259 lines
33 KiB
XML
259 lines
33 KiB
XML
<document id="374855D1327F6EFAC76369D5B1EC1F1D" ID-CLB-Dataset="51458" ID-DOI="10.5281/zenodo.155781" ID-GBIF-Dataset="9f9ad059-7f30-42dd-91d3-324ef0ba9c24" ID-ISSN="11755326" ID-Zenodo-Dep="155781" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459432365550" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="C419FA02FFA96263CB19A1CCB06CFE52" docLanguage="en" docName="zt00034.pdf" docOrigin="Zootaxa 34" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Microlia tetramera Gusarov, sp. nov." docType="treatment" docVersion="15" lastPageNumber="21" masterDocId="3820827AFFBB6277CA11A34FB41BFFFC" masterDocTitle="A revision of the genus Microlia Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae: Hoplandriini)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="19" updateTime="1698192266583" updateUser="ExternalLinkService">
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<mods:mods id="64B412D2DCC3DCC587579D41F69A8076" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4EF53C67C8A1B47EEEA2108A1EFDEC7B">
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<mods:title id="A12C83AD8D1F1037057ABBE653D015A2">A revision of the genus Microlia Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae: Hoplandriini)</mods:title>
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</mods:titleInfo>
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<mods:name id="EA493BEDCB13308CF28B71696E03372B" type="personal">
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<mods:role id="E9747DE4EA595291BB9BD698A7FEDFF0">
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<mods:roleTerm id="9323C93B99BB85BF94C4ACB6BE714C2A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D3FFAC542D25054CBAFA1F016D1365AB">Gusarov, Vladimir I.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="DFF52AB10127E291F3E5C243B2C03044">text</mods:typeOfResource>
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<mods:title id="07E7E76EE6BE6DB87E65B213EEC4C5EB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E3ADCF6299859F724FDC0DECECB83AEF">
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<mods:date id="93737C97D7CBF4C697A1B20E24C5F92E">2002</mods:date>
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<mods:detail id="76B4F154EAFCF55C38F131B5C08D7D42" type="volume">
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<mods:number id="2DE14A5C9FBC56A8677A7AFCB5906E99">34</mods:number>
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<mods:start id="35BF7A9100891C7014DFB448AFE45A51">1</mods:start>
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<mods:end id="0D3E0DCB816219BF9EF7DC129FD10A73">24</mods:end>
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</mods:extent>
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</mods:relatedItem>
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<mods:classification id="54A38A5943AB3838935B384D1B22CFB3">journal article</mods:classification>
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<mods:identifier id="CEDA7CB902FECEA6721ACD1DF39B20AD" type="CLB-Dataset">51458</mods:identifier>
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<mods:identifier id="C0B1DC4673593A1A82599DEF450830EC" type="DOI">10.5281/zenodo.155781</mods:identifier>
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<mods:identifier id="A8F563D660111A64DFD95DF7F8E9F0C2" type="GBIF-Dataset">9f9ad059-7f30-42dd-91d3-324ef0ba9c24</mods:identifier>
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<mods:identifier id="31FC53C39DB9FA15A9819BBA44AEFDBD" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="FEF999B6E9E513A97EBF9719C1F76D66" type="Zenodo-Dep">155781</mods:identifier>
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<treatment id="C419FA02FFA96263CB19A1CCB06CFE52" ID-DOI="http://doi.org/10.5281/zenodo.6277387" ID-Zenodo-Dep="6277387" LSID="urn:lsid:plazi:treatment:C419FA02FFA96263CB19A1CCB06CFE52" httpUri="http://treatment.plazi.org/id/C419FA02FFA96263CB19A1CCB06CFE52" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
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<subSubSection id="04AA189FFFA96265CB19A1CCB7C6FD61" box="[264,989,643,670]" pageId="18" pageNumber="19" type="key">
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<paragraph id="4C0F4B14FFA96265CB19A1CCB7C6FD61" blockId="18.[264,989,643,670]" box="[264,989,643,670]" pageId="18" pageNumber="19">
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<heading id="1747FC78FFA96265CB19A1CCB7C6FD61" bold="true" box="[264,989,643,670]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<emphasis id="7EC49706FFA96265CB19A1CCB7C6FD61" bold="true" box="[264,989,643,670]" pageId="18" pageNumber="19">
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<keyStep id="F74453B1FFA96265CB19A1CCB7C6FD61" box="[264,989,643,670]" pageId="18" pageNumber="19">
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<key id="38211C25FFA96265CB19A1CCB7C6FD61" box="[264,989,643,670]" pageId="18" pageNumber="19">
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<keyLead id="F741E821FFA96265CB19A1CCB7C6FD61" box="[264,989,643,670]" pageId="18" pageNumber="19">
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||
5.
