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<document id="3F338E038F1A9935D5673353002CBFDA" ID-CLB-Dataset="56252" ID-DOI="10.11646/zootaxa.5318.1.1" ID-GBIF-Dataset="955a5700-d953-4bad-89c4-6767df6c519f" ID-ISSN="1175-5326" ID-Zenodo-Dep="8158079" ID-ZooBank="DB9D6B30-5A8C-486A-BF67-0965E4C026DA" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1689662800392" checkinUser="plazi" docAuthor="Ng, Peter K. L., Forges, Bertrand Richer De &amp; Lee, Bee Yan" docDate="2023" docId="9B094F56FA45FFEAFF20C54AFEA2F87B" docLanguage="en" docName="zootaxa.5318.1.1.pdf" docOrigin="Zootaxa 5318 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5318.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Achaeus pholcus Ng &amp; Forges &amp; Lee 2023, n. sp." docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="6730372EFA48FFFAFFB7C401FF98FFA5" masterDocTitle="New and rare deep-sea majoid crabs (Crustacea: Brachyura) from Papua New Guinea" masterLastPageNumber="44" masterPageNumber="1" pageNumber="14" updateTime="1699541154989" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EAE24C43D381FCFCBE9E887836EC3818">New and rare deep-sea majoid crabs (Crustacea: Brachyura) from Papua New Guinea</mods:title>
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<mods:namePart id="2563E66AEE9423AC093A4DEA65D9751E">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="DB79BE0161D7513F3A6768EB377DDEB4">Forges, Bertrand Richer De</mods:namePart>
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<mods:namePart id="E780AE4E359EB867C19954EB1C3949E6">Lee, Bee Yan</mods:namePart>
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<mods:date id="0E26FB3B61F67FDBF4D0C67BF500EF69">2023</mods:date>
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<treatment id="9B094F56FA45FFEAFF20C54AFEA2F87B" ID-DOI="http://doi.org/10.5281/zenodo.8158109" ID-GBIF-Taxon="212415795" ID-Zenodo-Dep="8158109" LSID="urn:lsid:plazi:treatment:9B094F56FA45FFEAFF20C54AFEA2F87B" httpUri="http://treatment.plazi.org/id/9B094F56FA45FFEAFF20C54AFEA2F87B" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
<subSubSection id="5BBAADCBFA45FFF7FF20C54AFE37FEC0" box="[151,431,331,358]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="131FFE40FA45FFF7FF20C54AFE37FEC0" blockId="13.[151,431,331,393]" box="[151,431,331,358]" pageId="13" pageNumber="14">
<heading id="4857492CFA45FFF7FF20C54AFE37FEC0" bold="true" box="[151,431,331,358]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="21D42252FA45FFF7FF20C54AFE37FEC0" bold="true" box="[151,431,331,358]" pageId="13" pageNumber="14">
<taxonomicName id="D4A085C3FA45FFF7FF20C54AFEFEFEC3" ID-CoL="d26182f4-6d8d-4124-901b-eede81b2481f" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[151,358,331,358]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA45FFF7FF20C54AFEFEFEC3" bold="true" box="[151,358,331,358]" italics="true" pageId="13" pageNumber="14">Achaeus pholcus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AE79F29FA45FFF7FEDAC54AFE37FEC0" box="[365,431,331,357]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="5BBAADCBFA45FFF7FF20C56EFECAFE2C" box="[151,338,367,393]" pageId="13" pageNumber="14" type="description">
<paragraph id="131FFE40FA45FFF7FF20C56EFECAFE2C" blockId="13.[151,431,331,393]" box="[151,338,367,393]" pageId="13" pageNumber="14">
(
<figureCitation id="8B9BE2C5FA45FFF7FF28C56EFF6EFE2C" box="[159,246,367,393]" captionStart="FIGURE 1" captionStartId="3.[151,250,1772,1796]" captionTargetBox="[213,1375,179,1746]" captionTargetId="figure-18@3.[213,1375,179,1746]" captionTargetPageId="3" captionText="FIGURE 1. Colours in life. A, B, Dorhynchus profundus n. sp.. A, paratype male (pcl 10.3 mm) (ZRC 2022.0067), Budibudi Island; B, male (not examined), station CP3650, south of New Hanover Island, Papua New Guinea; C, Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland; D, Parapleisticantha aie n. sp., paratype ovigerous female (pcl 15.4 mm) (ZRC 2022.0073), New Ireland; E, Grypachaeus aff. tenuicollis Takeda, 1978, female (not examined) (MNHN- IU-2015-85), New Ireland; F, Grypachaeus aff. tenuicollis Takeda, 1978, ovigerous female (pcl 10.4 mm) (ZRC 2012.1160), Vanuatu; G, Orbicantha sacrebleu n. gen., n. sp., holotype female (pcl 7.2 mm) (MNHN-IU-2014-8158), New Ireland; H, Orbicantha sacrebleu n. gen., n. sp., female (pcl 6.7 mm) (ZRC 2022.0083), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.8158081" httpUri="https://zenodo.org/record/8158081/files/figure.png" pageId="13" pageNumber="14">Fig. 1C</figureCitation>
,
<figureCitation id="8B9BE2C5FA45FFF7FEB4C56EFED2FE2C" box="[259,330,367,393]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStartId-0="14.[151,250,1914,1938]" captionStartId-1="15.[151,250,1903,1927]" captionStartId-2="16.[151,250,1167,1191]" captionStartId-3="17.[151,250,1721,1745]" captionTargetBox-0="[152,1436,181,1890]" captionTargetBox-1="[151,1436,181,1879]" captionTargetBox-2="[151,1436,181,1142]" captionTargetBox-3="[153,1434,183,1694]" captionTargetId-0="figure-27@14.[152,1436,181,1890]" captionTargetId-1="figure-18@15.[151,1436,181,1879]" captionTargetId-2="figure-27@16.[151,1436,181,1143]" captionTargetId-3="figure-18@17.[151,1436,181,1697]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionText-0="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." captionText-1="FIGURE 11. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, subventral view of cephalothorax showing thoracic sternum, pleon, buccal cavity, epistome, antennae, antennules and eyes; third maxillipeds; B, ventral view of cephalothorax showing thoracic sternum, pleon and third maxillipeds; C, thoracic sternal crest on sternite 4; D, pleonal somites 3–5 and pleotelson; E, pleonal somites 1 and 2." captionText-2="FIGURE 12. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, dorsal view of right cheliped; B, right chela and carpus; C, left third maxilliped; D, E, right G1 (ventral views); F, right G1 (dorsal view); G, left G2. G1s broken medially and connected by photoshop." captionText-3="FIGURE 13. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, dorsal view of anterior part of carapace (setae not drawn); B, left third maxilliped (setae not drawn); C, left P2; D, right P3 (carpus to dactylus missing); E, left P4; F, left P4 dactylus; G, left P5; H, left P5 dactylus; I, right G1 (dorsal view) (medially broken); J, right G1 (ventral view) (medially broken); K, L, distal part of right G1 (dorsal view, slightly different angles); M, distal part of right G1 (mesial view); N, distal part of right G1 (ventral view); O, left G2. Scales: A, B, I, J, O = 0.5 mm; C–H = 1.0 mm; K, L, M, N = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8158107" figureDoi-1="http://doi.org/10.5281/zenodo.8158111" figureDoi-2="http://doi.org/10.5281/zenodo.8158113" figureDoi-3="http://doi.org/10.5281/zenodo.8158115" httpUri-0="https://zenodo.org/record/8158107/files/figure.png" httpUri-1="https://zenodo.org/record/8158111/files/figure.png" httpUri-2="https://zenodo.org/record/8158113/files/figure.png" httpUri-3="https://zenodo.org/record/8158115/files/figure.png" pageId="13" pageNumber="14">10–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5BBAADCBFA45FFF7FF20C5B6FF4FFDBC" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="131FFE40FA45FFF7FF20C5B6FF4FFDBC" blockId="13.[151,1437,439,2014]" pageId="13" pageNumber="14">
<emphasis id="21D42252FA45FFF7FF20C5B6FEDEFE74" bold="true" box="[151,326,439,465]" pageId="13" pageNumber="14">Type material.</emphasis>
<materialsCitation id="A3C8F41DFA45FFF7FEF9C5B6FF4FFDBC" ID-GBIF-Occurrence="4161833321" collectingDate="2014-08-28" collectorName="N. O. Alis" country="Papua New Guinea" elevation="600" latitude="-2.3333333" location="New Ireland" longLatPrecision="1307" longitude="150.63333" pageId="13" pageNumber="14" specimenCount="1" stateProvince="New Ireland" typeStatus="holotype">
<typeStatus id="CC1B40E2FA45FFF7FEF9C5B6FE23FE74" box="[334,443,439,465]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
: male (cl
<quantity id="D45853A5FA45FFF7FD9EC5B6FD1AFE77" box="[553,642,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="14" unit="mm" value="8.1">8.1 mm</quantity>
, pcl
<quantity id="D45853A5FA45FFF7FD0EC5B6FC8AFE77" box="[697,786,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="13" pageNumber="14" unit="mm" value="7.3">7.3 mm</quantity>
, cw
<quantity id="D45853A5FA45FFF7FCF1C5B6FC06FE74" box="[838,926,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="13" pageNumber="14" unit="mm" value="5.8">5.8 mm</quantity>
) (MNHN-IU-2014-8034), stn CP 4423,
<collectingRegion id="D16430A2FA45FFF7FAD0C5B9FF71FE50" country="Papua New Guinea" name="New Ireland" pageId="13" pageNumber="14">New Ireland</collectingRegion>
,
<collectingCountry id="6BB7BED0FA45FFF7FF44C5DDFE54FE50" box="[243,460,475,502]" name="Papua New Guinea" pageId="13" pageNumber="14">Papua New Guinea</collectingCountry>
,
<geoCoordinate id="76949887FA45FFF7FE61C5DAFDAAFE50" box="[470,562,475,501]" degrees="02" direction="south" minutes="20" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="-2.3333333">02°20’S</geoCoordinate>
<geoCoordinate id="76949887FA45FFF7FD8FC5DAFD3EFE53" box="[568,678,475,502]" degrees="150" direction="east" minutes="38" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="150.63333">150°38’E</geoCoordinate>
,
<quantity id="D45853A5FA45FFF7FD06C5DAFCA8FE53" box="[689,816,475,502]" metricMagnitude="2" metricUnit="m" metricValue="5.995" metricValueMax="6.49" metricValueMin="5.5" pageId="13" pageNumber="14" unit="m" value="599.5" valueMax="649.0" valueMin="550.0">
<elevation id="988D1973FA45FFF7FD06C5DAFCA8FE53" box="[689,816,475,502]" metricMagnitude="2" metricUnit="m" metricValue="5.995" metricValueMax="6.49" metricValueMin="5.5" pageId="13" pageNumber="14" unit="m" value="599.5" valueMax="649.0" valueMin="550.0">550–649 m</elevation>
</quantity>
, coll.
<collectorName id="BE559B96FA45FFF7FCC5C5DAFC46FE50" box="[882,990,475,501]" pageId="13" pageNumber="14">N.O. Alis</collectorName>
, KAVIENG 2014 expedition,
<date id="671ED880FA45FFF7FA9CC5DAFF4BFDBC" pageId="13" pageNumber="14" value="2014-08-28">
<collectingDate id="775A2168FA45FFF7FA9CC5DAFF4BFDBC" pageId="13" pageNumber="14" value="2014-08-28">28 August 2014</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5BBAADCBFA45FFF7FF70C622FC38FA94" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="131FFE40FA45FFF7FF70C622FB52FAA8" blockId="13.[151,1437,439,2014]" pageId="13" pageNumber="14">
<emphasis id="21D42252FA45FFF7FF70C622FEDAFD98" bold="true" box="[199,322,547,573]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
Carapace subpyriform, regions clearly demarcated; branchial and gastric regions appear almost globular, wide and long (
<figureCitation id="8B9BE2C5FA45FFF7FE01C646FDD8FDC4" box="[438,576,583,609]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10A, B</figureCitation>
); frontal and epigastric regions short, not elongate (
<figureCitation id="8B9BE2C5FA45FFF7FB27C646FAA3FDC4" box="[1168,1339,583,609]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10B, C, E</figureCitation>
); gastric regions not elongate, gently convex, without spine or sharp tubercles (
<figureCitation id="8B9BE2C5FA45FFF7FC18C66AFBCEFD20" box="[943,1110,619,645]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10B, C, E</figureCitation>
); cardiac region prominently swollen with sharp tubercle (
<figureCitation id="8B9BE2C5FA45FFF7FE50C68EFD0AFD0C" box="[487,658,655,682]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10B, E, F</figureCitation>
); intestinal region unarmed (
<figureCitation id="8B9BE2C5FA45FFF7FC57C68EFBF2FD0C" box="[992,1130,655,681]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10B, E</figureCitation>
); pseudorostrum relatively short, with 2 subtruncate lobes, separated by deep U-shaped cleft, margin of each lobe with 3 low tubercles (
<figureCitation id="8B9BE2C5FA45FFF7FAD3C6B2FF72FD54" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Figs. 10B–D</figureCitation>
,
<figureCitation id="8B9BE2C5FA45FFF7FF42C6D6FEB1FD54" box="[245,297,727,753]" captionStart="FIGURE 13" captionStartId="17.[151,250,1721,1745]" captionTargetBox="[153,1434,183,1694]" captionTargetId="figure-18@17.[151,1436,181,1697]" captionTargetPageId="17" captionText="FIGURE 13. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, dorsal view of anterior part of carapace (setae not drawn); B, left third maxilliped (setae not drawn); C, left P2; D, right P3 (carpus to dactylus missing); E, left P4; F, left P4 dactylus; G, left P5; H, left P5 dactylus; I, right G1 (dorsal view) (medially broken); J, right G1 (ventral view) (medially broken); K, L, distal part of right G1 (dorsal view, slightly different angles); M, distal part of right G1 (mesial view); N, distal part of right G1 (ventral view); O, left G2. Scales: A, B, I, J, O = 0.5 mm; C–H = 1.0 mm; K, L, M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8158115" httpUri="https://zenodo.org/record/8158115/files/figure.png" pageId="13" pageNumber="14">13A</figureCitation>
); supraorbital eave lined with small granules, without spines, antorbital angle visible but not dentiform or spiniform (
<figureCitation id="8B9BE2C5FA45FFF7FE98C6FAFE26FCB0" box="[303,446,763,789]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Figs. 10B, C</figureCitation>
,
<figureCitation id="8B9BE2C5FA45FFF7FE7FC6FAFE65FCB0" box="[456,509,763,789]" captionStart="FIGURE 13" captionStartId="17.[151,250,1721,1745]" captionTargetBox="[153,1434,183,1694]" captionTargetId="figure-18@17.[151,1436,181,1697]" captionTargetPageId="17" captionText="FIGURE 13. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, dorsal view of anterior part of carapace (setae not drawn); B, left third maxilliped (setae not drawn); C, left P2; D, right P3 (carpus to dactylus missing); E, left P4; F, left P4 dactylus; G, left P5; H, left P5 dactylus; I, right G1 (dorsal view) (medially broken); J, right G1 (ventral view) (medially broken); K, L, distal part of right G1 (dorsal view, slightly different angles); M, distal part of right G1 (mesial view); N, distal part of right G1 (ventral view); O, left G2. Scales: A, B, I, J, O = 0.5 mm; C–H = 1.0 mm; K, L, M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8158115" httpUri="https://zenodo.org/record/8158115/files/figure.png" pageId="13" pageNumber="14">13A</figureCitation>
); hepatic region prominently swollen, round (
<figureCitation id="8B9BE2C5FA45FFF7FC41C6FAFB2DFCB0" box="[1014,1205,763,790]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10B, C, E, F</figureCitation>
); surface and margin of carapace with numerous small rounded granules (
<figureCitation id="8B9BE2C5FA45FFF7FD52C71EFC14FC9C" box="[741,908,799,825]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10B, C, E</figureCitation>
); epistome quadrate (
<figureCitation id="8B9BE2C5FA45FFF7FBC8C71EFB70FC9C" box="[1151,1256,799,825]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10D</figureCitation>
); basal antennal article long, slender, distal margin with 3 longer sharp spines, outer and inner margins each with row of low spines, antennal article 3 two-thirds length of basal antennal article, antennal article 4 ca. half length of article 3; antennular fossa long, basal antennular article with sharp anterior spine (
<figureCitation id="8B9BE2C5FA45FFF7FCF3C78AFC0CFC00" box="[836,916,907,933]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
BC, D); interantennular spine large, triangular, lobiform (
<figureCitation id="8B9BE2C5FA45FFF7FEBEC7AEFE0BFC6C" box="[265,403,943,969]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10D, E</figureCitation>
). Inner margin of third maxilliped merus with 2 sharp spines, ischium with 2 rows of sharp tubercles (
<figureCitation id="8B9BE2C5FA45FFF7FEBAC7D2FEE7FC48" box="[269,383,979,1005]" captionStart="FIGURE 12" captionStartId="16.[151,250,1167,1191]" captionTargetBox="[151,1436,181,1142]" captionTargetId="figure-27@16.[151,1436,181,1143]" captionTargetPageId="16" captionText="FIGURE 12. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, dorsal view of right cheliped; B, right chela and carpus; C, left third maxilliped; D, E, right G1 (ventral views); F, right G1 (dorsal view); G, left G2. G1s broken medially and connected by photoshop." figureDoi="http://doi.org/10.5281/zenodo.8158113" httpUri="https://zenodo.org/record/8158113/files/figure.png" pageId="13" pageNumber="14">Figs. 12C</figureCitation>
,
