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<document id="F98AC59EAC7DF6F15D68F2FBA6FB75FF" ID-DOI="10.13133/2284-4880/765" ID-ISSN="2284-4880" ID-Zenodo-Dep="12762154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721238703073" checkinUser="felipe" docAuthor="Gardini, Giulio" docDate="2022" docId="D16187C89C7442115314FC038CCD7D94" docLanguage="en" docName="FragEntomol.54.1.119-132.pdf" docOrigin="Fragmenta entomologica (Frag. Entomol., Roma) 54 (1)" docSource="http://dx.doi.org/10.13133/2284-4880/765" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Acanthocreagris sardoa" docType="treatment" docVersion="2" lastPageNumber="131" masterDocId="2D58FFB09C7E421D502BFFD7895F7C5F" masterDocTitle="Two new species of Acanthocreagris from Corsica and mainland France, and notes on some congeneric species (Pseudoscorpiones: Neobisiidae)" masterLastPageNumber="132" masterPageNumber="119" pageNumber="129" updateTime="1721239365388" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="751C03F945D668AD3249A480E03453C9">Two new species of Acanthocreagris from Corsica and mainland France, and notes on some congeneric species (Pseudoscorpiones: Neobisiidae)</mods:title>
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<emphasis id="6BBCEACC9C7442175314FC038D197FB3" bold="true" box="[831,1094,980,1004]" italics="true" pageId="10" pageNumber="129">Acanthocreagris sardoa</emphasis>
(
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)
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<taxonomicName id="9EC84D5D9C7442175314FBC38D667873" ID-CoL="8VL7" baseAuthorityName="Beier" baseAuthorityYear="1959" box="[831,1081,1044,1068]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="129" phylum="Arthropoda" rank="species" species="sardoa">
<emphasis id="6BBCEACC9C7442175314FBC38D667873" box="[831,1081,1044,1068]" italics="true" pageId="10" pageNumber="129">Acanthocreagris sardoa</emphasis>
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:
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, figs 18 (in part: Dorgali, Grotta Pisanu; Orani, Grotta di San Francesco)
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<emphasis id="6BBCEACC9C7442175314FBA38D7678D3" bold="true" box="[831,1065,1140,1164]" pageId="10" pageNumber="129">Material examined.</emphasis>
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:
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:
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:
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,
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,
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,
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215
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/
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leg.
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<emphasis id="6BBCEACC9C7442175314FAC38D6E7973" bold="true" box="[831,1073,1300,1324]" pageId="10" pageNumber="129">Description of adults</emphasis>
[
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: according to
<bibRefCitation id="3D594B2F9C74421754D3FAC38CDA7973" author="Beier M." box="[1272,1413,1300,1324]" pageId="10" pageNumber="129" pagination="245 - 246" refId="ref12527" refString="Beier M. 1959. Neues uber Sardinische Hohlenpseudoscorpione. Annales des Speleologie, 14: 245 - 246." type="journal article" year="1959">Beier (1959)</bibRefCitation>
;
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: according to
<bibRefCitation id="3D594B2F9C74421753F9FAE38D257913" author="Gardini G." box="[978,1146,1332,1356]" pageId="10" pageNumber="129" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini (2018)</bibRefCitation>
]. Carapace, chelicera, pedipalps and palpal coxae red-brown pale. Carapace 1.22 (
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) or 1.37 (
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) times as long as broad, without eyes or eye-spots, anterior margin with slightly prominent epistome, apically rounded (
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, fig. 1); 22 macrosetae, anterior and posterior rows with 4 and 6 macrosetae, respectively. Chaetotaxy of tergites IX (
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) 6:6:7:9:9:10:10:11:10:9. Chaetotaxy of sternites IIXI (
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) 11:8:6:10:10:12:12:12:12:9; sternites III and IV (
