treatments-xml/data/22/2E/87/222E87ECAA1EFFFCFF1EFF5B30C2FC5B.xml

<document id="82C0553B1367224821FB7B8C597B7597" ID-DOI="10.11646/zootaxa.3737.4.5" ID-GBIF-Dataset="9d64f77d-ccf7-463e-98d3-1f898190c86d" ID-ISSN="1175-5326" ID-Zenodo-Dep="219354" ID-ZooBank="2499D127-1B3F-4868-A737-1274827D6806" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460701437723" checkinUser="plazi" docAuthor="Rowley, Jodi J. L., Dau, Vinh Quang &amp; Nguyen, Tao Thien" docDate="2013" docId="222E87ECAA1EFFFCFF1EFF5B30C2FC5B" docLanguage="en" docName="zt03737p428.pdf" docOrigin="Zootaxa 3737 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptolalax botsfordi Rowley, Dau &amp; Nguyen, 2013, sp. nov." docType="treatment" docVersion="14" lastPageNumber="425" masterDocId="DE17FF94AA1DFFF6FF89FFCC324DFF9C" masterDocTitle="A new species of Leptolalax (Anura: Megophryidae) from the highest mountain in Indochina" masterLastPageNumber="428" masterPageNumber="415" pageNumber="418" updateTime="1698354653046" updateUser="plazi">
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<mods:title id="6E1F0D4E6738095E24F7F758FE24A9B7">A new species of Leptolalax (Anura: Megophryidae) from the highest mountain in Indochina</mods:title>
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<mods:namePart id="CADE16049325005E0982CAB92C68F046">Rowley, Jodi J. L.</mods:namePart>
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<mods:namePart id="0135830BE0931A9AF4457CD10DC633F3">Dau, Vinh Quang</mods:namePart>
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<mods:namePart id="DAF75C8D30A490B74EEE065B3203C97C">Nguyen, Tao Thien</mods:namePart>
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<mods:date id="1CB71E28591DA5072FF8095238F9B311">2013</mods:date>
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<treatment id="222E87ECAA1EFFFCFF1EFF5B30C2FC5B" ID-DOI="http://doi.org/10.5281/zenodo.5622932" ID-GBIF-Taxon="119583410" ID-Zenodo-Dep="5622932" LSID="urn:lsid:plazi:treatment:222E87ECAA1EFFFCFF1EFF5B30C2FC5B" httpUri="http://treatment.plazi.org/id/222E87ECAA1EFFFCFF1EFF5B30C2FC5B" lastPageId="10" lastPageNumber="425" pageId="3" pageNumber="418">
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<paragraph id="AA3836FAAA1EFFF5FF1EFF5B33A2FF2E" blockId="3.[151,495,151,178]" box="[151,495,151,178]" pageId="3" pageNumber="418">
<heading id="F1708196AA1EFFF5FF1EFF5B33A2FF2E" bold="true" box="[151,495,151,178]" fontSize="11" level="1" pageId="3" pageNumber="418" reason="1">
<emphasis id="98F3EAE8AA1EFFF5FF1EFF5B33A2FF2E" bold="true" box="[151,495,151,178]" pageId="3" pageNumber="418">
<taxonomicName id="6D874D79AA1EFFF5FF1EFF5B33C7FF2D" ID-CoL="3TQPH" box="[151,394,151,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="418" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA1EFFF5FF1EFF5B33C7FF2D" bold="true" box="[151,394,151,177]" italics="true" pageId="3" pageNumber="418">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA1EFFF5FE18FF5433A2FF2E" box="[401,495,152,178]" pageId="3" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="E29D6571AA1EFFF5FF1EFF123085FDA7" pageId="3" pageNumber="418" type="materials_examined">
<paragraph id="AA3836FAAA1EFFF5FF1EFF123085FDA7" blockId="3.[151,1436,222,1003]" pageId="3" pageNumber="418">
<emphasis id="98F3EAE8AA1EFFF5FF1EFF123349FF6B" bold="true" box="[151,260,222,247]" pageId="3" pageNumber="418">
<typeStatus id="753C8858AA1EFFF5FF1EFF123349FF6B" box="[151,260,222,247]" pageId="3" pageNumber="418" type="holotype">Holotype</typeStatus>
</emphasis>
: VNMN 0 3682, adult male, calling under leaf litter, 
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from small trickle of water running through mossy rocks to a 1 m-wide cascading stream in upper montane forests on Mount Fansipan, Hoang Lien National Park, 
<collectingCountry id="D290766AAA1EFFF5FF50FEEB3348FEA2" box="[217,261,295,318]" name="Laos" pageId="3" pageNumber="418">Lao</collectingCountry>
Cai Province, 
<collectingCountry id="D290766AAA1EFFF5FE25FEEA3046FEA3" box="[428,523,294,319]" name="Vietnam" pageId="3" pageNumber="418">Vietnam</collectingCountry>
(
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, 
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, 
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elevation; 
<figureCitation id="32BC2A7FAA1EFFF5FB94FEEA3636FEA3" box="[1053,1147,294,319]" captionStart="FIGURE 1" captionStartId="1.[151,255,2003,2026]" captionTargetBox="[275,1317,1023,1985]" captionTargetId="figure@1.[275,1317,1023,1985]" captionTargetPageId="1" captionText="FIGURE 1. Type locality of Leptolalax botsfordi sp. nov. on Mount Fansipan, northern Vietnam." httpUri="https://zenodo.org/record/219355/files/figure.png" pageId="3" pageNumber="418">Figure 1</figureCitation>
). Collected at 19:35 h on 
<date id="DE39103AAA1EFFF5FF1EFE873361FEFF" box="[151,300,331,355]" pageId="3" pageNumber="418" value="2012-06-10">10 June 2012</date>
by Jodi J. L. Rowley, Vinh Quang Dau, Sang Van Pham, Tu Van Tran, Su A Hang, and 
<collectingCountry id="D290766AAA1EFFF5FA8CFE86376CFEFF" box="[1285,1313,330,355]" name="Iceland" pageId="3" pageNumber="418">Di</collectingCountry>
A Hoang. 
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<emphasis id="98F3EAE8AA1EFFF5FF1EFEA23342FE1B" bold="true" box="[151,271,366,391]" pageId="3" pageNumber="418">Paratypes</emphasis>
</typeStatus>
: AMS R 176540 (adult female) climbing into mud hole in seep at 19:55 h, and AMS R 176534–176538 (five adult males), collected at same geographic locality and date as 
<typeStatus id="753C8858AA1EFFF5FC29FE5E364BFE37" box="[928,1030,402,427]" pageId="3" pageNumber="418" type="holotype">holotype</typeStatus>
. AMS R 176541 (adult female) and AMS R 176539 (adult male), collected on 
<date id="DE39103AAA1EFFF5FDF5FE7B315BFE53" box="[636,790,439,463]" pageId="3" pageNumber="418" value="2012-06-11">11 June 2012</date>
, adjacent to a 2 m-wide, swift cascading stream in upper montane forest on Mount Fansipan, Hoang Lien Son National Park, 
<collectingCountry id="D290766AAA1EFFF5FC3FFE1731AFFE6E" box="[950,994,475,498]" name="Laos" pageId="3" pageNumber="418">Lao</collectingCountry>
Cai Province, 
<collectingCountry id="D290766AAA1EFFF5FB1BFE1636BCFE6F" box="[1170,1265,474,499]" name="Vietnam" pageId="3" pageNumber="418">Vietnam</collectingCountry>
at 
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<geoCoordinate id="CFB3503DAA1EFFF5FF1EFE323369FD8A" box="[151,292,510,535]" direction="east" orientation="longitude" pageId="3" pageNumber="418" precision="5" value="103.7654">103.7654º E</geoCoordinate>
, 
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elevation). All specimens were collected by Jodi J. L. Rowley, Vinh Quang Dau, Sang Van Pham, Tu Van Tran, Su A Hang, and 
<collectingCountry id="D290766AAA1EFFF5FDB1FDEE3019FDA7" box="[568,596,546,571]" name="Iceland" pageId="3" pageNumber="418">Di</collectingCountry>
A Hoang.
</paragraph>
</subSubSection>
<subSubSection id="E29D6571AA1EFFF5FF4EFD8A3196FD1F" pageId="3" pageNumber="418" type="etymology">
<paragraph id="AA3836FAAA1EFFF5FF4EFD8A3196FD1F" blockId="3.[151,1436,222,1003]" pageId="3" pageNumber="418">
<emphasis id="98F3EAE8AA1EFFF5FF4EFD8A3305FDC3" bold="true" box="[199,328,582,607]" pageId="3" pageNumber="418">Etymology</emphasis>
: Specific epithet is a patronym honouring Christopher Botsford, for his support of amphibian biodiversity conservation research and scientific capacity building in Asia.
</paragraph>
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<subSubSection id="E29D6571AA1EFFF5FF4EFD42314FFC77" pageId="3" pageNumber="418" type="diagnosis">
<paragraph id="AA3836FAAA1EFFF5FF4EFD42314FFC77" blockId="3.[151,1436,222,1003]" pageId="3" pageNumber="418">
<emphasis id="98F3EAE8AA1EFFF5FF4EFD423377FD3B" bold="true" box="[199,314,654,679]" pageId="3" pageNumber="418">Diagnosis</emphasis>
: Assigned to the genus 
<taxonomicName id="6D874D79AA1EFFF5FDC0FD42308DFD3A" box="[585,704,654,678]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="98F3EAE8AA1EFFF5FDC0FD42308DFD3A" box="[585,704,654,678]" italics="true" pageId="3" pageNumber="418">Leptolalax</emphasis>
</taxonomicName>
on the basis of the following characters: small (&lt;~ 
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SVL) size, rounded finger tips, the presence of an elevated inner palmar tubercle not continuous to the thumb, presence of macroglands on body (including supra-axillary, pectoral, femoral and ventrolateral glands), the absence of vomerine teeth, the presence of tubercles on eyelids, and anterior tip of snout with pale vertical bar (Dubois 1983; Lathrop 
<emphasis id="98F3EAE8AA1EFFF5FF71FCD33367FCAA" box="[248,298,798,822]" italics="true" pageId="3" pageNumber="418">et al</emphasis>
. 1998; Delorme 
<emphasis id="98F3EAE8AA1EFFF5FE67FCD3306FFCAA" box="[494,546,798,822]" italics="true" pageId="3" pageNumber="418">et al</emphasis>
. 2006). 
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<emphasis id="98F3EAE8AA1EFFF5FD0BFCD2312AFCAA" box="[642,871,798,822]" italics="true" pageId="3" pageNumber="418">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA1EFFF5FCF9FCD23186FCAB" box="[880,971,798,823]" pageId="3" pageNumber="418" rank="species">
<emphasis id="98F3EAE8AA1EFFF5FCF9FCD23186FCAB" bold="true" box="[880,971,798,823]" pageId="3" pageNumber="418">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from its congeners by a combination of (1) supra-axillary and ventrolateral glands present; (2) dark brownish red ventral surface with white speckling; (3) medium SVL for the genus (
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in 
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males, 30.0– 
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in 
<specimenCount id="BC81FD73AA1EFFF5FB3BFCAB376EFCE3" box="[1202,1315,870,895]" pageId="3" pageNumber="418" type="female">2 females</specimenCount>
); (4) black markings on the flanks absent; (5) toes with rudimentary webbing and weak lateral fringing; (6) large pectoral glands (
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; 4–6% SVL) and femoral glands (
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; 7–14% SVL); and (7) an advertisement call with a dominant frequency of 2.6–3.2 kHz (at 14.0 ºC).
</paragraph>
</subSubSection>
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<paragraph id="AA3836FAAA1EFFF5FF1EF8593026F850" blockId="3.[151,1436,1940,1996]" pageId="3" pageNumber="418">
<emphasis id="98F3EAE8AA1EFFF5FF1EF8593354F830" bold="true" box="[151,281,1941,1964]" pageId="3" pageNumber="418">FIGURE 3.</emphasis>
Adult female 
<taxonomicName id="6D874D79AA1EFFF5FE3DF85830C1F830" box="[436,652,1940,1964]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="418" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA1EFFF5FE3DF85830C1F830" box="[436,652,1940,1964]" italics="true" pageId="3" pageNumber="418">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA1EFFF5FD1AF85930A5F830" box="[659,744,1941,1964]" pageId="3" pageNumber="418" rank="species">
<emphasis id="98F3EAE8AA1EFFF5FD1AF85930AFF830" bold="true" box="[659,738,1941,1964]" pageId="3" pageNumber="418">sp. nov</emphasis>
.
</taxonomicNameLabel>
AMS R 176540 in life. Note metallic copper flecks, particularly along snout, supratympanic fold and flanks.
</paragraph>
</caption>
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Description of 
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</emphasis>
: Head width equal to head length; snout rounded in dorsal view and rounded in profile, projecting slightly beyond margin of the lower jaw; nostril equidistant between snout and eye; canthus rostralis indistinct, gently rounded; lores slightly concave; vertical pupil; eye diameter smaller than snout length; tympanum barely distinct, round, diameter smaller than that of the eye; tympanic rim elevated relative to skin of temporal region on ventral side only; vomerine teeth absent; pineal ocellus absent; vocal sac openings slit-like, located posterolaterally on floor of mouth in close proximity to the margins of the mandible; tongue long, wide, with broad, shallow notch at posterior tip; supratympanic ridge distinct, running from eye towards axillary gland, with raised tubercles. Tips of fingers rounded, slightly swollen; relative finger lengths I &lt;II &lt;IV &lt;III; nuptial pad absent; subarticular tubercles absent; a large, round inner palmar tubercle distinctly separated from small, laterally compressed outer palmar tubercle; finger webbing and dermal fringes absent (
<figureCitation id="32BC2A7FAA19FFF2FB89FE10362DFE69" box="[1024,1120,476,501]" captionStart="FIGURE 5" captionStartId="5.[151,255,1802,1825]" captionTargetBox="[172,1423,206,1780]" captionTargetId="figure@5.[151,1436,194,1781]" captionTargetPageId="5" captionText="FIGURE 5. Palmar surface of (A) left hand and (B) right foot of adult male holotype Leptolalax botsfordi sp. nov. VNMN 0 3682. Scale bar = 1 mm." httpUri="https://zenodo.org/record/219359/files/figure.png" pageId="4" pageNumber="419">Figure 5</figureCitation>
A). Tips of toes like fingers; relative toe length I &lt;II &lt;V &lt;III &lt;IV; subarticular tubercles absent, replaced by dermal ridges; large, oval inner metatarsal tubercle present, outer metatarsal tubercle absent; toe webbing rudimentary; narrow lateral fringes on all toes (
<figureCitation id="32BC2A7FAA19FFF2FF5AFD8B3379FDFC" box="[211,308,583,608]" captionStart="FIGURE 5" captionStartId="5.[151,255,1802,1825]" captionTargetBox="[172,1423,206,1780]" captionTargetId="figure@5.[151,1436,194,1781]" captionTargetPageId="5" captionText="FIGURE 5. Palmar surface of (A) left hand and (B) right foot of adult male holotype Leptolalax botsfordi sp. nov. VNMN 0 3682. Scale bar = 1 mm." httpUri="https://zenodo.org/record/219359/files/figure.png" pageId="4" pageNumber="419">Figure 5</figureCitation>
B). Tibia 46% of snout-vent length; tibiotarsal articulation reaches to middle of eye. Skin on dorsum shagreened; ventral skin smooth; pectoral gland large, oval, 
<quantity id="6D7F9B1FAA19FFF2FCF5FDA03196FD18" box="[892,987,620,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="419" unit="mm" value="1.5">1.5 mm</quantity>
diameter; femoral gland large, oval, approximately 
<quantity id="6D7F9B1FAA19FFF2FECAFD5C33D0FD3B" box="[323,413,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="419" unit="mm" value="3.4">3.4 mm</quantity>
diameter, on posteroventral surface of thigh, closer to knee than to vent; supra-axillary gland raised, 
<quantity id="6D7F9B1FAA19FFF2FEBAFD7833C1FD50" box="[307,396,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="419" unit="mm" value="1.8">1.8 mm</quantity>
diameter. Ventrolateral glands just visible as small white dots forming an incomplete line anteriorally, barely visible in preservative.
</paragraph>
<caption id="FEF86672AA19FFF2FF1EF88236D3F819" httpUri="https://zenodo.org/record/219358/files/figure.png" pageId="4" pageNumber="419" targetBox="[151,1407,817,1828]" targetPageId="4">
<paragraph id="AA3836FAAA19FFF2FF1EF88236D3F819" blockId="4.[151,1436,1869,1925]" pageId="4" pageNumber="419">
<emphasis id="98F3EAE8AA19FFF2FF1EF8823351F8F9" bold="true" box="[151,284,1870,1893]" pageId="4" pageNumber="419">FIGURE 4.</emphasis>
<taxonomicName id="6D874D79AA19FFF2FEAEF881304FF8F9" box="[295,514,1869,1893]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="419" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA19FFF2FEAEF881304FF8F9" box="[295,514,1869,1893]" italics="true" pageId="4" pageNumber="419">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA19FFF2FD85F8823028F8F9" box="[524,613,1870,1893]" pageId="4" pageNumber="419" rank="species">
<emphasis id="98F3EAE8AA19FFF2FD85F8823013F8F9" bold="true" box="[524,606,1870,1893]" pageId="4" pageNumber="419">sp. nov</emphasis>
.
</taxonomicNameLabel>
in life. (A) adult male holotype VNMN 03682; (B) adult female AMS R 176540; (C) adult male AMS R 176539; (D) female AMS R 176541. Note robust arms of males.