<figureCitation id="8B9BE2C5FA45FFF7FE3AC7D2FE27FC48" box="[397,447,979,1005]" captionStart="FIGURE 13" captionStartId="17.[151,250,1721,1745]" captionTargetBox="[153,1434,183,1694]" captionTargetId="figure-18@17.[151,1436,181,1697]" captionTargetPageId="17" captionText="FIGURE 13. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, dorsal view of anterior part of carapace (setae not drawn); B, left third maxilliped (setae not drawn); C, left P2; D, right P3 (carpus to dactylus missing); E, left P4; F, left P4 dactylus; G, left P5; H, left P5 dactylus; I, right G1 (dorsal view) (medially broken); J, right G1 (ventral view) (medially broken); K, L, distal part of right G1 (dorsal view, slightly different angles); M, distal part of right G1 (mesial view); N, distal part of right G1 (ventral view); O, left G2. Scales: A, B, I, J, O = 0.5 mm; C–H = 1.0 mm; K, L, M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8158115" httpUri="https://zenodo.org/record/8158115/files/figure.png" pageId="13" pageNumber="14">13B</figureCitation>
). Chelae slender, dorsal and ventral margins with short spines, fingers as long as palm (
<figureCitation id="8B9BE2C5FA45FFF7FF28C7F6FEB0FBB4" box="[159,296,1015,1041]" captionStart="FIGURE 12" captionStartId="16.[151,250,1167,1191]" captionTargetBox="[151,1436,181,1142]" captionTargetId="figure-27@16.[151,1436,181,1143]" captionTargetPageId="16" captionText="FIGURE 12. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, dorsal view of right cheliped; B, right chela and carpus; C, left third maxilliped; D, E, right G1 (ventral views); F, right G1 (dorsal view); G, left G2. G1s broken medially and connected by photoshop." figureDoi="http://doi.org/10.5281/zenodo.8158113" httpUri="https://zenodo.org/record/8158113/files/figure.png" pageId="13" pageNumber="14">Fig. 12A, B</figureCitation>
). P2–P5 long, slender, merus, carpus and propodus unarmed, P2 merus 7.8 times pcl, dactylus gently curved, without subterminal spine, ventral margin with evenly spaced 10 or 11 spinules (
<figureCitation id="8B9BE2C5FA45FFF7FBCBC01AFB76FB90" box="[1148,1262,1051,1077]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Figs. 10A</figureCitation>
,
<figureCitation id="8B9BE2C5FA45FFF7FB4DC01AFAD7FB90" box="[1274,1359,1051,1077]" captionStart="FIGURE 13" captionStartId="17.[151,250,1721,1745]" captionTargetBox="[153,1434,183,1694]" captionTargetId="figure-18@17.[151,1436,181,1697]" captionTargetPageId="17" captionText="FIGURE 13. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, dorsal view of anterior part of carapace (setae not drawn); B, left third maxilliped (setae not drawn); C, left P2; D, right P3 (carpus to dactylus missing); E, left P4; F, left P4 dactylus; G, left P5; H, left P5 dactylus; I, right G1 (dorsal view) (medially broken); J, right G1 (ventral view) (medially broken); K, L, distal part of right G1 (dorsal view, slightly different angles); M, distal part of right G1 (mesial view); N, distal part of right G1 (ventral view); O, left G2. Scales: A, B, I, J, O = 0.5 mm; C–H = 1.0 mm; K, L, M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8158115" httpUri="https://zenodo.org/record/8158115/files/figure.png" pageId="13" pageNumber="14">13C–H</figureCitation>
). Male thoracic sternites 3 and 4 fused; sternite 3 with 2 median depressions; sternite 4 arched ventrally to form median transverse convex crest anterior to sternopleonal cavity, crest medially clefted, serrate (
<figureCitation id="8B9BE2C5FA45FFF7FBCDC062FA9EFBD8" box="[1146,1286,1123,1149]" captionStart="FIGURE 11" captionStartId="15.[151,250,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-18@15.[151,1436,181,1879]" captionTargetPageId="15" captionText="FIGURE 11. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, subventral view of cephalothorax showing thoracic sternum, pleon, buccal cavity, epistome, antennae, antennules and eyes; third maxillipeds; B, ventral view of cephalothorax showing thoracic sternum, pleon and third maxillipeds; C, thoracic sternal crest on sternite 4; D, pleonal somites 3–5 and pleotelson; E, pleonal somites 1 and 2." figureDoi="http://doi.org/10.5281/zenodo.8158111" httpUri="https://zenodo.org/record/8158111/files/figure.png" pageId="13" pageNumber="14">Fig. 11A, C</figureCitation>
). Male pleon short, triangular; outer surfaces and margins of somites 1–4 with sharp granules; somite 4 trapezoidal; somite 5 subrectangular, as wide as pleotelson; somite 6 and telson completely fused with only lateral suture just visible, median part swollen to form subrectangular convexity (
<figureCitation id="8B9BE2C5FA45FFF7FCBAC0CEFC20FB4C" box="[781,952,1231,1258]" captionStart="FIGURE 11" captionStartId="15.[151,250,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-18@15.[151,1436,181,1879]" captionTargetPageId="15" captionText="FIGURE 11. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, subventral view of cephalothorax showing thoracic sternum, pleon, buccal cavity, epistome, antennae, antennules and eyes; third maxillipeds; B, ventral view of cephalothorax showing thoracic sternum, pleon and third maxillipeds; C, thoracic sternal crest on sternite 4; D, pleonal somites 3–5 and pleotelson; E, pleonal somites 1 and 2." figureDoi="http://doi.org/10.5281/zenodo.8158111" httpUri="https://zenodo.org/record/8158111/files/figure.png" pageId="13" pageNumber="14">Fig. 11B, D, E</figureCitation>
). G1 slender, broadly C-shaped, distal part with broadly rounded tip, small dorsal fold marking subterminal aperture (
<figureCitation id="8B9BE2C5FA45FFF7FC6EC0F2FBFEFAA8" box="[985,1126,1267,1293]" captionStart="FIGURE 12" captionStartId="16.[151,250,1167,1191]" captionTargetBox="[151,1436,181,1142]" captionTargetId="figure-27@16.[151,1436,181,1143]" captionTargetPageId="16" captionText="FIGURE 12. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, dorsal view of right cheliped; B, right chela and carpus; C, left third maxilliped; D, E, right G1 (ventral views); F, right G1 (dorsal view); G, left G2. G1s broken medially and connected by photoshop." figureDoi="http://doi.org/10.5281/zenodo.8158113" httpUri="https://zenodo.org/record/8158113/files/figure.png" pageId="13" pageNumber="14">Figs. 12D–F</figureCitation>
,
<figureCitation id="8B9BE2C5FA45FFF7FBC6C0F2FB25FAA8" box="[1137,1213,1267,1293]" captionStart="FIGURE 13" captionStartId="17.[151,250,1721,1745]" captionTargetBox="[153,1434,183,1694]" captionTargetId="figure-18@17.[151,1436,181,1697]" captionTargetPageId="17" captionText="FIGURE 13. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, dorsal view of anterior part of carapace (setae not drawn); B, left third maxilliped (setae not drawn); C, left P2; D, right P3 (carpus to dactylus missing); E, left P4; F, left P4 dactylus; G, left P5; H, left P5 dactylus; I, right G1 (dorsal view) (medially broken); J, right G1 (ventral view) (medially broken); K, L, distal part of right G1 (dorsal view, slightly different angles); M, distal part of right G1 (mesial view); N, distal part of right G1 (ventral view); O, left G2. Scales: A, B, I, J, O = 0.5 mm; C–H = 1.0 mm; K, L, M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8158115" httpUri="https://zenodo.org/record/8158115/files/figure.png" pageId="13" pageNumber="14">13I–N</figureCitation>
).