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) each with 3 microsetae in front on each stigma, sternites VI-VIII with 2 discal setae; anal cone with 2+2 setae. Chelicera (
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,
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, fig. 2) 1.9 times as long as broad, palm with 6 setae (
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, 7 setae in left chelicera of
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); fixed finger with about 12 teeth; movable finger with 9 teeth;
<emphasis id="6BBCEACC9C74421755BFF9028CF37AB3" box="[1428,1452,1749,1772]" italics="true" pageId="10" pageNumber="129">gs</emphasis>
ratio 0.73, galea squat, slightly squared apically, with 3 or 4 silk ducts; rallum with 7 blades, the three distal ones laterally pinnate, the two median ones serrate, the two proximal ones finely dentate; serrulae interior and exteri- or respectively with about 22 and 25 blades. Manducatory process with 3 setae. Coxal setae: pedipalp 6, I 4, II 5, III 45, IV 7; anterolateral process of coxa I sharp, apically pointed. Pedipalp (
<bibRefCitation id="3D594B2F9C754216517CFB23888B7953" author="Beier M." box="[343,468,1268,1292]" pageId="11" pageNumber="130" pagination="245 - 246" refId="ref12527" refString="Beier M. 1959. Neues uber Sardinische Hohlenpseudoscorpione. Annales des Speleologie, 14: 245 - 246." type="journal article" year="1959">Beier 1959</bibRefCitation>
, fig. 1;
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, figs 36): trochanter with granular surface, 2.74 (
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) times as long as broad, without tubercles on paraxial face, with 4 spiniform setae with simple and acuminate tips; femur gradually widened in the proximal third, then parallel in the distal two thirds, 4.2 (
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) or 4.25 (
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) times as long as broad, with granular surface as in
<bibRefCitation id="3D594B2F9C7542165229FA638BC27993" author="Gardini G." box="[514,669,1460,1484]" pageId="11" pageNumber="130" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini (2018</bibRefCitation>
, fig. 3); patella 3.1 (
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) or 3.36 (
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) times as long as broad, club elongate and granular on paraxial face, pedicel granular, ratio between club and pedicel about 2.1 (
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) or 1.88 (
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); chela with pedicel 4.0 (
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) or 4.05 (
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) times as long as broad; hand of chela with pedicel 1.8 (
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<specimenCount id="4FCEFD579C754216526FF9838B0B7A33" box="[580,596,1620,1644]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
) times as long as broad, briefly oval in dorsal view, with a very convex paraxial profile; granular surface as in
<bibRefCitation id="3D594B2F9C7542165203F9438B9D7AF3" author="Gardini G." box="[552,706,1684,1708]" pageId="11" pageNumber="130" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini (2018</bibRefCitation>
, figs 45); fixed chelal finger with 79 (
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) small contiguous teeth with dental canals (
<bibRefCitation id="3D594B2F9C75421651B1F9038B717AB3" author="Gardini G." box="[410,558,1748,1772]" pageId="11" pageNumber="130" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini 2018</bibRefCitation>
, figs 56), venom duct short, nodus ramosus subterminal, subterminal sensillum as in
<bibRefCitation id="3D594B2F9C754216513BF8C388F57B73" author="Gardini G." box="[272,426,1812,1836]" pageId="11" pageNumber="130" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini (2018</bibRefCitation>
, fig. 6); movable chelal finger with 75 (
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) small contiguous, rounded teeth with dental canals (
<bibRefCitation id="3D594B2F9C75421650F1F88388317B33" author="Gardini G." box="[218,366,1876,1900]" pageId="11" pageNumber="130" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini 2018</bibRefCitation>
, figs 56) reaching back near
<emphasis id="6BBCEACC9C754216529CF8838B9B7B33" box="[695,708,1876,1900]" italics="true" pageId="11" pageNumber="130">b</emphasis>
, the distal six teeth pointed; movable chelal finger with four sensilla, two of which subterminal (
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, fig. 6), the others just distal and proximal to trichobothrium
<emphasis id="6BBCEACC9C7542165556FB038CCA78B3" box="[1405,1429,1236,1260]" italics="true" pageId="11" pageNumber="130">sb</emphasis>
, respectively (
<bibRefCitation id="3D594B2F9C75421653E3FB238D3D7953" author="Gardini G." box="[968,1122,1268,1292]" pageId="11" pageNumber="130" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini 2018</bibRefCitation>
, fig. 5); trichobothria as in
<bibRefCitation id="3D594B2F9C754216531AFAC38AEA7973" author="Beier M." box="[817,949,1300,1324]" pageId="11" pageNumber="130" pagination="245 - 246" refId="ref12527" refString="Beier M. 1959. Neues uber Sardinische Hohlenpseudoscorpione. Annales des Speleologie, 14: 245 - 246." type="journal article" year="1959">Beier (1959</bibRefCitation>
, fig. 1) and
<bibRefCitation id="3D594B2F9C754216541DFAC38D8D7973" author="Gardini G." box="[1078,1234,1300,1324]" pageId="11" pageNumber="130" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini (2018</bibRefCitation>
, figs 45), relative position of trichobothria along chelal axis (
<specimenCount id="4FCEFD579C754216553BFAE38C7C7913" box="[1296,1315,1332,1356]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
):
<emphasis id="6BBCEACC9C7542165510FAE28C107913" box="[1339,1359,1333,1356]" italics="true" pageId="11" pageNumber="130">et</emphasis>
0.15/
<emphasis id="6BBCEACC9C75421655A6FAE28CC37913" box="[1421,1436,1333,1356]" italics="true" pageId="11" pageNumber="130">it</emphasis>
0.26/
<emphasis id="6BBCEACC9C7542165345FA828AD37933" box="[878,908,1365,1388]" italics="true" pageId="11" pageNumber="130">est</emphasis>
0.31/
<emphasis id="6BBCEACC9C75421653E0FA828ABA7933" box="[971,997,1365,1388]" italics="true" pageId="11" pageNumber="130">ist</emphasis>
0.40/
<emphasis id="6BBCEACC9C7542165408FA838D1C7933" box="[1059,1091,1364,1388]" italics="true" pageId="11" pageNumber="130">isb</emphasis>
0.68/
<emphasis id="6BBCEACC9C75421654AAFA838DC97933" box="[1153,1174,1364,1388]" italics="true" pageId="11" pageNumber="130">ib</emphasis>
0.82/
<emphasis id="6BBCEACC9C75421654F0FA838DA07933" box="[1243,1279,1364,1388]" italics="true" pageId="11" pageNumber="130">esb</emphasis>
0.90/
<emphasis id="6BBCEACC9C7542165516FA838C097933" box="[1341,1366,1364,1388]" italics="true" pageId="11" pageNumber="130">eb</emphasis>
0.95/
<emphasis id="6BBCEACC9C75421655BEFA828CC37933" box="[1429,1436,1365,1388]" italics="true" pageId="11" pageNumber="130">t</emphasis>
0.23/
<emphasis id="6BBCEACC9C7542165343FAA28A2579D3" box="[872,890,1397,1420]" italics="true" pageId="11" pageNumber="130">st</emphasis>
0.39/
<emphasis id="6BBCEACC9C7542165393FAA38A8F79D3" box="[952,976,1396,1420]" italics="true" pageId="11" pageNumber="130">sb</emphasis>
0.63/
<emphasis id="6BBCEACC9C7542165425FAA38D4479D3" box="[1038,1051,1396,1420]" italics="true" pageId="11" pageNumber="130">b</emphasis>
0.85; ratio between movable finger and hand of chela with pedicel about 1.3 (
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) or 1.36 (
<specimenCount id="4FCEFD579C7542165552FA438CD479F3" box="[1401,1419,1428,1452]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
); ratio between pedipalpal femur and movable finger 0.95 (
<specimenCount id="4FCEFD579C7542165312FA038A1179B3" box="[825,846,1492,1516]" count="1" pageId="11" pageNumber="130" type="male"></specimenCount>
) 0.91 (
<specimenCount id="4FCEFD579C754216538FFA038AE979B3" box="[932,950,1492,1516]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
); ratio between pedipalpal femur and carapace 1.11 (
<specimenCount id="4FCEFD579C7542165387FA238A9E7A53" box="[940,961,1524,1548]" count="1" pageId="11" pageNumber="130" type="male"></specimenCount>
) 1.12 (
<specimenCount id="4FCEFD579C754216543DFA238D777A53" box="[1046,1064,1524,1548]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
). Leg IV (
<specimenCount id="4FCEFD579C754216548FFA238DEA7A53" box="[1188,1205,1524,1548]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
) (
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, figs 78): femur + patella 3.68 times as long as deep, tibia 6.0 times, basitarsus 3.0 times, telotarsus 6.15 times as long as deep, tactile seta in the proximal half (TS = 0.30), ratio between basitarsus and telotarsus 0.6, subterminal seta furcate, slightly dentate, claws with a very small dorsal tooth.
</paragraph>
<caption id="0DB766569C75421650A3FBB38BCC78CD" ID-DOI="http://doi.org/10.5281/zenodo.12762164" ID-Zenodo-Dep="12762164" httpUri="https://zenodo.org/record/12762164/files/figure.png" pageId="11" pageNumber="130" startId="11.[136,167,1124,1143]" targetBox="[136,1436,168,1096]" targetPageId="11" targetType="figure">
<paragraph id="597736DE9C75421650A3FBB38BCC78CD" blockId="11.[135,1436,1124,1170]" pageId="11" pageNumber="130">
<emphasis id="6BBCEACC9C75421650A3FBB3898B7828" bold="true" box="[136,212,1124,1143]" pageId="11" pageNumber="130">Fig. 37 </emphasis>
Map depicting known distribution of
<taxonomicName id="9EC84D5D9C7542165239FBB38BBC7828" box="[530,739,1124,1143]" pageId="11" pageNumber="130">
<emphasis id="6BBCEACC9C7542165239FBB38BC07828" box="[530,671,1124,1143]" italics="true" pageId="11" pageNumber="130">Acanthocreagris</emphasis>
species
</taxonomicName>
from southeastern France, Italy, Corsica, Sardinia, Sicily and Maltese Archipelago (localities taken from
<bibRefCitation id="3D594B2F9C7542165168FBA888E878CD" author="Gardini G." box="[323,439,1151,1170]" pageId="11" pageNumber="130" pagination="1 - 73" refId="ref12652" refString="Gardini G. 1998. Il genere Acanthocreagris in Italia (Pseudoscorpionida, Neobisiidae). Fragmenta entomologica, 30 (1): 1 - 73." type="journal article" year="1998">Gardini 1998</bibRefCitation>
,
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and present paper).
</paragraph>
</caption>
<paragraph id="597736DE9C754211537DF9028BD57D73" blockId="11.[815,1437,1204,1932]" lastBlockId="12.[151,772,180,300]" lastPageId="12" lastPageNumber="131" pageId="11" pageNumber="130">
<emphasis id="6BBCEACC9C754216537DF9028AAE7AB3" box="[854,1009,1749,1772]" italics="true" pageId="11" pageNumber="130">Measurements</emphasis>
(mm). Body length 2.2 (
<specimenCount id="4FCEFD579C7542165529F9038C487AB3" box="[1282,1303,1748,1772]" count="1" pageId="11" pageNumber="130" type="male"></specimenCount>
) or 2.5 (
<specimenCount id="4FCEFD579C7542165550F9038CD27AB3" box="[1403,1421,1748,1772]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
). Carapace 0.60 × 0.49 (
<specimenCount id="4FCEFD579C7542165404F9238D1B7B53" box="[1071,1092,1780,1804]" count="1" pageId="11" pageNumber="130" type="male"></specimenCount>
) or 0.66 × 0.48 anteriorly (
<specimenCount id="4FCEFD579C7542165550F9238CD27B53" box="[1403,1421,1780,1804]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
). Chelicera 0.42 × 0.22 (
<specimenCount id="4FCEFD579C754216540EF8C38D687B73" box="[1061,1079,1812,1836]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
); movable finger length 0.28 (
<specimenCount id="4FCEFD579C7542165550F8C38CD27B73" box="[1403,1421,1812,1836]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
). Pedipalp: trochanter 0.48 × 0.175 (
<specimenCount id="4FCEFD579C7542165485F8E38D9F7B13" box="[1198,1216,1844,1868]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
); femur 0.67 × 0.16 (
<specimenCount id="4FCEFD579C7542165312F8838A117B33" box="[825,846,1876,1900]" count="1" pageId="11" pageNumber="130" type="male"></specimenCount>
) or 0.85 × 0.20 (
<specimenCount id="4FCEFD579C754216543DF8838D777B33" box="[1046,1064,1876,1900]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
); patella 0.61 × 0.20 (
<specimenCount id="4FCEFD579C754216550DF8838C647B33" box="[1318,1339,1876,1900]" count="1" pageId="11" pageNumber="130" type="male"></specimenCount>
) or 0.74 × 0.22 (
<specimenCount id="4FCEFD579C75421653A1F8A38AC37BD3" box="[906,924,1908,1932]" count="1" pageId="11" pageNumber="130" type="female"></specimenCount>
); chela with pedicel 1.21 × 0.30 (
<specimenCount id="4FCEFD579C754216553AF8A38C797BD3" box="[1297,1318,1908,1932]" count="1" pageId="11" pageNumber="130" type="male"></specimenCount>
) or 1.50 × 0.37 (depth 0.33) (
<specimenCount id="4FCEFD579C724211514DFF6388277C93" box="[358,376,180,204]" count="1" pageId="12" pageNumber="131" type="female"></specimenCount>
); hand with pedicel length 0.55 (
<specimenCount id="4FCEFD579C72421152CDFF638BA47C93" box="[742,763,180,204]" count="1" pageId="12" pageNumber="131" type="male"></specimenCount>
) or 0.68 (
<specimenCount id="4FCEFD579C72421150D3FF0388557CB3" box="[248,266,212,236]" count="1" pageId="12" pageNumber="131" type="female"></specimenCount>
); movable finger length 0.70 (
<specimenCount id="4FCEFD579C7242115271FF038B307CB3" box="[602,623,212,236]" count="1" pageId="12" pageNumber="131" type="male"></specimenCount>
) or 0.93 (
<specimenCount id="4FCEFD579C72421152CAFF038BAC7CB3" box="[737,755,212,236]" count="1" pageId="12" pageNumber="131" type="female"></specimenCount>
). Leg IV (
<specimenCount id="4FCEFD579C72421150DFFF2388597D53" box="[244,262,244,268]" count="1" pageId="12" pageNumber="131" type="female"></specimenCount>
): femur + patella 0.70 × 0.19; tibia 0.60 × 0.10; basitarsus 0.24 × 0.08; telotarsus 0.40 × 0.065.
</paragraph>
</subSubSection>
<subSubSection id="11D265559C72421150BCFE838CCD7D94" pageId="12" pageNumber="131" type="discussion">
<paragraph id="597736DE9C72421150BCFE8388D57F53" blockId="12.[150,773,340,1804]" pageId="12" pageNumber="131">
<emphasis id="6BBCEACC9C72421150BCFE8388577D33" bold="true" box="[151,264,340,364]" pageId="12" pageNumber="131">Remarks.</emphasis>
<bibRefCitation id="3D594B2F9C7242115138FE8388E57D33" author="Gardini G." box="[275,442,340,364]" pageId="12" pageNumber="131" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini (2018)</bibRefCitation>
redescribed
<taxonomicName id="9EC84D5D9C7242115264FE8388207DD3" authority="(Beier, 1959)" baseAuthorityName="Beier" baseAuthorityYear="1959" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="sardoa">
<emphasis id="6BBCEACC9C7242115264FE8389BE7DD3" italics="true" pageId="12" pageNumber="131">Acanthocreagris sardoa</emphasis>
(
<bibRefCitation id="3D594B2F9C72421150D8FEA388287DD3" author="Beier M." box="[243,375,372,396]" pageId="12" pageNumber="131" pagination="245 - 246" refId="ref12527" refString="Beier M. 1959. Neues uber Sardinische Hohlenpseudoscorpione. Annales des Speleologie, 14: 245 - 246." type="journal article" year="1959">Beier, 1959</bibRefCitation>
)
</taxonomicName>
on the basis of a female from the Grotta Pisanu near Dorgali, central-eastern
<collectingRegion id="9B0CF83C9C7242115258FE438B8E7DF3" box="[627,721,404,428]" country="Italy" name="Sardegna" pageId="12" pageNumber="131">Sardinia</collectingRegion>
, and a male—damaged, with broken chelae—from the Grotta Su Pittiolu de Gospuru near Armungia, southeastern
<collectingRegion id="9B0CF83C9C72421152FCFE03898E7E53" country="Italy" name="Sardegna" pageId="12" pageNumber="131">Sardinia</collectingRegion>
. Its redescription partially incorporated the original one of
<bibRefCitation id="3D594B2F9C72421150CAFDC388337E73" author="Beier M." box="[225,364,532,556]" pageId="12" pageNumber="131" pagination="245 - 246" refId="ref12527" refString="Beier M. 1959. Neues uber Sardinische Hohlenpseudoscorpione. Annales des Speleologie, 14: 245 - 246." type="journal article" year="1959">Beier (1959)</bibRefCitation>
, based on a male from the cave Pozzo di Tuttavista 216 Sa/NU near Galtellì, about
<quantity id="9E309B3B9C72421152A6FDE38B9A7E13" box="[653,709,564,588]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="131" unit="km" value="9.0">9 km</quantity>
from Dorgali. The species was considered by
<bibRefCitation id="3D594B2F9C724211524BFD838A5A7E33" author="Gardini G." box="[608,773,596,620]" pageId="12" pageNumber="131" pagination="1 - 73" refId="ref12652" refString="Gardini G. 1998. Il genere Acanthocreagris in Italia (Pseudoscorpionida, Neobisiidae). Fragmenta entomologica, 30 (1): 1 - 73." type="journal article" year="1998">Gardini (1998)</bibRefCitation>
phenetically similar to
<taxonomicName id="9EC84D5D9C7242115183FDA38B9F7ED3" authority="(Beier, 1966)" baseAuthorityName="Beier" baseAuthorityYear="1966" box="[424,704,628,652]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="agazzii">
<emphasis id="6BBCEACC9C7242115183FDA38B457ED3" box="[424,538,628,652]" italics="true" pageId="12" pageNumber="131">A. agazzii</emphasis>
(Beier, 1966)
</taxonomicName>
from Venetian pre-Alps, but probably related to
<taxonomicName id="9EC84D5D9C72421152AEFD438BA27EF3" authorityName="Gardini" authorityYear="2018" box="[645,765,660,684]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="foghesa">
<emphasis id="6BBCEACC9C72421152AEFD438BA27EF3" box="[645,765,660,684]" italics="true" pageId="12" pageNumber="131">A. foghesa</emphasis>
</taxonomicName>
, described in the same paper (
<bibRefCitation id="3D594B2F9C72421151CFFD638B247E93" author="Gardini G." box="[484,635,692,716]" pageId="12" pageNumber="131" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini 2018</bibRefCitation>
) on
<specimenCount id="4FCEFD579C7242115299FD6289897EB3" count="2" pageId="12" pageNumber="131" type="female">two females</specimenCount>
from the Grotta de Su Fenugu near Perdasdefogu, southeastern
<collectingRegion id="9B0CF83C9C7242115103FD2388D97F53" box="[296,390,756,780]" country="Italy" name="Sardegna" pageId="12" pageNumber="131">Sardinia</collectingRegion>
.