</paragraph>
</caption>
<caption id="FEF86672AA18FFF3FF1EF8C6304EF8DD" httpUri="https://zenodo.org/record/219359/files/figure.png" pageId="5" pageNumber="420" targetBox="[172,1423,206,1780]" targetPageId="5">
<paragraph id="AA3836FAAA18FFF3FF1EF8C6304EF8DD" blockId="5.[151,1436,1801,1857]" pageId="5" pageNumber="420">
<emphasis id="98F3EAE8AA18FFF3FF1EF8C63350F8BD" bold="true" box="[151,285,1802,1825]" pageId="5" pageNumber="420">FIGURE 5.</emphasis>
Palmar surface of (A) left hand and (B) right foot of adult male holotype 
<taxonomicName id="6D874D79AA18FFF3FBD6F8C53777F8BD" box="[1119,1338,1801,1825]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="420" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA18FFF3FBD6F8C53777F8BD" box="[1119,1338,1801,1825]" italics="true" pageId="5" pageNumber="420">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA18FFF3FACDF8C637D1F8BD" box="[1348,1436,1802,1825]" pageId="5" pageNumber="420" rank="species">
<emphasis id="98F3EAE8AA18FFF3FACDF8C637D8F8BD" bold="true" box="[1348,1429,1802,1825]" pageId="5" pageNumber="420">sp. nov</emphasis>
.
</taxonomicNameLabel>
VNMN 0 3682. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="AA3836FAAA1BFFF0FF4EFF5B31E8FE14" blockId="6.[151,1436,151,392]" pageId="6" pageNumber="421">
<emphasis id="98F3EAE8AA1BFFF0FF4EFF5B33B7FF2C" bold="true" box="[199,506,151,176]" pageId="6" pageNumber="421">
Colour of 
<typeStatus id="753C8858AA1BFFF0FECDFF5B33E6FF2C" box="[324,427,151,176]" pageId="6" pageNumber="421" type="holotype">holotype</typeStatus>
in life
</emphasis>
(
<figureCitation id="32BC2A7FAA1BFFF0FD83FF5B3021FF2C" box="[522,620,151,176]" captionStart="FIGURE 2" captionStartId="2.[151,255,1693,1716]" captionTargetBox="[151,1429,781,1670]" captionTargetId="figure@2.[151,1436,774,1671]" captionTargetPageId="2" captionText="FIGURE 2. Leptolalax botsfordi sp. nov. (A) dorsal and (B) ventral view of adult male holotype VNMN 0 3682 in life, (C) ventral view of gravid female paratype AMS R 176540 in life, (D) posterior view of adult male holotype VNMN 0 3682 in life, showing large femoral glands in (not to scale), (E) dorsal and (F) ventral view of adult male holotype VNMN 0 3682 in preservative, and (G) ventral view of adult female paratype AMS R 176540 in preservative. Scale bars = 5 mm." httpUri="https://zenodo.org/record/219356/files/figure.png" pageId="6" pageNumber="421">Figure 2</figureCitation>
A–B): Dorsal surface dark brown with gold edging dorsolaterally; gold interorbital bar; very faint transverse dark brown bars on the dorsal surface of the thighs, tibia, tarsus, lower arms, fingers and toes. Supratympannic ridge and vertical bar at anterior tip of snout metallic copper. Flanks marbled metallic gold and pale brown. Ventral surfaces slightly transparent reddish brown with faint paler flecks most obvious on slightly darker throat and thighs. Supra-axillary gland white edged in pale copper; pectoral glands opaque white; femoral glands white (
<figureCitation id="32BC2A7FAA1BFFF0FDD9FE8030FBFEF8" box="[592,694,332,357]" captionStart="FIGURE 2" captionStartId="2.[151,255,1693,1716]" captionTargetBox="[151,1429,781,1670]" captionTargetId="figure@2.[151,1436,774,1671]" captionTargetPageId="2" captionText="FIGURE 2. Leptolalax botsfordi sp. nov. (A) dorsal and (B) ventral view of adult male holotype VNMN 0 3682 in life, (C) ventral view of gravid female paratype AMS R 176540 in life, (D) posterior view of adult male holotype VNMN 0 3682 in life, showing large femoral glands in (not to scale), (E) dorsal and (F) ventral view of adult male holotype VNMN 0 3682 in preservative, and (G) ventral view of adult female paratype AMS R 176540 in preservative. Scale bars = 5 mm." httpUri="https://zenodo.org/record/219356/files/figure.png" pageId="6" pageNumber="421">Figure 2</figureCitation>
D). Iris dark brownish gold with densely-packed minute, black reticulations throughout. Iris periphery lined with black. Sclera white.
</paragraph>
<caption id="FEF86672AA1BFFF0FF1EFA56305DF9AD" httpUri="https://zenodo.org/record/219360/files/figure.png" pageId="6" pageNumber="421" targetBox="[167,1422,454,1403]" targetPageId="6">
<paragraph id="AA3836FAAA1BFFF0FF1EFA56305DF9AD" blockId="6.[151,1436,1433,1585]" pageId="6" pageNumber="421">
<emphasis id="98F3EAE8AA1BFFF0FF1EFA563356FA2D" bold="true" box="[151,283,1434,1457]" pageId="6" pageNumber="421">FIGURE 6.</emphasis>
Advertisement call of 
<taxonomicName id="6D874D79AA1BFFF0FD9BFA5530A6FA2D" box="[530,747,1433,1457]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="421" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA1BFFF0FD9BFA5530A6FA2D" box="[530,747,1433,1457]" italics="true" pageId="6" pageNumber="421">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA1BFFF0FD7AFA563104FA2D" box="[755,841,1434,1457]" pageId="6" pageNumber="421" rank="species">
<emphasis id="98F3EAE8AA1BFFF0FD7AFA563104FA2D" bold="true" box="[755,841,1434,1457]" pageId="6" pageNumber="421">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype VNMN 0 3682 (A) 14 s waveform of relative amplitude (Rel. amp.) over time for 16 “primary” calls, (B) waveform and (C) corresponding spectrogram of single representative “secondary” call expanded from first section shown in A, and (D) waveform and (E) corresponding spectrogram of single representative “primary” call expanded from second section shown in A; all recorded at an ambient air temperature of 14.0 ºC.
</paragraph>
</caption>
<paragraph id="AA3836FAAA1BFFF0FF4EF94E306EF89B" blockId="6.[151,1436,1666,1871]" pageId="6" pageNumber="421">
<emphasis id="98F3EAE8AA1BFFF0FF4EF94E3024F907" bold="true" box="[199,617,1666,1691]" pageId="6" pageNumber="421">
Colour of 
<typeStatus id="753C8858AA1BFFF0FECCF94E33E1F907" box="[325,428,1666,1691]" pageId="6" pageNumber="421" type="holotype">holotype</typeStatus>
in preservative
</emphasis>
(
<figureCitation id="32BC2A7FAA1BFFF0FDF3F94E3091F907" box="[634,732,1666,1691]" captionStart="FIGURE 2" captionStartId="2.[151,255,1693,1716]" captionTargetBox="[151,1429,781,1670]" captionTargetId="figure@2.[151,1436,774,1671]" captionTargetPageId="2" captionText="FIGURE 2. Leptolalax botsfordi sp. nov. (A) dorsal and (B) ventral view of adult male holotype VNMN 0 3682 in life, (C) ventral view of gravid female paratype AMS R 176540 in life, (D) posterior view of adult male holotype VNMN 0 3682 in life, showing large femoral glands in (not to scale), (E) dorsal and (F) ventral view of adult male holotype VNMN 0 3682 in preservative, and (G) ventral view of adult female paratype AMS R 176540 in preservative. Scale bars = 5 mm." httpUri="https://zenodo.org/record/219356/files/figure.png" pageId="6" pageNumber="421">Figure 2</figureCitation>
E–F): Dorsum dark brown; flanks marbled with paler brown; interorbital bar and dorsal surface of fingers and toes paler brown. Ventral surface of chest, belly, and thighs slightly mottled brown; throat dark brown speckled with white; arms and tibiotarsus mottled dark brown and cream. Macrogrands creamy white.
</paragraph>
<paragraph id="AA3836FAAA1BFFF0FF4EF8DE3010F8D3" blockId="6.[151,1436,1666,1871]" pageId="6" pageNumber="421">
<emphasis id="98F3EAE8AA1BFFF0FF4EF8DE333BF8B7" bold="true" box="[199,374,1810,1835]" pageId="6" pageNumber="421">Measurements</emphasis>
: 
<typeStatus id="753C8858AA1BFFF0FE0CF8DE33BFF8B7" box="[389,498,1810,1835]" pageId="6" pageNumber="421" type="holotype">Holotype</typeStatus>
: SVL 29.1, HDL 10.1, HDW 10.1, SNT 4.0, EYE 3.4, IOD 3.4, TMP 1.7, TEY 1.2, TIB 13.4, ML 7.4, PL 14.3, weight 
<quantity id="6D7F9B1FAA1BFFF0FDAFF8FB3010F8D3" box="[550,605,1847,1871]" metricMagnitude="-3" metricUnit="kg" metricValue="2.4" pageId="6" pageNumber="421" unit="g" value="2.4">2.4 g</quantity>
</paragraph>
<caption id="FEF86672AA1AFFF1FF1EF8AA36D4F801" httpUri="https://zenodo.org/record/219361/files/figure.png" pageId="7" pageNumber="422" targetBox="[151,1436,209,1872]" targetPageId="7">
<paragraph id="AA3836FAAA1AFFF1FF1EF8AA36D4F801" blockId="7.[151,1436,1893,1949]" pageId="7" pageNumber="422">
<emphasis id="98F3EAE8AA1AFFF1FF1EF8AA3357F8E1" bold="true" box="[151,282,1894,1917]" pageId="7" pageNumber="422">FIGURE 7.</emphasis>
Habitat of 
<taxonomicName id="6D874D79AA1AFFF1FE1CF8A93020F8E1" box="[405,621,1893,1917]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="422" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA1AFFF1FE1CF8A93020F8E1" box="[405,621,1893,1917]" italics="true" pageId="7" pageNumber="422">Leptolalax botsfordi</emphasis>
</taxonomicName>
<emphasis id="98F3EAE8AA1AFFF1FDFCF8AA3081F8E1" bold="true" box="[629,716,1894,1917]" pageId="7" pageNumber="422">
<taxonomicNameLabel id="83C05793AA1AFFF1FDFCF8AA3081F8E1" box="[629,716,1894,1917]" pageId="7" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) View of Mount Fansipan and surrounding mountains, Lao Cai Province, Vietnam. (B) Microhabitat of calling male 
<taxonomicName id="6D874D79AA1AFFF1FD45F84A310AF801" box="[716,839,1925,1949]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="422" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA1AFFF1FD45F84A310AF801" box="[716,839,1925,1949]" italics="true" pageId="7" pageNumber="422">L. botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA1AFFF1FCC4F84A31EEF801" box="[845,931,1926,1949]" pageId="7" pageNumber="422" rank="species">
<emphasis id="98F3EAE8AA1AFFF1FCC4F84A31EEF801" bold="true" box="[845,931,1926,1949]" pageId="7" pageNumber="422">sp. nov.</emphasis>
</taxonomicNameLabel>
at ~2,800 m elevation.
</paragraph>
</caption>
<caption id="FEF86672AA15FFFEFF1EFF5436C6FF33" ID-Table-UUID="FEF86672AA15FFFEFF1EFF5436C6FF33" box="[151,1163,151,175]" httpUri="http://table.plazi.org/id/FEF86672AA15FFFEFF1EFF5436C6FF33" pageId="8" pageNumber="423" targetBox="[165,1415,205,1053]" targetIsTable="true" targetPageId="8">
<paragraph id="AA3836FAAA15FFFEFF1EFF5436C6FF33" blockId="8.[151,1163,151,175]" box="[151,1163,151,175]" pageId="8" pageNumber="423">
<emphasis id="98F3EAE8AA15FFFEFF1EFF54334FFF33" bold="true" box="[151,258,152,175]" pageId="8" pageNumber="423">TABLE 1</emphasis>
. Measurements (mm) of 
<taxonomicName id="6D874D79AA15FFFEFD98FF5B30A4FF33" box="[529,745,151,175]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="423" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA15FFFEFD98FF5B30A4FF33" box="[529,745,151,175]" italics="true" pageId="8" pageNumber="423">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA15FFFEFD79FF543108FF33" box="[752,837,152,175]" pageId="8" pageNumber="423" rank="species">
<emphasis id="98F3EAE8AA15FFFEFD79FF543108FF33" bold="true" box="[752,837,152,175]" pageId="8" pageNumber="423">sp. nov.</emphasis>
</taxonomicNameLabel>
Abbreviations defined in text.