</paragraph>
<paragraph id="131FFE40FA45FFF7FF70C116FC38FA94" blockId="13.[151,1437,439,2014]" box="[199,928,1303,1330]" pageId="13" pageNumber="14">
<emphasis id="21D42252FA45FFF7FF70C116FEB8FA94" bold="true" box="[199,288,1303,1329]" pageId="13" pageNumber="14">Colour.</emphasis>
The overall colour is yellowish-white to white (
<figureCitation id="8B9BE2C5FA45FFF7FC8CC116FC0BFA94" box="[827,915,1303,1329]" captionStart="FIGURE 1" captionStartId="3.[151,250,1772,1796]" captionTargetBox="[213,1375,179,1746]" captionTargetId="figure-18@3.[213,1375,179,1746]" captionTargetPageId="3" captionText="FIGURE 1. Colours in life. A, B, Dorhynchus profundus n. sp.. A, paratype male (pcl 10.3 mm) (ZRC 2022.0067), Budibudi Island; B, male (not examined), station CP3650, south of New Hanover Island, Papua New Guinea; C, Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland; D, Parapleisticantha aie n. sp., paratype ovigerous female (pcl 15.4 mm) (ZRC 2022.0073), New Ireland; E, Grypachaeus aff. tenuicollis Takeda, 1978, female (not examined) (MNHN- IU-2015-85), New Ireland; F, Grypachaeus aff. tenuicollis Takeda, 1978, ovigerous female (pcl 10.4 mm) (ZRC 2012.1160), Vanuatu; G, Orbicantha sacrebleu n. gen., n. sp., holotype female (pcl 7.2 mm) (MNHN-IU-2014-8158), New Ireland; H, Orbicantha sacrebleu n. gen., n. sp., female (pcl 6.7 mm) (ZRC 2022.0083), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.8158081" httpUri="https://zenodo.org/record/8158081/files/figure.png" pageId="13" pageNumber="14">Fig. 1C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5BBAADCBFA45FFF7FF70C13AFB76FADF" pageId="13" pageNumber="14" type="etymology">
<paragraph id="131FFE40FA45FFF7FF70C13AFB76FADF" blockId="13.[151,1437,439,2014]" pageId="13" pageNumber="14">
<emphasis id="21D42252FA45FFF7FF70C13AFED6FAF0" bold="true" box="[199,334,1339,1365]" pageId="13" pageNumber="14">Etymology.</emphasis>
The species name is derived from the Greek “pholkos” for bow-legged; alluding to its resemblance to arachnids of the genus
<taxonomicName id="D4A085C3FA45FFF7FE01C15EFD88FADC" authorityName="Walckenaer" authorityYear="1805" box="[438,528,1375,1401]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="21D42252FA45FFF7FE01C15EFD88FADC" box="[438,528,1375,1401]" italics="true" pageId="13" pageNumber="14">Pholcus</emphasis>
</taxonomicName>
, better known as “daddy-long-legs”. The name is used as a noun.
</paragraph>
</subSubSection>
<subSubSection id="5BBAADCBFA45FFEAFF70C182FEA2F87B" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14" type="discussion">
<paragraph id="131FFE40FA45FFEAFF70C182FC32F947" blockId="13.[151,1437,439,2014]" lastBlockId="16.[151,1437,1304,2014]" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
<emphasis id="21D42252FA45FFF7FF70C182FEA3FA38" bold="true" box="[199,315,1411,1437]" pageId="13" pageNumber="14">Remarks.</emphasis>
The most distinctive character of
<taxonomicName id="D4A085C3FA45FFF7FD79C182FCDFFA38" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[718,839,1411,1438]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA45FFF7FD79C182FD47FA38" box="[718,735,1411,1437]" italics="true" pageId="13" pageNumber="14">A</emphasis>
.
<emphasis id="21D42252FA45FFF7FD47C182FCDFFA38" box="[752,839,1411,1437]" italics="true" pageId="13" pageNumber="14">pholcus</emphasis>
</taxonomicName>
<emphasis id="21D42252FA45FFF7FCE5C182FC0CFA38" bold="true" box="[850,916,1411,1437]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="3AE79F29FA45FFF7FCE5C182FC0CFA38" box="[850,916,1411,1437]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is the long and wide branchial, cardiac, and intestinal regions, together with the distinctly inflated hepatic regions, which give the carapace a subglobular appearance; with the orbital and frontal regions appearing disproportionately short (
<figureCitation id="8B9BE2C5FA45FFF7FB83C1CAFB24FA40" box="[1076,1212,1483,1509]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10A–C</figureCitation>
). The cardiac region is also prominently raised and capped by a sharp tubercle (
<figureCitation id="8B9BE2C5FA45FFF7FC9AC1EEFC2EF9AC" box="[813,950,1519,1545]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10D, E</figureCitation>
). All other species of
<taxonomicName id="D4A085C3FA45FFF7FB1BC1EEFA93F9AC" authorityName="Leach" authorityYear="1817" box="[1196,1291,1519,1545]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="21D42252FA45FFF7FB1BC1EEFA93F9AC" box="[1196,1291,1519,1545]" italics="true" pageId="13" pageNumber="14">Achaeus</emphasis>
</taxonomicName>
have a more pyriform and slender carapace, or carapace that appears subquadrate; most species have a less prominent and sharp cardiac region. Following the key of
<bibRefCitation id="773183B1FA45FFF7FD88C236FCE0F9F4" author="Griffin, D. J. G. &amp; Tranter, H. A." box="[575,888,1591,1618]" pageId="13" pageNumber="14" pagination="1 - 335" refId="ref20688" refString="Griffin, D. J. G. &amp; Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie Monograph, 39 (c 4), 1 - 335." type="journal article" year="1986">Griffin &amp; Tranter (1986: 4)</bibRefCitation>
,
<taxonomicName id="D4A085C3FA45FFF7FC32C236FC65F9F4" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[901,1021,1591,1617]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA45FFF7FC32C236FC65F9F4" box="[901,1021,1591,1617]" italics="true" pageId="13" pageNumber="14">A. pholcus</emphasis>
</taxonomicName>
<emphasis id="21D42252FA45FFF7FBB2C236FBDEF9F4" bold="true" box="[1029,1094,1591,1617]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="3AE79F29FA45FFF7FBB2C236FBDEF9F4" box="[1029,1094,1591,1617]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
belongs to a group of species in which the supraorbital eave has only small spinules lining the margin, the dorsal surface of the carapace does not have a transverse ridge or any large spines or tubercles (except for the cardiac region), the hepatic region is prominently produced with the epistome well-developed:
<taxonomicName id="D4A085C3FA45FFF7FCA8C2A2FBF2F91B" authority="Stimpson, 1857" authorityName="Stimpson" authorityYear="1857" box="[799,1130,1699,1726]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacertosus">
<emphasis id="21D42252FA45FFF7FCA8C2A2FCA8F918" box="[799,816,1699,1725]" italics="true" pageId="13" pageNumber="14">A</emphasis>
.
<emphasis id="21D42252FA45FFF7FC89C2A2FC29F918" box="[830,945,1699,1725]" italics="true" pageId="13" pageNumber="14">lacertosus</emphasis>
<bibRefCitation id="773183B1FA45FFF7FC0FC2A2FBF2F91B" author="Stimpson, W." box="[952,1130,1699,1726]" pageId="13" pageNumber="14" pagination="216 - 221" refId="ref23624" refString="Stimpson, W. (1857) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars III. Crustacea Maioidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 9, 216 - 221 (23 - 28). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1857">Stimpson, 1857</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="D4A085C3FA45FFF7FBC2C2A2FA00F918" authority="Rathbun, 1916" authorityName="Rathbun" authorityYear="1916" box="[1141,1432,1699,1726]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="21D42252FA45FFF7FBC2C2A2FB1EF918" box="[1141,1158,1699,1725]" italics="true" pageId="13" pageNumber="14">A</emphasis>
.
<emphasis id="21D42252FA45FFF7FB23C2A2FB72F918" box="[1172,1258,1699,1725]" italics="true" pageId="13" pageNumber="14">villosus</emphasis>
<bibRefCitation id="773183B1FA45FFF7FB47C2A2FA00F918" author="Rathbun, M. J." box="[1264,1432,1699,1725]" pageId="13" pageNumber="14" pagination="527 - 559" refId="ref22659" refString="Rathbun, M. J. (1916) New species of crabs of the families Inachidae and Parthenopidae. [Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34]. Proceedings of the United States National Museum, 50 (2135), 527 - 559." type="journal article" year="1916">Rathbun, 1916</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="D4A085C3FA45FFF7FF72C2C6FD9AF944" authority="Sakai, 1938" authorityName="Sakai" authorityYear="1938" box="[197,514,1735,1762]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevidactylus">
<emphasis id="21D42252FA45FFF7FF72C2C6FF4EF944" box="[197,214,1735,1761]" italics="true" pageId="13" pageNumber="14">A</emphasis>
.
