418 lines
35 KiB
XML
418 lines
35 KiB
XML
<document ID-DOI="10.15560/15.5.773" ID-ISSN="1809-127X" approvalRequired="58" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="55" approvalRequired_for_treatments="2" checkinTime="1630357722281" checkinUser="marcus" docAuthor="Llanqui, Irbin B., Y. Salas, Cinthya & Oblitas, Melissa P." docDate="2019" docId="900E87F7FFA5960FFCD8F95DB70C36DC" docLanguage="en" docName="CheckList.2019.15.5.773-796.pdf" docOrigin="Check List 15 (5)" docSource="http://dx.doi.org/10.15560/15.5.773" docStyle="DocumentStyle:BB30DA0EB5D11E1D6FE264CD32192DE2.1:CheckList.2017-.journal_article.annotatedlist" docStyleId="BB30DA0EB5D11E1D6FE264CD32192DE2" docStyleName="CheckList.2017-.journal_article.annotatedlist" docStyleVersion="1" docTitle="Pristimantis ockendeni Boulenger 1912" docType="treatment" docVersion="1" lastPageId="8" lastPageNumber="781" masterDocId="6C37FF8FFFA39607FFF5FF8EB52D3030" masterDocTitle="A preliminary checklist of amphibians and reptiles from the vicinity of La Nube Biological Station, Bahuaja-Sonene National Park, Peru" masterLastPageNumber="796" masterPageNumber="773" pageId="6" pageNumber="779" updateTime="1630357740796" updateUser="marcus" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A preliminary checklist of amphibians and reptiles from the vicinity of La Nube Biological Station, Bahuaja-Sonene National Park, Peru</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Llanqui, Irbin B.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Y. Salas, Cinthya</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Oblitas, Melissa P.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Check List</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2019-09-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>15</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>773</mods:start>
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<mods:end>796</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.15560/15.5.773</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.15560/15.5.773</mods:identifier>
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<mods:identifier type="ISSN">1809-127X</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:900E87F7FFA5960FFCD8F95DB70C36DC" httpUri="http://treatment.plazi.org/id/900E87F7FFA5960FFCD8F95DB70C36DC" lastPageId="8" lastPageNumber="781" pageId="6" pageNumber="779">
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<subSubSection box="[813,1260,1746,1771]" pageId="6" pageNumber="779" type="nomenclature">
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<paragraph blockId="6.[813,1260,1746,1771]" box="[813,1260,1746,1771]" pageId="6" pageNumber="779">
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||
<heading box="[813,1260,1746,1771]" fontSize="10" level="2" pageId="6" pageNumber="779" reason="2">
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<taxonomicName authority="Boulenger, 1912" authorityName="Boulenger" authorityYear="1912" box="[813,1260,1746,1771]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="779" phylum="Chordata" rank="species" species="ockendeni">
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<emphasis bold="true" box="[813,1260,1746,1771]" inLineHeading="true" pageId="6" pageNumber="779" reason="1">
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<emphasis bold="true" box="[813,1071,1746,1771]" italics="true" pageId="6" pageNumber="779">Pristimantis ockendeni</emphasis>
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Boulenger, 1912
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[813,922,1780,1805]" pageId="6" pageNumber="779" type="description">
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<paragraph blockId="6.[813,922,1780,1805]" box="[813,922,1780,1805]" pageId="6" pageNumber="779">
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<figureCitation box="[813,922,1780,1805]" captionStart="Figure 4" captionStartId="7.[168,231,2012,2032]" captionTargetBox="[184,1393,380,1942]" captionTargetId="figure-75@7.[168,1419,352,1959]" captionTargetPageId="7" captionText="Figure 4. A. Pristimantis cf. ockendeni (MUSA 4963, SVL: 21.8 mm), dorsal view. B. MUSA 4963, ventral view. C. Pristimantis cf. ockendeni (MUSA 4965, SVL: 22.7), dorsal view. D. Pristimantis ockendeni (MUSA 5808, SVL: 24.2 mm), dorsal view. E. Pristimantis reichlei (MUSA 4950, SVL:29.1 mm), dorsal view.F. Ameerega simulans (MUSA 4930,SVL: 25.53 mm),dorsal view.G. MUSA 4930,ventral view.Scale bars = 10 mm." pageId="6" pageNumber="779">Figure 4D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="781" pageId="6" pageNumber="779" type="materials_examined">
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||
<paragraph blockId="6.[813,1420,1829,2091]" lastBlockId="7.[168,775,160,284]" lastPageId="7" lastPageNumber="780" pageId="6" pageNumber="779">
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<emphasis bold="true" box="[813,1041,1830,1854]" pageId="6" pageNumber="779">Material examined.</emphasis>
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<collectionCode box="[1051,1128,1830,1854]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="6" pageNumber="779">MUSA</collectionCode>
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4956, 4958, 5794; (
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<geoCoordinate degrees="13" direction="south" minutes="25" orientation="latitude" pageId="6" pageNumber="779" precision="15" seconds="20" value="-13.422222">13°25′ 20″S</geoCoordinate>
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,
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||
<geoCoordinate box="[879,1027,1863,1888]" degrees="069" direction="west" minutes="36" orientation="longitude" pageId="6" pageNumber="779" precision="15" seconds="36" value="-69.61">069°36′36″W</geoCoordinate>
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), 22,23.
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<date box="[1113,1202,1864,1888]" pageId="6" pageNumber="779" value="2013-09">IX.2013</date>
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.
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<collectionCode box="[1212,1289,1864,1888]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="6" pageNumber="779">MUSA</collectionCode>
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4961, 5799; Transect TN21,
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<date box="[980,1102,1898,1922]" pageId="6" pageNumber="779" value="2013-09-25">25.IX.2013</date>
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.
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<collectionCode box="[1109,1186,1898,1922]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="6" pageNumber="779">MUSA</collectionCode>
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4962; Transect TN20,
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<date box="[813,934,1932,1956]" pageId="6" pageNumber="779" value="2013-09-25">25.IX.2013</date>
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.
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<collectionCode box="[950,1027,1932,1956]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="6" pageNumber="779">MUSA</collectionCode>
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5793; Transect TN16,
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<date box="[1294,1416,1932,1956]" pageId="6" pageNumber="779" value="2013-09-26">26.IX.2013</date>
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.
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<collectionCode box="[813,890,1966,1990]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="6" pageNumber="779">MUSA</collectionCode>
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5795, 5800; (
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<geoCoordinate box="[1109,1226,1964,1989]" degrees="13" direction="south" minutes="25" orientation="latitude" pageId="6" pageNumber="779" precision="15" seconds="41" value="-13.428056">13°25′41″S</geoCoordinate>
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,
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<geoCoordinate box="[1262,1408,1964,1989]" degrees="069" direction="west" minutes="36" orientation="longitude" pageId="6" pageNumber="779" precision="15" seconds="45" value="-69.612495">069°36′45″W</geoCoordinate>
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),
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<date box="[813,934,1999,2023]" pageId="6" pageNumber="779" value="2013-09-23">23.IX.2013</date>
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.
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<collectionCode box="[953,1030,1999,2023]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="6" pageNumber="779">MUSA</collectionCode>
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5798; TN13,
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<date box="[1201,1324,1999,2023]" pageId="6" pageNumber="779" value="2013-09-26">26.IX.2013</date>
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.
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<collectionCode box="[1342,1419,1999,2023]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="6" pageNumber="779">MUSA</collectionCode>
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5802; Transect TN11,
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<date box="[1066,1188,2033,2057]" pageId="6" pageNumber="779" value="2013-09-26">26.IX.2013</date>
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.
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<collectionCode box="[1203,1280,2033,2057]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="6" pageNumber="779">MUSA</collectionCode>
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5803, 5807, 5810; (
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<geoCoordinate box="[891,1012,2065,2090]" degrees="13" direction="south" minutes="25" orientation="latitude" pageId="6" pageNumber="779" precision="15" seconds="25" value="-13.423612">13°25′25″S</geoCoordinate>
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,
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<geoCoordinate box="[1030,1178,2065,2090]" degrees="069" direction="west" minutes="36" orientation="longitude" pageId="6" pageNumber="779" precision="15" seconds="36" value="-69.61">069°36′36″W</geoCoordinate>
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),
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<date box="[1203,1325,2067,2091]" pageId="6" pageNumber="779" value="2013-09-22">22.IX.2013</date>
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.
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<collectionCode box="[1342,1419,2067,2091]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="6" pageNumber="779">MUSA</collectionCode>
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5804, 5806; Transect TN21, 21,25.
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<date box="[536,624,160,184]" pageId="7" pageNumber="780" value="2013-09">IX.2013</date>
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.
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<collectionCode box="[633,710,160,184]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="7" pageNumber="780">MUSA</collectionCode>
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5808; (
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<geoCoordinate box="[176,297,192,217]" degrees="13" direction="south" minutes="25" orientation="latitude" pageId="7" pageNumber="780" precision="15" seconds="26" value="-13.423889">13°25′26″S</geoCoordinate>
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,
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<geoCoordinate box="[314,462,192,217]" degrees="069" direction="west" minutes="36" orientation="longitude" pageId="7" pageNumber="780" precision="15" seconds="36" value="-69.61">069°36′36″W</geoCoordinate>
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),
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<date box="[487,609,194,218]" pageId="7" pageNumber="780" value="2013-09-22">22.IX.2013</date>
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.
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<collectionCode box="[625,702,194,218]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="7" pageNumber="780">MUSA</collectionCode>
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5809; Transect TN20,
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<date box="[351,472,227,251]" pageId="7" pageNumber="780" value="2013-09-25">25.IX.2013</date>
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.
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<collectionCode box="[486,563,227,251]" country="0" httpUri="http://grbio.org/cool/42td-7yhx" name="Universidad Nacional de San Agustin, Museo de Historia Natural (Peru)" pageId="7" pageNumber="780">MUSA</collectionCode>
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5812; (
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<geoCoordinate box="[649,771,226,251]" degrees="13" direction="south" minutes="25" orientation="latitude" pageId="7" pageNumber="780" precision="15" seconds="24" value="-13.423334">13°25′24″S</geoCoordinate>
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,
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<geoCoordinate box="[168,316,259,284]" degrees="069" direction="west" minutes="36" orientation="longitude" pageId="7" pageNumber="780" precision="15" seconds="35" value="-69.60972">069°36′35″W</geoCoordinate>
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),
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<date box="[334,456,260,284]" pageId="7" pageNumber="780" value="2013-09-22">22.IX.2013</date>
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.
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</paragraph>
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<paragraph blockId="7.[813,1421,160,285]" lastBlockId="8.[167,777,1061,1465]" lastPageId="8" lastPageNumber="781" pageId="7" pageNumber="780">
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<emphasis bold="true" box="[813,974,160,184]" pageId="7" pageNumber="780">Identification.</emphasis>
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According to
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<bibRefCitation author="Hedges SB & Duellman WE & Heinicke MP" box="[1150,1390,160,184]" pageId="7" pageNumber="780" pagination="1 - 182" refId="ref15309" refString="Hedges SB, Duellman WE, Heinicke MP (2008) New World directdeveloping frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737: 1 - 182." type="journal article" year="2008">Hedges et al. (2008)</bibRefCitation>
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, a member of the
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<taxonomicName baseAuthorityName="Werner" baseAuthorityYear="1899" box="[993,1201,194,218]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="780" phylum="Chordata" rank="species" species="frater">
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<emphasis box="[993,1201,194,218]" italics="true" pageId="7" pageNumber="780">Pristimantis frater</emphasis>
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</taxonomicName>
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species group, but unassigned to any species group by
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<bibRefCitation author="Padial JM & Grant T & Frost DR" box="[1209,1416,227,251]" pageId="7" pageNumber="780" pagination="1 - 132" refId="ref16817" refString="Padial JM, Grant T, Frost DR (2014) Molecular systematics of terrara- nas (Anura: Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria. Zootaxa 3825 (1): 1 - 132. https: // doi. org / 10.11646 / zootaxa. 3825.1.1" type="journal article" year="2014">Padial et al. (2014)</bibRefCitation>
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. A small frog, SVL
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<quantity box="[1018,1170,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" metricValueMax="2.13" metricValueMin="1.25" pageId="7" pageNumber="780" unit="mm" value="16.9" valueMax="21.3" valueMin="12.5">12.5–21.3 mm</quantity>
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in males and
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<quantity lastPageId="8" lastPageNumber="781" metricMagnitude="-2" metricUnit="m" metricValue="2.685" metricValueMax="3.15" metricValueMin="2.2199999999999998" pageId="7" pageNumber="780" unit="mm" value="26.85" valueMax="31.5" valueMin="22.2">22.2–31.5 mm</quantity>
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in females. It can be identified by dorsum shagreen; venter coarsely areolate. Discoidal fold evident. Dorsolateral folds present or absent. Upper eyelids usually with small tubercles. Finger I <II. Toe
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<collectionCode box="[586,605,1165,1188]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="781" type="Museum">V</collectionCode>
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>> III. Dorsum varies from pale tan to reddish brown with dark brown marking, sometimes with an H- or W-shaped mark on the scapular region; venter pale gray. Labial bars present. Ventral and dorsal surfaces of thighs are brown to rosered (
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<bibRefCitation author="Duellman WE" box="[223,399,1337,1361]" pageId="8" pageNumber="781" pagination="352" refId="ref14598" refString="Duellman WE (1978) The biology of an equatorial herpetofauna in amazonian Ecuador. Miscellaneous Publications. Museum of Natural History University of Kansas 65: 352." type="journal article" year="1978">Duellman 1978</bibRefCitation>
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,
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<bibRefCitation author="Duellman WE" box="[417,473,1337,1361]" pageId="8" pageNumber="781" refId="ref14627" refString="Duellman WE (2005) Cusco Amazonico. The lives of amphibians and reptiles in an Amazonian Rianforest. Comstock Publishing, Cornell University Press, Ithaca, New York, 433 pp." type="book" year="2005">2005</bibRefCitation>
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,
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<bibRefCitation author="Rodriguez LO & Duellman WE" pageId="8" pageNumber="781" pagination="1 - 80" refId="ref17254" refString="Rodriguez LO, Duellman WE (1994) Guide to the frogs of the Iquitos Region, Amazonian Peru. The University of Kansas Natural History Museum, Special Publication 22: 1 - 80. https: // doi. org / 10.5962 / bhl. title. 7937" type="journal article" year="1994">Rodriguez and Duellman 1994</bibRefCitation>
|
||
,
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<bibRefCitation author="Duellman WE & Lehr E" box="[233,506,1372,1396]" pageId="8" pageNumber="781" refId="ref14661" refString="Duellman WE, Lehr E (2009) Terrestrial-breeding Frogs (Straboman- tidae) in Peru. Natur und Tier-Verlag, Munster, Germany, 382 pp." type="book" year="2009">Duellman and Lehr 2009</bibRefCitation>
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). There are no other similar
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<taxonomicName box="[211,430,1406,1430]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="781" phylum="Chordata" rank="species" species="undetermined">
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<emphasis box="[211,345,1407,1430]" italics="true" pageId="8" pageNumber="781">Pristimantis</emphasis>
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species
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</taxonomicName>
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in the area with small tubercles on the upper eyelids.
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</paragraph>
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</subSubSection>
|
||
<caption pageId="7" pageNumber="780" startId="7.[168,231,2012,2032]" targetBox="[184,1393,380,1942]" targetPageId="7">
|
||
<paragraph blockId="7.[168,1421,2012,2091]" pageId="7" pageNumber="780">
|
||
<emphasis bold="true" box="[168,282,2012,2032]" pageId="7" pageNumber="780">Figure 4. A.</emphasis>
|
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<taxonomicName authorityName="Boulenger" authorityYear="1912" box="[288,518,2012,2032]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="780" phylum="Chordata" rank="species" species="ockendeni">
|
||
<emphasis box="[288,392,2012,2032]" italics="true" pageId="7" pageNumber="780">Pristimantis</emphasis>
|
||
cf.
|
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<emphasis box="[427,518,2012,2032]" italics="true" pageId="7" pageNumber="780">ockendeni</emphasis>
|
||
</taxonomicName>
|
||
(MUSA 4963, SVL: 21.8 mm), dorsal view.
|
||
<emphasis bold="true" box="[902,921,2012,2032]" pageId="7" pageNumber="780">B.</emphasis>
|
||
MUSA 4963, ventral view.
|
||
<emphasis bold="true" box="[1167,1186,2012,2032]" pageId="7" pageNumber="780">C.</emphasis>
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<taxonomicName box="[1191,1295,2012,2032]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="780" phylum="Chordata" rank="genus">
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<emphasis box="[1191,1295,2012,2032]" italics="true" pageId="7" pageNumber="780">Pristimantis</emphasis>
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||
</taxonomicName>
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cf. o
|
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<emphasis box="[1340,1419,2012,2032]" italics="true" pageId="7" pageNumber="780">ckendeni</emphasis>
|
||
(MUSA 4965, SVL: 22.7), dorsal view.
|
||
<emphasis bold="true" box="[500,520,2041,2061]" pageId="7" pageNumber="780">D.</emphasis>
|
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<taxonomicName authorityName="Boulenger" authorityYear="1912" box="[525,725,2041,2061]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="780" phylum="Chordata" rank="species" species="ockendeni">
|
||
<emphasis box="[525,725,2041,2061]" italics="true" pageId="7" pageNumber="780">Pristimantis ockendeni</emphasis>
|
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</taxonomicName>
|
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(MUSA 5808, SVL: 24.2 mm), dorsal view.
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<emphasis bold="true" box="[1105,1123,2041,2061]" pageId="7" pageNumber="780">E.</emphasis>
|
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<taxonomicName authorityName="Padial & de la Riva" authorityYear="2009" box="[1127,1299,2041,2061]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="780" phylum="Chordata" rank="species" species="reichlei">
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<emphasis box="[1127,1299,2041,2061]" italics="true" pageId="7" pageNumber="780">Pristimantis reichlei</emphasis>
|
||
</taxonomicName>
|
||
(MUSA 4950, SVL:29.1 mm), dorsal view.
|
||
<emphasis bold="true" box="[412,429,2071,2091]" pageId="7" pageNumber="780">F.</emphasis>
|
||
<taxonomicName authorityName="Myers, Rodriguez, & Icochea" authorityYear="1998" box="[433,605,2071,2091]" class="Amphibia" family="Dendrobatidae" genus="Ameerega" kingdom="Animalia" order="Anura" pageId="7" pageNumber="780" phylum="Chordata" rank="species" species="simulans">
|
||
<emphasis box="[433,605,2071,2091]" italics="true" pageId="7" pageNumber="780">Ameerega simulans</emphasis>
|
||
</taxonomicName>
|
||
(MUSA 4930,SVL: 25.53 mm),dorsal view.
|
||
<emphasis bold="true" box="[984,1005,2071,2091]" pageId="7" pageNumber="780">G.</emphasis>
|
||
MUSA 4930,ventral view.Scale bars = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption box="[167,1401,162,182]" pageId="8" pageNumber="781" startId="8.[167,220,162,182]">
|
||
<paragraph blockId="8.[167,1401,162,182]" box="[167,1401,162,182]" pageId="8" pageNumber="781">
|
||
<emphasis bold="true" box="[167,243,162,182]" pageId="8" pageNumber="781">Table 3.</emphasis>
|
||
Comparison of characters of
|
||
<taxonomicName authorityName="Boulenger" authorityYear="1912" box="[512,737,162,182]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="781" phylum="Chordata" rank="species" species="ockendeni">
|
||
<emphasis box="[512,616,162,182]" italics="true" pageId="8" pageNumber="781">Pristimantis</emphasis>
|
||
cf.
|
||
<emphasis box="[646,737,162,182]" italics="true" pageId="8" pageNumber="781">ockendeni</emphasis>
|
||
</taxonomicName>
|
||
with those given by
|
||
<bibRefCitation author="Duellman WE & Lehr E" box="[927,1164,162,182]" pageId="8" pageNumber="781" refId="ref14661" refString="Duellman WE, Lehr E (2009) Terrestrial-breeding Frogs (Straboman- tidae) in Peru. Natur und Tier-Verlag, Munster, Germany, 382 pp." type="book" year="2009">Duellman and Lehr (2009)</bibRefCitation>
|
||
and Ocampo et al. (2017).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="8" pageNumber="781">
|
||
<table box="[176,1370,208,1011]" gridcols="3" gridrows="27" pageId="8" pageNumber="781">
|
||
<tr box="[176,1370,208,226]" gridrow="0" pageId="8" pageNumber="781">
|
||
<td box="[176,459,208,226]" gridcol="0" gridrow="0" pageId="8" pageNumber="781">
|
||
<emphasis bold="true" box="[176,265,208,226]" pageId="8" pageNumber="781">Character</emphasis>
|
||
</td>
|
||
<td box="[505,825,208,226]" gridcol="1" gridrow="0" pageId="8" pageNumber="781">
|
||
<emphasis bold="true" box="[505,825,208,226]" pageId="8" pageNumber="781">MUSA 4957, 4963, 4964, 4965, 4967</emphasis>
|
||
</td>
|
||
<td box="[904,1370,208,226]" gridcol="2" gridrow="0" pageId="8" pageNumber="781">
|
||
<emphasis bold="true" box="[904,1370,208,226]" pageId="8" pageNumber="781">
|
||
<bibRefCitation author="Duellman WE & Lehr E" box="[904,1140,208,226]" pageId="8" pageNumber="781" refId="ref14661" refString="Duellman WE, Lehr E (2009) Terrestrial-breeding Frogs (Straboman- tidae) in Peru. Natur und Tier-Verlag, Munster, Germany, 382 pp." type="book" year="2009">Duellman and Lehr (2009)</bibRefCitation>
|
||
, and Ocampo et al. (2017)
|
||
</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr box="[176,1370,240,258]" gridrow="1" pageId="8" pageNumber="781">
|
||
<td box="[176,459,240,258]" gridcol="0" gridrow="1" pageId="8" pageNumber="781">Dorsal view of head</td>
|
||
<td box="[505,825,240,258]" gridcol="1" gridrow="1" pageId="8" pageNumber="781">Subacuminate</td>
|
||
<td box="[904,1370,240,258]" gridcol="2" gridrow="1" pageId="8" pageNumber="781">Subacuminate</td>
|
||
</tr>
|
||
<tr box="[176,1370,270,288]" gridrow="2" pageId="8" pageNumber="781">
|
||
<td box="[176,459,270,288]" gridcol="0" gridrow="2" pageId="8" pageNumber="781">Lateral view of head</td>
|
||
<td box="[505,825,270,288]" gridcol="1" gridrow="2" pageId="8" pageNumber="781">Rounded</td>
|
||
<td box="[904,1370,270,288]" gridcol="2" gridrow="2" pageId="8" pageNumber="781">Rounded</td>
|
||
</tr>
|
||
<tr box="[176,1370,300,318]" gridrow="3" pageId="8" pageNumber="781">
|
||
<td box="[176,459,300,318]" gridcol="0" gridrow="3" pageId="8" pageNumber="781">Snout</td>
|
||
<td box="[505,825,300,318]" gridcol="1" gridrow="3" pageId="8" pageNumber="781">Short</td>
|
||
<td box="[904,1370,300,318]" gridcol="2" gridrow="3" pageId="8" pageNumber="781">Short</td>
|
||
</tr>
|
||
<tr box="[176,1370,330,348]" gridrow="4" pageId="8" pageNumber="781">
|
||
<td box="[176,459,330,348]" gridcol="0" gridrow="4" pageId="8" pageNumber="781">Skin dorsum</td>
|
||
<td box="[505,825,330,348]" gridcol="1" gridrow="4" pageId="8" pageNumber="781">Shagreen</td>
|
||
<td box="[904,1370,330,348]" gridcol="2" gridrow="4" pageId="8" pageNumber="781">Shagreen</td>
|
||
</tr>
|
||
<tr box="[176,1370,360,378]" gridrow="5" pageId="8" pageNumber="781">
|
||
<td box="[176,459,360,378]" gridcol="0" gridrow="5" pageId="8" pageNumber="781">Discoidal fold</td>
|
||
<td box="[505,825,360,378]" gridcol="1" gridrow="5" pageId="8" pageNumber="781">Present</td>
|
||
<td box="[904,1370,360,378]" gridcol="2" gridrow="5" pageId="8" pageNumber="781">Present</td>
|
||
</tr>
|
||
<tr box="[176,1370,391,409]" gridrow="6" pageId="8" pageNumber="781">
|
||
<td box="[176,459,391,409]" gridcol="0" gridrow="6" pageId="8" pageNumber="781">W–shaped occipital ridges</td>
|
||
<td box="[505,825,391,409]" gridcol="1" gridrow="6" pageId="8" pageNumber="781">yes/not</td>
|
||
<td box="[904,1370,391,409]" gridcol="2" gridrow="6" pageId="8" pageNumber="781">yes/not</td>
|
||
</tr>
|
||
<tr box="[176,1370,421,439]" gridrow="7" pageId="8" pageNumber="781">
|
||
<td box="[176,459,421,439]" gridcol="0" gridrow="7" pageId="8" pageNumber="781">Skin venter</td>
|
||
<td box="[505,825,421,439]" gridcol="1" gridrow="7" pageId="8" pageNumber="781">Weakly areolate</td>
|
||
<td box="[904,1370,421,439]" gridcol="2" gridrow="7" pageId="8" pageNumber="781">Areolate</td>
|
||
</tr>
|
||
<tr box="[176,1370,451,469]" gridrow="8" pageId="8" pageNumber="781">
|
||
<td box="[176,459,451,469]" gridcol="0" gridrow="8" pageId="8" pageNumber="781">Cranial crests</td>
|
||
<td box="[505,825,451,469]" gridcol="1" gridrow="8" pageId="8" pageNumber="781">Absent</td>
|
||
<td box="[904,1370,451,469]" gridcol="2" gridrow="8" pageId="8" pageNumber="781">Absent</td>
|
||
</tr>
|
||
<tr box="[176,1370,481,499]" gridrow="9" pageId="8" pageNumber="781">
|
||
<td box="[176,459,481,499]" gridcol="0" gridrow="9" pageId="8" pageNumber="781">Tympanic annulus</td>
|
||
<td box="[505,825,481,499]" gridcol="1" gridrow="9" pageId="8" pageNumber="781">Evident</td>
|
||
<td box="[904,1370,481,499]" gridcol="2" gridrow="9" pageId="8" pageNumber="781">Usually evident</td>
|
||
</tr>
|
||
<tr box="[176,1370,511,529]" gridrow="10" pageId="8" pageNumber="781">
|
||
<td box="[176,459,511,529]" gridcol="0" gridrow="10" pageId="8" pageNumber="781">Tympanic membrane</td>
|
||
<td box="[505,825,511,529]" gridcol="1" gridrow="10" pageId="8" pageNumber="781">Evident</td>
|
||
<td box="[904,1370,511,529]" gridcol="2" gridrow="10" pageId="8" pageNumber="781">Usually evident</td>
|
||
</tr>
|
||
<tr box="[176,1370,541,559]" gridrow="11" pageId="8" pageNumber="781">
|
||
<td box="[176,459,541,559]" gridcol="0" gridrow="11" pageId="8" pageNumber="781">Tympanum length</td>
|
||
<td box="[505,825,541,559]" gridcol="1" gridrow="11" pageId="8" pageNumber="781">0.16–0.32 of the eye length</td>
|
||
<td box="[904,1370,541,559]" gridcol="2" gridrow="11" pageId="8" pageNumber="781">0.25–0.4 of the eye length</td>
|
||
</tr>
|
||
<tr box="[176,1370,571,589]" gridrow="12" pageId="8" pageNumber="781">
|
||
<td box="[176,459,571,589]" gridcol="0" gridrow="12" pageId="8" pageNumber="781">Eye length</td>
|
||
<td box="[505,825,571,589]" gridcol="1" gridrow="12" pageId="8" pageNumber="781">> eye nostril distance</td>
|
||
<td box="[904,1370,571,589]" gridcol="2" gridrow="12" pageId="8" pageNumber="781">> eye nostril distance</td>
|
||
</tr>
|
||
<tr box="[176,1370,601,619]" gridrow="13" pageId="8" pageNumber="781">
|
||
<td box="[176,459,601,619]" gridcol="0" gridrow="13" pageId="8" pageNumber="781">Prevomerine odontophores</td>
|
||
<td box="[505,825,601,619]" gridcol="1" gridrow="13" pageId="8" pageNumber="781">Oblique</td>
|
||
<td box="[904,1370,601,619]" gridcol="2" gridrow="13" pageId="8" pageNumber="781">Oblique</td>
|
||
</tr>
|
||
<tr box="[176,1370,631,649]" gridrow="14" pageId="8" pageNumber="781">
|
||
<td box="[176,459,631,649]" gridcol="0" gridrow="14" pageId="8" pageNumber="781">Canthal stripe from nostril to eye</td>
|
||
<td box="[505,825,631,649]" gridcol="1" gridrow="14" pageId="8" pageNumber="781">Absent</td>
|
||
<td box="[904,1370,631,649]" gridcol="2" gridrow="14" pageId="8" pageNumber="781">Absent</td>
|
||
</tr>
|
||
<tr box="[176,1370,662,680]" gridrow="15" pageId="8" pageNumber="781">
|
||
<td box="[176,459,662,680]" gridcol="0" gridrow="15" pageId="8" pageNumber="781">Upper eyelid tubercles</td>
|
||
<td box="[505,825,662,680]" gridcol="1" gridrow="15" pageId="8" pageNumber="781">Yes/not</td>
|
||
<td box="[904,1370,662,680]" gridcol="2" gridrow="15" pageId="8" pageNumber="781">Present</td>
|
||
</tr>
|
||
<tr box="[176,1370,692,710]" gridrow="16" pageId="8" pageNumber="781">
|
||
<td box="[176,459,692,710]" gridcol="0" gridrow="16" pageId="8" pageNumber="781">Supratympanic stripe</td>
|
||
<td box="[505,825,692,710]" gridcol="1" gridrow="16" pageId="8" pageNumber="781">Present</td>
|
||
<td box="[904,1370,692,710]" gridcol="2" gridrow="16" pageId="8" pageNumber="781">Present</td>
|
||
</tr>
|
||
<tr box="[176,1370,722,740]" gridrow="17" pageId="8" pageNumber="781">
|
||
<td box="[176,459,722,740]" gridcol="0" gridrow="17" pageId="8" pageNumber="781">Finger</td>
|
||
<td box="[505,825,722,740]" gridcol="1" gridrow="17" pageId="8" pageNumber="781">I <II</td>
|
||
<td box="[904,1370,722,740]" gridcol="2" gridrow="17" pageId="8" pageNumber="781">I <II</td>
|
||
</tr>
|
||
<tr box="[176,1370,752,770]" gridrow="18" pageId="8" pageNumber="781">
|
||
<td box="[176,459,752,770]" gridcol="0" gridrow="18" pageId="8" pageNumber="781">All fingers bearing pads and discs</td>
|
||
<td box="[505,825,752,770]" gridcol="1" gridrow="18" pageId="8" pageNumber="781">Yes</td>
|
||
<td box="[904,1370,752,770]" gridcol="2" gridrow="18" pageId="8" pageNumber="781">Yes</td>
|
||
</tr>
|
||
<tr box="[176,1370,782,800]" gridrow="19" pageId="8" pageNumber="781">
|
||
<td box="[176,459,782,800]" gridcol="0" gridrow="19" pageId="8" pageNumber="781">Largest pad on finger III or IV</td>
|
||
<td box="[505,825,782,800]" gridcol="1" gridrow="19" pageId="8" pageNumber="781">Slightly larger/yes</td>
|
||
<td box="[904,1370,782,800]" gridcol="2" gridrow="19" pageId="8" pageNumber="781">Yes</td>
|
||
</tr>
|
||
<tr box="[176,1370,812,830]" gridrow="20" pageId="8" pageNumber="781">
|
||
<td box="[176,459,812,830]" gridcol="0" gridrow="20" pageId="8" pageNumber="781">Toes</td>
|
||
<td box="[505,825,812,830]" gridcol="1" gridrow="20" pageId="8" pageNumber="781">III ≈ V</td>
|
||
<td box="[904,1370,812,830]" gridcol="2" gridrow="20" pageId="8" pageNumber="781">III << V</td>
|
||
</tr>
|
||
<tr box="[176,1370,842,860]" gridrow="21" pageId="8" pageNumber="781">
|
||
<td box="[176,459,842,860]" gridcol="0" gridrow="21" pageId="8" pageNumber="781">Ulnar tubercles</td>
|
||
<td box="[505,825,842,860]" gridcol="1" gridrow="21" pageId="8" pageNumber="781">Conspicuous</td>
|
||
<td box="[904,1370,842,860]" gridcol="2" gridrow="21" pageId="8" pageNumber="781">Minute</td>
|
||
</tr>
|
||
<tr box="[176,1370,872,890]" gridrow="22" pageId="8" pageNumber="781">
|
||
<td box="[176,459,872,890]" gridcol="0" gridrow="22" pageId="8" pageNumber="781">SVL males (mm)</td>
|
||
<td box="[505,825,872,890]" gridcol="1" gridrow="22" pageId="8" pageNumber="781">18.75–23.45</td>
|
||
<td box="[904,1370,872,890]" gridcol="2" gridrow="22" pageId="8" pageNumber="781">15.3–21.3</td>
|
||
</tr>
|
||
<tr box="[176,1370,903,921]" gridrow="23" pageId="8" pageNumber="781">
|
||
<td box="[176,459,903,921]" gridcol="0" gridrow="23" pageId="8" pageNumber="781">SVL females (mm)</td>
|
||
<td box="[505,825,903,921]" gridcol="1" gridrow="23" pageId="8" pageNumber="781">No available</td>
|
||
<td box="[904,1370,903,921]" gridcol="2" gridrow="23" pageId="8" pageNumber="781">22.2–31.5</td>
|
||
</tr>
|
||
<tr box="[176,1370,933,951]" gridrow="24" pageId="8" pageNumber="781">
|
||
<td box="[176,459,933,951]" gridcol="0" gridrow="24" pageId="8" pageNumber="781">Lumbar bar</td>
|
||
<td box="[505,825,933,951]" gridcol="1" gridrow="24" pageId="8" pageNumber="781">Absent/Present</td>
|
||
<td box="[904,1370,933,951]" gridcol="2" gridrow="24" pageId="8" pageNumber="781">Present</td>
|
||
</tr>
|
||
<tr box="[176,1370,963,981]" gridrow="25" pageId="8" pageNumber="781">
|
||
<td box="[176,459,963,981]" gridcol="0" gridrow="25" pageId="8" pageNumber="781">Dorsal design</td>
|
||
<td box="[505,825,963,981]" gridcol="1" gridrow="25" pageId="8" pageNumber="781">Some with longitudinal lines</td>
|
||
<td box="[904,1370,963,981]" gridcol="2" gridrow="25" pageId="8" pageNumber="781">Dark brown markings</td>
|
||
</tr>
|
||
<tr box="[176,1370,993,1011]" gridrow="26" pageId="8" pageNumber="781">
|
||
<td box="[176,459,993,1011]" gridcol="0" gridrow="26" pageId="8" pageNumber="781">Tights coloration</td>
|
||
<td box="[505,825,993,1011]" gridcol="1" gridrow="26" pageId="8" pageNumber="781">Brown to orange-red</td>
|
||
<td box="[904,1370,993,1011]" gridcol="2" gridrow="26" pageId="8" pageNumber="781">Brown to rose-red</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection pageId="8" pageNumber="781" type="distribution">
|
||
<paragraph blockId="8.[167,777,1491,1653]" pageId="8" pageNumber="781">
|
||
<emphasis box="[168,775,1491,1515]" inLineHeading="true" pageId="8" pageNumber="781" reason="1">
|
||
<emphasis bold="true" box="[168,317,1491,1515]" pageId="8" pageNumber="781">Distribution.</emphasis>
|
||
<taxonomicName authorityName="Boulenger" authorityYear="1912" box="[322,571,1491,1515]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="781" phylum="Chordata" rank="species" species="ockendeni">
|
||
<emphasis box="[322,571,1491,1515]" italics="true" pageId="8" pageNumber="781">Pristimantis ockendeni</emphasis>
|
||
</taxonomicName>
|
||
has a controversial
|
||
</emphasis>
|
||
distribution (see
|
||
<bibRefCitation author="Elmer KR & Cannatella DC" box="[351,654,1526,1550]" pageId="8" pageNumber="781" pagination="11 - 38" refId="ref14831" refString="Elmer KR, Cannatella DC (2008) Three new species of leaflitter frogs from the upper Amazon forest: cryptic diversity within Pristimantis " ockendeni " (Anura: Strabomantidae) in Ecuador. Zootaxa 1784: 11 - 38." type="journal article" year="2008">Elmer and Cannatella 2008</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Duellman WE & Lehr E" pageId="8" pageNumber="781" refId="ref14661" refString="Duellman WE, Lehr E (2009) Terrestrial-breeding Frogs (Straboman- tidae) in Peru. Natur und Tier-Verlag, Munster, Germany, 382 pp." type="book" year="2009">Duellman and Lehr 2009</bibRefCitation>
|
||
), but currently is restricted to Amazonia of
|
||
<collectingCountry box="[201,283,1594,1618]" name="Bolivia" pageId="8" pageNumber="781">Bolivia</collectingCountry>
|
||
,
|
||
<collectingCountry box="[297,366,1594,1618]" name="Brazil" pageId="8" pageNumber="781">Brazil</collectingCountry>
|
||
,
|
||
<collectingCountry box="[380,488,1594,1618]" name="Colombia" pageId="8" pageNumber="781">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry box="[501,593,1594,1618]" name="Ecuador" pageId="8" pageNumber="781">Ecuador</collectingCountry>
|
||
, and
|
||
<collectingCountry box="[656,709,1595,1618]" name="Peru" pageId="8" pageNumber="781">Peru</collectingCountry>
|
||
, from
|
||
<quantity box="[168,323,1629,1653]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.8" metricValueMin="0.3" pageId="8" pageNumber="781" unit="m" value="1050.0" valueMax="1800.0" valueMin="300.0">300 to 1800 m</quantity>
|
||
a.s.l. (Ocampo et al. 2017,
|
||
<bibRefCitation author="Frost DR" box="[611,727,1629,1653]" pageId="8" pageNumber="781" refId="ref15004" refString="Frost DR (2019) Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History, New York, USA. http: // research. amnh. org / herpetology / amphibia / index. html. Accessed on: 2019 - 05 - 01" type="url" year="2019">Frost 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="781" type="discussion">
|
||
<paragraph blockId="8.[168,777,1679,1772]" pageId="8" pageNumber="781">
|
||
<emphasis box="[168,746,1679,1704]" inLineHeading="true" pageId="8" pageNumber="781" reason="1">
|
||
<emphasis bold="true" box="[168,279,1680,1704]" pageId="8" pageNumber="781">Remarks.</emphasis>
|
||
In respect to coloration and dorsal pattern,
|
||
</emphasis>
|
||
<taxonomicName authorityName="Boulenger" authorityYear="1912" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="781" phylum="Chordata" rank="species" species="ockendeni">
|
||
<emphasis box="[753,777,1680,1703]" inLineHeading="true" italics="true" pageId="8" pageNumber="781" reason="1">P.</emphasis>
|
||
ockendeni
|
||
</taxonomicName>
|
||
can be highly variable, and this character is insufficient to identify this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |