301" lastPageNumber="309" pageId="0" pageNumber="309" scope_family="Metasarcidae" scope_order="Opiliones" scope_subOrder="Laniatores">
<subSubSection id="2F0C54E853AE128B6662FF1CC2E53E8F" pageId="0" pageNumber="309" type="nomenclature">
<paragraph id="D85FC3D9934CF58EBB0ED9B56FA39850" pageId="0" pageNumber="309">
3.13.
<taxonomicName id="4587656A512E25D7A10436AAA593A15F" ID-CoL="9X669" LSID="1B054189-2046-59C9-9211-985EBCC44301" authority="(Roewer, 1957)" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho inermis" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="inermis" status="comb. nov.">Ayacucho inermis (Roewer, 1957)</taxonomicName>
<taxonomicNameLabel id="934F9F781E413AE1F0868D2F38877578" pageId="0" pageNumber="309">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2F414BB5F4068DED6D859226495E080D" pageId="0" pageNumber="309" type="description">
<paragraph id="81BC1B667F70CF0BECED267EEC9AD085" pageId="0" pageNumber="309">
<figureCitation id="FAAA2BA1CDF7D8C67515938F0FDDB3E0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">Figs 2C</figureCitation>
<figureCitation id="2AB2B5B31CDEBC50FFB32516139486E5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">, 7E, F</figureCitation>
<figureCitation id="86F3B665649609F29938515865CE4D0F" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">, 12C, D</figureCitation>
<figureCitation id="666499F65DC963FD59FBA1F87F4D4D9F" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F4EC8468EA4DAE18FE87C66C9DA56769" pageId="0" pageNumber="309" type="reference_group">
<paragraph id="C6CBBFC9F7E5D1201AAB90B68C6FA38C" pageId="0" pageNumber="309">
<taxonomicName id="882839187834A00581CCF37945F3045D" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Benedetti &amp; Pinto-da-Rocha" baseAuthorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Pinocchio" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinocchio inermis" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="inermis" status="comb. nov.">Pinocchio inermis</taxonomicName>
Roewer, 1957: 70 (desc.), fig. 3 (male coxa-patella IV).
</paragraph>
<paragraph id="41061D9D09DC1CB3FBC3940E08E7EBD4" pageId="0" pageNumber="309">
<taxonomicName id="1456D23F732B172CF9319105A7551E10" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Palcares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Palcares inermis" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="inermis" status="comb. nov.">Palcares inermis</taxonomicName>
:
<bibRefCitation id="476590FA10DE72DF40433FE77A70211B" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
: 145 (cat., syst.).
</paragraph>
</subSubSection>
<subSubSection id="41659D087CDE3354664720F2EDF103F7" pageId="0" pageNumber="309" type="redescription">
<paragraph id="E2289D6385FA7ADAAF96358B91B688EB" pageId="0" pageNumber="309">Redescription.</paragraph>
<paragraph id="A1E08A608052383D377C1666DCCB8D5E" pageId="0" pageNumber="309">
<emphasis id="C9A9CF8194C08668EED1F1B8537EF088" bold="true" pageId="0" pageNumber="309">
MALE:
<emphasis id="8359A3B5DA27002D8D13096EBC2E60FB" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis id="61CC6B179590431A4633B252754119E5" italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) DSW: 2.8; DSL: 3.2; CL: 1.0. FIVL: 2.0. ChL: 1.2.
<emphasis id="59818618CD9EC670A0A96309355BA576" bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
(in ethanol): Uniformly yellowish. In the original description: rusty-yellow body, dorsally darker than ventrally; legs rusty-yellow, slightly blackish.
<emphasis id="46E60F2B5A880809F8FA768FC8A7A427" bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig.
<figureCitation id="9A6AE14D80CA62C7123BD3F247A32A6F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">2C</figureCitation>
) Alpha-type DSS, with long coda. Anterior margin densely covered with granules, with median elevation. Ocularium with a pair of spines; densely granulate. Carapace densely granulate at lateral and posterior regions. Areas I-IV densely covered with granules; I unarmed; II-IV with two pairs of small paramedian tubercles, slightly larger than the granules. Posterior margin of dorsal scutum with a row of acuminate and scattered tiny tubercles. Free tergites I-III with a row of large acuminate tubercles, much larger than tubercles of the areas of dorsal scutum and interspersed with small tubercles. Lateral margins of dorsal scutum densely covered with granules.
<emphasis id="36A164AC7B822019D67DDC526F30C2D0" bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig.