<emphasis id="21D42252FA45FFF7FF55C2C6FEE0F944" box="[226,376,1735,1761]" italics="true" pageId="13" pageNumber="14">brevidactylus</emphasis>
<bibRefCitation id="773183B1FA45FFF7FECBC2C6FD9AF944" author="Sakai, T." box="[380,514,1735,1761]" pageId="13" pageNumber="14" pagination="193 - 364" refId="ref23249" refString="Sakai, T. (1938) s. n. In: Studies on the Crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo, Tokyo, pp. 193 - 364, figs. 1 - 55, pls. 20 - 41, tab. I." type="book chapter" year="1938">Sakai, 1938</bibRefCitation>
</taxonomicName>
. These three species can easily be distinguished from
<taxonomicName id="D4A085C3FA45FFF7FBFBC2C6FB58F944" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[1100,1216,1735,1761]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA45FFF7FBFBC2C6FB58F944" box="[1100,1216,1735,1761]" italics="true" pageId="13" pageNumber="14">A. pholcus</emphasis>
</taxonomicName>
<emphasis id="21D42252FA45FFF7FB72C2C6FA9AF944" bold="true" box="[1221,1282,1735,1761]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="3AE79F29FA45FFF7FB72C2C6FA9AF944" box="[1221,1282,1735,1761]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the latter’s longer and more inflated carapace (
<figureCitation id="8B9BE2C5FA45FFF7FD91C2EAFD12F8A0" box="[550,650,1771,1797]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Fig. 10B</figureCitation>
) (cf.
<bibRefCitation id="773183B1FA45FFF7FD70C2EAFCD1F8A0" author="Sakai, T." box="[711,841,1771,1797]" pageId="13" pageNumber="14" pagination="193 - 364" refId="ref23249" refString="Sakai, T. (1938) s. n. In: Studies on the Crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo, Tokyo, pp. 193 - 364, figs. 1 - 55, pls. 20 - 41, tab. I." type="book chapter" year="1938">Sakai 1938</bibRefCitation>
: pl. 21 fig. 4).
<taxonomicName id="D4A085C3FA45FFF7FC58C2EAFB50F8A0" authorityName="Stimpson" authorityYear="1857" box="[1007,1224,1771,1797]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacertosus">
<emphasis id="21D42252FA45FFF7FC58C2EAFB50F8A0" box="[1007,1224,1771,1797]" italics="true" pageId="13" pageNumber="14">Achaeus lacertosus</emphasis>
</taxonomicName>
has the antennular fossa and basal antennal article relatively short, the ambulatory dactylus semilunate with strong teeth along the ventral margin, and the male pleon is prominently broader, with somite 5 and the pleotelson very wide (
<bibRefCitation id="773183B1FA45FFF7FAFAC332FF4BF8D4" author="Griffin, D. J. G." pageId="13" pageNumber="14" pagination="97 - 119" refId="ref20518" refString="Griffin, D. J. G. (1970) The Australian majid spider crabs of the genus Achaeus (Crustacea, Brachyura). Journal of the Royal Society of Western Australia, 53 (4), 97 - 119, figs. 1 - 15." type="journal article" year="1970">Griffin 1970</bibRefCitation>
: fig. 5a, c, d) (versus the antennular fossa and basal antennal article distinctly longer, the ambulatory dactylus is gently curved with scattered low spinules along the ventral margin, and the male pleon is narrower with somite 5 subquadrate and the pleotelson less wide in
<taxonomicName id="D4A085C3FA45FFF7FDCCC39EFD68F81C" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[635,752,1951,1977]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA45FFF7FDCCC39EFD68F81C" box="[635,752,1951,1977]" italics="true" pageId="13" pageNumber="14">A. pholcus</emphasis>
</taxonomicName>
<emphasis id="21D42252FA45FFF7FD42C39EFCABF81C" bold="true" box="[757,819,1951,1977]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="3AE79F29FA45FFF7FD42C39EFCABF81C" box="[757,819,1951,1977]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="8B9BE2C5FA45FFF7FCF6C39EFC4AF81F" box="[833,978,1951,1978]" captionStart="FIGURE 10" captionStartId="14.[151,250,1914,1938]" captionTargetBox="[152,1436,181,1890]" captionTargetId="figure-27@14.[152,1436,181,1890]" captionTargetPageId="14" captionText="FIGURE 10. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="13" pageNumber="14">Figs. 10A, D</figureCitation>
,
<figureCitation id="8B9BE2C5FA45FFF7FC6BC39EFB95F81C" box="[988,1037,1951,1977]" captionStart="FIGURE 11" captionStartId="15.[151,250,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-18@15.[151,1436,181,1879]" captionTargetPageId="15" captionText="FIGURE 11. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, subventral view of cephalothorax showing thoracic sternum, pleon, buccal cavity, epistome, antennae, antennules and eyes; third maxillipeds; B, ventral view of cephalothorax showing thoracic sternum, pleon and third maxillipeds; C, thoracic sternal crest on sternite 4; D, pleonal somites 3–5 and pleotelson; E, pleonal somites 1 and 2." figureDoi="http://doi.org/10.5281/zenodo.8158111" httpUri="https://zenodo.org/record/8158111/files/figure.png" pageId="13" pageNumber="14">11D</figureCitation>
,
<figureCitation id="8B9BE2C5FA45FFF7FBAFC39EFBFEF81C" box="[1048,1126,1951,1977]" captionStart="FIGURE 13" captionStartId="17.[151,250,1721,1745]" captionTargetBox="[153,1434,183,1694]" captionTargetId="figure-18@17.[151,1436,181,1697]" captionTargetPageId="17" captionText="FIGURE 13. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, dorsal view of anterior part of carapace (setae not drawn); B, left third maxilliped (setae not drawn); C, left P2; D, right P3 (carpus to dactylus missing); E, left P4; F, left P4 dactylus; G, left P5; H, left P5 dactylus; I, right G1 (dorsal view) (medially broken); J, right G1 (ventral view) (medially broken); K, L, distal part of right G1 (dorsal view, slightly different angles); M, distal part of right G1 (mesial view); N, distal part of right G1 (ventral view); O, left G2. Scales: A, B, I, J, O = 0.5 mm; C–H = 1.0 mm; K, L, M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8158115" httpUri="https://zenodo.org/record/8158115/files/figure.png" pageId="13" pageNumber="14">13F, H</figureCitation>
).
<taxonomicName id="D4A085C3FA45FFF7FBCFC39EFAABF81C" authorityName="Rathbun" authorityYear="1916" box="[1144,1331,1951,1977]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="21D42252FA45FFF7FBCFC39EFAABF81C" box="[1144,1331,1951,1977]" italics="true" pageId="13" pageNumber="14">Achaeus villosus</emphasis>
</taxonomicName>
differs in having a more entire frontal margin with only a low concavity separating the two lobes, there are many small spinules along the surfaces of the hepatic and branchial regions, the chela is stouter and shorter with the margins smooth, the ambulatory dactylus is relatively shorter, and the male pleotelson is more triangular (
<bibRefCitation id="773183B1FA58FFEAFBE9C13DFAF2FAF3" author="Griffin, D. J. G. &amp; Tranter, H. A." box="[1118,1386,1340,1366]" pageId="16" pageNumber="17" pagination="1 - 335" refId="ref20688" refString="Griffin, D. J. G. &amp; Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie Monograph, 39 (c 4), 1 - 335." type="journal article" year="1986">Griffin &amp; Tranter 1986</bibRefCitation>
: fig. 4b–d) (versus two large frontal lobes distinctly separated by a deep U-shaped cleft, the hepatic and branchial regions covered with granules, the chela is more slender and longer with the margins lined with spinules, the ambulatory dactylus is relatively longer, and the male pleotelson is relatively wider in
<taxonomicName id="D4A085C3FA58FFEAFC5DC1A9FBFAFA67" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[1002,1122,1448,1474]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA58FFEAFC5DC1A9FC63FA67" box="[1002,1019,1448,1474]" italics="true" pageId="16" pageNumber="17">A</emphasis>
.