</paragraph>
</caption>
<paragraph id="AA3836FAAA15FFFEFEDDFF02371DFB81" pageId="8" pageNumber="423">
<table id="D887C45AAA150009FF2CFF0137CAFB81" box="[165,1415,205,1053]" gridcols="10" gridrows="19" pageId="8" pageNumber="423">
<tr id="14B734B8AA150009FF2CFF0137CAFE99" box="[165,1415,205,261]" gridrow="0" pageId="8" pageNumber="423" rowspan-0="1">
<th id="57665DC4AA150009FEC2FF0133ECFE99" box="[331,417,205,261]" gridcol="1" gridrow="0" pageId="8" pageNumber="423">VNMN 03682*</th>
<th id="57665DC4AA150009FE4EFF013053FE99" box="[455,542,205,261]" gridcol="2" gridrow="0" pageId="8" pageNumber="423">AMS R 176534</th>
<th id="57665DC4AA150009FDCDFF0130D6FE99" box="[580,667,205,261]" gridcol="3" gridrow="0" pageId="8" pageNumber="423">AMS R 176535</th>
<th id="57665DC4AA150009FD48FF013155FE99" box="[705,792,205,261]" gridcol="4" gridrow="0" pageId="8" pageNumber="423">AMS R 176536</th>
<th id="57665DC4AA150009FCB4FF0131D9FE99" box="[829,916,205,261]" gridcol="5" gridrow="0" pageId="8" pageNumber="423">AMS R 176537</th>
<th id="57665DC4AA150009FC33FF01365CFE99" box="[954,1041,205,261]" gridcol="6" gridrow="0" pageId="8" pageNumber="423">AMS R 176538</th>
<th id="57665DC4AA150009FBBFFF0136C3FE99" box="[1078,1166,205,261]" gridcol="7" gridrow="0" pageId="8" pageNumber="423">AMS R 176539</th>
<th id="57665DC4AA150009FB3AFF013747FE99" box="[1203,1290,205,261]" gridcol="8" gridrow="0" pageId="8" pageNumber="423">AMS R 176540</th>
<th id="57665DC4AA150009FAB9FF0137CAFE99" box="[1328,1415,205,261]" gridcol="9" gridrow="0" pageId="8" pageNumber="423">AMS R 176541</th>
</tr>
<tr id="14B734B8AA150009FF2CFEDB37CAFEB1" box="[165,1415,279,301]" gridrow="1" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFEDB3351FEB1" box="[165,284,279,301]" gridcol="0" gridrow="1" pageId="8" pageNumber="423">SEX</th>
<td id="57665DC4AA150009FEC2FEDB33ECFEB1" box="[331,417,279,301]" gridcol="1" gridrow="1" pageId="8" pageNumber="423">Male</td>
<td id="57665DC4AA150009FE4EFEDB3053FEB1" box="[455,542,279,301]" gridcol="2" gridrow="1" pageId="8" pageNumber="423">Male</td>
<td id="57665DC4AA150009FDCDFEDB30D6FEB1" box="[580,667,279,301]" gridcol="3" gridrow="1" pageId="8" pageNumber="423">Male</td>
<td id="57665DC4AA150009FD48FEDB3155FEB1" box="[705,792,279,301]" gridcol="4" gridrow="1" pageId="8" pageNumber="423">Male</td>
<td id="57665DC4AA150009FCB4FEDB31D9FEB1" box="[829,916,279,301]" gridcol="5" gridrow="1" pageId="8" pageNumber="423">Male</td>
<td id="57665DC4AA150009FC33FEDB365CFEB1" box="[954,1041,279,301]" gridcol="6" gridrow="1" pageId="8" pageNumber="423">Male</td>
<td id="57665DC4AA150009FBBFFEDB36C3FEB1" box="[1078,1166,279,301]" gridcol="7" gridrow="1" pageId="8" pageNumber="423">Male</td>
<td id="57665DC4AA150009FB3AFEDB3747FEB1" box="[1203,1290,279,301]" gridcol="8" gridrow="1" pageId="8" pageNumber="423">Female</td>
<td id="57665DC4AA150009FAB9FEDB37CAFEB1" box="[1328,1415,279,301]" gridcol="9" gridrow="1" pageId="8" pageNumber="423">Female</td>
</tr>
<tr id="14B734B8AA150009FF2CFEF337CAFEC9" box="[165,1415,319,341]" gridrow="2" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFEF33351FEC9" box="[165,284,319,341]" gridcol="0" gridrow="2" pageId="8" pageNumber="423">SVL</th>
<td id="57665DC4AA150009FEC2FEF333ECFEC9" box="[331,417,319,341]" gridcol="1" gridrow="2" pageId="8" pageNumber="423">29.1</td>
<td id="57665DC4AA150009FE4EFEF33053FEC9" box="[455,542,319,341]" gridcol="2" gridrow="2" pageId="8" pageNumber="423">32.6</td>
<td id="57665DC4AA150009FDCDFEF330D6FEC9" box="[580,667,319,341]" gridcol="3" gridrow="2" pageId="8" pageNumber="423">29.5</td>
<td id="57665DC4AA150009FD48FEF33155FEC9" box="[705,792,319,341]" gridcol="4" gridrow="2" pageId="8" pageNumber="423">30.4</td>
<td id="57665DC4AA150009FCB4FEF331D9FEC9" box="[829,916,319,341]" gridcol="5" gridrow="2" pageId="8" pageNumber="423">28.7</td>
<td id="57665DC4AA150009FC33FEF3365CFEC9" box="[954,1041,319,341]" gridcol="6" gridrow="2" pageId="8" pageNumber="423">28.8</td>
<td id="57665DC4AA150009FBBFFEF336C3FEC9" box="[1078,1166,319,341]" gridcol="7" gridrow="2" pageId="8" pageNumber="423">30.5</td>
<td id="57665DC4AA150009FB3AFEF33747FEC9" box="[1203,1290,319,341]" gridcol="8" gridrow="2" pageId="8" pageNumber="423">31.8</td>
<td id="57665DC4AA150009FAB9FEF337CAFEC9" box="[1328,1415,319,341]" gridcol="9" gridrow="2" pageId="8" pageNumber="423">30.0</td>
</tr>
<tr id="14B734B8AA150009FF2CFEAB37CAFEE1" box="[165,1415,359,381]" gridrow="3" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFEAB3351FEE1" box="[165,284,359,381]" gridcol="0" gridrow="3" pageId="8" pageNumber="423">HDL</th>
<td id="57665DC4AA150009FEC2FEAB33ECFEE1" box="[331,417,359,381]" gridcol="1" gridrow="3" pageId="8" pageNumber="423">10.1</td>
<td id="57665DC4AA150009FE4EFEAB3053FEE1" box="[455,542,359,381]" gridcol="2" gridrow="3" pageId="8" pageNumber="423">10.7</td>
<td id="57665DC4AA150009FDCDFEAB30D6FEE1" box="[580,667,359,381]" gridcol="3" gridrow="3" pageId="8" pageNumber="423">10.4</td>
<td id="57665DC4AA150009FD48FEAB3155FEE1" box="[705,792,359,381]" gridcol="4" gridrow="3" pageId="8" pageNumber="423">10.1</td>
<td id="57665DC4AA150009FCB4FEAB31D9FEE1" box="[829,916,359,381]" gridcol="5" gridrow="3" pageId="8" pageNumber="423">10.3</td>
<td id="57665DC4AA150009FC33FEAB365CFEE1" box="[954,1041,359,381]" gridcol="6" gridrow="3" pageId="8" pageNumber="423">10.8</td>
<td id="57665DC4AA150009FBBFFEAB36C3FEE1" box="[1078,1166,359,381]" gridcol="7" gridrow="3" pageId="8" pageNumber="423">10.5</td>
<td id="57665DC4AA150009FB3AFEAB3747FEE1" box="[1203,1290,359,381]" gridcol="8" gridrow="3" pageId="8" pageNumber="423">11.0</td>
<td id="57665DC4AA150009FAB9FEAB37CAFEE1" box="[1328,1415,359,381]" gridcol="9" gridrow="3" pageId="8" pageNumber="423">10.2</td>
</tr>
<tr id="14B734B8AA150009FF2CFE4337CAFE39" box="[165,1415,399,421]" gridrow="4" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFE433351FE39" box="[165,284,399,421]" gridcol="0" gridrow="4" pageId="8" pageNumber="423">HDW</th>
<td id="57665DC4AA150009FEC2FE4333ECFE39" box="[331,417,399,421]" gridcol="1" gridrow="4" pageId="8" pageNumber="423">10.1</td>
<td id="57665DC4AA150009FE4EFE433053FE39" box="[455,542,399,421]" gridcol="2" gridrow="4" pageId="8" pageNumber="423">10.9</td>
<td id="57665DC4AA150009FDCDFE4330D6FE39" box="[580,667,399,421]" gridcol="3" gridrow="4" pageId="8" pageNumber="423">10.4</td>
<td id="57665DC4AA150009FD48FE433155FE39" box="[705,792,399,421]" gridcol="4" gridrow="4" pageId="8" pageNumber="423">10.6</td>
<td id="57665DC4AA150009FCB4FE4331D9FE39" box="[829,916,399,421]" gridcol="5" gridrow="4" pageId="8" pageNumber="423">11.0</td>
<td id="57665DC4AA150009FC33FE43365CFE39" box="[954,1041,399,421]" gridcol="6" gridrow="4" pageId="8" pageNumber="423">10.3</td>
<td id="57665DC4AA150009FBBFFE4336C3FE39" box="[1078,1166,399,421]" gridcol="7" gridrow="4" pageId="8" pageNumber="423">10.6</td>
<td id="57665DC4AA150009FB3AFE433747FE39" box="[1203,1290,399,421]" gridcol="8" gridrow="4" pageId="8" pageNumber="423">10.5</td>
<td id="57665DC4AA150009FAB9FE4337CAFE39" box="[1328,1415,399,421]" gridcol="9" gridrow="4" pageId="8" pageNumber="423">10.3</td>
</tr>
<tr id="14B734B8AA150009FF2CFE7B37CAFE51" box="[165,1415,439,461]" gridrow="5" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFE7B3351FE51" box="[165,284,439,461]" gridcol="0" gridrow="5" pageId="8" pageNumber="423">SNT</th>
<td id="57665DC4AA150009FEC2FE7B33ECFE51" box="[331,417,439,461]" gridcol="1" gridrow="5" pageId="8" pageNumber="423">4.0</td>
<td id="57665DC4AA150009FE4EFE7B3053FE51" box="[455,542,439,461]" gridcol="2" gridrow="5" pageId="8" pageNumber="423">4.2</td>
<td id="57665DC4AA150009FDCDFE7B30D6FE51" box="[580,667,439,461]" gridcol="3" gridrow="5" pageId="8" pageNumber="423">3.8</td>
<td id="57665DC4AA150009FD48FE7B3155FE51" box="[705,792,439,461]" gridcol="4" gridrow="5" pageId="8" pageNumber="423">4.1</td>
<td id="57665DC4AA150009FCB4FE7B31D9FE51" box="[829,916,439,461]" gridcol="5" gridrow="5" pageId="8" pageNumber="423">4.2</td>
<td id="57665DC4AA150009FC33FE7B365CFE51" box="[954,1041,439,461]" gridcol="6" gridrow="5" pageId="8" pageNumber="423">4.3</td>
<td id="57665DC4AA150009FBBFFE7B36C3FE51" box="[1078,1166,439,461]" gridcol="7" gridrow="5" pageId="8" pageNumber="423">4.1</td>
<td id="57665DC4AA150009FB3AFE7B3747FE51" box="[1203,1290,439,461]" gridcol="8" gridrow="5" pageId="8" pageNumber="423">4.7</td>
<td id="57665DC4AA150009FAB9FE7B37CAFE51" box="[1328,1415,439,461]" gridcol="9" gridrow="5" pageId="8" pageNumber="423">4.3</td>
</tr>
<tr id="14B734B8AA150009FF2CFE1337CAFE69" box="[165,1415,479,501]" gridrow="6" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFE133351FE69" box="[165,284,479,501]" gridcol="0" gridrow="6" pageId="8" pageNumber="423">EYE</th>
<td id="57665DC4AA150009FEC2FE1333ECFE69" box="[331,417,479,501]" gridcol="1" gridrow="6" pageId="8" pageNumber="423">3.4</td>
<td id="57665DC4AA150009FE4EFE133053FE69" box="[455,542,479,501]" gridcol="2" gridrow="6" pageId="8" pageNumber="423">2.9</td>
<td id="57665DC4AA150009FDCDFE1330D6FE69" box="[580,667,479,501]" gridcol="3" gridrow="6" pageId="8" pageNumber="423">3.2</td>
<td id="57665DC4AA150009FD48FE133155FE69" box="[705,792,479,501]" gridcol="4" gridrow="6" pageId="8" pageNumber="423">3.4</td>
<td id="57665DC4AA150009FCB4FE1331D9FE69" box="[829,916,479,501]" gridcol="5" gridrow="6" pageId="8" pageNumber="423">3.0</td>
<td id="57665DC4AA150009FC33FE13365CFE69" box="[954,1041,479,501]" gridcol="6" gridrow="6" pageId="8" pageNumber="423">2.8</td>
<td id="57665DC4AA150009FBBFFE1336C3FE69" box="[1078,1166,479,501]" gridcol="7" gridrow="6" pageId="8" pageNumber="423">2.8</td>
<td id="57665DC4AA150009FB3AFE133747FE69" box="[1203,1290,479,501]" gridcol="8" gridrow="6" pageId="8" pageNumber="423">3.5</td>
<td id="57665DC4AA150009FAB9FE1337CAFE69" box="[1328,1415,479,501]" gridcol="9" gridrow="6" pageId="8" pageNumber="423">2.9</td>
</tr>
<tr id="14B734B8AA150009FF2CFDCB37CAFD81" box="[165,1415,519,541]" gridrow="7" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFDCB3351FD81" box="[165,284,519,541]" gridcol="0" gridrow="7" pageId="8" pageNumber="423">IOD</th>
<td id="57665DC4AA150009FEC2FDCB33ECFD81" box="[331,417,519,541]" gridcol="1" gridrow="7" pageId="8" pageNumber="423">3.4</td>
<td id="57665DC4AA150009FE4EFDCB3053FD81" box="[455,542,519,541]" gridcol="2" gridrow="7" pageId="8" pageNumber="423">-</td>
<td id="57665DC4AA150009FDCDFDCB30D6FD81" box="[580,667,519,541]" gridcol="3" gridrow="7" pageId="8" pageNumber="423">3.0</td>
<td id="57665DC4AA150009FD48FDCB3155FD81" box="[705,792,519,541]" gridcol="4" gridrow="7" pageId="8" pageNumber="423">3.2</td>
<td id="57665DC4AA150009FCB4FDCB31D9FD81" box="[829,916,519,541]" gridcol="5" gridrow="7" pageId="8" pageNumber="423">3.5</td>
<td id="57665DC4AA150009FC33FDCB365CFD81" box="[954,1041,519,541]" gridcol="6" gridrow="7" pageId="8" pageNumber="423">3.1</td>
<td id="57665DC4AA150009FBBFFDCB36C3FD81" box="[1078,1166,519,541]" gridcol="7" gridrow="7" pageId="8" pageNumber="423">3.3</td>
<td id="57665DC4AA150009FB3AFDCB3747FD81" box="[1203,1290,519,541]" gridcol="8" gridrow="7" pageId="8" pageNumber="423">3.0</td>
<td id="57665DC4AA150009FAB9FDCB37CAFD81" box="[1328,1415,519,541]" gridcol="9" gridrow="7" pageId="8" pageNumber="423">3.4</td>
</tr>
<tr id="14B734B8AA150009FF2CFDE337CAFDD9" box="[165,1415,559,581]" gridrow="8" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFDE33351FDD9" box="[165,284,559,581]" gridcol="0" gridrow="8" pageId="8" pageNumber="423">TMP</th>
<td id="57665DC4AA150009FEC2FDE333ECFDD9" box="[331,417,559,581]" gridcol="1" gridrow="8" pageId="8" pageNumber="423">1.7</td>
<td id="57665DC4AA150009FE4EFDE33053FDD9" box="[455,542,559,581]" gridcol="2" gridrow="8" pageId="8" pageNumber="423">1.8</td>
<td id="57665DC4AA150009FDCDFDE330D6FDD9" box="[580,667,559,581]" gridcol="3" gridrow="8" pageId="8" pageNumber="423">1.4</td>
<td id="57665DC4AA150009FD48FDE33155FDD9" box="[705,792,559,581]" gridcol="4" gridrow="8" pageId="8" pageNumber="423">1.8</td>
<td id="57665DC4AA150009FCB4FDE331D9FDD9" box="[829,916,559,581]" gridcol="5" gridrow="8" pageId="8" pageNumber="423">1.5</td>
<td id="57665DC4AA150009FC33FDE3365CFDD9" box="[954,1041,559,581]" gridcol="6" gridrow="8" pageId="8" pageNumber="423">1.8</td>
<td id="57665DC4AA150009FBBFFDE336C3FDD9" box="[1078,1166,559,581]" gridcol="7" gridrow="8" pageId="8" pageNumber="423">2.0</td>
<td id="57665DC4AA150009FB3AFDE33747FDD9" box="[1203,1290,559,581]" gridcol="8" gridrow="8" pageId="8" pageNumber="423">1.8</td>
<td id="57665DC4AA150009FAB9FDE337CAFDD9" box="[1328,1415,559,581]" gridcol="9" gridrow="8" pageId="8" pageNumber="423">1.7</td>
</tr>
<tr id="14B734B8AA150009FF2CFD9B37CAFDF1" box="[165,1415,599,621]" gridrow="9" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFD9B3351FDF1" box="[165,284,599,621]" gridcol="0" gridrow="9" pageId="8" pageNumber="423">TEY</th>
<td id="57665DC4AA150009FEC2FD9B33ECFDF1" box="[331,417,599,621]" gridcol="1" gridrow="9" pageId="8" pageNumber="423">1.2</td>
<td id="57665DC4AA150009FE4EFD9B3053FDF1" box="[455,542,599,621]" gridcol="2" gridrow="9" pageId="8" pageNumber="423">1.6</td>
<td id="57665DC4AA150009FDCDFD9B30D6FDF1" box="[580,667,599,621]" gridcol="3" gridrow="9" pageId="8" pageNumber="423">1.6</td>
<td id="57665DC4AA150009FD48FD9B3155FDF1" box="[705,792,599,621]" gridcol="4" gridrow="9" pageId="8" pageNumber="423">-</td>
<td id="57665DC4AA150009FCB4FD9B31D9FDF1" box="[829,916,599,621]" gridcol="5" gridrow="9" pageId="8" pageNumber="423">1.6</td>
<td id="57665DC4AA150009FC33FD9B365CFDF1" box="[954,1041,599,621]" gridcol="6" gridrow="9" pageId="8" pageNumber="423">1.4</td>
<td id="57665DC4AA150009FBBFFD9B36C3FDF1" box="[1078,1166,599,621]" gridcol="7" gridrow="9" pageId="8" pageNumber="423">1.5</td>
<td id="57665DC4AA150009FB3AFD9B3747FDF1" box="[1203,1290,599,621]" gridcol="8" gridrow="9" pageId="8" pageNumber="423">1.2</td>
<td id="57665DC4AA150009FAB9FD9B37CAFDF1" box="[1328,1415,599,621]" gridcol="9" gridrow="9" pageId="8" pageNumber="423">1.5</td>
</tr>
<tr id="14B734B8AA150009FF2CFDB337CAFD09" box="[165,1415,639,661]" gridrow="10" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFDB33351FD09" box="[165,284,639,661]" gridcol="0" gridrow="10" pageId="8" pageNumber="423">TIB</th>
<td id="57665DC4AA150009FEC2FDB333ECFD09" box="[331,417,639,661]" gridcol="1" gridrow="10" pageId="8" pageNumber="423">13.4</td>
<td id="57665DC4AA150009FE4EFDB33053FD09" box="[455,542,639,661]" gridcol="2" gridrow="10" pageId="8" pageNumber="423">14.2</td>
<td id="57665DC4AA150009FDCDFDB330D6FD09" box="[580,667,639,661]" gridcol="3" gridrow="10" pageId="8" pageNumber="423">13.5</td>
<td id="57665DC4AA150009FD48FDB33155FD09" box="[705,792,639,661]" gridcol="4" gridrow="10" pageId="8" pageNumber="423">14.3</td>
<td id="57665DC4AA150009FCB4FDB331D9FD09" box="[829,916,639,661]" gridcol="5" gridrow="10" pageId="8" pageNumber="423">13.6</td>
<td id="57665DC4AA150009FC33FDB3365CFD09" box="[954,1041,639,661]" gridcol="6" gridrow="10" pageId="8" pageNumber="423">14.5</td>
<td id="57665DC4AA150009FBBFFDB336C3FD09" box="[1078,1166,639,661]" gridcol="7" gridrow="10" pageId="8" pageNumber="423">15.0</td>
<td id="57665DC4AA150009FB3AFDB33747FD09" box="[1203,1290,639,661]" gridcol="8" gridrow="10" pageId="8" pageNumber="423">14.1</td>
<td id="57665DC4AA150009FAB9FDB337CAFD09" box="[1328,1415,639,661]" gridcol="9" gridrow="10" pageId="8" pageNumber="423">15.1</td>
</tr>
<tr id="14B734B8AA150009FF2CFD6B37CAFD21" box="[165,1415,679,701]" gridrow="11" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFD6B3351FD21" box="[165,284,679,701]" gridcol="0" gridrow="11" pageId="8" pageNumber="423">EN</th>
<td id="57665DC4AA150009FEC2FD6B33ECFD21" box="[331,417,679,701]" gridcol="1" gridrow="11" pageId="8" pageNumber="423">1.9</td>
<td id="57665DC4AA150009FE4EFD6B3053FD21" box="[455,542,679,701]" gridcol="2" gridrow="11" pageId="8" pageNumber="423">2.2</td>
<td id="57665DC4AA150009FDCDFD6B30D6FD21" box="[580,667,679,701]" gridcol="3" gridrow="11" pageId="8" pageNumber="423">1.9</td>
<td id="57665DC4AA150009FD48FD6B3155FD21" box="[705,792,679,701]" gridcol="4" gridrow="11" pageId="8" pageNumber="423">2.0</td>
<td id="57665DC4AA150009FCB4FD6B31D9FD21" box="[829,916,679,701]" gridcol="5" gridrow="11" pageId="8" pageNumber="423">2.1</td>
<td id="57665DC4AA150009FC33FD6B365CFD21" box="[954,1041,679,701]" gridcol="6" gridrow="11" pageId="8" pageNumber="423">2.4</td>
<td id="57665DC4AA150009FBBFFD6B36C3FD21" box="[1078,1166,679,701]" gridcol="7" gridrow="11" pageId="8" pageNumber="423">2.3</td>
<td id="57665DC4AA150009FB3AFD6B3747FD21" box="[1203,1290,679,701]" gridcol="8" gridrow="11" pageId="8" pageNumber="423">2.1</td>
<td id="57665DC4AA150009FAB9FD6B37CAFD21" box="[1328,1415,679,701]" gridcol="9" gridrow="11" pageId="8" pageNumber="423">2.1</td>
</tr>
<tr id="14B734B8AA150009FF2CFD0337CAFD79" box="[165,1415,719,741]" gridrow="12" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFD033351FD79" box="[165,284,719,741]" gridcol="0" gridrow="12" pageId="8" pageNumber="423">IN</th>
<td id="57665DC4AA150009FEC2FD0333ECFD79" box="[331,417,719,741]" gridcol="1" gridrow="12" pageId="8" pageNumber="423">3.1</td>
<td id="57665DC4AA150009FE4EFD033053FD79" box="[455,542,719,741]" gridcol="2" gridrow="12" pageId="8" pageNumber="423">2.9</td>
<td id="57665DC4AA150009FDCDFD0330D6FD79" box="[580,667,719,741]" gridcol="3" gridrow="12" pageId="8" pageNumber="423">2.8</td>
<td id="57665DC4AA150009FD48FD033155FD79" box="[705,792,719,741]" gridcol="4" gridrow="12" pageId="8" pageNumber="423">2.9</td>
<td id="57665DC4AA150009FCB4FD0331D9FD79" box="[829,916,719,741]" gridcol="5" gridrow="12" pageId="8" pageNumber="423">2.8</td>
<td id="57665DC4AA150009FC33FD03365CFD79" box="[954,1041,719,741]" gridcol="6" gridrow="12" pageId="8" pageNumber="423">3.0</td>
<td id="57665DC4AA150009FBBFFD0336C3FD79" box="[1078,1166,719,741]" gridcol="7" gridrow="12" pageId="8" pageNumber="423">2.8</td>
<td id="57665DC4AA150009FB3AFD033747FD79" box="[1203,1290,719,741]" gridcol="8" gridrow="12" pageId="8" pageNumber="423">3.0</td>
<td id="57665DC4AA150009FAB9FD0337CAFD79" box="[1328,1415,719,741]" gridcol="9" gridrow="12" pageId="8" pageNumber="423">3.0</td>
</tr>
<tr id="14B734B8AA150009FF2CFD3B37CAFC91" box="[165,1415,759,781]" gridrow="13" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFD3B3351FC91" box="[165,284,759,781]" gridcol="0" gridrow="13" pageId="8" pageNumber="423">NS</th>
<td id="57665DC4AA150009FEC2FD3B33ECFC91" box="[331,417,759,781]" gridcol="1" gridrow="13" pageId="8" pageNumber="423">1.9</td>
<td id="57665DC4AA150009FE4EFD3B3053FC91" box="[455,542,759,781]" gridcol="2" gridrow="13" pageId="8" pageNumber="423">2.0</td>
<td id="57665DC4AA150009FDCDFD3B30D6FC91" box="[580,667,759,781]" gridcol="3" gridrow="13" pageId="8" pageNumber="423">2.1</td>
<td id="57665DC4AA150009FD48FD3B3155FC91" box="[705,792,759,781]" gridcol="4" gridrow="13" pageId="8" pageNumber="423">2.1</td>
<td id="57665DC4AA150009FCB4FD3B31D9FC91" box="[829,916,759,781]" gridcol="5" gridrow="13" pageId="8" pageNumber="423">1.9</td>
<td id="57665DC4AA150009FC33FD3B365CFC91" box="[954,1041,759,781]" gridcol="6" gridrow="13" pageId="8" pageNumber="423">2.0</td>
<td id="57665DC4AA150009FBBFFD3B36C3FC91" box="[1078,1166,759,781]" gridcol="7" gridrow="13" pageId="8" pageNumber="423">1.9</td>
<td id="57665DC4AA150009FB3AFD3B3747FC91" box="[1203,1290,759,781]" gridcol="8" gridrow="13" pageId="8" pageNumber="423">2.2</td>
<td id="57665DC4AA150009FAB9FD3B37CAFC91" box="[1328,1415,759,781]" gridcol="9" gridrow="13" pageId="8" pageNumber="423">2.1</td>
</tr>
<tr id="14B734B8AA150009FF2CFCD337CAFCA9" box="[165,1415,799,821]" gridrow="14" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFCD33351FCA9" box="[165,284,799,821]" gridcol="0" gridrow="14" pageId="8" pageNumber="423">ML</th>
<td id="57665DC4AA150009FEC2FCD333ECFCA9" box="[331,417,799,821]" gridcol="1" gridrow="14" pageId="8" pageNumber="423">7.4</td>
<td id="57665DC4AA150009FE4EFCD33053FCA9" box="[455,542,799,821]" gridcol="2" gridrow="14" pageId="8" pageNumber="423">7.5</td>
<td id="57665DC4AA150009FDCDFCD330D6FCA9" box="[580,667,799,821]" gridcol="3" gridrow="14" pageId="8" pageNumber="423">6.9</td>
<td id="57665DC4AA150009FD48FCD33155FCA9" box="[705,792,799,821]" gridcol="4" gridrow="14" pageId="8" pageNumber="423">7.9</td>
<td id="57665DC4AA150009FCB4FCD331D9FCA9" box="[829,916,799,821]" gridcol="5" gridrow="14" pageId="8" pageNumber="423">7.3</td>
<td id="57665DC4AA150009FC33FCD3365CFCA9" box="[954,1041,799,821]" gridcol="6" gridrow="14" pageId="8" pageNumber="423">7.4</td>
<td id="57665DC4AA150009FBBFFCD336C3FCA9" box="[1078,1166,799,821]" gridcol="7" gridrow="14" pageId="8" pageNumber="423">8.3</td>
<td id="57665DC4AA150009FB3AFCD33747FCA9" box="[1203,1290,799,821]" gridcol="8" gridrow="14" pageId="8" pageNumber="423">7.8</td>
<td id="57665DC4AA150009FAB9FCD337CAFCA9" box="[1328,1415,799,821]" gridcol="9" gridrow="14" pageId="8" pageNumber="423">8.1</td>
</tr>
<tr id="14B734B8AA150009FF2CFC8B37CAFCC1" box="[165,1415,839,861]" gridrow="15" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFC8B3351FCC1" box="[165,284,839,861]" gridcol="0" gridrow="15" pageId="8" pageNumber="423">PL</th>
<td id="57665DC4AA150009FEC2FC8B33ECFCC1" box="[331,417,839,861]" gridcol="1" gridrow="15" pageId="8" pageNumber="423">14.3</td>
<td id="57665DC4AA150009FE4EFC8B3053FCC1" box="[455,542,839,861]" gridcol="2" gridrow="15" pageId="8" pageNumber="423">14.4</td>
<td id="57665DC4AA150009FDCDFC8B30D6FCC1" box="[580,667,839,861]" gridcol="3" gridrow="15" pageId="8" pageNumber="423">13.2</td>
<td id="57665DC4AA150009FD48FC8B3155FCC1" box="[705,792,839,861]" gridcol="4" gridrow="15" pageId="8" pageNumber="423">14.1</td>
<td id="57665DC4AA150009FCB4FC8B31D9FCC1" box="[829,916,839,861]" gridcol="5" gridrow="15" pageId="8" pageNumber="423">14.8</td>
<td id="57665DC4AA150009FC33FC8B365CFCC1" box="[954,1041,839,861]" gridcol="6" gridrow="15" pageId="8" pageNumber="423">14.3</td>
<td id="57665DC4AA150009FBBFFC8B36C3FCC1" box="[1078,1166,839,861]" gridcol="7" gridrow="15" pageId="8" pageNumber="423">15.2</td>
<td id="57665DC4AA150009FB3AFC8B3747FCC1" box="[1203,1290,839,861]" gridcol="8" gridrow="15" pageId="8" pageNumber="423">14.5</td>
<td id="57665DC4AA150009FAB9FC8B37CAFCC1" box="[1328,1415,839,861]" gridcol="9" gridrow="15" pageId="8" pageNumber="423">15.0</td>
</tr>
<tr id="14B734B8AA150009FF2CFCA337CAFC19" box="[165,1415,879,901]" gridrow="16" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFCA33351FC19" box="[165,284,879,901]" gridcol="0" gridrow="16" pageId="8" pageNumber="423">HDL/HDW</th>
<td id="57665DC4AA150009FEC2FCA333ECFC19" box="[331,417,879,901]" gridcol="1" gridrow="16" pageId="8" pageNumber="423">1.00</td>
<td id="57665DC4AA150009FE4EFCA33053FC19" box="[455,542,879,901]" gridcol="2" gridrow="16" pageId="8" pageNumber="423">0.99</td>
<td id="57665DC4AA150009FDCDFCA330D6FC19" box="[580,667,879,901]" gridcol="3" gridrow="16" pageId="8" pageNumber="423">1.00</td>
<td id="57665DC4AA150009FD48FCA33155FC19" box="[705,792,879,901]" gridcol="4" gridrow="16" pageId="8" pageNumber="423">0.95</td>
<td id="57665DC4AA150009FCB4FCA331D9FC19" box="[829,916,879,901]" gridcol="5" gridrow="16" pageId="8" pageNumber="423">0.94</td>
<td id="57665DC4AA150009FC33FCA3365CFC19" box="[954,1041,879,901]" gridcol="6" gridrow="16" pageId="8" pageNumber="423">1.04</td>
<td id="57665DC4AA150009FBBFFCA336C3FC19" box="[1078,1166,879,901]" gridcol="7" gridrow="16" pageId="8" pageNumber="423">0.99</td>
<td id="57665DC4AA150009FB3AFCA33747FC19" box="[1203,1290,879,901]" gridcol="8" gridrow="16" pageId="8" pageNumber="423">1.05</td>
<td id="57665DC4AA150009FAB9FCA337CAFC19" box="[1328,1415,879,901]" gridcol="9" gridrow="16" pageId="8" pageNumber="423">0.99</td>
</tr>
<tr id="14B734B8AA150009FF2CFC5B37CAFC31" box="[165,1415,919,941]" gridrow="17" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFC5B3351FC31" box="[165,284,919,941]" gridcol="0" gridrow="17" pageId="8" pageNumber="423">HDL/SVL</th>
<td id="57665DC4AA150009FEC2FC5B33ECFC31" box="[331,417,919,941]" gridcol="1" gridrow="17" pageId="8" pageNumber="423">0.35</td>
<td id="57665DC4AA150009FE4EFC5B3053FC31" box="[455,542,919,941]" gridcol="2" gridrow="17" pageId="8" pageNumber="423">0.33</td>
<td id="57665DC4AA150009FDCDFC5B30D6FC31" box="[580,667,919,941]" gridcol="3" gridrow="17" pageId="8" pageNumber="423">0.35</td>
<td id="57665DC4AA150009FD48FC5B3155FC31" box="[705,792,919,941]" gridcol="4" gridrow="17" pageId="8" pageNumber="423">0.33</td>
<td id="57665DC4AA150009FCB4FC5B31D9FC31" box="[829,916,919,941]" gridcol="5" gridrow="17" pageId="8" pageNumber="423">0.36</td>
<td id="57665DC4AA150009FC33FC5B365CFC31" box="[954,1041,919,941]" gridcol="6" gridrow="17" pageId="8" pageNumber="423">0.37</td>
<td id="57665DC4AA150009FBBFFC5B36C3FC31" box="[1078,1166,919,941]" gridcol="7" gridrow="17" pageId="8" pageNumber="423">0.34</td>
<td id="57665DC4AA150009FB3AFC5B3747FC31" box="[1203,1290,919,941]" gridcol="8" gridrow="17" pageId="8" pageNumber="423">0.35</td>
<td id="57665DC4AA150009FAB9FC5B37CAFC31" box="[1328,1415,919,941]" gridcol="9" gridrow="17" pageId="8" pageNumber="423">0.34</td>
</tr>
<tr id="14B734B8AA150009FF2CFC7337CAFB81" box="[165,1415,959,1053]" gridrow="18" pageId="8" pageNumber="423">
<th id="57665DC4AA150009FF2CFC733351FB81" box="[165,284,959,1053]" gridcol="0" gridrow="18" pageId="8" pageNumber="423">TIB/SVL Weight in life (g)</th>
<td id="57665DC4AA150009FEC2FC7333ECFB81" box="[331,417,959,1053]" gridcol="1" gridrow="18" pageId="8" pageNumber="423">0.46 2.4</td>
<td id="57665DC4AA150009FE4EFC733053FB81" box="[455,542,959,1053]" gridcol="2" gridrow="18" pageId="8" pageNumber="423">0.44 2.8</td>
<td id="57665DC4AA150009FDCDFC7330D6FB81" box="[580,667,959,1053]" gridcol="3" gridrow="18" pageId="8" pageNumber="423">0.46 2.2</td>
<td id="57665DC4AA150009FD48FC733155FB81" box="[705,792,959,1053]" gridcol="4" gridrow="18" pageId="8" pageNumber="423">0.47 2.5</td>
<td id="57665DC4AA150009FCB4FC7331D9FB81" box="[829,916,959,1053]" gridcol="5" gridrow="18" pageId="8" pageNumber="423">0.47 2.4</td>
<td id="57665DC4AA150009FC33FC73365CFB81" box="[954,1041,959,1053]" gridcol="6" gridrow="18" pageId="8" pageNumber="423">0.50 2.7</td>
<td id="57665DC4AA150009FBBFFC7336C3FB81" box="[1078,1166,959,1053]" gridcol="7" gridrow="18" pageId="8" pageNumber="423">0.49 2.3</td>
<td id="57665DC4AA150009FB3AFC733747FB81" box="[1203,1290,959,1053]" gridcol="8" gridrow="18" pageId="8" pageNumber="423">0.44 3.0</td>
<td id="57665DC4AA150009FAB9FC7337CAFB81" box="[1328,1415,959,1053]" gridcol="9" gridrow="18" pageId="8" pageNumber="423">0.50 2.3</td>
</tr>
</table>
</paragraph>
<paragraph id="AA3836FAAA15FFFEFF4EFB94312EFB40" blockId="8.[151,1436,1112,2037]" pageId="8" pageNumber="423">
<emphasis id="98F3EAE8AA15FFFEFF4EFB94337AFBED" bold="true" box="[199,311,1112,1137]" pageId="8" pageNumber="423">Variation</emphasis>
: Measurements of the 
<typeStatus id="753C8858AA15FFFEFDB3FB953027FBED" box="[570,618,1113,1137]" pageId="8" pageNumber="423">type</typeStatus>
series are shown in 
<tableCitation id="E7050341AA15FFFEFCD8FB9431E9FBEC" box="[849,932,1112,1137]" captionStart="TABLE 1" captionStartId="8.[151,239,152,175]" captionTargetBox="[165,1415,205,1053]" captionTargetPageId="8" captionText="TABLE 1. Measurements (mm) of Leptolalax botsfordi sp. nov. Abbreviations defined in text." httpUri="http://table.plazi.org/id/FEF86672AA15FFFEFF1EFF5436C6FF33" pageId="8" pageNumber="423" tableUuid="FEF86672AA15FFFEFF1EFF5436C6FF33">Table 1</tableCitation>
and representative photographs of 
<typeStatus id="753C8858AA15FFFEFAB8FB9537D1FBED" box="[1329,1436,1113,1137]" pageId="8" pageNumber="423" type="paratype">paratypes</typeStatus>
are shown in 
<figureCitation id="32BC2A7FAA15FFFEFEA6FBB033F4FB08" box="[303,441,1148,1173]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="2.[151,255,1693,1716]" captionStartId-1="3.[151,255,1941,1964]" captionStartId-2="4.[151,255,1870,1893]" captionTargetBox-0="[151,1429,781,1670]" captionTargetBox-1="[151,1436,1045,1919]" captionTargetBox-2="[151,1407,817,1828]" captionTargetId-0="figure@2.[151,1436,774,1671]" captionTargetId-1="figure@3.[151,1436,1045,1920]" captionTargetId-2="figure@4.[151,1436,807,1846]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 2. Leptolalax botsfordi sp. nov. (A) dorsal and (B) ventral view of adult male holotype VNMN 0 3682 in life, (C) ventral view of gravid female paratype AMS R 176540 in life, (D) posterior view of adult male holotype VNMN 0 3682 in life, showing large femoral glands in (not to scale), (E) dorsal and (F) ventral view of adult male holotype VNMN 0 3682 in preservative, and (G) ventral view of adult female paratype AMS R 176540 in preservative. Scale bars = 5 mm." captionText-1="FIGURE 3. Adult female Leptolalax botsfordi sp. nov. AMS R 176540 in life. Note metallic copper flecks, particularly along snout, supratympanic fold and flanks." captionText-2="FIGURE 4. Leptolalax botsfordi sp. nov. in life. (A) adult male holotype VNMN 03682; (B) adult female AMS R 176540; (C) adult male AMS R 176539; (D) female AMS R 176541. Note robust arms of males." httpUri-0="https://zenodo.org/record/219356/files/figure.png" httpUri-1="https://zenodo.org/record/219357/files/figure.png" httpUri-2="https://zenodo.org/record/219358/files/figure.png" pageId="8" pageNumber="423">Figures 2–4</figureCitation>
. Specimens vary only slightly in the colour and extent of marbling on the dorsum; male AMS R 176537 was slightly paler pinkish brown in life. Males have more robust forearms than females (compare males in figures 4A and 4C with females in figures 4B and 4D).