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<taxonomicName id="8BB03097FFA96265CB37A1CBB665FD61" ID-CoL="73GRV" authority="Gusarov" authorityName="Gusarov" box="[294,638,643,670]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tetramera" status="sp. nov.">
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<emphasis id="7EC49706FFA96265CB37A1CBB611FD62" bold="true" box="[294,522,644,670]" italics="true" pageId="18" pageNumber="19">Microlia tetramera</emphasis>
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Gusarov
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</taxonomicName>
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,
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<taxonomicNameLabel id="65F72A7DFFA96265C89BA1CCB6F3FD61" box="[650,744,643,669]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="D48B5791FFA96265C8E9A1CCB799FD61" box="[760,898,643,669]" captionStart="FIGURES 13 20" captionStartId="5.[264,383,1693,1717]" captionText="FIGURES 13 20. Details of Microlia meticola (Casey) (13 15, Arizona), M. azteca (Casey) (16, paralectotype), M. cf. tetramera Gusarov, sp. nov. (17 18, female from Costa Rica) and M. silacea (Erichson) (19 20, paralectotype of Homalota silacea). 13 – pronotum; 14 – mesometathorax, ventral view; 15 16 – right mesotarsus, posterior view; 17, 19 – right mesotarsus, dorsal view; 18, 20 – right mesotarsus, ventral view; d – dorsal setae of mesotarsus; v – ventral setae; l – lateral setae. Scale bar 0.4 mm (13 14), 0.28 mm (15 16), 0.1 mm (17 20)." pageId="18" pageNumber="19">Figs. 1718</figureCitation>
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,
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<figureCitation id="D48B5791FFA96265C99EA1CCB7CEFD61" box="[911,981,643,669]" captionStart-0="FIGURES 62 67" captionStart-1="FIGURES 68 76" captionStartId-0="19.[264,383,1428,1452]" captionStartId-1="20.[264,383,1277,1301]" captionTargetBox-0="[360,1260,320,1405]" captionTargetBox-1="[351,1226,513,1257]" captionTargetId-0="figure@19.[354,1262,320,1409]" captionTargetId-1="figure@20.[348,1227,513,1257]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 62 67. Details of Microlia tetramera Gusarov, sp. nov. (62 63 – holotype; 64 67 – possible female of M. tetramera from Costa Rica, 64 65 – type A, 66 67 – type B). 62 – male sternum 8; 63 – male tergum 8; 64, 66 – female sternum 8; 65, 67 – female tergum 8. Scale bar 0.2 mm." captionText-1="FIGURES 68 76. Details of Microlia tetramera Gusarov, sp. nov. (68 72, 75 76 – holotype; 73 74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73 74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73 74), 0.1 mm (69, 71 72, 75), 0.55 mm (76)." httpUri-0="https://zenodo.org/record/155790/files/figure.png" httpUri-1="https://zenodo.org/record/155791/files/figure.png" pageId="18" pageNumber="19">6276</figureCitation>
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)
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</keyLead>
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</key>
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||
</keyStep>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="04AA189FFFA96265CB19A19CB0BCFC72" pageId="18" pageNumber="19" type="materials_examined">
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<paragraph id="4C0F4B14FFA96265CB19A19CB6B0FCEA" blockId="18.[264,1324,723,1870]" pageId="18" pageNumber="19">
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<emphasis id="7EC49706FFA96265CB19A19CB5A5FD11" bold="true" box="[264,446,723,749]" pageId="18" pageNumber="19">
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<typeStatus id="930BF5B6FFA96265CB19A19CB55EFD11" box="[264,325,723,749]" pageId="18" pageNumber="19">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="930BF5B6FFA96265CBDFA19BB65BFD12" box="[462,576,724,750]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
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:,
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<emphasis id="7EC49706FFA96265C87CA19CB730FD11" bold="true" box="[621,811,723,749]" pageId="18" pageNumber="19">
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<collectingCountry id="34A70B84FFA96265C87CA19CB73DFD11" box="[621,806,723,749]" name="Costa Rica" pageId="18" pageNumber="19">COSTA RICA</collectingCountry>
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:
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</emphasis>
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San Jose, Zurquí de Moravia,
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<quantity id="8B48E6F1FFA96265CE8CA19BB0EAFD12" box="[1181,1265,724,750]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="18" pageNumber="19" unit="m" value="1600.0">1600m</quantity>
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, malaise (P. Hanson),
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<date id="380E6DD4FFA96265CBCDA1B3B62CFCEA" box="[476,567,764,790]" pageId="18" pageNumber="19" value="1994-03">iii.1994</date>
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(
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<collectionCode id="2AA1D3D1FFA96265C857A1B3B685FCEA" box="[582,670,764,790]" pageId="18" pageNumber="19">KSEM</collectionCode>
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).
