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<document id="08C43A81F453CE576A934EC0A202A139" ID-DOI="10.11646/zootaxa.3716.4.2" ID-GBIF-Dataset="a9b85d9e-a8fb-4ceb-9f20-65d2cd3c84df" ID-ISSN="1175-5326" ID-Zenodo-Dep="220316" ID-ZooBank="F1AD3425-314F-4B5F-AE7A-2D683135DBE6" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460682586901" checkinUser="plazi" docAuthor="Pyrcz, Tomasz W., Prieto, Carlos, Viloria, Angel L. &amp; Andrade, Gonzalo" docDate="2013" docId="03B29E2BFFE71877FF0FCFDFFCA9F870" docLanguage="en" docName="zt03716p538.pdf" docOrigin="Zootaxa 3716 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pedaliodes rodriguezi Pyrcz &amp; Andrade, n. sp." docType="treatment" docVersion="8" lastPageNumber="532" masterDocId="FF8BE653FFE41873FF98C97BFFC1FF9A" masterDocTitle="New species of high elevation cloud forest butterflies of the genus Pedaliodes Butler from the northern Colombian Andes (Lepidoptera, Nymphalidae, Satyrinae)" masterLastPageNumber="538" masterPageNumber="528" pageNumber="531" updateTime="1698350629498" updateUser="plazi">
<mods:mods id="A770EB3FE8E8268EA079ADA2F71C6626" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4329D7C889CD94237AD661DE70F3AAD7">
<mods:title id="125177FCF8FF7273EB2860D686CAD027">New species of high elevation cloud forest butterflies of the genus Pedaliodes Butler from the northern Colombian Andes (Lepidoptera, Nymphalidae, Satyrinae)</mods:title>
</mods:titleInfo>
<mods:name id="60CC9ECAC8981E4248A2A0AA4CF3DD80" type="personal">
<mods:role id="B42BEA21B66E6D970B823E7E97D113E1">
<mods:roleTerm id="DB47F2D3CF5E5BA89525274BD7B07FC1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C4BFB22C69D9C97FCF05131D3C98A185">Pyrcz, Tomasz W.</mods:namePart>
</mods:name>
<mods:name id="1F30C93F0085F419C610FCE29EADC86E" type="personal">
<mods:role id="9532142A771DC91DEAFAA323FCBE884C">
<mods:roleTerm id="EE5839DCBCDD93754B7108612011C646">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7DCBB70B110E9160EB6D5BD219963583">Prieto, Carlos</mods:namePart>
</mods:name>
<mods:name id="0E3F397867A8E5CA60F42DCADA07C09F" type="personal">
<mods:role id="EA57FEA58CFDFF06F807550E1BAA4914">
<mods:roleTerm id="B7B74F94B0ECFD2BEF9F74CE31AE5A18">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9A512151F31BDAB9D3E5A512165EE89A">Viloria, Angel L.</mods:namePart>
</mods:name>
<mods:name id="7CC2583363201A4B950759D33EA5FE4B" type="personal">
<mods:role id="69B12A37A8B0846ABD6547084E9D39F6">
<mods:roleTerm id="999CA2D5312B10DB14ECAA5FC8236383">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3777593251226EEF12DD157E31A5EE6A">Andrade, Gonzalo</mods:namePart>
</mods:name>
<mods:typeOfResource id="E604FF9341B3C0ADBA558DE200AC297B">text</mods:typeOfResource>
<mods:relatedItem id="BEE5916542D004EBE15399CEC8D90734" type="host">
<mods:titleInfo id="DBAB8B5E74CA5C9C6D1A9BE7CD4EDE19">
<mods:title id="7873C7662CC7CFF5C21DB15817B83CB3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="66E14C64A0E6783180E220ABA581F138">
<mods:date id="D770A018CA44FB05AC7D49B010D6B107">2013</mods:date>
<mods:detail id="7677EF97442C151A8299F11758896068" type="volume">
<mods:number id="795CB124C10D8923A02DB4A50EF8D742">3716</mods:number>
</mods:detail>
<mods:detail id="7FCB385D671A295589546F1A458123C1" type="issue">
<mods:number id="1BF31418079FD95E4DE25533FF3AB810">4</mods:number>
</mods:detail>
<mods:extent id="6C13F358AD78572BCFB09F6BD2E4A4AB" unit="page">
<mods:start id="6EEE3D29702E510522BFA654A950FCE8">528</mods:start>
<mods:end id="A3D67D63D2A6CFE99A9F71B132BCC7B9">538</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="167946963577451E5BC2DCB6C2D46AC8">journal article</mods:classification>
<mods:identifier id="2D992FA30E399A5FB2AB9C06C05C527A" type="DOI">10.11646/zootaxa.3716.4.2</mods:identifier>
<mods:identifier id="735821B8A6707C2F061CA459BAB1B568" type="GBIF-Dataset">a9b85d9e-a8fb-4ceb-9f20-65d2cd3c84df</mods:identifier>
<mods:identifier id="14C341C81DEFB622F4AFAAEB84AEB65C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="05C65EAA4DBFE13A1187BB0E6997D3F9" type="Zenodo-Dep">220316</mods:identifier>
<mods:identifier id="8DB3F36BF1F5D493BF012694AE198981" type="ZooBank">F1AD3425-314F-4B5F-AE7A-2D683135DBE6</mods:identifier>
</mods:mods>
<treatment id="03B29E2BFFE71877FF0FCFDFFCA9F870" ID-DOI="http://doi.org/10.5281/zenodo.6151279" ID-GBIF-Taxon="119580930" ID-Zenodo-Dep="6151279" LSID="urn:lsid:plazi:treatment:03B29E2BFFE71877FF0FCFDFFCA9F870" httpUri="http://treatment.plazi.org/id/03B29E2BFFE71877FF0FCFDFFCA9F870" lastPageId="4" lastPageNumber="532" pageId="3" pageNumber="531">
<subSubSection id="C3017CB6FFE71870FF0FCFDFFEA7F945" pageId="3" pageNumber="531" type="nomenclature">
<paragraph id="8BA42F3DFFE71870FF0FCFDFFD10F924" blockId="3.[151,721,1700,1759]" box="[151,721,1700,1726]" pageId="3" pageNumber="531">
<heading id="D0EC9851FFE71870FF0FCFDFFD10F924" bold="true" box="[151,721,1700,1726]" fontSize="11" level="1" pageId="3" pageNumber="531" reason="1">
<emphasis id="B96FF32FFFE71870FF0FCFDFFD10F924" bold="true" box="[151,721,1700,1726]" pageId="3" pageNumber="531">
<taxonomicName id="4C1B54BEFFE71870FF0FCFDFFD40F924" authority="Pyrcz &amp; Andrade" authorityName="Pyrcz &amp; Andrade" box="[151,641,1700,1726]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="531" phylum="Arthropoda" rank="species" species="rodriguezi" status="sp. nov.">
<emphasis id="B96FF32FFFE71870FF0FCFDFFE5DF924" bold="true" box="[151,412,1700,1726]" italics="true" pageId="3" pageNumber="531">Pedaliodes rodriguezi</emphasis>
Pyrcz &amp; Andrade
</taxonomicName>
,
<taxonomicNameLabel id="A25C4E54FFE71870FD16CFDFFD10F924" box="[654,721,1700,1726]" pageId="3" pageNumber="531" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA42F3DFFE71870FF0FCFBDFEA7F945" blockId="3.[151,721,1700,1759]" box="[151,358,1734,1759]" pageId="3" pageNumber="531">
(
<figureCitation id="132033B8FFE71870FF07CFBDFECBF945" box="[159,266,1734,1759]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1932,1954]" captionTargetBox="[262,1342,215,1896]" captionTargetId="figure@5.[221,1381,195,1934]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Pedaliodes adults. 1, P. haydoni male (holotype); 2, P. porcia male; 3, P. adrianae male (holotype); 4, P. adrianae female (paratype); 5, P. rodriguezi male (holotype); 6, P. ro dr i gu e z i female (paratype); 7, P. philinae male (holotype); 8, P. philinae female (paratype)." httpUri="https://zenodo.org/record/220317/files/figure.png" pageId="3" pageNumber="531">Figs. 5, 6</figureCitation>
,
<figureCitation id="132033B8FFE71870FE8ECFBCFEF5F945" box="[278,308,1735,1759]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1891,1913]" captionTargetBox="[247,1333,233,1790]" captionTargetId="figure@6.[243,1347,193,1849]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Male genitalia. Pedaliodes haydoni (paratype): 9 A, genitalia in lateral view; 9 B, aedeagus in lateral view; 9 C, aedeagus in dorsal view; Pedaliodes adrianae (paratype): 10 A, genitalia in lateral view; 10 B, aedeagus in lateral view; 10 C, aedeagus in dorsal view; Pedaliodes philinae (paratype): 11 A, genitalia in lateral view; 11 B, aedeagus in lateral view; 11 C, aedeagus in dorsal view; Pedaliodes rodriguezi (paratype): 12 A, genitalia in lateral view; 12 B, aedeagus in lateral view; 12 C, aedeagus in dorsal view." httpUri="https://zenodo.org/record/220318/files/figure.png" pageId="3" pageNumber="531">12</figureCitation>
,
<figureCitation id="132033B8FFE71870FED8CFBDFE9CF945" box="[320,349,1734,1759]" captionStart="FIGURES 13 – 15. 13" captionStartId="7.[151,264,1953,1975]" captionTargetBox="[378,1185,207,1911]" captionTargetId="figure@7.[214,1373,193,1932]" captionTargetPageId="7" captionText="FIGURES 13 – 15. 13, Pedaliodes adrianae (paratype) female genitalia; 14, Pedaliodes philinae (paratype) female genitalia; 15, Pedaliodes rodriguezi male (Cerro San Felix, Antioquia, Colombia, 2900 m)" httpUri="https://zenodo.org/record/220319/files/figure.png" pageId="3" pageNumber="531">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3017CB6FFE71877FF0FCE75FC42FE44" lastPageId="4" lastPageNumber="532" pageId="3" pageNumber="531" type="materials_examined">
<paragraph id="8BA42F3DFFE71870FF0FCE75FA5DF844" blockId="3.[151,1437,1806,2014]" pageId="3" pageNumber="531">
<emphasis id="B96FF32FFFE71870FF0FCE75FE40F8BD" bold="true" box="[151,385,1806,1831]" pageId="3" pageNumber="531">Material examined.</emphasis>
<typeStatus id="54A0919FFFE71870FE11CE74FDDCF8BD" box="[393,541,1807,1831]" pageId="3" pageNumber="531" type="holotype">HOLOTYPE</typeStatus>
(male):
<collectingCountry id="F30C6FADFFE71870FDE6CE75FD2EF8BD" box="[638,751,1806,1831]" name="Colombia" pageId="3" pageNumber="531">Colombia</collectingCountry>
, Departamento Antioquia, El Retiro, Reserva La Castellana,
<geoCoordinate id="EE2F49FAFFE71870FF0FCE4EFED4F8D4" box="[151,277,1842,1870]" direction="north" orientation="latitude" pageId="3" pageNumber="531" precision="15" value="6.108333">6o06’30’’N</geoCoordinate>
,
<geoCoordinate id="EE2F49FAFFE71870FEB0CE4EFE76F8D4" box="[296,439,1842,1870]" direction="west" orientation="longitude" pageId="3" pageNumber="531" precision="15" value="-75.549164">75o32’57’’W</geoCoordinate>
,
<quantity id="4CE382D8FFE71870FE52CE4EFDAFF8D7" box="[458,622,1845,1870]" metricMagnitude="3" metricUnit="m" metricValue="2.825" metricValueMax="2.85" metricValueMin="2.8" pageId="3" pageNumber="531" unit="m" value="2825.0" valueMax="2850.0" valueMin="2800.0">2800–2850 m</quantity>
,
<date id="FFA509FDFFE71870FD19CE4DFCD7F8D4" box="[641,790,1846,1870]" pageId="3" pageNumber="531" value="2007-08-08">08.VIII.2007</date>
, T. Pyrcz leg., MZUJ, to be deposited in MHN-UN;
<typeStatus id="54A0919FFFE71870FF0FCE21FEF3F8E8" box="[151,306,1882,1906]" pageId="3" pageNumber="531" type="paratype">PARATYPES</typeStatus>
(
<specimenCount id="9D1DE4B4FFE71870FED9CE21FE67F8E8" box="[321,422,1881,1906]" pageId="3" pageNumber="531" type="male">18 males</specimenCount>
and
<specimenCount id="9D1DE4B4FFE71870FE45CE21FD85F8E8" box="[477,580,1881,1906]" pageId="3" pageNumber="531" type="female">1 female</specimenCount>
):
<specimenCount id="9D1DE4B4FFE71870FDCFCE21FD73F8E8" box="[599,690,1881,1906]" pageId="3" pageNumber="531" type="male">2 males</specimenCount>
: same data as the
<typeStatus id="54A0919FFFE71870FCE3CE22FC21F8E8" box="[891,992,1881,1906]" pageId="3" pageNumber="531" type="holotype">holotype</typeStatus>
, MZUJ;
<specimenCount id="9D1DE4B4FFE71870FBDBCE21FB5FF8E8" box="[1091,1182,1881,1906]" pageId="3" pageNumber="531" type="male">2 males</specimenCount>
: El Retiro, Reserva La Castellana,
<quantity id="4CE382D8FFE71870FE81CE06FE77F80F" box="[281,438,1917,1942]" metricMagnitude="3" metricUnit="m" metricValue="2.725" metricValueMax="2.75" metricValueMin="2.7" pageId="3" pageNumber="531" unit="m" value="2725.0" valueMax="2750.0" valueMin="2700.0">2700–2750 m</quantity>
,
<date id="FFA509FDFFE71870FE5ACE05FD85F80C" box="[450,580,1918,1942]" pageId="3" pageNumber="531" value="2003-09-12">12.IX.2003</date>
, T. Pyrcz leg., MZUJ;
<specimenCount id="9D1DE4B4FFE71870FCDACE05FC53F80C" box="[834,914,1917,1942]" pageId="3" pageNumber="531" type="male">1 male</specimenCount>
: Antioquia, Envigado,
<quantity id="4CE382D8FFE71870FB0CCE06FB2AF80F" box="[1172,1259,1917,1942]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="531" unit="m" value="2600.0">2600 m</quantity>
,
<date id="FFA509FDFFE71870FB6ECE05FAB8F80C" box="[1270,1401,1918,1942]" pageId="3" pageNumber="531" value="2002-11-09">09.XI.2002</date>
, G. Rodríguez leg., MZUJ;
<specimenCount id="9D1DE4B4FFE71870FE3ACED9FE3DF820" box="[418,508,1953,1978]" pageId="3" pageNumber="531" type="male">3 males</specimenCount>
: same data but
<quantity id="4CE382D8FFE71870FD31CEDAFC87F823" box="[681,838,1953,1978]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="3" pageNumber="531" unit="m" value="2700.0" valueMax="2800.0" valueMin="2600.0">2600–2800 m</quantity>
,
<date id="FFA509FDFFE71870FCCACED9FC1AF820" box="[850,987,1954,1978]" pageId="3" pageNumber="531" value="2002-12-20">20.XII.2002</date>
(prep. genit. 08/
<date id="FFA509FDFFE71870FB0DCED9FAD6F820" box="[1173,1303,1954,1978]" pageId="3" pageNumber="531" value="2010-10-22">22.10.2010</date>
, A. Zubek), MZUJ;
<specimenCount id="9D1DE4B4FFE71870FF77CEBDFEFEF844" box="[239,319,1989,2014]" pageId="3" pageNumber="531" type="male">1 male</specimenCount>
: same data but
<quantity id="4CE382D8FFE71870FE72CEBEFD81F847" box="[490,576,1989,2014]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="531" unit="m" value="2600.0">2600 m</quantity>
,
<date id="FFA509FDFFE71870FDD4CEBDFD0FF844" box="[588,718,1990,2014]" pageId="3" pageNumber="531" value="2002-11-09">09.XI.2002</date>
, MZUJ;
<specimenCount id="9D1DE4B4FFE71870FCA9CEBDFC40F844" box="[817,897,1989,2014]" pageId="3" pageNumber="531" type="male">1 male</specimenCount>
: same data but
<quantity id="4CE382D8FFE71870FBB4CEBEFB42F847" box="[1068,1155,1989,2014]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="531" unit="m" value="2600.0">2600 m</quantity>
,
<date id="FFA509FDFFE71870FB17CEBDFACCF844" box="[1167,1293,1990,2014]" pageId="3" pageNumber="531" value="2002-03-31">31.III.2002</date>
(prep. genit.
</paragraph>
<paragraph id="8BA42F3DFFE01877FF0FC9E0FC42FE44" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="532">
09/
<date id="FFA509FDFFE01877FF23C9E0FEFCFF29" box="[187,317,155,179]" pageId="4" pageNumber="532" value="2010-09-11">09.11.2010</date>
), MZUJ;
<specimenCount id="9D1DE4B4FFE01877FE2EC9E0FDCCFF29" box="[438,525,154,179]" pageId="4" pageNumber="532" type="male">1 male</specimenCount>
: same data but
<date id="FFA509FDFFE01877FD4EC9E0FCF0FF29" box="[726,817,155,179]" pageId="4" pageNumber="532" value="2002-04">IV.2002</date>
, MZUJ;
<specimenCount id="9D1DE4B4FFE01877FC3BC9E0FC3AFF29" box="[931,1019,154,179]" pageId="4" pageNumber="532" type="male">1 male</specimenCount>
: Antioquia, San Felix,
<geoCoordinate id="EE2F49FAFFE01877FA81C9E1FA56FF29" box="[1305,1431,151,179]" direction="north" orientation="latitude" pageId="4" pageNumber="532" precision="15" value="6.338889">6o20’20’’N</geoCoordinate>
,
<geoCoordinate id="EE2F49FAFFE01877FF0FC9BAFEE7FF40" box="[151,294,190,218]" direction="west" orientation="longitude" pageId="4" pageNumber="532" precision="15" value="-75.64083">75o38’27’’W</geoCoordinate>
,
<quantity id="4CE382D8FFE01877FEAEC9B9FE50FF43" box="[310,401,194,218]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="532" unit="m" value="3000.0">3000 m</quantity>
,
<date id="FFA509FDFFE01877FE39C9B9FDE2FF40" box="[417,547,194,218]" pageId="4" pageNumber="532" value="2003-09-13">13.IX.2003</date>
, T. Pyrcz leg., MZUJ;
<specimenCount id="9D1DE4B4FFE01877FCAEC9B9FC4BFF40" box="[822,906,193,218]" pageId="4" pageNumber="532" type="male">1 male</specimenCount>
: same data but
<quantity id="4CE382D8FFE01877FBDDC9BAFB27FF43" box="[1093,1254,193,218]" metricMagnitude="3" metricUnit="m" metricValue="2.625" metricValueMax="2.65" metricValueMin="2.6" pageId="4" pageNumber="532" unit="m" value="2625.0" valueMax="2650.0" valueMin="2600.0">2600–2650 m</quantity>
,
<date id="FFA509FDFFE01877FB6EC9BAFAB4FF40" box="[1270,1397,193,218]" pageId="4" pageNumber="532" value="2006-04-09">09.IV.2006</date>
, G. Rodriguez leg., MZUJ;
<specimenCount id="9D1DE4B4FFE01877FE3CC992FE35FE9B" box="[420,500,232,257]" pageId="4" pageNumber="532" type="male">1 male</specimenCount>
: Antioquia, Carretera Santa Rosa – Yarumal,
<geoCoordinate id="EE2F49FAFFE01877FC6FC993FBB4FE9B" box="[1015,1141,229,257]" direction="north" orientation="latitude" pageId="4" pageNumber="532" precision="15" value="6.691111">6o41’28’’N</geoCoordinate>
,
<geoCoordinate id="EE2F49FAFFE01877FB1AC993FAD3FE9B" box="[1154,1298,229,257]" direction="west" orientation="longitude" pageId="4" pageNumber="532" precision="15" value="-75.467224">75o28’02’’W</geoCoordinate>
,
<quantity id="4CE382D8FFE01877FA86C993FF6EFEBE" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="4" pageNumber="532" unit="m" value="2750.0" valueMax="2900.0" valueMin="2600.0">2600–2900 m</quantity>
,
<date id="FFA509FDFFE01877FF24C876FEE8FEBF" box="[188,297,269,293]" pageId="4" pageNumber="532" value="2003-01-04">04.I.2003</date>
, G. Rodríguez leg., MZUJ;
<specimenCount id="9D1DE4B4FFE01877FDFCC876FD74FEBF" box="[612,693,268,293]" pageId="4" pageNumber="532" type="male">1 male</specimenCount>
: Antioquia, Santa Rosa via Llanos de San Andrés,
<quantity id="4CE382D8FFE01877FB61C877FA56FEBE" box="[1273,1431,268,293]" metricMagnitude="3" metricUnit="m" metricValue="2.725" metricValueMax="2.75" metricValueMin="2.7" pageId="4" pageNumber="532" unit="m" value="2725.0" valueMax="2750.0" valueMin="2700.0">2700–2750 m</quantity>
,
<date id="FFA509FDFFE01877FF0FC84AFEEDFED3" box="[151,300,305,329]" pageId="4" pageNumber="532" value="2007-08-17">17.VIII.2007</date>
, T. Pyrcz leg., MZUJ;
<specimenCount id="9D1DE4B4FFE01877FDB4C84AFD46FED3" box="[556,647,304,329]" pageId="4" pageNumber="532" type="male">2 males</specimenCount>
: Antioquia, Loma Escobero, via al Retiro, Envigado, Medellin,
<quantity id="4CE382D8FFE01877FACEC84BFF3BFEF6" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="4" pageNumber="532" unit="m" value="2700.0" valueMax="2900.0" valueMin="2500.0">2500– 2900 m</quantity>
,
<date id="FFA509FDFFE01877FE8AC82EFE66FEF7" box="[274,423,341,365]" pageId="4" pageNumber="532" value="2007-08-04">04.VIII.2007</date>
, Pierre Boyer
<emphasis id="B96FF32FFFE01877FDF5C82FFD4EFEF6" box="[621,655,340,364]" italics="true" pageId="4" pageNumber="532">leg</emphasis>
.;
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: Antioquia, Envigado,
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,
<date id="FFA509FDFFE01877FB37C82EFAF1FEF7" box="[1199,1328,341,365]" pageId="4" pageNumber="532" value="2002-03-31">31.III.2002</date>
, Gabriel Rodriguez
<emphasis id="B96FF32FFFE01877FE8AC800FEF5FE09" box="[274,308,379,403]" italics="true" pageId="4" pageNumber="532">leg</emphasis>
., PBF;
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: Antioquia, La Estrella, El Romeral,
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,
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,
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, jama,
<date id="FFA509FDFFE01877FF0FC8D9FED9FE20" box="[151,280,418,442]" pageId="4" pageNumber="532" value="2010-02-28">28.02.2010</date>
, C. F. Alvarez leg., 256, BMC 20-1187, MUCM;
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: Caldas, Manizales,
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,
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,
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,
<date id="FFA509FDFFE01877FF0FC8BDFED9FE44" box="[151,280,454,478]" pageId="4" pageNumber="532" value="2007-09-28">28.09.2007</date>
, H. Warren-Gash leg., ICN-MHN-L 26865, MHN-UN.
</paragraph>
</subSubSection>
<subSubSection id="C3017CB6FFE01877FF5FC891FEB2FD41" pageId="4" pageNumber="532" type="diagnosis">
<paragraph id="8BA42F3DFFE01877FF5FC891FEB2FD41" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="532">
<emphasis id="B96FF32FFFE01877FF5FC891FE80FD99" bold="true" box="[199,321,490,515]" pageId="4" pageNumber="532">Diagnosis.</emphasis>
The only conspicuous HWV marking is a golden yellow mid-costal streak extending posteriorly to M1, which sets the species apart from all other sympatric congeners, in particular from the local population of
<taxonomicName id="4C1B54BEFFE01877FA1FCB74FEA9FDD1" authority="Hewitson" authorityName="Hewitson" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="porcia">
<emphasis id="B96FF32FFFE01877FA1FCB74FF1EFDD0" italics="true" pageId="4" pageNumber="532">P. porcia</emphasis>
(Hewitson)
</taxonomicName>
, which appears to be the most closely related species. The Antioquian population of
<taxonomicName id="4C1B54BEFFE01877FAAECB48FA5DFDD0" box="[1334,1436,563,586]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="porcia">
<emphasis id="B96FF32FFFE01877FAAECB48FA5DFDD0" box="[1334,1436,563,586]" italics="true" pageId="4" pageNumber="532">P. porcia</emphasis>
</taxonomicName>
(
<figureCitation id="132033B8FFE01877FF07CB2EFF25FDF4" box="[159,228,597,622]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1932,1954]" captionTargetBox="[262,1342,215,1896]" captionTargetId="figure@5.[221,1381,195,1934]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Pedaliodes adults. 1, P. haydoni male (holotype); 2, P. porcia male; 3, P. adrianae male (holotype); 4, P. adrianae female (paratype); 5, P. rodriguezi male (holotype); 6, P. ro dr i gu e z i female (paratype); 7, P. philinae male (holotype); 8, P. philinae female (paratype)." httpUri="https://zenodo.org/record/220317/files/figure.png" pageId="4" pageNumber="532">Fig. 2</figureCitation>
), however, is characterized by a long costal streak extending into the median area of the wing. A somewhat similar HWV pattern is found in
<taxonomicName id="4C1B54BEFFE01877FD90CB00FD72FD09" authority="Pyrcz" authorityName="Pyrcz" box="[520,691,634,659]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="rudnyi">
<emphasis id="B96FF32FFFE01877FD90CB00FDADFD08" box="[520,620,634,658]" italics="true" pageId="4" pageNumber="532">P. rudnyi</emphasis>
Pyrcz
</taxonomicName>
in southern
<collectingCountry id="F30C6FADFFE01877FCA5CB01FC6AFD09" box="[829,939,634,659]" name="Colombia" pageId="4" pageNumber="532">Colombia</collectingCountry>
and a few other Peruvian species, such as
<taxonomicName id="4C1B54BEFFE01877FA1FCB00FE99FD2C" authority="Pyrcz, P." authorityName="Pyrcz, P." class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="aureola">
<emphasis id="B96FF32FFFE01877FA1FCB00FF2CFD2C" italics="true" pageId="4" pageNumber="532">P. aureola</emphasis>
Pyrcz,
<emphasis id="B96FF32FFFE01877FEDACBE4FE99FD2C" box="[322,344,671,694]" italics="true" pageId="4" pageNumber="532">P.</emphasis>
</taxonomicName>
auraria Thieme and
<taxonomicName id="4C1B54BEFFE01877FDDFCBE4FCD3FD2D" authority="Pyrcz" authorityName="Pyrcz" box="[583,786,670,695]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="boettgeri">
<emphasis id="B96FF32FFFE01877FDDFCBE4FD09FD2C" box="[583,712,670,694]" italics="true" pageId="4" pageNumber="532">P. boettgeri</emphasis>
Pyrcz
</taxonomicName>
, all differing noticeably in male genitalia, which rules out their closer affinity.
</paragraph>
</subSubSection>
<subSubSection id="C3017CB6FFE01877FF5FCB9EFBDBF9A1" pageId="4" pageNumber="532" type="description">
<paragraph id="8BA42F3DFFE01877FF5FCB9EFBAAFA69" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="532">
<emphasis id="B96FF32FFFE01877FF5FCB9EFE94FD64" bold="true" box="[199,341,741,766]" pageId="4" pageNumber="532">Description.</emphasis>
<emphasis id="B96FF32FFFE01877FEC5CB9DFE57FD64" box="[349,406,742,766]" italics="true" pageId="4" pageNumber="532">Male</emphasis>
(
<figureCitation id="132033B8FFE01877FE3FCB9EFE2AFD64" box="[423,491,741,766]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1932,1954]" captionTargetBox="[262,1342,215,1896]" captionTargetId="figure@5.[221,1381,195,1934]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Pedaliodes adults. 1, P. haydoni male (holotype); 2, P. porcia male; 3, P. adrianae male (holotype); 4, P. adrianae female (paratype); 5, P. rodriguezi male (holotype); 6, P. ro dr i gu e z i female (paratype); 7, P. philinae male (holotype); 8, P. philinae female (paratype)." httpUri="https://zenodo.org/record/220317/files/figure.png" pageId="4" pageNumber="532">Fig. 5</figureCitation>
,
<figureCitation id="132033B8FFE01877FE61CB9EFDD6FD64" box="[505,535,741,766]" captionStart="FIGURES 13 – 15. 13" captionStartId="7.[151,264,1953,1975]" captionTargetBox="[378,1185,207,1911]" captionTargetId="figure@7.[214,1373,193,1932]" captionTargetPageId="7" captionText="FIGURES 13 – 15. 13, Pedaliodes adrianae (paratype) female genitalia; 14, Pedaliodes philinae (paratype) female genitalia; 15, Pedaliodes rodriguezi male (Cerro San Felix, Antioquia, Colombia, 2900 m)" httpUri="https://zenodo.org/record/220319/files/figure.png" pageId="4" pageNumber="532">15</figureCitation>
). Head: Antennae slender, reaching nearly half length of the costa, covered with sparse scales, dorsally brown, ventrally chestnut, with orange brown club formed gradually, slightly thicker than shaft, composed of 11 flagellomeres. Eyes blackish brown, lustrous, covered with sparse and short (compared to other species of
<taxonomicName id="4C1B54BEFFE01877FE95CA29FE46FCF0" box="[269,391,850,874]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="genus">
<emphasis id="B96FF32FFFE01877FE95CA29FE46FCF0" box="[269,391,850,874]" italics="true" pageId="4" pageNumber="532">Pedaliodes</emphasis>
</taxonomicName>
described here) hair. Labial palpi two times as long as the head, ventrally covered with long, black hairy scales, laterally and dorsally with short hair and greyish-brown scales. Frons with a tuft of rather short, black hair. Thorax: Dorsally black, patagium, tegulae and prothorax covered with long, brown and golden yellow hairy scales, otherwise mostly naked; ventrally black; legs brown, tibiae covered with chestnut hair, femora and tarsi with chestnut and sandy yellow scales, tarsi with numerous blackish, short spines. FW (length:
<quantity id="4CE382D8FFE01877FB23CA99FAE1FC60" box="[1211,1312,994,1019]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="4" pageNumber="532" unit="mm" value="27.5">27.5 mm</quantity>
) with apex subacute, outer margin straight; fringes intermittently milky white and brown. FWD uniform blackish brown, lustrous; a rectangular scent patch in median area, roughly
<quantity id="4CE382D8FFE01877FCB6CD51FC4DFBD8" box="[814,908,1066,1090]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="532" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
wide, extending from subcosta to anal margin. HW rounded with a slightly undulating outer margin; fringes short, dark brown. HWD uniform blakish brown, lustrous, basal one-third hairy. FWV medium brown, a shade lighter in distal one-third; a faint postdiscal costal streak made up of milky white scales, gradually fading away before reaching vein M2; distal margin from apex to vein M2 auburn. HWV ground color medium brown, liberally covered with dark brown ripple-like pattern, more prominent towards costal margin, and along apex, with a short, golden yellow costal streak in median area; a single, minute, milky white submarginal dot in Cu1–Cu2. Abdomen: Black, dorsally and laterally naked covered with chestnut, ventrally with chestnut and sandy yellow scales. Male genitalia (
<figureCitation id="132033B8FFE01877FCC3CC5EFC6BFAA4" box="[859,938,1317,1342]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1891,1913]" captionTargetBox="[247,1333,233,1790]" captionTargetId="figure@6.[243,1347,193,1849]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Male genitalia. Pedaliodes haydoni (paratype): 9 A, genitalia in lateral view; 9 B, aedeagus in lateral view; 9 C, aedeagus in dorsal view; Pedaliodes adrianae (paratype): 10 A, genitalia in lateral view; 10 B, aedeagus in lateral view; 10 C, aedeagus in dorsal view; Pedaliodes philinae (paratype): 11 A, genitalia in lateral view; 11 B, aedeagus in lateral view; 11 C, aedeagus in dorsal view; Pedaliodes rodriguezi (paratype): 12 A, genitalia in lateral view; 12 B, aedeagus in lateral view; 12 C, aedeagus in dorsal view." httpUri="https://zenodo.org/record/220318/files/figure.png" pageId="4" pageNumber="532">Fig. 12</figureCitation>
) with dorsum of tegumen flat, uncus slender, gently curving downwards, approximately same length of tegumen dorsum; subunci slender, approximately 2/3 the length of uncus; pedunculus small; valvae roughly the length of tegumen+uncus, ending with sharp, dorsal process aligned with the blunt distal extremity; saccus wide and rather shallow; aedeagus as long as saccus+valve, stout, contorted, slightly flattened dorsoventrally, proximal opening 2/5 the length of the entire aedeagus, approximately the same width as the remaining part of it, without any lateral cogs, and a subacute extremity.
</paragraph>
<paragraph id="8BA42F3DFFE01877FF5FCC85FBDBF9A1" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="532">
<emphasis id="B96FF32FFFE01877FF5FCC85FEDAF98C" box="[199,283,1534,1558]" italics="true" pageId="4" pageNumber="532">Female</emphasis>
(
<figureCitation id="132033B8FFE01877FEB5CC86FEB2F98C" box="[301,371,1533,1558]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1932,1954]" captionTargetBox="[262,1342,215,1896]" captionTargetId="figure@5.[221,1381,195,1934]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Pedaliodes adults. 1, P. haydoni male (holotype); 2, P. porcia male; 3, P. adrianae male (holotype); 4, P. adrianae female (paratype); 5, P. rodriguezi male (holotype); 6, P. ro dr i gu e z i female (paratype); 7, P. philinae male (holotype); 8, P. philinae female (paratype)." httpUri="https://zenodo.org/record/220317/files/figure.png" pageId="4" pageNumber="532">Fig. 6</figureCitation>
). Head and Thorax: Sexual dimorphism slight, expressed mostly in paler color of female on both dorsal and ventral wing surface. Abdomen: Female genitalia not examined.
</paragraph>
</subSubSection>
<subSubSection id="C3017CB6FFE01877FF5FCF3EFDC2F93C" pageId="4" pageNumber="532" type="etymology">
<paragraph id="8BA42F3DFFE01877FF5FCF3EFDC2F93C" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="532">
<emphasis id="B96FF32FFFE01877FF5FCF3EFE8FF9C4" bold="true" box="[199,334,1605,1630]" pageId="4" pageNumber="532">Etymology.</emphasis>
The specific epithet is a patronym for Gabriel Rodríguez, a Colombian geologist and lepidopterist from Medellín who has contributed significantly over the years to the knowledge of Colombian butterflies, and was the first collector of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3017CB6FFE01877FF5FCFC9FCA9F870" pageId="4" pageNumber="532" type="discussion">
<paragraph id="8BA42F3DFFE01877FF5FCFC9FCA9F870" blockId="4.[151,1437,151,2027]" pageId="4" pageNumber="532">
<emphasis id="B96FF32FFFE01877FF5FCFC9FE8EF951" bold="true" box="[199,335,1714,1739]" pageId="4" pageNumber="532">Comments.</emphasis>
This species could have been mistaken in the past with the widespread and polytypic
<taxonomicName id="4C1B54BEFFE01877FA87CFC9FF1EF974" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="porcia">
<emphasis id="B96FF32FFFE01877FA87CFC9FF1EF974" italics="true" pageId="4" pageNumber="532">Pedaliodes porcia</emphasis>
</taxonomicName>
. Nevertheless, the two occur sympatrically in Antioquia which immediately rules out their conspecifity.
<taxonomicName id="4C1B54BEFFE01877FA1FCFACFECAF888" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="rodriguezi">
<emphasis id="B96FF32FFFE01877FA1FCFACFECAF888" italics="true" pageId="4" pageNumber="532">P. rodriguezi</emphasis>
</taxonomicName>
is found in a number of localities in central Antioquia and is not uncommon at
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. It has not been reported so far from the areas of Central Cordillera situated farther south, for the example in the Nevado del Ruíz area, despite rather long-term and thorough sampling. This suggests that it may be the only endemic species of
<taxonomicName id="4C1B54BEFFE01877FF21CE1DFEF4F8E4" box="[185,309,1894,1918]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="genus">
<emphasis id="B96FF32FFFE01877FF21CE1DFEF4F8E4" box="[185,309,1894,1918]" italics="true" pageId="4" pageNumber="532">Pedaliodes</emphasis>
</taxonomicName>
in the Central Cordillera in Antioquia. One specimen from the collection of H. Warren-Gash is labeled as from Manizales, Caldas, but this record needs confirmation since no other individuals of
<taxonomicName id="4C1B54BEFFE01877FA93CEF0FA5DF838" box="[1291,1436,1930,1954]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="532" phylum="Arthropoda" rank="species" species="rodriguezi">
<emphasis id="B96FF32FFFE01877FA93CEF0FA5DF838" box="[1291,1436,1930,1954]" italics="true" pageId="4" pageNumber="532">P. rodriguezi</emphasis>
</taxonomicName>
from Caldas have been located, even in the Universidad de Caldas collection or during a long-term sampling project in the area of Manizales (C. Ríos Malaver, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
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