</paragraph>
<paragraph id="AA3836FAAA15FFFEFF4EFB24303EF965" blockId="8.[151,1436,1112,2037]" pageId="8" pageNumber="423">
<emphasis id="98F3EAE8AA15FFFEFF4EFB2433E6FA9D" bold="true" box="[199,427,1256,1281]" pageId="8" pageNumber="423">Advertisement call</emphasis>
: Call descriptions are based on the calls of the 
<typeStatus id="753C8858AA15FFFEFC68FB24360BFA9D" box="[993,1094,1256,1281]" pageId="8" pageNumber="423" type="holotype">holotype</typeStatus>
, recorded at 14.0 ºC ambient temperature. Calls were of two distinct 
<typeStatus id="753C8858AA15FFFEFDEDFAC130D2FAB9" box="[612,671,1293,1317]" pageId="8" pageNumber="423">types</typeStatus>
(
<figureCitation id="32BC2A7FAA15FFFEFD38FAC0315BFAB9" box="[689,790,1292,1317]" captionStart="FIGURE 6" captionStartId="6.[151,255,1434,1457]" captionTargetBox="[167,1422,454,1403]" captionTargetId="figure@6.[151,1436,446,1413]" captionTargetPageId="6" captionText="FIGURE 6. Advertisement call of Leptolalax botsfordi sp. nov. holotype VNMN 0 3682 (A) 14 s waveform of relative amplitude (Rel. amp.) over time for 16 “ primary ” calls, (B) waveform and (C) corresponding spectrogram of single representative “ secondary ” call expanded from first section shown in A, and (D) waveform and (E) corresponding spectrogram of single representative “ primary ” call expanded from second section shown in A; all recorded at an ambient air temperature of 14.0 ºC." httpUri="https://zenodo.org/record/219360/files/figure.png" pageId="8" pageNumber="423">Figure 6</figureCitation>
), the more frequent and higher amplitude clicking calls referred to here as ‘primary calls’, and low amplitude ‘rasps’ referred to as ‘secondary calls’. Primary calls were an average of 267 ms in duration and consistently comprised of three notes (
<tableCitation id="E7050341AA15FFFEFC67FA98360AFAF0" box="[1006,1095,1364,1389]" captionStart="TABLE 2" captionStartId="9.[151,239,580,603]" captionTargetBox="[165,1162,667,1089]" captionTargetPageId="9" captionText="TABLE 2. Measurements of advertisement call parameters for adult male holotype (VNMN 03682) of Leptolalax botsfordi sp. nov. recorded at 14.0 ºC ambient temperature. Parameter values are given as means (and ranges)." httpUri="http://table.plazi.org/id/FEF86672AA14FFFFFF1EFD88377DFDE7" pageId="8" pageNumber="423" tableUuid="FEF86672AA14FFFFFF1EFD88377DFDE7">Table 2</tableCitation>
, 
<figureCitation id="32BC2A7FAA15FFFEFBDFFA9836F4FAF1" box="[1110,1209,1364,1389]" captionStart="FIGURE 6" captionStartId="6.[151,255,1434,1457]" captionTargetBox="[167,1422,454,1403]" captionTargetId="figure@6.[151,1436,446,1413]" captionTargetPageId="6" captionText="FIGURE 6. Advertisement call of Leptolalax botsfordi sp. nov. holotype VNMN 0 3682 (A) 14 s waveform of relative amplitude (Rel. amp.) over time for 16 “ primary ” calls, (B) waveform and (C) corresponding spectrogram of single representative “ secondary ” call expanded from first section shown in A, and (D) waveform and (E) corresponding spectrogram of single representative “ primary ” call expanded from second section shown in A; all recorded at an ambient air temperature of 14.0 ºC." httpUri="https://zenodo.org/record/219360/files/figure.png" pageId="8" pageNumber="423">Figure 6</figureCitation>
). Calls were highly amplitude modulated, with amplitude peaking at the start of each note. Notes contained a variable number of pulses repeated at a highly variable rate (average ~86 pulses/s, range ~40–146 pulses/s). The number of pulses decreased from note one to note three, with notes two and three almost always containing 4 and 2 pulses respectively. The dominant frequency of the primary call was 2.6–3.2 kHz, and harmonics were weakly present at approximately 5.0–5.5 kHz. A fundamental frequency was not visible. Primary calls were repeated at a rate of approximately 1.3 calls per second, and had an average intercall interval of 
<geoCoordinate id="CFB3503DAA15FFFEFCBFF9E0313DF9D8" box="[822,880,1580,1605]" direction="south" orientation="latitude" pageId="8" pageNumber="423" precision="5555" value="-0.5">0.5 s</geoCoordinate>
. Secondary calls were an average of 299 ms in duration and consisted of a single note with 14–19 irregularly-spaced pulses (
<tableCitation id="E7050341AA15FFFEFB82F99C3629F9F4" box="[1035,1124,1616,1641]" captionStart="TABLE 2" captionStartId="9.[151,239,580,603]" captionTargetBox="[165,1162,667,1089]" captionTargetPageId="9" captionText="TABLE 2. Measurements of advertisement call parameters for adult male holotype (VNMN 03682) of Leptolalax botsfordi sp. nov. recorded at 14.0 ºC ambient temperature. Parameter values are given as means (and ranges)." httpUri="http://table.plazi.org/id/FEF86672AA14FFFFFF1EFD88377DFDE7" pageId="8" pageNumber="423" tableUuid="FEF86672AA14FFFFFF1EFD88377DFDE7">Table 2</tableCitation>
, 
<figureCitation id="32BC2A7FAA15FFFEFBF8F99C369EF9F5" box="[1137,1235,1616,1641]" captionStart="FIGURE 6" captionStartId="6.[151,255,1434,1457]" captionTargetBox="[167,1422,454,1403]" captionTargetId="figure@6.[151,1436,446,1413]" captionTargetPageId="6" captionText="FIGURE 6. Advertisement call of Leptolalax botsfordi sp. nov. holotype VNMN 0 3682 (A) 14 s waveform of relative amplitude (Rel. amp.) over time for 16 “ primary ” calls, (B) waveform and (C) corresponding spectrogram of single representative “ secondary ” call expanded from first section shown in A, and (D) waveform and (E) corresponding spectrogram of single representative “ primary ” call expanded from second section shown in A; all recorded at an ambient air temperature of 14.0 ºC." httpUri="https://zenodo.org/record/219360/files/figure.png" pageId="8" pageNumber="423">Figure 6</figureCitation>
). Secondary calls were of less amplitude than primary calls, and not strongly amplitude modulated. The dominant frequency of the secondary call was slightly higher than the primary call (2.8–3.4 kHz). Due to background noise in the recording, full analysis of secondary calls was not possible. To the human ear, the advertisement call of 
<taxonomicName id="6D874D79AA15FFFEFB2BF971376EF948" box="[1186,1315,1724,1748]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="423" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA15FFFEFB2BF971376EF948" box="[1186,1315,1724,1748]" italics="true" pageId="8" pageNumber="423">L. botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA15FFFEFAA3F97037CEF949" box="[1322,1411,1724,1749]" pageId="8" pageNumber="423" rank="species">
<emphasis id="98F3EAE8AA15FFFEFAA3F97037CEF949" bold="true" box="[1322,1411,1724,1749]" pageId="8" pageNumber="423">sp. nov.</emphasis>
</taxonomicNameLabel>
is a weak chirping, similar to an orthopteran.
</paragraph>
<paragraph id="AA3836FAAA15FFFFFF4EF8C83100FE80" blockId="8.[151,1436,1112,2037]" lastBlockId="9.[151,1436,151,536]" lastPageId="9" lastPageNumber="424" pageId="8" pageNumber="423">
<emphasis id="98F3EAE8AA15FFFEFF4EF8C83362F881" bold="true" box="[199,303,1796,1821]" pageId="8" pageNumber="423">Ecology:</emphasis>
All specimens of 
<taxonomicName id="6D874D79AA15FFFEFE77F8C830AFF880" box="[510,738,1796,1820]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="423" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA15FFFEFE77F8C830AFF880" box="[510,738,1796,1820]" italics="true" pageId="8" pageNumber="423">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA15FFFEFD63F8C83108F881" box="[746,837,1796,1821]" pageId="8" pageNumber="423" rank="species">
<emphasis id="98F3EAE8AA15FFFEFD63F8C83108F881" bold="true" box="[746,837,1796,1821]" pageId="8" pageNumber="423">sp. nov.</emphasis>
</taxonomicNameLabel>
were found in upper montane forest between 
<quantity id="6D7F9B1FAA15FFFEFAC6F8C832B9F8DC" metricMagnitude="3" metricUnit="m" metricValue="2.805" metricValueMax="2.815" metricValueMin="2.795" pageId="8" pageNumber="423" unit="m" value="2805.0" valueMax="2815.0" valueMin="2795.0">2,795– 2,815 m</quantity>
elevation on Mount Fansipan (
<figureCitation id="32BC2A7FAA15FFFEFDD3F8E430F0F8DC" box="[602,701,1832,1857]" captionStart="FIGURE 7" captionStartId="7.[151,255,1894,1917]" captionTargetBox="[151,1436,209,1872]" captionTargetId="figure@7.[151,1436,194,1872]" captionTargetPageId="7" captionText="FIGURE 7. Habitat of Leptolalax botsfordi sp. nov. (A) View of Mount Fansipan and surrounding mountains, Lao Cai Province, Vietnam. (B) Microhabitat of calling male L. botsfordi sp. nov. at ~ 2,800 m elevation." httpUri="https://zenodo.org/record/219361/files/figure.png" pageId="8" pageNumber="423">Figure 7</figureCitation>
A). The climate of Mount Fansipan is almost temperate, with an average temperature of about 15 ºC (range -3–20 ºC), with frequent freezing in the coldest months (Nguyen &amp; Harder 1996). The mountain has 
<quantity id="6D7F9B1FAA15FFFEFD9CF8BC30C7F814" box="[533,650,1904,1929]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="423" unit="mm" value="3500.0">3,500 mm</quantity>
of rain and 2–3 snow days per year, and no dry months (Nguyen &amp; Harder 1996; Nguyen 
<emphasis id="98F3EAE8AA15FFFEFE15F859339CF830" box="[412,465,1940,1964]" italics="true" pageId="8" pageNumber="423">et al</emphasis>
. 2000). In addition to the heavy rainfall, fog is common and evaporation rates low (Nguyen 1998). Several males were heard faintly calling under leaf litter adjacent to a small stream (
<figureCitation id="32BC2A7FAA15FFFEFB7BF874371CF84C" box="[1266,1361,1976,2001]" captionStart="FIGURE 7" captionStartId="7.[151,255,1894,1917]" captionTargetBox="[151,1436,209,1872]" captionTargetId="figure@7.[151,1436,194,1872]" captionTargetPageId="7" captionText="FIGURE 7. Habitat of Leptolalax botsfordi sp. nov. (A) View of Mount Fansipan and surrounding mountains, Lao Cai Province, Vietnam. (B) Microhabitat of calling male L. botsfordi sp. nov. at ~ 2,800 m elevation." httpUri="https://zenodo.org/record/219361/files/figure.png" pageId="8" pageNumber="423">Figure 7</figureCitation>
B) and females were located whilst moving towards calling males. The species was very difficult to locate, even when calling. During surveys conducted within the same week at lower elevations (
<quantity id="6D7F9B1FAA14FFFFFB8DFF5B36FDFF2C" box="[1028,1200,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.7975" metricValueMax="2.2399999999999998" metricValueMin="1.355" pageId="9" pageNumber="424" unit="m" value="1797.5" valueMax="2240.0" valueMin="1355.0">1,355–2,240 m</quantity>
) on Mount Fansipan and surrounding habitat in Hoang Lien National Park and Sa Pa, District, 
<taxonomicName id="6D874D79AA14FFFFFC66FF703699FF48" box="[1007,1236,188,212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA14FFFFFC66FF703699FF48" box="[1007,1236,188,212]" italics="true" pageId="9" pageNumber="424">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA14FFFFFB54FF703777FF49" box="[1245,1338,188,213]" pageId="9" pageNumber="424" rank="species">
<emphasis id="98F3EAE8AA14FFFFFB54FF703777FF49" bold="true" box="[1245,1338,188,213]" pageId="9" pageNumber="424">sp. nov.</emphasis>
</taxonomicNameLabel>
was not detected, although the two other 
<taxonomicName id="6D874D79AA14FFFFFD9DFF2C30C1FF64" box="[532,652,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="genus">
<emphasis id="98F3EAE8AA14FFFFFD9DFF2C30C1FF64" box="[532,652,224,248]" italics="true" pageId="9" pageNumber="424">Leptolalax</emphasis>
</taxonomicName>
species known from Mount Fansipan and surrounds (Bourret 1937; Ohler 
<emphasis id="98F3EAE8AA14FFFFFF54FEC9335BFE80" box="[221,278,260,284]" italics="true" pageId="9" pageNumber="424">et al.</emphasis>
2000), 
<taxonomicName id="6D874D79AA14FFFFFEE5FEC933A9FE80" box="[364,484,260,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="bourreti">
<emphasis id="98F3EAE8AA14FFFFFEE5FEC933A9FE80" box="[364,484,260,284]" italics="true" pageId="9" pageNumber="424">L. bourreti</emphasis>
</taxonomicName>
and 
<emphasis id="98F3EAE8AA14FFFFFD92FEC930EDFE80" box="[539,672,260,284]" italics="true" pageId="9" pageNumber="424">
<taxonomicName id="6D874D79AA14FFFFFD92FEC930D1FE80" box="[539,668,260,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="pluvialis">L. pluvialis</taxonomicName>
,
</emphasis>
were common.
</paragraph>
</subSubSection>
<subSubSection id="E29D6571AA14FFFFFF4EFEEB30D3FA9C" pageId="9" pageNumber="424" type="distribution">
<paragraph id="AA3836FAAA14FFFFFF4EFEEB3158FD84" blockId="9.[151,1436,151,536]" pageId="9" pageNumber="424">
<emphasis id="98F3EAE8AA14FFFFFF4EFEEB3314FEDC" bold="true" box="[199,345,295,320]" pageId="9" pageNumber="424">Distribution</emphasis>
: 
<taxonomicName id="6D874D79AA14FFFFFEE1FEE43007FEDC" box="[360,586,296,320]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA14FFFFFEE1FEE43007FEDC" box="[360,586,296,320]" italics="true" pageId="9" pageNumber="424">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA14FFFFFDD8FEEB30E6FEDC" box="[593,683,295,320]" pageId="9" pageNumber="424" rank="species">
<emphasis id="98F3EAE8AA14FFFFFDD8FEEB30E6FEDC" bold="true" box="[593,683,295,320]" pageId="9" pageNumber="424">sp. nov.</emphasis>
</taxonomicNameLabel>
is only known from relatively near the summit of Mount Fansipan, northern 
<collectingCountry id="D290766AAA14FFFFFF76FE80332FFEF9" box="[255,354,332,357]" name="Vietnam" pageId="9" pageNumber="424">Vietnam</collectingCountry>
. Mount Fansipan is the highest point in Indochina (
<quantity id="6D7F9B1FAA14FFFFFC3DFE80365AFEF8" box="[948,1047,332,356]" metricMagnitude="3" metricUnit="m" metricValue="3.143" pageId="9" pageNumber="424" unit="m" value="3143.0">3,143 m</quantity>
), and part of the Hoang Lien Son Mountain Range, which extends south from the Ailao Shan mountains in 
<collectingCountry id="D290766AAA14FFFFFC41FEA33643FE14" box="[968,1038,367,392]" name="China" pageId="9" pageNumber="424">China</collectingCountry>
, the south-easternmost extension of the Himalayan chain. The actual distribution of the new species is unknown but may extend along the range northwest through 
<collectingCountry id="D290766AAA14FFFFFEE7FE7433D7FE53" box="[366,410,440,463]" name="Laos" pageId="9" pageNumber="424">Lao</collectingCountry>
Cai and Lai Chai provinces and into Yunnan Province in 
<collectingCountry id="D290766AAA14FFFFFBBEFE7B3631FE4C" box="[1079,1148,439,464]" name="China" pageId="9" pageNumber="424">China</collectingCountry>
, or southeast further into 
<collectingCountry id="D290766AAA14FFFFFF1EFE1032B7FE69" box="[151,250,476,501]" name="Vietnam" pageId="9" pageNumber="424">Vietnam</collectingCountry>
. However, there are only a handful of small, isolated areas along the range above 
<quantity id="6D7F9B1FAA14FFFFFB71FE103716FE68" box="[1272,1371,476,500]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="424" unit="m" value="2700.0">2,700 m</quantity>
until approximately 
<quantity id="6D7F9B1FAA14FFFFFEB6FE3333D8FD84" box="[319,405,511,536]" metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="424" unit="km" value="250.0">250 km</quantity>
further north in Yunnan Province.
</paragraph>
<caption id="FEF86672AA14FFFFFF1EFD88377DFDE7" ID-Table-UUID="FEF86672AA14FFFFFF1EFD88377DFDE7" httpUri="http://table.plazi.org/id/FEF86672AA14FFFFFF1EFD88377DFDE7" pageId="9" pageNumber="424" targetBox="[165,1162,667,1089]" targetIsTable="true" targetPageId="9">
<paragraph id="AA3836FAAA14FFFFFF1EFD88377DFDE7" blockId="9.[151,1436,579,635]" pageId="9" pageNumber="424">
<emphasis id="98F3EAE8AA14FFFFFF1EFD883340FDC7" bold="true" box="[151,269,580,603]" pageId="9" pageNumber="424">TABLE 2.</emphasis>
Measurements of advertisement call parameters for adult male holotype (VNMN 03682) of 
<taxonomicName id="6D874D79AA14FFFFFAA3FD8F32BBFDE7" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA14FFFFFAA3FD8F32BBFDE7" italics="true" pageId="9" pageNumber="424">Leptolalax botsfordi</emphasis>
</taxonomicName>
<emphasis id="98F3EAE8AA14FFFFFF74FDA8331FFDE7" bold="true" box="[253,338,612,635]" pageId="9" pageNumber="424">
<taxonomicNameLabel id="83C05793AA14FFFFFF74FDA8331FFDE7" box="[253,338,612,635]" pageId="9" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
recorded at 14.0 ºC ambient temperature. Parameter values are given as means (and ranges).
</paragraph>
</caption>
<paragraph id="AA3836FAAA14FFFFFDE5FD573085FBDD" pageId="9" pageNumber="424">
<table id="D887C45AAA140009FF2CFD5736C7FBDD" box="[165,1162,667,1089]" gridcols="3" gridrows="11" pageId="9" pageNumber="424">
<tr id="14B734B8AA140009FF2CFD5736C7FD2D" box="[165,1162,667,689]" gridrow="0" pageId="9" pageNumber="424" rowspan-0="1">
<th id="57665DC4AA140009FDE5FD573143FD2D" box="[620,782,667,689]" gridcol="1" gridrow="0" pageId="9" pageNumber="424">
<emphasis id="98F3EAE8AA14FFFFFDE5FD5730BEFD2D" bold="true" box="[620,755,667,689]" pageId="9" pageNumber="424">Primary call</emphasis>
</th>
<th id="57665DC4AA140009FC65FD5736C7FD2D" box="[1004,1162,667,689]" gridcol="2" gridrow="0" pageId="9" pageNumber="424">
<emphasis id="98F3EAE8AA14FFFFFC65FD5736C7FD2D" bold="true" box="[1004,1162,667,689]" pageId="9" pageNumber="424">Secondary call</emphasis>
</th>
</tr>
<tr id="14B734B8AA140009FF2CFD0E36C7FD45" box="[165,1162,706,729]" gridrow="1" pageId="9" pageNumber="424">
<th id="57665DC4AA140009FF2CFD0E33F4FD45" box="[165,441,706,729]" gridcol="0" gridrow="1" pageId="9" pageNumber="424">Number of calls measured</th>
<td id="57665DC4AA140009FDE5FD0E3143FD45" box="[620,782,706,729]" gridcol="1" gridrow="1" pageId="9" pageNumber="424">10</td>
<td id="57665DC4AA140009FC65FD0E36C7FD45" box="[1004,1162,706,729]" gridcol="2" gridrow="1" pageId="9" pageNumber="424">5</td>
</tr>
<tr id="14B734B8AA140009FF2CFD2636C7FC9D" box="[165,1162,746,769]" gridrow="2" pageId="9" pageNumber="424">
<th id="57665DC4AA140009FF2CFD2633F4FC9D" box="[165,441,746,769]" gridcol="0" gridrow="2" pageId="9" pageNumber="424">Call duration (ms)</th>
<td id="57665DC4AA140009FDE5FD263143FC9D" box="[620,782,746,769]" gridcol="1" gridrow="2" pageId="9" pageNumber="424">267 (239–303)</td>
<td id="57665DC4AA140009FC65FD2636C7FC9D" box="[1004,1162,746,769]" gridcol="2" gridrow="2" pageId="9" pageNumber="424">229 (194–283)</td>
</tr>
<tr id="14B734B8AA140009FF2CFCDE36C7FCB5" box="[165,1162,786,809]" gridrow="3" pageId="9" pageNumber="424" rowspan-2="1">
<th id="57665DC4AA140009FF2CFCDE33F4FCB5" box="[165,441,786,809]" gridcol="0" gridrow="3" pageId="9" pageNumber="424">Call repetition rate (calls/s)</th>
<td id="57665DC4AA140009FDE5FCDE3143FCB5" box="[620,782,786,809]" gridcol="1" gridrow="3" pageId="9" pageNumber="424">1.3</td>
</tr>
<tr id="14B734B8AA140009FF2CFCF636C7FCCD" box="[165,1162,826,849]" gridrow="4" pageId="9" pageNumber="424" rowspan-2="1">
<th id="57665DC4AA140009FF2CFCF633F4FCCD" box="[165,441,826,849]" gridcol="0" gridrow="4" pageId="9" pageNumber="424">Intercall interval (ms)</th>
<td id="57665DC4AA140009FDE5FCF63143FCCD" box="[620,782,826,849]" gridcol="1" gridrow="4" pageId="9" pageNumber="424">536 (406–1028)</td>
</tr>
<tr id="14B734B8AA140009FF2CFCAE36C7FCE5" box="[165,1162,866,889]" gridrow="5" pageId="9" pageNumber="424">
<th id="57665DC4AA140009FF2CFCAE33F4FCE5" box="[165,441,866,889]" gridcol="0" gridrow="5" pageId="9" pageNumber="424">Notes/call</th>
<td id="57665DC4AA140009FDE5FCAE3143FCE5" box="[620,782,866,889]" gridcol="1" gridrow="5" pageId="9" pageNumber="424">3</td>
<td id="57665DC4AA140009FC65FCAE36C7FCE5" box="[1004,1162,866,889]" gridcol="2" gridrow="5" pageId="9" pageNumber="424">1</td>
</tr>
<tr id="14B734B8AA140009FF2CFC4636C7FC3D" box="[165,1162,906,929]" gridrow="6" pageId="9" pageNumber="424">
<th id="57665DC4AA140009FF2CFC4633F4FC3D" box="[165,441,906,929]" gridcol="0" gridrow="6" pageId="9" pageNumber="424">Dominant frequency (kHz)</th>
<td id="57665DC4AA140009FDE5FC463143FC3D" box="[620,782,906,929]" gridcol="1" gridrow="6" pageId="9" pageNumber="424">3.0 (2.6–3.2)</td>
<td id="57665DC4AA140009FC65FC4636C7FC3D" box="[1004,1162,906,929]" gridcol="2" gridrow="6" pageId="9" pageNumber="424">3.1 (2.8–3.4)</td>
</tr>
<tr id="14B734B8AA140009FF2CFC7E36C7FC54" box="[165,1162,946,968]" gridrow="7" pageId="9" pageNumber="424" rowspan-1="1" rowspan-2="1">
<th id="57665DC4AA140009FF2CFC7E33F4FC54" box="[165,441,946,968]" gridcol="0" gridrow="7" pageId="9" pageNumber="424">Pulses/note</th>
</tr>
<tr id="14B734B8AA140009FF2CFC1636C7FC6D" box="[165,1162,986,1009]" gridrow="8" pageId="9" pageNumber="424">
<th id="57665DC4AA140009FF2CFC1633F4FC6D" box="[165,441,986,1009]" gridcol="0" gridrow="8" pageId="9" pageNumber="424">Note 1</th>
<td id="57665DC4AA140009FDE5FC163143FC6D" box="[620,782,986,1009]" gridcol="1" gridrow="8" pageId="9" pageNumber="424">10.8 (7–15)</td>
<td id="57665DC4AA140009FC65FC1636C7FC6D" box="[1004,1162,986,1009]" gridcol="2" gridrow="8" pageId="9" pageNumber="424">15.8 (14–19)</td>
</tr>
<tr id="14B734B8AA140009FF2CFBCE36C7FB85" box="[165,1162,1026,1049]" gridrow="9" pageId="9" pageNumber="424" rowspan-2="1">
<th id="57665DC4AA140009FF2CFBCE33F4FB85" box="[165,441,1026,1049]" gridcol="0" gridrow="9" pageId="9" pageNumber="424">Note 2</th>
<td id="57665DC4AA140009FDE5FBCE3143FB85" box="[620,782,1026,1049]" gridcol="1" gridrow="9" pageId="9" pageNumber="424">4.1 (4–5)</td>
</tr>
<tr id="14B734B8AA140009FF2CFBE636C7FBDD" box="[165,1162,1066,1089]" gridrow="10" pageId="9" pageNumber="424" rowspan-2="1">
<th id="57665DC4AA140009FF2CFBE633F4FBDD" box="[165,441,1066,1089]" gridcol="0" gridrow="10" pageId="9" pageNumber="424">Note 3</th>
<td id="57665DC4AA140009FDE5FBE63143FBDD" box="[620,782,1066,1089]" gridcol="1" gridrow="10" pageId="9" pageNumber="424">2.4 (1–4)</td>
</tr>
</table>
</paragraph>
<paragraph id="AA3836FAAA14FFFFFF4EFBB030D3FA9C" blockId="9.[151,1437,1148,2037]" pageId="9" pageNumber="424">
<emphasis id="98F3EAE8AA14FFFFFF4EFBB03323FB09" bold="true" box="[199,366,1148,1173]" pageId="9" pageNumber="424">Comparisons:</emphasis>
In having supra-axillary and ventrolateral glands, 
<taxonomicName id="6D874D79AA14FFFFFC29FBB1366CFB08" box="[928,1057,1148,1172]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA14FFFFFC29FBB1366CFB08" box="[928,1057,1148,1172]" italics="true" pageId="9" pageNumber="424">L. botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA14FFFFFBA1FBB036CDFB09" box="[1064,1152,1148,1173]" pageId="9" pageNumber="424" rank="species">
<emphasis id="98F3EAE8AA14FFFFFBA1FBB036CDFB09" bold="true" box="[1064,1152,1148,1173]" pageId="9" pageNumber="424">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from the southern species 
<taxonomicName id="6D874D79AA14FFFFFF67FB6D3319FB2B" box="[238,340,1184,1208]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="arayai">
<emphasis id="98F3EAE8AA14FFFFFF67FB6D3319FB2B" box="[238,340,1184,1208]" italics="true" pageId="9" pageNumber="424">L. arayai</emphasis>
</taxonomicName>
, 
<emphasis id="98F3EAE8AA14FFFFFEE8FB6D3066FB40" italics="true" pageId="9" pageNumber="424">
<taxonomicName id="6D874D79AA14FFFFFEE8FB6D3388FB24" box="[353,453,1184,1208]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFE59FB6D3026FB24" box="[464,619,1184,1208]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="fuliginosus">L. fuliginosus</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFDFFFB6D3145FB24" box="[630,776,1184,1208]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="fritinniens">L. fritinniens</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFC9AFB6D31C5FB24" box="[787,904,1184,1208]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFC1AFB6D364CFB24" box="[915,1025,1184,1208]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFB82FB6D36C7FB24" box="[1035,1162,1184,1208]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="heterops">L. heterops</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFB1CFB6D3774FB24" box="[1173,1337,1184,1208]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="kajangensis">L. kajangensis</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFACDFB6D37DAFB24" box="[1348,1431,1184,1208]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFF1EFB093346FB40" box="[151,267,1221,1244]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="maurus">L. maurus</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFE9CFB09338CFB40" box="[277,449,1220,1244]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="melanolecus">L. melanolecus</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFE45FB093066FB40" box="[460,555,1221,1244]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
</emphasis>
and 
<taxonomicName id="6D874D79AA14FFFFFDE8FB0930FAFB40" box="[609,695,1220,1244]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="solus">
<emphasis id="98F3EAE8AA14FFFFFDE8FB0930FAFB40" box="[609,695,1220,1244]" italics="true" pageId="9" pageNumber="424">L. solus</emphasis>
</taxonomicName>
(all of which are considered to belong in the subgenus 
<taxonomicName id="6D874D79AA14FFFFFA95FB0837D9FB40" box="[1308,1428,1220,1244]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="genus">
<emphasis id="98F3EAE8AA14FFFFFA95FB0837D9FB40" box="[1308,1428,1220,1244]" italics="true" pageId="9" pageNumber="424">Leptolalax</emphasis>
</taxonomicName>
; Delorme, Dubois, Grosjean, and Ohler, 2006).
</paragraph>
</subSubSection>
<subSubSection id="E29D6571AA14FFFCFF4EFAC030C2FC5B" lastPageId="10" lastPageNumber="425" pageId="9" pageNumber="424" type="description">
<paragraph id="AA3836FAAA14FFFFFF4EFAC037C2F9F4" blockId="9.[151,1437,1148,2037]" pageId="9" pageNumber="424">
<taxonomicName id="6D874D79AA14FFFFFF4EFAC033FCFAB8" box="[199,433,1292,1316]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA14FFFFFF4EFAC033FCFAB8" box="[199,433,1292,1316]" italics="true" pageId="9" pageNumber="424">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA14FFFFFE36FAC0306DFAB9" box="[447,544,1292,1317]" pageId="9" pageNumber="424" rank="species">
<emphasis id="98F3EAE8AA14FFFFFE36FAC0306DFAB9" bold="true" box="[447,544,1292,1317]" pageId="9" pageNumber="424">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from all species in the genus except for 
<emphasis id="98F3EAE8AA14FFFFFB62FAC1336EFAD4" italics="true" pageId="9" pageNumber="424">
<taxonomicName id="6D874D79AA14FFFFFB62FAC1373EFAB8" box="[1259,1395,1292,1316]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFA0CFAC1336DFAD4" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="bidoupensis">L. bidoupensis</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="6D874D79AA14FFFFFEE8FAFD3395FAD4" box="[353,472,1328,1352]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="melicus">
<emphasis id="98F3EAE8AA14FFFFFEE8FAFD3395FAD4" box="[353,472,1328,1352]" italics="true" pageId="9" pageNumber="424">L. melicus</emphasis>
</taxonomicName>
by having a dark brownish red ventral surface with white speckling (
<emphasis id="98F3EAE8AA14FFFFFA80FAFD3099FA28" italics="true" pageId="9" pageNumber="424">
<taxonomicName id="6D874D79AA14FFFFFA80FAFD373BFAD4" box="[1289,1398,1329,1352]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="aereus">L. aereus</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFA0CFAFD32ABFAF0" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="alpinis">L. alpinis</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFF71FA993325FAF0" box="[248,360,1365,1388]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFEF3FA9933B6FAF0" box="[378,507,1364,1388]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="bourreti">L. bourreti</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFD87FA993034FAF0" box="[526,633,1364,1388]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFD05FA9930A6FAF0" box="[652,747,1364,1388]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="firthi">L. firthi</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFD74FA9931D2FAF0" box="[765,927,1364,1388]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="fuliginosus">L. fuliginosus</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFC38FA993613FAF0" box="[945,1118,1364,1388]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="fritinniensis">L. fritinniensis</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFBF9FA9936A0FAF0" box="[1136,1261,1364,1388]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFB76FA99373EFAF0" box="[1279,1395,1364,1388]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFA0FFA99335DFA0C" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="khasiorum">L. khasiorum</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFE96FAB533ECFA0C" box="[287,417,1400,1424]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="lateralis">L. lateralis</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFE38FAB533B6FA0C" box="[433,507,1400,1424]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="liui">L. liui</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFD83FAB530C3FA13" box="[522,654,1400,1424]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="minimus">L. minimus</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFD14FAB53103FA0C" box="[669,846,1400,1424]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="nahangensis">L. nahangensis</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFCD7FAB5364BFA0C" box="[862,1030,1400,1424]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="nokrekensis">L. nokrekensis</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFB9CFAB536F8FA0C" box="[1045,1205,1400,1424]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="oshanensis">L. oshanensis</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFB4DFAB5373BFA0C" box="[1220,1398,1400,1424]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFA0FFAB53290FA28" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFF6EFA5133D6FA28" box="[231,411,1436,1460]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="playcephalus">L. playcephalus</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFE2CFA5133B3FA28" box="[421,510,1436,1460]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFD80FA513028FA28" box="[521,613,1437,1460]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="sungi">L. sungi</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFDF9FA513099FA28" box="[624,724,1436,1460]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="tamdil">L. tamdil</taxonomicName>
</emphasis>
and 
<taxonomicName id="6D874D79AA14FFFFFC80FA5131DFFA28" box="[777,914,1436,1460]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="98F3EAE8AA14FFFFFC80FA5131DFFA28" box="[777,914,1436,1460]" italics="true" pageId="9" pageNumber="424">L. tuberosus</emphasis>
</taxonomicName>
have mostly white or pale grey/brown venters, with or without dark spots or mottling; 
<taxonomicName id="6D874D79AA14FFFFFDEDFA0D3097FA44" box="[612,730,1473,1496]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="croceus">
<emphasis id="98F3EAE8AA14FFFFFDEDFA0D3097FA44" box="[612,730,1473,1496]" italics="true" pageId="9" pageNumber="424">L. croceus</emphasis>
</taxonomicName>
has a bright orange belly; 
<taxonomicName id="6D874D79AA14FFFFFB9CFA0D36DBFA44" box="[1045,1174,1472,1496]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="98F3EAE8AA14FFFFFB9CFA0D36DBFA44" box="[1045,1174,1472,1496]" italics="true" pageId="9" pageNumber="424">L. pluvialis</emphasis>
</taxonomicName>
has a dirty white/grey venter with dark brown/grey marbling, and uniform pale dirty white/grey throat with pale speckling only around the margins; 
<taxonomicName id="6D874D79AA14FFFFFEBAF9C533BDF9BC" box="[307,496,1544,1568]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="98F3EAE8AA14FFFFFEBAF9C533BDF9BC" box="[307,496,1544,1568]" italics="true" pageId="9" pageNumber="424">L. melanoleucus</emphasis>
</taxonomicName>
and 
<taxonomicName id="6D874D79AA14FFFFFDB9F9C5314CF983" box="[560,769,1544,1568]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="98F3EAE8AA14FFFFFDB9F9C5314CF983" box="[560,769,1544,1568]" italics="true" pageId="9" pageNumber="424">L. ventripunctatus</emphasis>
</taxonomicName>
display large patches of distinct brown/black and white marbling, 
<taxonomicName id="6D874D79AA14FFFFFE85F9E133D5F9D8" box="[268,408,1580,1604]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="98F3EAE8AA14FFFFFE85F9E133D5F9D8" box="[268,408,1580,1604]" italics="true" pageId="9" pageNumber="424">L. heteropus</emphasis>
</taxonomicName>
has a grey venter, speckled with black; 
<taxonomicName id="6D874D79AA14FFFFFCEFF9E13197F9D8" box="[870,986,1581,1604]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="maurus">
<emphasis id="98F3EAE8AA14FFFFFCEFF9E13197F9D8" box="[870,986,1581,1604]" italics="true" pageId="9" pageNumber="424">L. maurus</emphasis>
</taxonomicName>
has a black or dark grey brown venter, with indistinct small light areas, and 
<taxonomicName id="6D874D79AA14FFFFFDBDF99D30CBF9F4" box="[564,646,1616,1640]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="kecil">
<emphasis id="98F3EAE8AA14FFFFFDBDF99D30CBF9F4" box="[564,646,1616,1640]" italics="true" pageId="9" pageNumber="424">L. kecil</emphasis>
</taxonomicName>
has a uniformly dark venter with large, dark orange pectoral glands).