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</paragraph>
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<paragraph id="4C0F4B14FFA96265CB29A06CB0BCFC72" blockId="18.[264,1324,723,1870]" pageId="18" pageNumber="19">
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<emphasis id="7EC49706FFA96265CB29A06CB70FFCC1" bold="true" box="[312,788,803,829]" pageId="18" pageNumber="19">
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Additional material:
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<collectingCountry id="34A70B84FFA96265C843A06CB714FCC1" box="[594,783,803,829]" name="Costa Rica" pageId="18" pageNumber="19">COSTA RICA</collectingCountry>
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:
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</emphasis>
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possible females of
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<taxonomicName id="8BB03097FFA96265CE0EA06BB0A6FCC1" box="[1055,1213,804,829]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tetramera">
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<emphasis id="7EC49706FFA96265CE0EA06BB0A6FCC1" box="[1055,1213,804,829]" italics="true" pageId="18" pageNumber="19">M. tetramera</emphasis>
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</taxonomicName>
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,
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<typeStatus id="930BF5B6FFA96265CEC1A06BB119FCC2" box="[1232,1282,804,830]" pageId="18" pageNumber="19">type</typeStatus>
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A: 3, same data as the
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<typeStatus id="930BF5B6FFA96265C835A003B691FC9A" box="[548,650,844,870]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
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but vivii.1994;, ditto but
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<date id="380E6DD4FFA96265C9E1A003B053FC9A" box="[1008,1096,844,870]" pageId="18" pageNumber="19" value="1993-06">vi.1993</date>
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||
(
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||
<collectionCode id="2AA1D3D1FFA96265CE4DA003B0AFFC9A" box="[1116,1204,844,870]" pageId="18" pageNumber="19">KSEM</collectionCode>
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); possible females of
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<taxonomicName id="8BB03097FFA96265CB9BA03BB63EFC71" box="[394,549,884,909]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tetramera">
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<emphasis id="7EC49706FFA96265CB9BA03BB63EFC71" box="[394,549,884,909]" italics="true" pageId="18" pageNumber="19">M. tetramera</emphasis>
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||
</taxonomicName>
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,
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<typeStatus id="930BF5B6FFA96265C825A03BB67DFC72" box="[564,614,884,910]" pageId="18" pageNumber="19">type</typeStatus>
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B: 2, as the
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<typeStatus id="930BF5B6FFA96265C936A03BB796FC72" box="[807,909,884,910]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
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||
but ixx.1993 (
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||
<collectionCode id="2AA1D3D1FFA96265CE53A03BB081FC72" box="[1090,1178,884,910]" pageId="18" pageNumber="19">KSEM</collectionCode>
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||
).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="04AA189FFFA96265CB29A0D4B5F9FBAA" pageId="18" pageNumber="19" type="diagnosis">
|
||
<paragraph id="4C0F4B14FFA96265CB29A0D4B5F9FBAA" blockId="18.[264,1324,723,1870]" pageId="18" pageNumber="19">
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<emphasis id="7EC49706FFA96265CB29A0D4B5ABFC49" bold="true" box="[312,432,923,949]" pageId="18" pageNumber="19">Diagnosis</emphasis>
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(based on the male
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||
<typeStatus id="930BF5B6FFA96265C8A8A0D3B73BFC4A" box="[697,800,924,950]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
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||
only):