<emphasis id="21D42252FA58FFEAFBBCC1A9FBFAFA67" box="[1035,1122,1448,1474]" italics="true" pageId="16" pageNumber="17">pholcus</emphasis>
</taxonomicName>
<emphasis id="21D42252FA58FFEAFBDDC1A9FB32FA67" bold="true" box="[1130,1194,1448,1474]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="3AE79F29FA58FFEAFBDDC1A9FB32FA67" box="[1130,1194,1448,1474]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="8B9BE2C5FA58FFEAFB0CC1A9FAD7FA67" box="[1211,1359,1448,1474]" captionStart="FIGURE 11" captionStartId="15.[151,250,1903,1927]" captionTargetBox="[151,1436,181,1879]" captionTargetId="figure-18@15.[151,1436,181,1879]" captionTargetPageId="15" captionText="FIGURE 11. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, subventral view of cephalothorax showing thoracic sternum, pleon, buccal cavity, epistome, antennae, antennules and eyes; third maxillipeds; B, ventral view of cephalothorax showing thoracic sternum, pleon and third maxillipeds; C, thoracic sternal crest on sternite 4; D, pleonal somites 3–5 and pleotelson; E, pleonal somites 1 and 2." figureDoi="http://doi.org/10.5281/zenodo.8158111" httpUri="https://zenodo.org/record/8158111/files/figure.png" pageId="16" pageNumber="17">Figs. 11A–D</figureCitation>
,
<figureCitation id="8B9BE2C5FA58FFEAFAEBC1A9FA08FA67" box="[1372,1424,1448,1474]" captionStart="FIGURE 13" captionStartId="17.[151,250,1721,1745]" captionTargetBox="[153,1434,183,1694]" captionTargetId="figure-18@17.[151,1436,181,1697]" captionTargetPageId="17" captionText="FIGURE 13. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, dorsal view of anterior part of carapace (setae not drawn); B, left third maxilliped (setae not drawn); C, left P2; D, right P3 (carpus to dactylus missing); E, left P4; F, left P4 dactylus; G, left P5; H, left P5 dactylus; I, right G1 (dorsal view) (medially broken); J, right G1 (ventral view) (medially broken); K, L, distal part of right G1 (dorsal view, slightly different angles); M, distal part of right G1 (mesial view); N, distal part of right G1 (ventral view); O, left G2. Scales: A, B, I, J, O = 0.5 mm; C–H = 1.0 mm; K, L, M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8158115" httpUri="https://zenodo.org/record/8158115/files/figure.png" pageId="16" pageNumber="17">13A</figureCitation>
).
<taxonomicName id="D4A085C3FA58FFEAFF20C1CDFE0CFA43" authorityName="Sakai" authorityYear="1938" box="[151,404,1484,1510]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brevidactylus">
<emphasis id="21D42252FA58FFEAFF20C1CDFE0CFA43" box="[151,404,1484,1510]" italics="true" pageId="16" pageNumber="17">Achaeus brevidactylus</emphasis>
</taxonomicName>
can be separated by its distinctive P2 and P3 dactyli, which have the distal part dilated and spade-like (
<bibRefCitation id="773183B1FA58FFEAFEAEC1F1FE04F9AF" author="Sakai, T." box="[281,412,1520,1546]" pageId="16" pageNumber="17" pagination="193 - 364" refId="ref23249" refString="Sakai, T. (1938) s. n. In: Studies on the Crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo, Tokyo, pp. 193 - 364, figs. 1 - 55, pls. 20 - 41, tab. I." type="book chapter" year="1938">Sakai 1938</bibRefCitation>
: text-fig. 8a, b), and its male pleotelson is distinctly triangular in shape with almost straight lateral margins (
<bibRefCitation id="773183B1FA58FFEAFEFAC215FE49F98B" author="Sakai, T." box="[333,465,1556,1582]" pageId="16" pageNumber="17" pagination="193 - 364" refId="ref23249" refString="Sakai, T. (1938) s. n. In: Studies on the Crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo, Tokyo, pp. 193 - 364, figs. 1 - 55, pls. 20 - 41, tab. I." type="book chapter" year="1938">Sakai 1938</bibRefCitation>
: text-fig. 8f) (versus P2 and P3 dactyli tapering to a sharp tip with the male pleotelson possessing strongly sinuous lateral margins in
<taxonomicName id="D4A085C3FA58FFEAFD28C239FC8DF9F7" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[671,789,1592,1618]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA58FFEAFD28C239FD28F9F7" box="[671,688,1592,1618]" italics="true" pageId="16" pageNumber="17">A</emphasis>
.
<emphasis id="21D42252FA58FFEAFD09C239FC8DF9F7" box="[702,789,1592,1618]" italics="true" pageId="16" pageNumber="17">pholcus</emphasis>
</taxonomicName>
<emphasis id="21D42252FA58FFEAFCABC239FCC3F9F7" bold="true" box="[796,859,1592,1618]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="3AE79F29FA58FFEAFCABC239FCC3F9F7" box="[796,859,1592,1618]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="8B9BE2C5FA58FFEAFCDDC239FC5BF9F7" box="[874,963,1592,1618]" captionStart="FIGURE 9" captionStartId="12.[151,250,1706,1730]" captionTargetBox="[177,1411,345,1651]" captionTargetId="figure-117@12.[177,1411,345,1682]" captionTargetPageId="12" captionText="FIGURE 9. Dorhynchus rostratus (Sakai, 1932), ovigerous female (pcl 5.0 mmm) (ZRC 2020.0386), Philippines. A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, sternopleonal cavity and vulvae; E–H, left P2–P5, respectively (all same scale, right P5 tip broken)." figureDoi="http://doi.org/10.5281/zenodo.8158105" httpUri="https://zenodo.org/record/8158105/files/figure.png" pageId="16" pageNumber="17">Fig. 9D</figureCitation>
). The G1 of
<taxonomicName id="D4A085C3FA58FFEAFBE7C239FA9DF9F7" authorityName="Sakai" authorityYear="1938" box="[1104,1285,1592,1618]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brevidactylus">
<emphasis id="21D42252FA58FFEAFBE7C239FBF9F9F7" box="[1104,1121,1592,1618]" italics="true" pageId="16" pageNumber="17">A</emphasis>
.
<emphasis id="21D42252FA58FFEAFBD8C239FA9DF9F7" box="[1135,1285,1592,1618]" italics="true" pageId="16" pageNumber="17">brevidactylus</emphasis>
</taxonomicName>
is not known but those of the other two species are quite different:
<taxonomicName id="D4A085C3FA58FFEAFD47C25DFCFDF9D3" authorityName="Rathbun" authorityYear="1916" box="[752,869,1628,1654]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="21D42252FA58FFEAFD47C25DFC99F9D3" box="[752,769,1628,1654]" italics="true" pageId="16" pageNumber="17">A</emphasis>
.
<emphasis id="21D42252FA58FFEAFCB8C25DFCFDF9D3" box="[783,869,1628,1654]" italics="true" pageId="16" pageNumber="17">villosus</emphasis>
</taxonomicName>
has a relatively shorter and more sinuous G1 with the tip wide and appearing bilobed (
<bibRefCitation id="773183B1FA58FFEAFD82C281FCDAF93F" author="Griffin, D. J. G. &amp; Tranter, H. A." box="[565,834,1664,1690]" pageId="16" pageNumber="17" pagination="1 - 335" refId="ref20688" refString="Griffin, D. J. G. &amp; Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie Monograph, 39 (c 4), 1 - 335." type="journal article" year="1986">Griffin &amp; Tranter 1986</bibRefCitation>
: fig.
<quantity id="D45853A5FA58FFEAFCC9C281FC04F93E" box="[894,924,1664,1691]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="16" pageNumber="17" unit="g" value="5.0">5g</quantity>
, h) while that of
<taxonomicName id="D4A085C3FA58FFEAFBD3C281FB60F93F" authorityName="Stimpson" authorityYear="1857" box="[1124,1272,1664,1690]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lacertosus">
<emphasis id="21D42252FA58FFEAFBD3C281FBEDF93F" box="[1124,1141,1664,1690]" italics="true" pageId="16" pageNumber="17">A</emphasis>
.
<emphasis id="21D42252FA58FFEAFB32C281FB60F93F" box="[1157,1272,1664,1690]" italics="true" pageId="16" pageNumber="17">lacertosus</emphasis>
</taxonomicName>
is shorter and weakly C-shaped with the distal part subconical (
<bibRefCitation id="773183B1FA58FFEAFD0CC2A5FCD5F91B" author="Griffin, D. J. G." box="[699,845,1700,1726]" pageId="16" pageNumber="17" pagination="97 - 119" refId="ref20518" refString="Griffin, D. J. G. (1970) The Australian majid spider crabs of the genus Achaeus (Crustacea, Brachyura). Journal of the Royal Society of Western Australia, 53 (4), 97 - 119, figs. 1 - 15." type="journal article" year="1970">Griffin 1970</bibRefCitation>
: fig. 14a, b). The G1 of
<taxonomicName id="D4A085C3FA58FFEAFBEFC2A5FB55F91B" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[1112,1229,1700,1726]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA58FFEAFBEFC2A5FBF1F91B" box="[1112,1129,1700,1726]" italics="true" pageId="16" pageNumber="17">A</emphasis>
.
<emphasis id="21D42252FA58FFEAFBC1C2A5FB55F91B" box="[1142,1229,1700,1726]" italics="true" pageId="16" pageNumber="17">pholcus</emphasis>
</taxonomicName>
<emphasis id="21D42252FA58FFEAFB63C2A5FA8AF91B" bold="true" box="[1236,1298,1700,1726]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="3AE79F29FA58FFEAFB63C2A5FA8AF91B" box="[1236,1298,1700,1726]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is relatively long, slender, C-shaped with the distal part wide (
<figureCitation id="8B9BE2C5FA58FFEAFD74C2C9FCDEF947" box="[707,838,1736,1762]" captionStart="FIGURE 12" captionStartId="16.[151,250,1167,1191]" captionTargetBox="[151,1436,181,1142]" captionTargetId="figure-27@16.[151,1436,181,1143]" captionTargetPageId="16" captionText="FIGURE 12. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland.A, dorsal view of right cheliped; B, right chela and carpus; C, left third maxilliped; D, E, right G1 (ventral views); F, right G1 (dorsal view); G, left G2. G1s broken medially and connected by photoshop." figureDoi="http://doi.org/10.5281/zenodo.8158113" httpUri="https://zenodo.org/record/8158113/files/figure.png" pageId="16" pageNumber="17">Fig. 12D–F</figureCitation>
,
<figureCitation id="8B9BE2C5FA58FFEAFCE6C2C9FC05F947" box="[849,925,1736,1762]" captionStart="FIGURE 13" captionStartId="17.[151,250,1721,1745]" captionTargetBox="[153,1434,183,1694]" captionTargetId="figure-18@17.[151,1436,181,1697]" captionTargetPageId="17" captionText="FIGURE 13. Achaeus pholcus n. sp., holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, dorsal view of anterior part of carapace (setae not drawn); B, left third maxilliped (setae not drawn); C, left P2; D, right P3 (carpus to dactylus missing); E, left P4; F, left P4 dactylus; G, left P5; H, left P5 dactylus; I, right G1 (dorsal view) (medially broken); J, right G1 (ventral view) (medially broken); K, L, distal part of right G1 (dorsal view, slightly different angles); M, distal part of right G1 (mesial view); N, distal part of right G1 (ventral view); O, left G2. Scales: A, B, I, J, O = 0.5 mm; C–H = 1.0 mm; K, L, M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8158115" httpUri="https://zenodo.org/record/8158115/files/figure.png" pageId="16" pageNumber="17">13I–N</figureCitation>
).
</paragraph>
<caption id="47DFAEC8FA46FFF4FF20C37BFA01F87F" ID-DOI="http://doi.org/10.5281/zenodo.8158107" ID-Zenodo-Dep="8158107" httpUri="https://zenodo.org/record/8158107/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1914,1938]" targetBox="[152,1436,181,1890]" targetPageId="14" targetType="figure">
<paragraph id="131FFE40FA46FFF4FF20C37BFA01F87F" blockId="14.[151,1436,1914,2010]" pageId="14" pageNumber="15">
<emphasis id="21D42252FA46FFF4FF20C37BFE86F837" bold="true" box="[151,286,1914,1938]" pageId="14" pageNumber="15">FIGURE 10.</emphasis>
<taxonomicName id="D4A085C3FA46FFF4FE94C37BFE55F837" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[291,461,1914,1938]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA46FFF4FE94C37BFE55F837" box="[291,461,1914,1938]" italics="true" pageId="14" pageNumber="15">Achaeus pholcus</emphasis>
</taxonomicName>
<emphasis id="21D42252FA46FFF4FE65C37BFD92F837" bold="true" box="[466,522,1914,1938]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="3AE79F29FA46FFF4FE65C37BFD92F837" box="[466,522,1914,1938]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, overall dorsal view; B, dorsal view of carapace; C, dorsal view of anterior part of carapace showing pseudorostrum and orbits; D, ventral view of epistome, antennae, antenules, orbit and pseudorostrum; E, lateral view of cephalothorax; F, subfrontal view of cephalothorax.
</paragraph>
</caption>
<caption id="47DFAEC8FA47FFF5FF20C36EFC83F856" ID-DOI="http://doi.org/10.5281/zenodo.8158111" ID-Zenodo-Dep="8158111" httpUri="https://zenodo.org/record/8158111/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1903,1927]" targetBox="[151,1436,181,1879]" targetPageId="15" targetType="figure">
<paragraph id="131FFE40FA47FFF5FF20C36EFC83F856" blockId="15.[151,1437,1903,2035]" pageId="15" pageNumber="16">
<emphasis id="21D42252FA47FFF5FF20C36EFE86F822" bold="true" box="[151,286,1903,1927]" pageId="15" pageNumber="16">FIGURE 11.</emphasis>
<taxonomicName id="D4A085C3FA47FFF5FE93C36EFE56F822" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[292,462,1903,1927]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA47FFF5FE93C36EFE56F822" box="[292,462,1903,1927]" italics="true" pageId="15" pageNumber="16">Achaeus pholcus</emphasis>
</taxonomicName>
<emphasis id="21D42252FA47FFF5FE63C36EFD95F822" bold="true" box="[468,525,1903,1927]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="3AE79F29FA47FFF5FE63C36EFD95F822" box="[468,525,1903,1927]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, subventral view of cephalothorax showing thoracic sternum, pleon, buccal cavity, epistome, antennae, antennules and eyes; third maxillipeds; B, ventral view of cephalothorax showing thoracic sternum, pleon and third maxillipeds; C, thoracic sternal crest on sternite 4; D, pleonal somites 3–5 and pleotelson; E, pleonal somites 1 and 2.
</paragraph>
</caption>
<caption id="47DFAEC8FA58FFEAFF20C08EFD03FB4A" ID-DOI="http://doi.org/10.5281/zenodo.8158113" ID-Zenodo-Dep="8158113" httpUri="https://zenodo.org/record/8158113/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1167,1191]" targetBox="[151,1436,181,1142]" targetPageId="16" targetType="figure">
<paragraph id="131FFE40FA58FFEAFF20C08EFD03FB4A" blockId="16.[151,1437,1167,1263]" pageId="16" pageNumber="17">
<emphasis id="21D42252FA58FFEAFF20C08EFE87FB02" bold="true" box="[151,287,1167,1191]" pageId="16" pageNumber="17">FIGURE 12.</emphasis>
<taxonomicName id="D4A085C3FA58FFEAFE93C08EFE56FB02" authority="Ng &amp; Forges &amp; Lee, 2023" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[292,462,1167,1191]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pholcus" status="sp. nov.">
<emphasis id="21D42252FA58FFEAFE93C08EFE56FB02" box="[292,462,1167,1191]" italics="true" pageId="16" pageNumber="17">Achaeus pholcus</emphasis>
</taxonomicName>
<emphasis id="21D42252FA58FFEAFE64C08EFD93FB02" bold="true" box="[467,523,1167,1191]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="3AE79F29FA58FFEAFE64C08EFD93FB02" box="[467,523,1167,1191]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (pcl 7.3 mm) (MNHN-IU-2014-8034), New Ireland. A, dorsal view of right cheliped; B, right chela and carpus; C, left third maxilliped; D, E, right G1 (ventral views); F, right G1 (dorsal view); G, left G2. G1s broken medially and connected by photoshop.
</paragraph>
</caption>
<paragraph id="131FFE40FA58FFEAFF70C2EDFEA2F87B" blockId="16.[151,1437,1304,2014]" pageId="16" pageNumber="17">
<taxonomicName id="D4A085C3FA58FFEAFF70C2EDFE1EF8A3" authorityName="Rathbun" authorityYear="1916" box="[199,390,1772,1798]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="21D42252FA58FFEAFF70C2EDFE1EF8A3" box="[199,390,1772,1798]" italics="true" pageId="16" pageNumber="17">Achaeus villosus</emphasis>
</taxonomicName>
is known from the southern
<collectingCountry id="6BB7BED0FA58FFEAFD6FC2EDFCCEF8A3" box="[728,854,1772,1798]" name="Philippines" pageId="16" pageNumber="17">Philippines</collectingCountry>
(
<collectingRegion id="D16430A2FA58FFEAFCDFC2EDFC05F8A3" box="[872,925,1772,1798]" country="Philippines" name="Sulu" pageId="16" pageNumber="17">Sulu</collectingRegion>
islands), northern
<collectingCountry id="6BB7BED0FA58FFEAFBC0C2EDFB7FF8A3" box="[1143,1255,1772,1798]" name="Indonesia" pageId="16" pageNumber="17">Moluccas</collectingCountry>
,
<collectingCountry id="6BB7BED0FA58FFEAFB44C2EDFAD2F8A3" box="[1267,1354,1772,1798]" name="Indonesia" pageId="16" pageNumber="17">Ambon</collectingCountry>
, Timor and Sunda Strait (
<collectingCountry id="6BB7BED0FA58FFEAFEE8C311FE57F88F" box="[351,463,1808,1834]" name="Indonesia" pageId="16" pageNumber="17">Indonesia</collectingCountry>
) (
<bibRefCitation id="773183B1FA58FFEAFE52C311FD10F88F" author="Rathbun, M. J." box="[485,648,1808,1834]" pageId="16" pageNumber="17" pagination="527 - 559" refId="ref22659" refString="Rathbun, M. J. (1916) New species of crabs of the families Inachidae and Parthenopidae. [Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34]. Proceedings of the United States National Museum, 50 (2135), 527 - 559." type="journal article" year="1916">Rathbun 1916</bibRefCitation>
;
<bibRefCitation id="773183B1FA58FFEAFD24C311FCBBF88F" author="Griffin, D. J. G." box="[659,803,1808,1834]" pageId="16" pageNumber="17" pagination="179 - 222" refId="ref20625" refString="Griffin, D. J. G. (1976) Spider crabs of the family Majidae (Crustacea: Brachyura) from the Philippine Islands. Journal of Natural History, 10 (2), 179 - 222, figs. 1 - 11, tabs. 1 + 2. https: // doi. org / 10.1080 / 00222937600770131" type="journal article" year="1976">Griffin 1976</bibRefCitation>
;
<bibRefCitation id="773183B1FA58FFEAFC99C311FBABF88F" author="Griffin, D. J. G. &amp; Tranter, H. A." box="[814,1075,1808,1834]" pageId="16" pageNumber="17" pagination="1 - 335" refId="ref20688" refString="Griffin, D. J. G. &amp; Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie Monograph, 39 (c 4), 1 - 335." type="journal article" year="1986">Griffin &amp; Tranter 1986</bibRefCitation>
).
<taxonomicName id="D4A085C3FA58FFEAFBF1C311FA87F88F" authorityName="Stimpson" authorityYear="1857" box="[1094,1311,1808,1834]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lacertosus">
<emphasis id="21D42252FA58FFEAFBF1C311FA87F88F" box="[1094,1311,1808,1834]" italics="true" pageId="16" pageNumber="17">Achaeus lacertosus</emphasis>
</taxonomicName>
has a wide range across the Indo-West Pacific, ranging from
<collectingCountry id="6BB7BED0FA58FFEAFD77C335FCD7F8EB" box="[704,847,1844,1870]" name="South Africa" pageId="16" pageNumber="17">South Africa</collectingCountry>
and Iranian Gulf to southern
<collectingCountry id="6BB7BED0FA58FFEAFB20C335FB4BF8EB" box="[1175,1235,1844,1870]" name="India" pageId="16" pageNumber="17">India</collectingCountry>
,
<collectingRegion id="D16430A2FA58FFEAFB69C335FA8AF8EB" box="[1246,1298,1844,1870]" country="Indonesia" name="Jawa Timur" pageId="16" pageNumber="17">Java</collectingRegion>
, Kai Islands in the
<collectingCountry id="6BB7BED0FA58FFEAFF6BC359FED4F8D7" box="[220,332,1880,1906]" name="Indonesia" pageId="16" pageNumber="17">Moluccas</collectingCountry>
, Aru islands, northern
<collectingCountry id="6BB7BED0FA58FFEAFDF2C359FD37F8D7" box="[581,687,1880,1906]" name="Australia" pageId="16" pageNumber="17">Australia</collectingCountry>
, and
<collectingCountry id="6BB7BED0FA58FFEAFD5FC359FCB0F8D7" box="[744,808,1880,1906]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
(
<bibRefCitation id="773183B1FA58FFEAFC81C359FC71F8D6" author="Stimpson, W." box="[822,1001,1880,1907]" pageId="16" pageNumber="17" pagination="216 - 221" refId="ref23624" refString="Stimpson, W. (1857) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars III. Crustacea Maioidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 9, 216 - 221 (23 - 28). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1857">Stimpson, 1857</bibRefCitation>
,
<bibRefCitation id="773183B1FA58FFEAFC43C359FBB7F8D7" author="Stimpson, W." box="[1012,1071,1880,1906]" pageId="16" pageNumber="17" pagination="1 - 240" refId="ref23696" refString="Stimpson, W. (1907) Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition 1853 - 1856. Smithsonian Miscellaneous Collections, Part of 49 (1717), 1 - 240, 26 pls. https: // doi. org / 10.5962 / bhl. title. 51448" type="journal article" year="1907">1907</bibRefCitation>
,
<bibRefCitation id="773183B1FA58FFEAFB8EC359FB20F8D7" author="Sakai, T." box="[1081,1208,1880,1906]" pageId="16" pageNumber="17" pagination="193 - 364" refId="ref23249" refString="Sakai, T. (1938) s. n. In: Studies on the Crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo, Tokyo, pp. 193 - 364, figs. 1 - 55, pls. 20 - 41, tab. I." type="book chapter" year="1938">Sakai 1938</bibRefCitation>
,
<bibRefCitation id="773183B1FA58FFEAFB74C359FB66F8D7" author="Sakai, T." box="[1219,1278,1880,1906]" pageId="16" pageNumber="17" refId="ref23379" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Vols. 1 - 3. Kodansha Ltd., Tokyo, pp. xxix + 773 pp. (English Txt), 461 pp. (Japanese Txt), 16 pp., 251 pls. (Pls. Vol.)" type="book" year="1976">1976</bibRefCitation>
;
<bibRefCitation id="773183B1FA58FFEAFABEC359FA0FF8D7" author="Griffin, D. J. G." box="[1289,1431,1880,1906]" pageId="16" pageNumber="17" pagination="97 - 119" refId="ref20518" refString="Griffin, D. J. G. (1970) The Australian majid spider crabs of the genus Achaeus (Crustacea, Brachyura). Journal of the Royal Society of Western Australia, 53 (4), 97 - 119, figs. 1 - 15." type="journal article" year="1970">Griffin 1970</bibRefCitation>
;
<bibRefCitation id="773183B1FA58FFEAFF20C37DFE3CF833" author="Griffin, D. J. G. &amp; Tranter, H. A." box="[151,420,1916,1942]" pageId="16" pageNumber="17" pagination="1 - 335" refId="ref20688" refString="Griffin, D. J. G. &amp; Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie Monograph, 39 (c 4), 1 - 335." type="journal article" year="1986">Griffin &amp; Tranter 1986</bibRefCitation>
).
<taxonomicName id="D4A085C3FA58FFEAFE0DC37DFD20F833" authorityName="Sakai" authorityYear="1938" box="[442,696,1916,1942]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brevidactylus">
<emphasis id="21D42252FA58FFEAFE0DC37DFD20F833" box="[442,696,1916,1942]" italics="true" pageId="16" pageNumber="17">Achaeus brevidactylus</emphasis>
</taxonomicName>
is known only from
<collectingCountry id="6BB7BED0FA58FFEAFC18C37DFC77F833" box="[943,1007,1916,1942]" name="Japan" pageId="16" pageNumber="17">Japan</collectingCountry>
(
<bibRefCitation id="773183B1FA58FFEAFC48C37DFB1CF833" author="Sakai, T." box="[1023,1156,1916,1942]" pageId="16" pageNumber="17" pagination="193 - 364" refId="ref23249" refString="Sakai, T. (1938) s. n. In: Studies on the Crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo, Tokyo, pp. 193 - 364, figs. 1 - 55, pls. 20 - 41, tab. I." type="book chapter" year="1938">Sakai 1938</bibRefCitation>
,
<bibRefCitation id="773183B1FA58FFEAFB26C37DFB55F833" author="Sakai, T." box="[1169,1229,1916,1942]" pageId="16" pageNumber="17" refId="ref23379" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Vols. 1 - 3. Kodansha Ltd., Tokyo, pp. xxix + 773 pp. (English Txt), 461 pp. (Japanese Txt), 16 pp., 251 pls. (Pls. Vol.)" type="book" year="1976">1976</bibRefCitation>
). In addition, it is noteworthy that
<taxonomicName id="D4A085C3FA58FFEAFEE7C3A1FE5FF81F" authorityName="Ng &amp; Forges &amp; Lee" authorityYear="2023" box="[336,455,1952,1978]" class="Malacostraca" family="Inachidae" genus="Achaeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pholcus">
<emphasis id="21D42252FA58FFEAFEE7C3A1FE5FF81F" box="[336,455,1952,1978]" italics="true" pageId="16" pageNumber="17">A. pholcus</emphasis>
</taxonomicName>
is a deep-water species (
<quantity id="D45853A5FA58FFEAFD54C3A1FCF0F81F" box="[739,872,1952,1979]" metricMagnitude="2" metricUnit="m" metricValue="5.995" metricValueMax="6.49" metricValueMin="5.5" pageId="16" pageNumber="17" unit="m" value="599.5" valueMax="649.0" valueMin="550.0">550–649 m</quantity>
); the other three species occur in depths of
<quantity id="D45853A5FA58FFEAFAE2C3A1FA04F81F" box="[1365,1436,1952,1978]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="m" value="200.0">200 m</quantity>
and shallower.
</paragraph>
</subSubSection>
</treatment>
</document>
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