</paragraph>
<paragraph id="597736DE9C7242115096FCC38BD27B53" blockId="12.[150,773,340,1804]" pageId="12" pageNumber="131">
According to
<bibRefCitation id="3D594B2F9C724211517BFCC388AB7F73" author="Gardini G." box="[336,500,788,812]" pageId="12" pageNumber="131" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini (2018)</bibRefCitation>
,
<taxonomicName id="9EC84D5D9C72421151D6FCC38A5D7F73" baseAuthorityName="Beier" baseAuthorityYear="1959" box="[509,770,788,812]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="sardoa">
<emphasis id="6BBCEACC9C72421151D6FCC38A5D7F73" box="[509,770,788,812]" italics="true" pageId="12" pageNumber="131">Acanthocreagris sardoa</emphasis>
</taxonomicName>
differs from
<taxonomicName id="9EC84D5D9C724211510EFCE388C67F13" authorityName="Gardini" authorityYear="2018" box="[293,409,820,844]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="foghesa">
<emphasis id="6BBCEACC9C724211510EFCE388C67F13" box="[293,409,820,844]" italics="true" pageId="12" pageNumber="131">A. foghesa</emphasis>
</taxonomicName>
in the lesser troglomorphic facies: length of pedipalpal femur
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(4.24.25 times as long as broad) in
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<emphasis id="6BBCEACC9C72421151FBFCA38B627FD3" box="[464,573,884,908]" italics="true" pageId="12" pageNumber="131">A. sardoa</emphasis>
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(
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),
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(5.065.33 ×) in
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<emphasis id="6BBCEACC9C72421151BBFC438B587FF3" box="[400,519,916,940]" italics="true" pageId="12" pageNumber="131">A. foghesa</emphasis>
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(
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); length of patella
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(3.13.35 ×) in
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<emphasis id="6BBCEACC9C72421151D9FC638B027F93" box="[498,605,948,972]" italics="true" pageId="12" pageNumber="131">A. sardoa</emphasis>
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(
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), 1.0
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(3.843.93 ×) in
<taxonomicName id="9EC84D5D9C7242115155FC0388AE7FB3" authorityName="Gardini" authorityYear="2018" box="[382,497,980,1004]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="foghesa">
<emphasis id="6BBCEACC9C7242115155FC0388AE7FB3" box="[382,497,980,1004]" italics="true" pageId="12" pageNumber="131">A. foghesa</emphasis>
</taxonomicName>
(
<specimenCount id="4FCEFD579C724211522AFC038B4C7FB3" box="[513,531,980,1004]" count="1" pageId="12" pageNumber="131" type="female"></specimenCount>
); length of chela with pedicel about
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(4.04.04 ×) in
<taxonomicName id="9EC84D5D9C72421152BCFC238A5C7853" baseAuthorityName="Beier" baseAuthorityYear="1959" box="[663,771,1012,1036]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="sardoa">
<emphasis id="6BBCEACC9C72421152BCFC238A5C7853" box="[663,771,1012,1036]" italics="true" pageId="12" pageNumber="131">A. sardoa</emphasis>
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(
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),
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(4.76 5.04 ×) in
<taxonomicName id="9EC84D5D9C724211527DFBC38B937873" authorityName="Gardini" authorityYear="2018" box="[598,716,1044,1068]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="foghesa">
<emphasis id="6BBCEACC9C724211527DFBC38B937873" box="[598,716,1044,1068]" italics="true" pageId="12" pageNumber="131">A. foghesa</emphasis>
</taxonomicName>
(
<specimenCount id="4FCEFD579C72421152CBFBC38BAD7873" box="[736,754,1044,1068]" count="1" pageId="12" pageNumber="131" type="female"></specimenCount>
); ratio between pedipalpal femur and carapace
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<taxonomicName id="9EC84D5D9C7242115098FB8388447833" baseAuthorityName="Beier" baseAuthorityYear="1959" box="[179,283,1108,1132]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="sardoa">
<emphasis id="6BBCEACC9C7242115098FB8388447833" box="[179,283,1108,1132]" italics="true" pageId="12" pageNumber="131">A. sardoa</emphasis>
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(
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),
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<taxonomicName id="9EC84D5D9C72421151D0FB838B307833" authorityName="Gardini" authorityYear="2018" box="[507,623,1108,1132]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="foghesa">
<emphasis id="6BBCEACC9C72421151D0FB838B307833" box="[507,623,1108,1132]" italics="true" pageId="12" pageNumber="131">A. foghesa</emphasis>
</taxonomicName>
(
<specimenCount id="4FCEFD579C72421152ABFB838BCD7833" box="[640,658,1108,1132]" count="1" pageId="12" pageNumber="131" type="female"></specimenCount>
). Also the shape of chelal hand—a character overlooked by
<bibRefCitation id="3D594B2F9C724211529BFBA3898278F3" author="Gardini G." pageId="12" pageNumber="131" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini (2018)</bibRefCitation>
—is different in the two species: briefly oval in dorsal view, with a very convex paraxial profile, in
<taxonomicName id="9EC84D5D9C72421152C7FB6389BE78B3" baseAuthorityName="Beier" baseAuthorityYear="1959" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="sardoa">
<emphasis id="6BBCEACC9C72421152C7FB6389BE78B3" italics="true" pageId="12" pageNumber="131">A. sardoa</emphasis>
</taxonomicName>
(
<bibRefCitation id="3D594B2F9C72421150DFFB03882B78B3" author="Beier M." box="[244,372,1236,1260]" pageId="12" pageNumber="131" pagination="245 - 246" refId="ref12527" refString="Beier M. 1959. Neues uber Sardinische Hohlenpseudoscorpione. Annales des Speleologie, 14: 245 - 246." type="journal article" year="1959">Beier 1959</bibRefCitation>
, fig. 1;
<bibRefCitation id="3D594B2F9C72421151FBFB038B3878B3" author="Gardini G." box="[464,615,1236,1260]" pageId="12" pageNumber="131" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini 2018</bibRefCitation>
, fig. 4), elongated oval, with a slightly convex paraxial profile, in
<taxonomicName id="9EC84D5D9C72421152C7FB2389B37973" authorityName="Gardini" authorityYear="2018" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="foghesa">
<emphasis id="6BBCEACC9C72421152C7FB2389B37973" italics="true" pageId="12" pageNumber="131">A. foghesa</emphasis>
</taxonomicName>
(
<bibRefCitation id="3D594B2F9C72421150D7FAC388CC7973" author="Gardini G." box="[252,403,1300,1324]" pageId="12" pageNumber="131" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini 2018</bibRefCitation>
, fig. 12). This last character cannot be evaluated in the male from the cave near Armungia, as both hands of its palps are broken, which makes its attribution to
<taxonomicName id="9EC84D5D9C724211517EFAA38B3179D3" authority="(Gardini 2018)" baseAuthorityName="Gardini" baseAuthorityYear="2018" box="[341,622,1396,1420]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="sardoa">
<emphasis id="6BBCEACC9C724211517EFAA388E079D3" box="[341,447,1396,1420]" italics="true" pageId="12" pageNumber="131">A. sardoa</emphasis>
(
<bibRefCitation id="3D594B2F9C72421151FAFAA38B3879D3" author="Gardini G." box="[465,615,1396,1420]" pageId="12" pageNumber="131" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini 2018</bibRefCitation>
)
</taxonomicName>
uncertain, to be evaluated only with the examination of further material. The cave near Armungia is about
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from the
<typeStatus id="8673887C9C72421150B2FA02899779B2" box="[153,200,1493,1517]" pageId="12" pageNumber="131">type</typeStatus>
locality of
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<emphasis id="6BBCEACC9C7242115165FA03889A79B3" box="[334,453,1492,1516]" italics="true" pageId="12" pageNumber="131">A. foghesa</emphasis>
</taxonomicName>
and about
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from that of
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<emphasis id="6BBCEACC9C724211509DFA23887F7A53" box="[182,288,1524,1548]" italics="true" pageId="12" pageNumber="131">A. sardoa</emphasis>
</taxonomicName>
, each in karst complexes independent from each other (G. Grafitti, in litt.:
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). The redescription of
<taxonomicName id="9EC84D5D9C7242115160F9E38B0F7A13" baseAuthorityName="Gardini" baseAuthorityYear="2018" box="[331,592,1588,1612]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="sardoa">
<emphasis id="6BBCEACC9C7242115160F9E38B0F7A13" box="[331,592,1588,1612]" italics="true" pageId="12" pageNumber="131">Acanthocreagris sardoa</emphasis>
</taxonomicName>
proposed above does not include the data relating to the male of Armungia and is therefore based only on the original description of
<taxonomicName id="9EC84D5D9C724211509EF94388EF7AF3" authority="(Beier 1959)" baseAuthorityName="Beier" baseAuthorityYear="1959" box="[181,432,1684,1708]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="sardoa">
<emphasis id="6BBCEACC9C724211509EF94388427AF3" box="[181,285,1684,1708]" italics="true" pageId="12" pageNumber="131">A. sardoa</emphasis>
(
<bibRefCitation id="3D594B2F9C7242115107F94388F77AF3" author="Beier M." box="[300,424,1684,1708]" pageId="12" pageNumber="131" pagination="245 - 246" refId="ref12527" refString="Beier M. 1959. Neues uber Sardinische Hohlenpseudoscorpione. Annales des Speleologie, 14: 245 - 246." type="journal article" year="1959">Beier 1959</bibRefCitation>
)
</taxonomicName>
and on that of the female from Dorgali (
<bibRefCitation id="3D594B2F9C72421150D3F96388D07A93" author="Gardini G." box="[248,399,1716,1740]" pageId="12" pageNumber="131" pagination="19 - 29" refId="ref12680" refString="Gardini G. 2018. New species and records of the pseudoscorpion genus Acanthocreagris from Italy (Pseudoscorpiones: Neobisiidae). Fragmenta entomologica, 50 (1): 19 - 29." type="journal article" year="2018">Gardini 2018</bibRefCitation>
). The male from the cave near Armungia is doubtfully attributed to
<taxonomicName id="9EC84D5D9C7242115239F9038BD87AB3" authorityName="Gardini" authorityYear="2018" box="[530,647,1748,1772]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="foghesa">
<emphasis id="6BBCEACC9C7242115239F9038BD87AB3" box="[530,647,1748,1772]" italics="true" pageId="12" pageNumber="131">A. foghesa</emphasis>
</taxonomicName>
and briefly described (see Remarks under
<taxonomicName id="9EC84D5D9C72421151CDF9238B227B53" authorityName="Gardini" authorityYear="2018" box="[486,637,1780,1804]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" isUncertain="true" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="foghesa">
<emphasis id="6BBCEACC9C72421151CDF92388A17B54" box="[486,510,1780,1803]" italics="true" pageId="12" pageNumber="131">A.</emphasis>
cf.
<emphasis id="6BBCEACC9C7242115203F9238B227B53" box="[552,637,1780,1804]" italics="true" pageId="12" pageNumber="131">foghesa</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="597736DE9C7242115314FF618CCD7D94" blockId="12.[831,1453,182,460]" pageId="12" pageNumber="131">
Noël Dubois (Fédération française de spéléologie), the biospeleologists of the Ligue Insulaire Spéléologique
<collectingRegion id="9B0CF83C9C72421154DAFF018C747CB4" box="[1265,1323,214,235]" country="France" name="Corse" pageId="12" pageNumber="131">Corse</collectingRegion>
, Josiane Lips (Villeurbanne), Carlo Onnis (Unione Speleologica Cagliaritana, Cagliari) and Cosimo Baviera (University of Messina). Giuseppe Grafitti (Gruppo Speleologico Sassarese, Sassari) kindly provided information on Sardinian caves. Helpful comments on the manuscript were made by Carlo Giusto (Genoa) and Pietro Gardini (Genoa) kindly provided invaluable expertise in digitizing plates of the drawings and production of the distribution map.
</paragraph>
</subSubSection>
</treatment>
</document>