<figureCitation id="C8A50ED278C1EE604A615E0FAD19EE58" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">2C</figureCitation>
) Similar to the female. Segment I densely granulate. Segment II with granules with a much lower density than on segment I; four teeth on finger. Segment III with two teeth.
<taxonomicName id="53CC45ADD1C1E38532FCBBD219709ED7" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis id="4930B23E886BA620E9C639D112D65BD9" bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: Trochanter with a ventroapical tubercle. Femur with a ventrobasal tubercle; a row of three small ventral tubercles, smaller than the ventrobasal tubercle. Tibia: prolateral IIi, retrolateral iIi. Tarsus: prolateral IIi, retrolateral iIii.
<emphasis id="117D331ACD84D68DFFB2185DADB4A049" bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxa I with a middle row of four small tubercles. Coxae II-IV densely covered with granules. The area between coxae II-III and III-IV with a small tubercle at the apex. Smooth genital area. Free sternites with a row of small granules. Anal operculum covered with granules.
<emphasis id="678055DC094790E5BED363EE2A11E224" bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs
<figureCitation id="1A2BCD16162E25BB42F4108569D007DB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">2C</figureCitation>
,
<figureCitation id="5622FF7C788E628D5EFF1CD17905A584" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7E, F</figureCitation>
) Coxae I-II with a prolateral apophyses; III unarmed; IV densely granulate throughout its length, with an apical spiniform apophysis. Trochanters I-IV granulate; IV with a retroapical acuminate tubercle. Femora I-III with granules scattered throughout their extension; IV densely granulate; a retroventral row of 15 large acuminate tubercles throughout its length except at the base and apex; a proventral row of 10-12 large acuminate tubercles (tiny at base) throughout its length, with variation in number of tubercles between right and left legs in the same specimen. Patellae I-IV densely granulate. Tibiae I-IV densely granulate. Tarsal segmentation (
<emphasis id="99B98B414A98C98AE2B72507F8214832" italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) 3, 8, 5-6, 6.
<emphasis id="3DF8C216BF6FDCD08F28CBBD6F2B8F05" bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig.
<figureCitation id="7BCA965F755E2A7C0BED6683685FCEA1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">12C, D</figureCitation>
) VP subrectangular with sides diverging towards the apex; distal margin straight; slightly sinuous on lateral view. MS C1-C3 subapical short and straight; MS A1 short and straight; placed more dorsally than MS C. Lateral sacs long, robust and apically blunt; with short T3-like microsetae. Stylus slightly thick, with long apex and small apical projections. Dorsal process present. -
<emphasis id="60B8A48636DDB32F575A802F4D125CCE" bold="true" pageId="0" pageNumber="309">
FEMALE:
<emphasis id="936093FE733919EA6BD095F59DCC80E9" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis id="3B9835B4370A6657676267A701073E89" italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) DSW: 3.2; DSL: 3.6; CL: 1.2. FIVL: 2.3. ChL: 1.5. Female very similar to male. Chelicerae of similar size. Femur IV with a retroventral and a proventral rows of eight and 11 tubercles respectively, being much smaller than the tubercles in males and tubercles in proventral row larger than that of the retroventral row. Tarsal segmentation: (
<emphasis id="039FA0223CB50882F021136627933535" italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) 5, 8, 6, 6.
</paragraph>
<caption id="3CAE74563BF4272CCA533A7D1A6BEC40" doi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309" start="Figure 7" startId="F7">
<paragraph id="4410216529E2C87B232D998ABA44DEEF" pageId="0" pageNumber="309">
<emphasis id="84E053707ADE648E96BD6458D916D214" bold="true" pageId="0" pageNumber="309">Figure 7.</emphasis>
Male trochanter-tibia IV of
<taxonomicName id="4A39493C323D3EE80799D604A9D362BB" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="9FE64C9F4D0C01A8B473F28C807223DB" italics="true" pageId="0" pageNumber="309">Ayacucho</emphasis>
</taxonomicName>
, dorsal and ventral view, respectively.