</paragraph>
<paragraph id="AA3836FAAA14FFFFFF4EF9B9329EF84C" blockId="9.[151,1437,1148,2037]" pageId="9" pageNumber="424">
From the three species with dark reddish brown ventral surfaces, 
<taxonomicName id="6D874D79AA14FFFFFC2DF9B93668F910" box="[932,1061,1652,1676]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA14FFFFFC2DF9B93668F910" box="[932,1061,1652,1676]" italics="true" pageId="9" pageNumber="424">L. botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA14FFFFFBA5F9B836CBF911" box="[1068,1158,1652,1677]" pageId="9" pageNumber="424" rank="species">
<emphasis id="98F3EAE8AA14FFFFFBA5F9B836CBF911" bold="true" box="[1068,1158,1652,1677]" pageId="9" pageNumber="424">sp. nov.</emphasis>
</taxonomicNameLabel>
differs by having a SVL of 
<quantity id="6D7F9B1FAA14FFFFFF3EF95B3311F92C" box="[183,348,1687,1712]" metricMagnitude="-2" metricUnit="m" metricValue="3.085" metricValueMax="3.26" metricValueMin="2.91" pageId="9" pageNumber="424" unit="mm" value="30.85" valueMax="32.6" valueMin="29.1">29.1–32.6 mm</quantity>
in 
<specimenCount id="BC81FD73AA14FFFFFE0AF955339CF92C" box="[387,465,1687,1712]" pageId="9" pageNumber="424" type="adult">7 adult</specimenCount>
males, 30.0– 
<quantity id="6D7F9B1FAA14FFFFFDE2F954309DF92C" box="[619,720,1688,1712]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="9" pageNumber="424" unit="mm" value="31.8">31.8 mm</quantity>
in 
<specimenCount id="BC81FD73AA14FFFFFD7EF9543128F92C" box="[759,869,1687,1712]" pageId="9" pageNumber="424" type="female">2 females</specimenCount>
(
<taxonomicName id="6D874D79AA14FFFFFCFFF95531BBF92C" box="[886,1014,1688,1712]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="98F3EAE8AA14FFFFFCFFF95531BBF92C" box="[886,1014,1688,1712]" italics="true" pageId="9" pageNumber="424">L. applebyi</emphasis>
</taxonomicName>
males 
<quantity id="6D7F9B1FAA14FFFFFBC0F95B36BDF92C" box="[1097,1264,1687,1712]" metricMagnitude="-2" metricUnit="m" metricValue="2.095" metricValueMax="2.23" metricValueMin="1.96" pageId="9" pageNumber="424" unit="mm" value="20.95" valueMax="22.3" valueMin="19.6">19.6–22.3 mm</quantity>
, females 
<quantity id="6D7F9B1FAA14FFFFFAD4F954334DF948" metricMagnitude="-2" metricUnit="m" metricValue="2.38" metricValueMax="2.59" metricValueMin="2.17" pageId="9" pageNumber="424" unit="mm" value="23.799999999999997" valueMax="25.9" valueMin="21.7">21.7– 25.9 mm</quantity>
; 
<taxonomicName id="6D874D79AA14FFFFFE84F97133FFF948" box="[269,434,1724,1748]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="98F3EAE8AA14FFFFFE84F97133FFF948" box="[269,434,1724,1748]" italics="true" pageId="9" pageNumber="424">L. bidoupensis</emphasis>
</taxonomicName>
males 
<quantity id="6D7F9B1FAA14FFFFFD8CF97030E1F948" box="[517,684,1724,1749]" metricMagnitude="-2" metricUnit="m" metricValue="2.41" metricValueMax="2.46" metricValueMin="2.36" pageId="9" pageNumber="424" unit="mm" value="24.1" valueMax="24.6" valueMin="23.6">23.6–24.6 mm</quantity>
, females 
<quantity id="6D7F9B1FAA14FFFFFC93F970318FF948" box="[794,962,1724,1748]" metricMagnitude="-2" metricUnit="m" metricValue="2.9299999999999997" metricValueMax="2.94" metricValueMin="2.92" pageId="9" pageNumber="424" unit="mm" value="29.299999999999997" valueMax="29.4" valueMin="29.2">29.2–29.4 mm</quantity>
; 
<taxonomicName id="6D874D79AA14FFFFFC46F971360EF948" box="[975,1091,1724,1748]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="melicus">
<emphasis id="98F3EAE8AA14FFFFFC46F971360EF948" box="[975,1091,1724,1748]" italics="true" pageId="9" pageNumber="424">L. melicus</emphasis>
</taxonomicName>
males 
<quantity id="6D7F9B1FAA14FFFFFB1EF970370DF948" box="[1175,1344,1724,1749]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" metricValueMax="2.27" metricValueMin="1.95" pageId="9" pageNumber="424" unit="mm" value="21.1" valueMax="22.7" valueMin="19.5">19.5–22.7 mm</quantity>
), and in having only faint white spotting (versus dense white spotting or marbling on the belly). Body size also distinguishes the new species from the smaller 
<taxonomicName id="6D874D79AA14FFFFFD3DF8C9316BF880" box="[692,806,1796,1820]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="alpinus">
<emphasis id="98F3EAE8AA14FFFFFD3DF8C9316BF880" box="[692,806,1796,1820]" italics="true" pageId="9" pageNumber="424">L. alpinus</emphasis>
</taxonomicName>
(
<quantity id="6D7F9B1FAA14FFFFFCBEF8C831EEF880" box="[823,931,1796,1821]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="9" pageNumber="424" unit="mm" value="26.3">26.3 mm</quantity>
), 
<taxonomicName id="6D874D79AA14FFFFFC31F8C93660F880" box="[952,1069,1797,1820]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="croceus">
<emphasis id="98F3EAE8AA14FFFFFC31F8C93660F880" box="[952,1069,1797,1820]" italics="true" pageId="9" pageNumber="424">L. croceus</emphasis>
</taxonomicName>
(males 
<quantity id="6D7F9B1FAA14FFFFFB03F8C8377EF880" box="[1162,1331,1796,1820]" metricMagnitude="-2" metricUnit="m" metricValue="2.475" metricValueMax="2.73" metricValueMin="2.2199999999999998" pageId="9" pageNumber="424" unit="mm" value="24.75" valueMax="27.3" valueMin="22.2">22.2–27.3 mm</quantity>
), 
<taxonomicName id="6D874D79AA14FFFFFACEF8C937D1F880" box="[1351,1436,1796,1820]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="kecil">
<emphasis id="98F3EAE8AA14FFFFFACEF8C937D1F880" box="[1351,1436,1796,1820]" italics="true" pageId="9" pageNumber="424">L. kecil</emphasis>
</taxonomicName>
(males 
<quantity id="6D7F9B1FAA14FFFFFF60F8EB33C3F8DC" box="[233,398,1831,1856]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.05" metricValueMin="1.93" pageId="9" pageNumber="424" unit="mm" value="19.9" valueMax="20.5" valueMin="19.3">19.3–20.5 mm</quantity>
, female 
<quantity id="6D7F9B1FAA14FFFFFE64F8EB300DF8DC" box="[493,576,1831,1856]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="424" unit="mm" value="25.0">25 mm</quantity>
), 
<taxonomicName id="6D874D79AA14FFFFFDDAF8E530AAF8DC" box="[595,743,1832,1856]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="khasiorum">
<emphasis id="98F3EAE8AA14FFFFFDDAF8E530AAF8DC" box="[595,743,1832,1856]" italics="true" pageId="9" pageNumber="424">L. khasiorum</emphasis>
</taxonomicName>
(
<quantity id="6D7F9B1FAA14FFFFFD7FF8EB31D2F8DC" box="[758,927,1831,1856]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" metricValueMax="2.73" metricValueMin="2.45" pageId="9" pageNumber="424" unit="mm" value="25.9" valueMax="27.3" valueMin="24.5">24.5–27.3 mm</quantity>
), and 
<taxonomicName id="6D874D79AA14FFFFFC6BF8E5362DF8DC" box="[994,1120,1832,1856]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="98F3EAE8AA14FFFFFC6BF8E5362DF8DC" box="[994,1120,1832,1856]" italics="true" pageId="9" pageNumber="424">L. pluvialis</emphasis>
</taxonomicName>
(males 
<quantity id="6D7F9B1FAA14FFFFFB30F8E4372DF8DC" box="[1209,1376,1832,1856]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" metricValueMax="2.23" metricValueMin="2.13" pageId="9" pageNumber="424" unit="mm" value="21.8" valueMax="22.3" valueMin="21.3">21.3–22.3 mm</quantity>
), and the larger 
<taxonomicName id="6D874D79AA14FFFFFE84F88133C5F8F8" box="[269,392,1868,1892]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="bourreti">
<emphasis id="98F3EAE8AA14FFFFFE84F88133C5F8F8" box="[269,392,1868,1892]" italics="true" pageId="9" pageNumber="424">L. bourreti</emphasis>
</taxonomicName>
(male 
<quantity id="6D7F9B1FAA14FFFFFE53F8803008F8F8" box="[474,581,1868,1893]" metricMagnitude="-2" metricUnit="m" metricValue="3.62" pageId="9" pageNumber="424" unit="mm" value="36.2">36.2 mm</quantity>
), 
<taxonomicName id="6D874D79AA14FFFFFDD3F88130D2F8F8" box="[602,671,1869,1892]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="eos">
<emphasis id="98F3EAE8AA14FFFFFDD3F88130D2F8F8" box="[602,671,1869,1892]" italics="true" pageId="9" pageNumber="424">L. eos</emphasis>
</taxonomicName>
(males 
<quantity id="6D7F9B1FAA14FFFFFD74F88031EBF8F8" box="[765,934,1868,1892]" metricMagnitude="-2" metricUnit="m" metricValue="3.39" metricValueMax="3.47" metricValueMin="3.31" pageId="9" pageNumber="424" unit="mm" value="33.9" valueMax="34.7" valueMin="33.1">33.1–34.7 mm</quantity>
), 
<taxonomicName id="6D874D79AA14FFFFFC35F881367CF8F8" box="[956,1073,1868,1892]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="98F3EAE8AA14FFFFFC35F881367CF8F8" box="[956,1073,1868,1892]" italics="true" pageId="9" pageNumber="424">L. gracilis</emphasis>
</taxonomicName>
(males 34.3–39.0 mm, females 
<quantity id="6D7F9B1FAA14FFFFFF1EF8BC3372F814" box="[151,319,1904,1928]" metricMagnitude="-2" metricUnit="m" metricValue="4.584999999999999" metricValueMax="4.93" metricValueMin="4.24" pageId="9" pageNumber="424" unit="mm" value="45.849999999999994" valueMax="49.3" valueMin="42.4">42.4–49.3 mm</quantity>
), 
<taxonomicName id="6D874D79AA14FFFFFEDAF8BD33BBF814" box="[339,502,1904,1928]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="kajangensis">
<emphasis id="98F3EAE8AA14FFFFFEDAF8BD33BBF814" box="[339,502,1904,1928]" italics="true" pageId="9" pageNumber="424">L. kajangensis</emphasis>
</taxonomicName>
(males 
<quantity id="6D7F9B1FAA14FFFFFDD9F8A33083F814" box="[592,718,1903,1928]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="9" pageNumber="424" unit="mm" value="34.5" valueMax="35.0" valueMin="34.0">34–35 mm</quantity>
), 
<taxonomicName id="6D874D79AA14FFFFFD6BF8BD31C0F814" box="[738,909,1904,1928]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="nahangensis">
<emphasis id="98F3EAE8AA14FFFFFD6BF8BD31C0F814" box="[738,909,1904,1928]" italics="true" pageId="9" pageNumber="424">L. nahangensis</emphasis>
</taxonomicName>
(male 
<quantity id="6D7F9B1FAA14FFFFFC54F8BC360BF814" box="[989,1094,1904,1928]" metricMagnitude="-2" metricUnit="m" metricValue="4.08" pageId="9" pageNumber="424" unit="mm" value="40.8">40.8 mm</quantity>
), 
<taxonomicName id="6D874D79AA14FFFFFBD3F8BD3759F814" box="[1114,1300,1904,1928]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="98F3EAE8AA14FFFFFBD3F8BD3759F814" box="[1114,1300,1904,1928]" italics="true" pageId="9" pageNumber="424">L. platycephalus</emphasis>
</taxonomicName>
(male 35.1, female 
<quantity id="6D7F9B1FAA14FFFFFF65F858330FF830" box="[236,322,1940,1965]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="9" pageNumber="424" unit="mm" value="46.0">46 mm</quantity>
), 
<taxonomicName id="6D874D79AA14FFFFFEDEF85933F8F830" box="[343,437,1941,1964]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="sungi">
<emphasis id="98F3EAE8AA14FFFFFEDEF85933F8F830" box="[343,437,1941,1964]" italics="true" pageId="9" pageNumber="424">L. sungi</emphasis>
</taxonomicName>
(males 
<quantity id="6D7F9B1FAA14FFFFFD9BF85830F7F830" box="[530,698,1940,1965]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.27" metricValueMin="4.83" pageId="9" pageNumber="424" unit="mm" value="50.5" valueMax="52.7" valueMin="48.3">48.3–52.7 mm</quantity>
, females 56.7–58.9), and 
<taxonomicName id="6D874D79AA14FFFFFC6DF8593601F830" box="[996,1100,1940,1964]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="tamdil">
<emphasis id="98F3EAE8AA14FFFFFC6DF8593601F830" box="[996,1100,1940,1964]" italics="true" pageId="9" pageNumber="424">L. tamdil</emphasis>
</taxonomicName>
(male 
<quantity id="6D7F9B1FAA14FFFFFB17F858374AF830" box="[1182,1287,1940,1964]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="9" pageNumber="424" unit="mm" value="32.3">32.3 mm</quantity>
, female 
<quantity id="6D7F9B1FAA14FFFFFAE2F8583285F84C" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="9" pageNumber="424" unit="mm" value="31.8">31.8 mm</quantity>
).
</paragraph>
<paragraph id="AA3836FAAA14FFFCFF4EF8103691FE14" blockId="9.[151,1437,1148,2037]" lastBlockId="10.[151,1437,152,968]" lastPageId="10" lastPageNumber="425" pageId="9" pageNumber="424">
An absence of black markings on the flanks further distinguishes the new species from 
<emphasis id="98F3EAE8AA14FFFFFB10F81137D8F868" box="[1177,1429,2012,2036]" italics="true" pageId="9" pageNumber="424">
<taxonomicName id="6D874D79AA14FFFFFB10F8113746F868" box="[1177,1291,2012,2036]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="alpinus">L. alpinus</taxonomicName>
, 
<taxonomicName id="6D874D79AA14FFFFFA9EF81137D8F868" box="[1303,1429,2012,2036]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="424" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
</emphasis>
, 
<emphasis id="98F3EAE8AA17FFFCFF1EFF55309CFF48" italics="true" pageId="10" pageNumber="425">
<taxonomicName id="6D874D79AA17FFFCFF1EFF55330DFF2C" box="[151,320,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="bidoupensis">L. bidoupensis</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFEC4FF553387FF2C" box="[333,458,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="bourreti">L. bourreti</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFE51FF55303BFF2C" box="[472,630,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="oshanensis">L. oshanensis</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFD0AFF55316CFF2C" box="[643,801,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="fuliginosus">L. fuliginosus</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFCA7FF55319AFF2C" box="[814,983,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="frittianensis">L. frittianensis</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFC6DFF553611FF2C" box="[996,1116,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFBE3FF553696FF2C" box="[1130,1243,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFB61FF553735FF2C" box="[1256,1400,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFA0FFF553283FF48" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFF52FF7133DAFF48" box="[219,407,188,212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="melanoleucus">L. melanoleucus</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFE2DFF713056FF48" box="[420,539,188,212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="melicus">L. melicus</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFDAEFF713081FF48" box="[551,716,188,212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="nokrekensis">L. nokrekensis</taxonomicName>
,
</emphasis>
and 
<emphasis id="98F3EAE8AA17FFFCFC83FF7131F3FF48" box="[778,958,188,212]" italics="true" pageId="10" pageNumber="425">
<taxonomicName id="6D874D79AA17FFFCFC83FF7131F7FF48" box="[778,954,188,212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
.
</emphasis>
In having toes with rudimentary webbing and weak lateral fringing, 
<taxonomicName id="6D874D79AA17FFFCFE4DFF2D3005FF64" box="[452,584,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA17FFFCFE4DFF2D3005FF64" box="[452,584,224,248]" italics="true" pageId="10" pageNumber="425">L. botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA17FFFCFDD8FF1330E1FF64" box="[593,684,223,248]" pageId="10" pageNumber="425" rank="species">
<emphasis id="98F3EAE8AA17FFFCFDD8FF1330E1FF64" bold="true" box="[593,684,223,248]" pageId="10" pageNumber="425">sp. nov.</emphasis>
</taxonomicNameLabel>
can also be readily differentiated from 
<taxonomicName id="6D874D79AA17FFFCFBFBFF2D3690FF64" box="[1138,1245,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="alpinis">
<emphasis id="98F3EAE8AA17FFFCFBFBFF2D3690FF64" box="[1138,1245,224,248]" italics="true" pageId="10" pageNumber="425">L. alpinis</emphasis>
</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFB64FF2D3709FF64" box="[1261,1348,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="firthi">
<emphasis id="98F3EAE8AA17FFFCFB64FF2D3709FF64" box="[1261,1348,224,248]" italics="true" pageId="10" pageNumber="425">L. firthi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="6D874D79AA17FFFCFA0CFF2D32F1FE80" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="liui">
<emphasis id="98F3EAE8AA17FFFCFA0CFF2D32F1FE80" italics="true" pageId="10" pageNumber="425">L. liui</emphasis>
</taxonomicName>
, which have wide lateral fringing on toes, and from 
<taxonomicName id="6D874D79AA17FFFCFC95FEC93184FE80" box="[796,969,260,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="98F3EAE8AA17FFFCFC95FEC93184FE80" box="[796,969,260,284]" italics="true" pageId="10" pageNumber="425">L. pelodytoides</emphasis>
</taxonomicName>
, which has more extensive webbing and wide lateral fringes between toes. In addition, 
<taxonomicName id="6D874D79AA17FFFCFD14FEE431CCFEDC" box="[669,897,296,320]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA17FFFCFD14FEE431CCFEDC" box="[669,897,296,320]" italics="true" pageId="10" pageNumber="425">Leptolalax botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA17FFFCFC0EFEEB31ACFEDC" box="[903,993,295,320]" pageId="10" pageNumber="425" rank="species">
<emphasis id="98F3EAE8AA17FFFCFC0EFEEB31ACFEDC" bold="true" box="[903,993,295,320]" pageId="10" pageNumber="425">sp. nov.</emphasis>
</taxonomicNameLabel>
has large pectoral glands (
<quantity id="6D7F9B1FAA17FFFCFA85FEE437D5FEA3" box="[1292,1432,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.9" metricValueMin="1.1" pageId="10" pageNumber="425" unit="mm" value="1.5" valueMax="1.9" valueMin="1.1">1.1–1.9 mm</quantity>
; 4–6% SVL) and femoral glands (
<quantity id="6D7F9B1FAA17FFFCFDA1FE8030F4FEF8" box="[552,697,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="4.3" metricValueMin="2.4" pageId="10" pageNumber="425" unit="mm" value="3.3499999999999996" valueMax="4.3" valueMin="2.4">2.4–4.3 mm</quantity>
; 7–14% SVL). The femoral glands in particular are the most prominent of all 
<taxonomicName id="6D874D79AA17FFFCFEDBFEBC3387FE14" box="[338,458,368,392]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="genus">
<emphasis id="98F3EAE8AA17FFFCFEDBFEBC3387FE14" box="[338,458,368,392]" italics="true" pageId="10" pageNumber="425">Leptolalax</emphasis>
</taxonomicName>
species measured by us (see Appendix I; femoral glands 2–8% SVL).
</paragraph>
<paragraph id="AA3836FAAA17FFFCFF4EFE5830C2FC5B" blockId="10.[151,1437,152,968]" pageId="10" pageNumber="425">
The advertisement call of 
<taxonomicName id="6D874D79AA17FFFCFE79FE59303FFE30" box="[496,626,404,428]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA17FFFCFE79FE59303FFE30" box="[496,626,404,428]" italics="true" pageId="10" pageNumber="425">L. botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA17FFFCFDF2FE583098FE31" box="[635,725,404,429]" pageId="10" pageNumber="425" rank="species">
<emphasis id="98F3EAE8AA17FFFCFDF2FE583098FE31" bold="true" box="[635,725,404,429]" pageId="10" pageNumber="425">sp. nov.</emphasis>
</taxonomicNameLabel>
is unique among congeners with known calls. In having a call with a dominant frequency of 2.6–3.2 kHz (at 14.0 ºC), 
<taxonomicName id="6D874D79AA17FFFCFC82FE7531C0FE4C" box="[779,909,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="botsfordi">
<emphasis id="98F3EAE8AA17FFFCFC82FE7531C0FE4C" box="[779,909,440,464]" italics="true" pageId="10" pageNumber="425">L. botsfordi</emphasis>
</taxonomicName>
differs from all but 
<taxonomicName id="6D874D79AA17FFFCFBE6FE75375EFE4C" box="[1135,1299,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="98F3EAE8AA17FFFCFBE6FE75375EFE4C" box="[1135,1299,440,464]" italics="true" pageId="10" pageNumber="425">L. bidoupensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFAA8FE7537D8FE4C" box="[1313,1429,441,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="croceus">
<emphasis id="98F3EAE8AA17FFFCFAA8FE7537D8FE4C" box="[1313,1429,441,464]" italics="true" pageId="10" pageNumber="425">L. croceus</emphasis>
</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFF1EFE11337DFE68" box="[151,304,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="98F3EAE8AA17FFFCFF1EFE11337DFE68" box="[151,304,476,500]" italics="true" pageId="10" pageNumber="425">L. fuliginosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFEB6FE1133FFFE68" box="[319,434,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="98F3EAE8AA17FFFCFEB6FE1133FFFE68" box="[319,434,476,500]" italics="true" pageId="10" pageNumber="425">L. gracilis</emphasis>
</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFE4BFE113000FE68" box="[450,589,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="98F3EAE8AA17FFFCFE4BFE113000FE68" box="[450,589,476,500]" italics="true" pageId="10" pageNumber="425">L. heteropus</emphasis>
</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFDD5FE1130E2FE68" box="[604,687,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="kecil">
<emphasis id="98F3EAE8AA17FFFCFDD5FE1130E2FE68" box="[604,687,476,500]" italics="true" pageId="10" pageNumber="425">L. kecil</emphasis>
</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFD37FE11313AFE68" box="[702,887,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="98F3EAE8AA17FFFCFD37FE11313AFE68" box="[702,887,476,500]" italics="true" pageId="10" pageNumber="425">L. melanoleucus</emphasis>
</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFC0CFE1131B4FE68" box="[901,1017,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="melicus">
<emphasis id="98F3EAE8AA17FFFCFC0CFE1131B4FE68" box="[901,1017,476,500]" italics="true" pageId="10" pageNumber="425">L. melicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="6D874D79AA17FFFCFB81FE11362CFE68" box="[1032,1121,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="solus">
<emphasis id="98F3EAE8AA17FFFCFB81FE11362CFE68" box="[1032,1121,476,500]" italics="true" pageId="10" pageNumber="425">L. solus</emphasis>
</taxonomicName>
and 
<taxonomicName id="6D874D79AA17FFFCFB11FE11376EFE68" box="[1176,1315,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="98F3EAE8AA17FFFCFB11FE11376EFE68" box="[1176,1315,476,500]" italics="true" pageId="10" pageNumber="425">L. tuberosus</emphasis>
</taxonomicName>
(
<taxonomicName id="6D874D79AA17FFFCFABDFE1137D1FE68" box="[1332,1436,477,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="aereus">
<emphasis id="98F3EAE8AA17FFFCFABDFE1137D1FE68" box="[1332,1436,477,500]" italics="true" pageId="10" pageNumber="425">L. aereus</emphasis>
</taxonomicName>
6.2–7.9 kHz at 22.4–25.7 ºC; 
<taxonomicName id="6D874D79AA17FFFCFE60FDCD3017FD84" box="[489,602,512,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="alpinus">
<emphasis id="98F3EAE8AA17FFFCFE60FDCD3017FD84" box="[489,602,512,536]" italics="true" pageId="10" pageNumber="425">L. alpinus</emphasis>
</taxonomicName>
~6.7 kHz at 16 ºC; 
<taxonomicName id="6D874D79AA17FFFCFCB7FDCD31F0FD84" box="[830,957,512,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="98F3EAE8AA17FFFCFCB7FDCD31F0FD84" box="[830,957,512,536]" italics="true" pageId="10" pageNumber="425">L. applebyi</emphasis>
</taxonomicName>
4.0–4.3 kHz at 21.5 ºC; 
<taxonomicName id="6D874D79AA17FFFCFB51FDCD3772FD8B" box="[1240,1343,512,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="arayai">
<emphasis id="98F3EAE8AA17FFFCFB51FDCD3772FD8B" box="[1240,1343,512,536]" italics="true" pageId="10" pageNumber="425">L. arayai</emphasis>
</taxonomicName>
5.4–5.8 kHz at 17.4 ºC; 
<taxonomicName id="6D874D79AA17FFFCFEC0FDE933E6FDA0" box="[329,427,548,572]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="dringi">
<emphasis id="98F3EAE8AA17FFFCFEC0FDE933E6FDA0" box="[329,427,548,572]" italics="true" pageId="10" pageNumber="425">L. dringi</emphasis>
</taxonomicName>
7–7.5 kHz at 24.3 ºC; 
<taxonomicName id="6D874D79AA17FFFCFD23FDE93158FDA0" box="[682,789,548,572]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="98F3EAE8AA17FFFCFD23FDE93158FDA0" box="[682,789,548,572]" italics="true" pageId="10" pageNumber="425">L. hamidi</emphasis>
</taxonomicName>
7.0 kHz at 22.9 ºC; 
<taxonomicName id="6D874D79AA17FFFCFC71FDE93676FDA0" box="[1016,1083,548,572]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="liui">
<emphasis id="98F3EAE8AA17FFFCFC71FDE93676FDA0" box="[1016,1083,548,572]" italics="true" pageId="10" pageNumber="425">L. liui</emphasis>
</taxonomicName>
5.3 kHz; 
<taxonomicName id="6D874D79AA17FFFCFB20FDE9370CFDA0" box="[1193,1345,548,572]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="98F3EAE8AA17FFFCFB20FDE9370CFDA0" box="[1193,1345,548,572]" italics="true" pageId="10" pageNumber="425">L. oshanensis</emphasis>
</taxonomicName>
4.4–4.6 kHz, 14 ºC, recorded from c. 
<quantity id="6D7F9B1FAA17FFFCFE57FD843069FDFC" box="[478,548,583,608]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="425" unit="km" value="40.0">40 km</quantity>
from 
<typeStatus id="753C8858AA17FFFCFDE1FD8430D5FDFC" box="[616,664,584,608]" pageId="10" pageNumber="425">type</typeStatus>
locality of 
<taxonomicName id="6D874D79AA17FFFCFC9EFD8531E2FDFC" box="[791,943,584,608]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="98F3EAE8AA17FFFCFC9EFD8531E2FDFC" box="[791,943,584,608]" italics="true" pageId="10" pageNumber="425">L. oshanensis</emphasis>
</taxonomicName>
; 
<taxonomicName id="6D874D79AA17FFFCFC34FD853650FDC3" box="[957,1053,584,608]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="pictus">
<emphasis id="98F3EAE8AA17FFFCFC34FD853650FDC3" box="[957,1053,584,608]" italics="true" pageId="10" pageNumber="425">L. pictus</emphasis>
</taxonomicName>
6.8–7.2 at 19–22 ºC; 
<taxonomicName id="6D874D79AA17FFFCFA86FD853729FDFC" box="[1295,1380,584,608]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="firthi">
<emphasis id="98F3EAE8AA17FFFCFA86FD853729FDFC" box="[1295,1380,584,608]" italics="true" pageId="10" pageNumber="425">L. firthi</emphasis>
</taxonomicName>
5.6– 6.4 kHz at 18.3–21.2; 
<taxonomicName id="6D874D79AA17FFFCFE15FDA1300EFD18" box="[412,579,620,644]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="fritinniensis">
<emphasis id="98F3EAE8AA17FFFCFE15FDA1300EFD18" box="[412,579,620,644]" italics="true" pageId="10" pageNumber="425">L. fritinniensis</emphasis>
</taxonomicName>
7.3–9.2 at 24.3–24.9 ºC). Although frequency can vary with temperature, differences among species of the scale reported here are unlikely to be attributed to temperature alone. From 
<taxonomicName id="6D874D79AA17FFFCFA0CFD5D3350FD50" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="98F3EAE8AA17FFFCFA0CFD5D3350FD50" italics="true" pageId="10" pageNumber="425">L. bidoupensis</emphasis>
</taxonomicName>
, the call differs in having variable note 
<typeStatus id="753C8858AA17FFFCFD6DFD793152FD51" box="[740,799,693,717]" pageId="10" pageNumber="425">types</typeStatus>
and in the number of notes per call (1 or 3 versus 6–9), from 
<taxonomicName id="6D874D79AA17FFFCFF5DFD15330AFD6C" box="[212,327,729,752]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="croceus">
<emphasis id="98F3EAE8AA17FFFCFF5DFD15330AFD6C" box="[212,327,729,752]" italics="true" pageId="10" pageNumber="425">L. croceus</emphasis>
</taxonomicName>
in the number of notes per call (1 or 3 versus 4–5), 
<taxonomicName id="6D874D79AA17FFFCFC03FD15366FFD6C" box="[906,1058,728,752]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="98F3EAE8AA17FFFCFC03FD15366FFD6C" box="[906,1058,728,752]" italics="true" pageId="10" pageNumber="425">L. fuliginosus</emphasis>
</taxonomicName>
in the number of notes per call (1 or 3 versus 6–9), 
<taxonomicName id="6D874D79AA17FFFCFED0FD313381FC88" box="[345,460,764,788]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="98F3EAE8AA17FFFCFED0FD313381FC88" box="[345,460,764,788]" italics="true" pageId="10" pageNumber="425">L. gracilis</emphasis>
</taxonomicName>
in the number of notes per call (1 or 3 versus 3–4), 
<taxonomicName id="6D874D79AA17FFFCFB86FD3136D4FC88" box="[1039,1177,764,788]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="98F3EAE8AA17FFFCFB86FD3136D4FC88" box="[1039,1177,764,788]" italics="true" pageId="10" pageNumber="425">L. heteropus</emphasis>
</taxonomicName>
in the number of notes per call (1 or 3 versus 6–9), 
<taxonomicName id="6D874D79AA17FFFCFE5BFCED3068FCA4" box="[466,549,800,824]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="kecil">
<emphasis id="98F3EAE8AA17FFFCFE5BFCED3068FCA4" box="[466,549,800,824]" italics="true" pageId="10" pageNumber="425">L. kecil</emphasis>
</taxonomicName>
in having variable note 
<typeStatus id="753C8858AA17FFFCFCBDFCEC3122FCA4" box="[820,879,800,824]" pageId="10" pageNumber="425">types</typeStatus>
and in the number of notes per call (1 or 3 versus 4), 
<taxonomicName id="6D874D79AA17FFFCFF35FC89333EFCC0" box="[188,371,836,860]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="98F3EAE8AA17FFFCFF35FC89333EFCC0" box="[188,371,836,860]" italics="true" pageId="10" pageNumber="425">L. melanoleucus</emphasis>
</taxonomicName>
in the number of notes per call (1 or 3 versus 3–4), 
<taxonomicName id="6D874D79AA17FFFCFC3BFC893669FCC0" box="[946,1060,836,860]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="melicus">
<emphasis id="98F3EAE8AA17FFFCFC3BFC893669FCC0" box="[946,1060,836,860]" italics="true" pageId="10" pageNumber="425">L. melicus</emphasis>
</taxonomicName>
in the number of notes per call (1 or 3 versus 4–11), 
<taxonomicName id="6D874D79AA17FFFCFEE4FCA5338BFC1C" box="[365,454,872,896]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="solus">
<emphasis id="98F3EAE8AA17FFFCFEE4FCA5338BFC1C" box="[365,454,872,896]" italics="true" pageId="10" pageNumber="425">L. solus</emphasis>
</taxonomicName>
in the number of notes per call (1 or 3 versus 4–8) and 
<taxonomicName id="6D874D79AA17FFFCFBC3FCA53698FC1C" box="[1098,1237,872,896]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="98F3EAE8AA17FFFCFBC3FCA53698FC1C" box="[1098,1237,872,896]" italics="true" pageId="10" pageNumber="425">L. tuberosus</emphasis>
</taxonomicName>
in the number of notes per call (1 or 3 versus 1 single pulsed call). In lacking strong frequency modulation, the call of 
<taxonomicName id="6D874D79AA17FFFCFA90FC4137D1FC38" box="[1305,1436,908,932]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="botsfordi" status="sp. nov.">
<emphasis id="98F3EAE8AA17FFFCFA90FC4137D1FC38" box="[1305,1436,908,932]" italics="true" pageId="10" pageNumber="425">L. botsfordi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="83C05793AA17FFFCFF1EFC6332BDFC54" box="[151,240,943,968]" pageId="10" pageNumber="425" rank="species">
<emphasis id="98F3EAE8AA17FFFCFF1EFC6332BDFC54" bold="true" box="[151,240,943,968]" pageId="10" pageNumber="425">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="6D874D79AA17FFFCFE0DFC7D33AAFC54" box="[388,487,944,968]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="dringi">
<emphasis id="98F3EAE8AA17FFFCFE0DFC7D33AAFC54" box="[388,487,944,968]" italics="true" pageId="10" pageNumber="425">L. dringi</emphasis>
</taxonomicName>
and 
<taxonomicName id="6D874D79AA17FFFCFD95FC7D30C5FC54" box="[540,648,944,968]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="425" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="98F3EAE8AA17FFFCFD95FC7D30C5FC54" box="[540,648,944,968]" italics="true" pageId="10" pageNumber="425">L. hamidi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>