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<taxonomicName id="8BB03097FFA96265C99CA0D3B068FC49" box="[909,1139,924,949]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tetramera">
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<emphasis id="7EC49706FFA96265C99CA0D3B068FC49" box="[909,1139,924,949]" italics="true" pageId="18" pageNumber="19">Microlia tetramera</emphasis>
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</taxonomicName>
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can be distinguished from other species of
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<taxonomicName id="8BB03097FFA96265C87EA08BB6CEFC21" box="[623,725,964,989]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="7EC49706FFA96265C87EA08BB6CEFC21" box="[623,725,964,989]" italics="true" pageId="18" pageNumber="19">Microlia</emphasis>
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</taxonomicName>
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by having mesotarsus with 4 segments, the lateral angles of male tergum 3 extended posteriorly as pointed triangular lobes (
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<figureCitation id="D48B5791FFA96265CE62A0A3B0D7FBFA" box="[1139,1228,1004,1030]" captionStart="FIGURES 68 76" captionStartId="20.[264,383,1277,1301]" captionTargetBox="[351,1226,513,1257]" captionTargetId="figure@20.[348,1227,513,1257]" captionTargetPageId="20" captionText="FIGURES 68 76. Details of Microlia tetramera Gusarov, sp. nov. (68 72, 75 76 – holotype; 73 74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73 74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73 74), 0.1 mm (69, 71 72, 75), 0.55 mm (76)." httpUri="https://zenodo.org/record/155791/files/figure.png" pageId="18" pageNumber="19">Fig. 76</figureCitation>
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), by the shape of the aedeagus (
|
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<figureCitation id="D48B5791FFA96265C832A75BB6ABFBD2" box="[547,688,1044,1070]" captionStart="FIGURES 68 76" captionStartId="20.[264,383,1277,1301]" captionTargetBox="[351,1226,513,1257]" captionTargetId="figure@20.[348,1227,513,1257]" captionTargetPageId="20" captionText="FIGURES 68 76. Details of Microlia tetramera Gusarov, sp. nov. (68 72, 75 76 – holotype; 73 74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73 74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73 74), 0.1 mm (69, 71 72, 75), 0.55 mm (76)." httpUri="https://zenodo.org/record/155791/files/figure.png" pageId="18" pageNumber="19">Figs. 6872</figureCitation>
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), particularly the hooklike apical process of median lobe (
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<figureCitation id="D48B5791FFA96265CB5AA773B5CEFBAA" box="[331,469,1084,1110]" captionStart="FIGURES 68 76" captionStartId="20.[264,383,1277,1301]" captionTargetBox="[351,1226,513,1257]" captionTargetId="figure@20.[348,1227,513,1257]" captionTargetPageId="20" captionText="FIGURES 68 76. Details of Microlia tetramera Gusarov, sp. nov. (68 72, 75 76 – holotype; 73 74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73 74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73 74), 0.1 mm (69, 71 72, 75), 0.55 mm (76)." httpUri="https://zenodo.org/record/155791/files/figure.png" pageId="18" pageNumber="19">Figs. 6869</figureCitation>
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||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04AA189FFFA96265CB29A72CB130F8B2" pageId="18" pageNumber="19" type="description">
|
||
<paragraph id="4C0F4B14FFA96265CB29A72CB6A4FB0A" blockId="18.[264,1324,723,1870]" pageId="18" pageNumber="19">
|
||
<emphasis id="7EC49706FFA96265CB29A72CB5D2FB81" bold="true" box="[312,457,1123,1149]" pageId="18" pageNumber="19">Description</emphasis>
|
||
(based on the male
|
||
<typeStatus id="930BF5B6FFA96265C8A3A72BB702FB82" box="[690,793,1124,1150]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
|
||
only): Length
|
||
<quantity id="8B48E6F1FFA96265C9DCA72BB030FB82" box="[973,1067,1124,1150]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="18" pageNumber="19" unit="mm" value="1.8">1.8 mm</quantity>
|
||
. Head, pronotum and abdominal tergum 3 brownish yellow; elytra brown, with lighter humeral angles; abdominal segments 47 dark brown; legs, mouthparts and seven basal antennal segments yellow; four apical antennal segments brown.
|
||
</paragraph>
|
||
<paragraph id="4C0F4B14FFA96265CB29A64BB10AFABA" blockId="18.[264,1324,723,1870]" pageId="18" pageNumber="19">Head surface mat, on disk with strong isodiametric microsculpture, puncturation fine, distance between punctures equal to their diameter. Eyes 2.8 times longer than temples.</paragraph>
|
||
<paragraph id="4C0F4B14FFA96265CB29A61BB640F9A2" blockId="18.[264,1324,723,1870]" pageId="18" pageNumber="19">
|
||
Pronotum strongly transverse, 1.4 times wider than head, width
|
||
<quantity id="8B48E6F1FFA96265CE3FA61BB081FA92" box="[1070,1178,1364,1390]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="mm" value="0.5">0.50 mm</quantity>
|
||
, length
|
||
<quantity id="8B48E6F1FFA96265CEE9A61BB522FA6A" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="mm" value="0.3">0.30 mm</quantity>
|
||
, width to length ratio 1.7; surface mat, with strong isodiametric microsculpture; puncturation as on head. Elytra wider (
|
||
<quantity id="8B48E6F1FFA96265C888A6EBB71FFA42" box="[665,772,1444,1470]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="mm" value="0.6">0.60 mm</quantity>
|
||
) and longer (
|
||
<quantity id="8B48E6F1FFA96265C98CA6EBB011FA42" box="[925,1034,1444,1470]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="18" pageNumber="19" unit="mm" value="0.47">0.47 mm</quantity>
|
||
, measured from humeral angle) than pronotum (pronotal length to elytral length ratio 0.64), 1.3 times wider than long, surface glossy, with weak transverse microsculpture, puncturation stronger than on pronotum and slightly asperate, distance between punctures equal to their diameter. Mesotarsus with 4 segments.
|
||
</paragraph>
|
||
<paragraph id="4C0F4B14FFA96265CB29A523B7B6F952" blockId="18.[264,1324,723,1870]" pageId="18" pageNumber="19">
|
||
Abdominal terga glossy, without visible (at
|
||
<date id="380E6DD4FFA96265C957A523B76FF97A" box="[838,884,1644,1670]" pageId="18" pageNumber="19" value="1970-10">x70</date>
|
||
) microsculpture, with fine puncturation, distance between punctures 23 times their diameter.
|
||
</paragraph>
|
||
<paragraph id="4C0F4B14FFA96265CB29A5F3B130F8B2" blockId="18.[264,1324,723,1870]" pageId="18" pageNumber="19">
|
||
Lateral angles of male tergum 3 extended posteriorly as pointed triangular lobes (
|
||
<figureCitation id="D48B5791FFA96265CEEDA5F3B532F902" captionStart="FIGURES 68 76" captionStartId="20.[264,383,1277,1301]" captionTargetBox="[351,1226,513,1257]" captionTargetId="figure@20.[348,1227,513,1257]" captionTargetPageId="20" captionText="FIGURES 68 76. Details of Microlia tetramera Gusarov, sp. nov. (68 72, 75 76 – holotype; 73 74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73 74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73 74), 0.1 mm (69, 71 72, 75), 0.55 mm (76)." httpUri="https://zenodo.org/record/155791/files/figure.png" pageId="18" pageNumber="19">Fig. 76</figureCitation>
|
||
). Male tergum 7 without tubercles. Posterior margin of male tergum 8 crenulate (
|
||
<figureCitation id="D48B5791FFA96265CEEDA5ABB532F8DA" captionStart="FIGURES 62 67" captionStartId="19.[264,383,1428,1452]" captionTargetBox="[360,1260,320,1405]" captionTargetId="figure@19.[354,1262,320,1409]" captionTargetPageId="19" captionText="FIGURES 62 67. Details of Microlia tetramera Gusarov, sp. nov. (62 63 – holotype; 64 67 – possible female of M. tetramera from Costa Rica, 64 65 – type A, 66 67 – type B). 62 – male sternum 8; 63 – male tergum 8; 64, 66 – female sternum 8; 65, 67 – female tergum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155790/files/figure.png" pageId="18" pageNumber="19">Fig. 63</figureCitation>
|
||
). Posterior margin of male sternum 8 with broad triangular lobe (
|
||
<figureCitation id="D48B5791FFA96265CE36A443B065F8DA" box="[1063,1150,1804,1830]" captionStart="FIGURES 62 67" captionStartId="19.[264,383,1428,1452]" captionTargetBox="[360,1260,320,1405]" captionTargetId="figure@19.[354,1262,320,1409]" captionTargetPageId="19" captionText="FIGURES 62 67. Details of Microlia tetramera Gusarov, sp. nov. (62 63 – holotype; 64 67 – possible female of M. tetramera from Costa Rica, 64 65 – type A, 66 67 – type B). 62 – male sternum 8; 63 – male tergum 8; 64, 66 – female sternum 8; 65, 67 – female tergum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/155790/files/figure.png" pageId="18" pageNumber="19">Fig. 62</figureCitation>
|
||
). Aedeagus as in
|
||
<figureCitation id="D48B5791FFA96265CB37A47BB5B4F8B2" box="[294,431,1844,1870]" captionStart="FIGURES 68 76" captionStartId="20.[264,383,1277,1301]" captionTargetBox="[351,1226,513,1257]" captionTargetId="figure@20.[348,1227,513,1257]" captionTargetPageId="20" captionText="FIGURES 68 76. Details of Microlia tetramera Gusarov, sp. nov. (68 72, 75 76 – holotype; 73 74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73 74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73 74), 0.1 mm (69, 71 72, 75), 0.55 mm (76)." httpUri="https://zenodo.org/record/155791/files/figure.png" pageId="18" pageNumber="19">Figs. 6872</figureCitation>
|
||
. Apical process of median lobe in lateral view bent like hook (
|
||
<figureCitation id="D48B5791FFA96265CE82A47BB105F8B2" box="[1171,1310,1844,1870]" captionStart="FIGURES 68 76" captionStartId="20.[264,383,1277,1301]" captionTargetBox="[351,1226,513,1257]" captionTargetId="figure@20.[348,1227,513,1257]" captionTargetPageId="20" captionText="FIGURES 68 76. Details of Microlia tetramera Gusarov, sp. nov. (68 72, 75 76 – holotype; 73 74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73 74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73 74), 0.1 mm (69, 71 72, 75), 0.55 mm (76)." httpUri="https://zenodo.org/record/155791/files/figure.png" pageId="18" pageNumber="19">Figs. 6869</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="18CF1B9CFFA86264CB19A6DBB11CFA0D" httpUri="https://zenodo.org/record/155790/files/figure.png" pageId="19" pageNumber="20" targetBox="[360,1260,320,1405]" targetPageId="19">
|
||
<paragraph id="4C0F4B14FFA86264CB19A6DBB11CFA0D" blockId="19.[264,1323,1428,1521]" pageId="19" pageNumber="20">
|
||
<emphasis id="7EC49706FFA86264CB19A6DBB5D8FA50" bold="true" box="[264,451,1428,1452]" pageId="19" pageNumber="20">FIGURES 6267</emphasis>
|
||
. Details of
|
||
<taxonomicName id="8BB03097FFA86264C854A6DAB768FA57" authority="Gusarov" authorityName="Gusarov" box="[581,883,1428,1452]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tetramera" status="sp. nov.">
|
||
<emphasis id="7EC49706FFA86264C854A6DAB70AFA50" box="[581,785,1429,1452]" italics="true" pageId="19" pageNumber="20">Microlia tetramera</emphasis>
|
||
Gusarov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="65F72A7DFFA86264C96CA6DBB7C8FA50" box="[893,979,1428,1452]" pageId="19" pageNumber="20" rank="species">
|
||
<emphasis id="7EC49706FFA86264C96CA6DBB7C8FA50" bold="true" box="[893,979,1428,1452]" pageId="19" pageNumber="20">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(6263 – holotype; 6467 – possible female of
|
||
<taxonomicName id="8BB03097FFA86264CBBCA6F8B622FA32" box="[429,569,1463,1486]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tetramera">
|
||
<emphasis id="7EC49706FFA86264CBBCA6F8B622FA32" box="[429,569,1463,1486]" italics="true" pageId="19" pageNumber="20">M. tetramera</emphasis>
|
||
</taxonomicName>
|
||
from Costa Rica, 6465 – type A, 6667 – type B). 62 – male sternum 8; 63 – male tergum 8; 64, 66 – female sternum 8; 65, 67 – female tergum 8. Scale bar 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="04AA189FFFA86263CB29A57BB7B8FEA2" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="discussion">
|
||
<paragraph id="4C0F4B14FFA86264CB29A57BB130F89A" blockId="19.[264,1323,1588,1895]" pageId="19" pageNumber="20">
|
||
<emphasis id="7EC49706FFA86264CB29A57BB5DDF9B2" bold="true" box="[312,454,1588,1614]" pageId="19" pageNumber="20">Discussion:</emphasis>
|
||
Two
|
||
<typeStatus id="930BF5B6FFA86264C80CA57AB640F9B3" box="[541,603,1589,1615]" pageId="19" pageNumber="20">types</typeStatus>
|
||
of females, apparently representing different species, both with tarsal formula 445, have been collected at the
|
||
<typeStatus id="930BF5B6FFA86264C996A512B7A1F98B" box="[903,954,1629,1655]" pageId="19" pageNumber="20">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="8BB03097FFA86264CE57A512B0F8F98A" box="[1094,1251,1629,1654]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tetramera">
|
||
<emphasis id="7EC49706FFA86264CE57A512B0F8F98A" box="[1094,1251,1629,1654]" italics="true" pageId="19" pageNumber="20">M. tetramera</emphasis>
|
||
</taxonomicName>
|
||
. The females of the
|
||
<typeStatus id="930BF5B6FFA86264CBAAA5CAB5F5F963" box="[443,494,1669,1695]" pageId="19" pageNumber="20">type</typeStatus>
|
||
A have thicker spermatheca with two coils and pointed capsule (
|
||
<figureCitation id="D48B5791FFA86264CEEAA5CAB53DF93B" captionStart="FIGURES 68 76" captionStartId="20.[264,383,1277,1301]" captionTargetBox="[351,1226,513,1257]" captionTargetId="figure@20.[348,1227,513,1257]" captionTargetPageId="20" captionText="FIGURES 68 76. Details of Microlia tetramera Gusarov, sp. nov. (68 72, 75 76 – holotype; 73 74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73 74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73 74), 0.1 mm (69, 71 72, 75), 0.55 mm (76)." httpUri="https://zenodo.org/record/155791/files/figure.png" pageId="19" pageNumber="20">Fig. 73</figureCitation>
|
||
) while in the
|
||
<typeStatus id="930BF5B6FFA86264CBDAA5E2B5E5F93B" box="[459,510,1709,1735]" pageId="19" pageNumber="20">type</typeStatus>
|
||
B females the spermatheca has about five coils and rounded capsule (
|
||
<figureCitation id="D48B5791FFA86264CB00A59AB571F913" box="[273,362,1749,1775]" captionStart="FIGURES 68 76" captionStartId="20.[264,383,1277,1301]" captionTargetBox="[351,1226,513,1257]" captionTargetId="figure@20.[348,1227,513,1257]" captionTargetPageId="20" captionText="FIGURES 68 76. Details of Microlia tetramera Gusarov, sp. nov. (68 72, 75 76 – holotype; 73 74 – possible female of M. tetramera from Costa Rica). 68 – median lobe of aedeagus, lateral view; 69 – apex of median lobe of aedeagus, lateral view; 70 – median lobe of aedeagus, ventral view; 71 – apex of median lobe of aedeagus, ventral view; 72 – apex of median lobe of aedeagus, apical ventral view; 73 74 – spermatheca; 75 – apical lobe of right paramere, internal view; 76 – abdominal segment 3, dorsal view. Scale bar 0.2 mm (68, 70, 73 74), 0.1 mm (69, 71 72, 75), 0.55 mm (76)." httpUri="https://zenodo.org/record/155791/files/figure.png" pageId="19" pageNumber="20">Fig. 74</figureCitation>
|
||
). The male
|
||
<typeStatus id="930BF5B6FFA86264CBEBA59AB67AF913" box="[506,609,1749,1775]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="8BB03097FFA86264C897A59AB73BF912" box="[646,800,1749,1774]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tetramera">
|
||
<emphasis id="7EC49706FFA86264C897A59AB73BF912" box="[646,800,1749,1774]" italics="true" pageId="19" pageNumber="20">M. tetramera</emphasis>
|
||
</taxonomicName>
|
||
differs from both
|
||