<emphasis id="EACD55A429D986DC0A4111758DE075A5" bold="true" pageId="0" pageNumber="309">A</emphasis>
-
<emphasis id="8CC835662BCD2797CBE4D1BE54A9E7C7" bold="true" pageId="0" pageNumber="309">B</emphasis>
<taxonomicName id="1C5F4307D6988F4C4362730A421B16A1" lsidName="A. bambamarca" pageId="0" pageNumber="309" rank="species" species="bambamarca">
<emphasis id="3B35E29D667705A013353A009C5C1C87" italics="true" pageId="0" pageNumber="309">A. bambamarca</emphasis>
</taxonomicName>
(Roewer, 1957)
<emphasis id="38201BDF1737DD60D907090F13CA54DE" bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
;
<emphasis id="F86A86E9A4093FB1EFFB1607916DE077" bold="true" pageId="0" pageNumber="309">C</emphasis>
-
<emphasis id="E692732BB16660437E0BE826CDAB43EA" bold="true" pageId="0" pageNumber="309">D</emphasis>
<taxonomicName id="27CF24B55E70095FCCB9269CAB21057C" lsidName="A. glauberrochai" pageId="0" pageNumber="309" rank="species" species="glauberrochai">
<emphasis id="959C659E436E9A7663DBB6928F49ACC7" italics="true" pageId="0" pageNumber="309">A. glauberrochai</emphasis>
</taxonomicName>
<emphasis id="7D87B8A9D11619941804754B51212E0B" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
;
<emphasis id="786AF8B88B81192943D4B7A2DB451F54" bold="true" pageId="0" pageNumber="309">E</emphasis>
-
<emphasis id="71E8EDD9AEE2E128DCD14B5E841C4C84" bold="true" pageId="0" pageNumber="309">F</emphasis>
<taxonomicName id="23B414DC858C121F838792208D442D7E" lsidName="A. inermis" pageId="0" pageNumber="309" rank="species" species="inermis">
<emphasis id="625EC38659B181D99BBDC432DEDBE146" italics="true" pageId="0" pageNumber="309">A. inermis</emphasis>
</taxonomicName>
(Roewer, 1957)
<emphasis id="B95209ABA69B095BDFFD1F6C40B5D6AC" bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
;
<emphasis id="990F0D2C3CAA5D6A5DE0BC0C95B0D7A8" bold="true" pageId="0" pageNumber="309">G</emphasis>
-
<emphasis id="763702B873F20A560623A0F896A669A0" bold="true" pageId="0" pageNumber="309">H</emphasis>
<taxonomicName id="399DBD2AFD2A6D7CEA19B289DA4CE71F" lsidName="A. pasolinii" pageId="0" pageNumber="309" rank="species" species="pasolinii">
<emphasis id="A4482CC7B27C773F25A6AF284C725B73" italics="true" pageId="0" pageNumber="309">A. pasolinii</emphasis>
</taxonomicName>
<emphasis id="D611639E6C4AB71F0D27DCCD87883108" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
;
<emphasis id="B9D1E4FFF94B02C01D015EC0E381DF11" bold="true" pageId="0" pageNumber="309">I</emphasis>
-
<emphasis id="105DA98A70504846522B68689551BEBC" bold="true" pageId="0" pageNumber="309">J</emphasis>
<taxonomicName id="13FF0B8467CC62D0B689A091A7ACC5F8" lsidName="A. pomacocha" pageId="0" pageNumber="309" rank="species" species="pomacocha">
<emphasis id="DD0A25B9E9C342BE3B4C0B7C2BAB06E9" italics="true" pageId="0" pageNumber="309">A. pomacocha</emphasis>
</taxonomicName>
<emphasis id="CCE412BAE2F0209AB3E433C0B83D6C29" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
;
<emphasis id="DF7C7E84DDCC0B3894B6024367523A6A" bold="true" pageId="0" pageNumber="309">K</emphasis>
-
<emphasis id="75609F01CCE9251E1401FA604A166AD5" bold="true" pageId="0" pageNumber="309">L</emphasis>
<taxonomicName id="FC02D8BB6FD4FA39B31E6B0318A5A63E" lsidName="A. querococha" pageId="0" pageNumber="309" rank="species" species="querococha">
<emphasis id="C841830E22D8D1A487EEF2BF3853DD09" italics="true" pageId="0" pageNumber="309">A. querococha</emphasis>
</taxonomicName>
<emphasis id="19D2CB12D5F5A88C08F41190D5D88244" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
;
<emphasis id="9CA342B8AE86BE4901F7A49F210884D3" bold="true" pageId="0" pageNumber="309">M</emphasis>
-
<emphasis id="A53847856055967C5EB1C6A505B5F90E" bold="true" pageId="0" pageNumber="309">N</emphasis>
<taxonomicName id="BE8620A805CEEE3CB980F6D48A359AE8" lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
<emphasis id="7F7FBD0B3B01FC581DE553CC31CB541D" italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
</taxonomicName>
<emphasis id="818CD3ECDA2A2738A396F34F810EE4C1" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
;
<emphasis id="4EB4C8FEDC6F3150D6C20B83EF4EF264" bold="true" pageId="0" pageNumber="309">O</emphasis>
-
<emphasis id="E2A64892C0B688619F1F96978D1BBD7B" bold="true" pageId="0" pageNumber="309">P</emphasis>
<taxonomicName id="1EFA1D8F15EF91C7A2BAB6E002BC2F67" lsidName="A. spielbergi" pageId="0" pageNumber="309" rank="species" species="spielbergi">
<emphasis id="A92BC580F6AF3DD6E86C649F3EE5F532" italics="true" pageId="0" pageNumber="309">A. spielbergi</emphasis>
</taxonomicName>
<emphasis id="6D6E813744FF1D0B15B768785E45FFEB" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
;
<emphasis id="79506A6D4B987DEEE96130BDA1EE7085" bold="true" pageId="0" pageNumber="309">Q</emphasis>
-
<emphasis id="16956F5C884297F8806E76E40B14AF97" bold="true" pageId="0" pageNumber="309">R</emphasis>
<taxonomicName id="3FE7C0E5A78FECDF7B36792D7E745621" lsidName="A. spiniger" pageId="0" pageNumber="309" rank="species" species="spiniger">
<emphasis id="0D5AED06616D9B56D0076A640CBCF208" italics="true" pageId="0" pageNumber="309">A. spiniger</emphasis>
</taxonomicName>
(Roewer, 1957)
<emphasis id="7BDD0E7A0EFBA4F183AE1A90D1ABBED9" bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
;
<emphasis id="4AAC85C5A95E27A1E9930410C1206C12" bold="true" pageId="0" pageNumber="309">S</emphasis>
-
<emphasis id="6E7B8B8C4CF21B5B0DF1BF374929E271" bold="true" pageId="0" pageNumber="309">T</emphasis>
:
<taxonomicName id="622FA3DA149E8C81972FEC0EE128FB26" lsidName="A. tapacocha" pageId="0" pageNumber="309" rank="species" species="tapacocha">
<emphasis id="FCE53C002BC52CE6AD49284B2216AADD" italics="true" pageId="0" pageNumber="309">A. tapacocha</emphasis>
</taxonomicName>
<emphasis id="9453E6DA87E516266A3EC15C56006E23" bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
;
<emphasis id="ED1F15797B338C192B151BAAFD4683AF" bold="true" pageId="0" pageNumber="309">U</emphasis>
-
<emphasis id="06A26D404F28F5935ACD7AC3E9C300BC" bold="true" pageId="0" pageNumber="309">V</emphasis>
<taxonomicName id="AD6EA243DA6CF4934951D9911CCE88B6" lsidName="A. titschacki" pageId="0" pageNumber="309" rank="species" species="titschacki">
<emphasis id="605FF4517D9B220CE2EB59D90045E1F0" italics="true" pageId="0" pageNumber="309">A. titschacki</emphasis>
</taxonomicName>
Roewer, 1949;
<emphasis id="D7ADD6180E0ACF9A1ED49F56720F4A67" bold="true" pageId="0" pageNumber="309">W</emphasis>
-
<emphasis id="3B3471508260CC139A3C68B0A1B5E30D" bold="true" pageId="0" pageNumber="309">X</emphasis>
:
<taxonomicName id="05C1B8D876D6BFB80D67A9BA61A7E0F1" lsidName="A. triarmatus" pageId="0" pageNumber="309" rank="species" species="triarmatus">
<emphasis id="4A66359E7E50506F24C839086D123118" italics="true" pageId="0" pageNumber="309">A. triarmatus</emphasis>
</taxonomicName>
<emphasis id="267AFC692DC0C4C7B2EC271939A1FF25" bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
; Legend bars = 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="269286A35D6B6928CB2E4E6EA7E5ADBB" pageId="0" pageNumber="309" type="diagnosis">
<paragraph id="B4C5C59E9C3B64E34C9A79A064693204" pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph id="E5C2A1A225B927F5ED030A3871437D60" pageId="0" pageNumber="309">
Resembles
<taxonomicName id="AE81172D932AC47436C11A52F605DBA6" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho spiniger" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="DCDA181C7344AF1AA3959A49352B12CA" italics="true" pageId="0" pageNumber="309">Ayacucho spiniger</emphasis>
</taxonomicName>
<emphasis id="B113111886E5813DB2ECC946A83C2D7F" bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
because two ventral rows of tubercles in femur IV (Fig.
<figureCitation id="3505F213E994B4D894E3408F83D7E8A8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7F</figureCitation>
). Differs from
<taxonomicName id="43DCEFBE1A1F0721A8F9D155DD818A6F" lsidName="A. spiniger" pageId="0" pageNumber="309" rank="species" species="spiniger">
<emphasis id="ACC4E7055F379D82DE821F0BDE0145D8" italics="true" pageId="0" pageNumber="309">A. spiniger</emphasis>
</taxonomicName>
<emphasis id="6B6DFACF04F79D7B27F39AC742FE0DBC" bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
because femur IV with spiniform tubercles (Fig.
<figureCitation id="13C339DB32848FAF76BBB9789BB01045" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7F</figureCitation>
); ventral plate of penis without lateral projections; VP with short macrosetae; lateral sacs with short microsetae; dorsal process shorter than the stylus (Fig.
<figureCitation id="E2922295C368DF21CE81B76C533977DA" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">12C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BEBB23514C554AE28FCF2FEA82586F6F" pageId="0" pageNumber="309" type="distribution">
<paragraph id="4EA14E8F4C90C4B9AFB82CDBB9DD9DA6" pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph id="7D1E3CF35F3B41E4C692A0868B37DFE8" pageId="0" pageNumber="309">
(Fig.
<figureCitation id="ED1AD1B92BA6A712C6FE0B0D66B5B652" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">28</figureCitation>
) PERU. La Libertad. Huamachuco.
</paragraph>
</subSubSection>
<subSubSection id="A4702743DFD50C6B557C185CE344498C" pageId="0" pageNumber="309" type="materials_examined">
<paragraph id="EFFA49927B9E90DDC610278F72BE51B4" pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph id="23D08CB6D611F5F43AA118E61D010BF6" pageId="0" pageNumber="309">
<emphasis id="642E3D8F120B47F63A1773D01B69DA83" bold="true" pageId="0" pageNumber="309">
<emphasis id="147A236C2A25E8195D72F2B6157520F2" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
:
<materialsCitation id="5BCD46B552C54B86AE62374DCE7C387C" ID-GBIF-Occurrence="3889641324" accessionNumber="RII 11393" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Weyrauch" country="PERU" elevation="3200" location="Huamachuco" specimenCount="♂" stateProvince="La Libertad" typeStatus="Holotype">
<typeStatus id="81B5781A2AF0988D21590BA9FC85A788">Holotype</typeStatus>
<specimenCount id="56E9F7461CCA0E85B1E563113A387DE9">♂</specimenCount>
, '
<collectingCountry id="F66DBCC749552083887B6ADBE9F60590" name="Peru">PERU</collectingCountry>
,
<collectingRegion id="AA89A0DC657EEA62E8504F1C58B27DC6" country="Peru" name="La Libertad">La Libertad</collectingRegion>
,
<location id="8D2DD3EC2842A29386E50D5967594BCA" LSID="urn:lsid:plazi:treatment:1B054189204659C99211985EBCC44301:8D2DD3EC2842A29386E50D5967594BCA" country="PERU" name="Huamachuco" stateProvince="La Libertad">Huamachuco</location>
,
<elevation id="BB848D6AB370A4520EC98CB7DF45E58C" metricMagnitude="3" metricUnit="m" metricValue="3.2" unit="m" value="3200.0">
<quantity id="3AF4F7512EC0800731CFA52A1E07CD9C" metricMagnitude="3" metricUnit="m" metricValue="3.2" unit="m" value="3200.0">3,200m</quantity>
a.s.l.
</elevation>
without date,
<collectorName id="DF8A3D46237765D4B9C6B373EC309B3C">Weyrauch</collectorName>
leg. (SMF
<accessionNumber id="54AA3D81A888285B2C13FC85267C9055" httpUri="http://www.ncbi.nlm.nih.gov/protein/RII11393">RII 11393</accessionNumber>
/23)
</materialsCitation>
</emphasis>
-
<materialsCitation id="B03ED8DC391EDED1E432940CE764FCF0" ID-GBIF-Occurrence="3889641309" accessionNumber="RII 11393" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Peru" specimenCount="♀" stateProvince="La Libertad" typeStatus="Paratype">
<emphasis id="E363C958576A69005D4DC6734E8F6D45" bold="true" pageId="0" pageNumber="309">
<typeStatus id="672F6FAD92D3A9EE869188CBB6BFED0D">Paratype</typeStatus>
</emphasis>
<specimenCount id="1C538BD7559EC0FD840E8FF067A923B2">♀</specimenCount>
,
<normalizedToken id="A56A1E0EA018CD5F722E3B7574310683" originalValue="‘ditto’">'ditto'</normalizedToken>
(SMF
<accessionNumber id="F8583132A526BF268CB094581EE46940" httpUri="http://www.ncbi.nlm.nih.gov/protein/RII11393">RII 11393</accessionNumber>
/23)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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