<typeStatus id="930BF5B6FFA86264C9E7A59AB02FF913" box="[1014,1076,1749,1775]" pageId="19" pageNumber="20">types</typeStatus>
|
||
of females in having darker coloration, stronger microsculpture of head and pronotum. It is impossible to reliably associate the male
|
||
<typeStatus id="930BF5B6FFA86264C80DA46AB698F8C3" box="[540,643,1829,1855]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
with one of the two
|
||
<typeStatus id="930BF5B6FFA86264C963A46AB7ABF8C3" box="[882,944,1829,1855]" pageId="19" pageNumber="20">types</typeStatus>
|
||
of females without having additional material. An alternative hypothesis, that the
|
||
<typeStatus id="930BF5B6FFA86264C970A402B788F89B" box="[865,915,1869,1895]" pageId="19" pageNumber="20">type</typeStatus>
|
||
B females are conspecific with
|
||
<emphasis id="7EC49706FFA86264CF1AA402B130F89A" box="[1291,1323,1869,1894]" italics="true" pageId="19" pageNumber="20">M.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4C0F4B14FFAF6263CB19A253B7B8FEA2" blockId="20.[264,1323,284,430]" pageId="20" pageNumber="21">
|
||
<taxonomicName id="8BB03097FFAF6263CB19A253B59EFEC9" box="[264,389,284,309]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pentamera">
|
||
<emphasis id="7EC49706FFAF6263CB19A253B59EFEC9" box="[264,389,284,309]" italics="true" pageId="20" pageNumber="21">pentamera</emphasis>
|
||
</taxonomicName>
|
||
(tarsal formula 455), seems implausible because no intraspecific variation of tarsal formula was observed in other species of
|
||
<taxonomicName id="8BB03097FFAF6263C926A20BB787FEA1" box="[823,924,324,349]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7EC49706FFAF6263C926A20BB787FEA1" box="[823,924,324,349]" italics="true" pageId="20" pageNumber="21">Microlia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04AA189FFFAF6263CB29A224B06CFE52" pageId="20" pageNumber="21" type="distribution">
|
||
<paragraph id="4C0F4B14FFAF6263CB29A224B047FE7A" blockId="20.[264,1323,284,430]" box="[312,1116,363,390]" pageId="20" pageNumber="21">
|
||
<emphasis id="7EC49706FFAF6263CB29A224B5C0FE79" bold="true" box="[312,475,363,389]" pageId="20" pageNumber="21">Distribution:</emphasis>
|
||
Known from a single locality in
|
||
<collectingCountry id="34A70B84FFAF6263C977A223B7FCFE7A" box="[870,999,364,390]" name="Costa Rica" pageId="20" pageNumber="21">Costa Rica</collectingCountry>
|
||
(
|
||
<figureCitation id="D48B5791FFAF6263C9E7A223B055FE7A" box="[1014,1102,364,390]" captionStart="FIGURE 80" captionStartId="22.[264,368,1184,1208]" captionTargetBox="[360,1265,358,1173]" captionTargetId="figure@22.[358,1266,353,1174]" captionTargetPageId="22" captionText="FIGURE 80. Geographical distribution of Microlia Casey. 1 – M. silacea (Erichson); 2 – M. meticola (Casey); 3 – M. azteca (Casey); 4 – M. pentamera Gusarov, sp. nov.; 5 – M. tetramera Gusarov, sp. nov.; 6 – M. panamensis Gusarov, sp. nov." httpUri="https://zenodo.org/record/155793/files/figure.png" pageId="20" pageNumber="21">Fig. 80</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4C0F4B14FFAF6263CB29A2DCB06CFE52" blockId="20.[264,1323,284,430]" box="[312,1143,403,430]" pageId="20" pageNumber="21">
|
||
<emphasis id="7EC49706FFAF6263CB29A2DCB612FE51" bold="true" box="[312,521,403,429]" pageId="20" pageNumber="21">Natural History:</emphasis>
|
||
<taxonomicName id="8BB03097FFAF6263C809A2DBB6E3FE51" box="[536,760,404,429]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tetramera">
|
||
<emphasis id="7EC49706FFAF6263C809A2DBB6E3FE51" box="[536,760,404,429]" italics="true" pageId="20" pageNumber="21">Microlia tetramera</emphasis>
|
||
</taxonomicName>
|
||
was collected with malaise trap.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |