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<document id="A150487CED094B3EF0A895C8FC5E0C0E" ID-CLB-Dataset="51311" ID-DOI="10.5281/zenodo.156493" ID-GBIF-Dataset="c3fd5e49-3b05-4a99-9bc8-455ae3056faa" ID-ISSN="1175­5326" ID-Zenodo-Dep="156493" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459617947450" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="0846107D83356D4AFEFEFEB4FAD6FC79" docLanguage="en" docName="zt00239.pdf" docOrigin="Zootaxa 239" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Stethusa" docType="key" docVersion="13" lastPageNumber="12" masterDocId="F47F6805833E6D41FFF6FFADFFFCFFAF" masterDocTitle="A revision of the Nearctic species of the genus Stethusa Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="43" masterPageNumber="1" pageNumber="12" updateTime="1698204628127" updateUser="ExternalLinkService">
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<mods:title id="7F6A4E18476D05054FB23728D5574DDD">A revision of the Nearctic species of the genus Stethusa Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="6FADCDE005F6BD3B05F5B74ECE3B9ACB">Gusarov, Vladimir I.</mods:namePart>
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<treatment id="0846107D83356D4AFEFEFEB4FAD6FC79" ID-DOI="http://doi.org/10.5281/zenodo.6275923" ID-GBIF-Taxon="119331777" ID-Zenodo-Dep="6275923" LSID="urn:lsid:plazi:treatment:0846107D83356D4AFEFEFEB4FAD6FC79" httpUri="http://treatment.plazi.org/id/0846107D83356D4AFEFEFEB4FAD6FC79" lastPageNumber="12" pageId="11" pageNumber="12">
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Key to Nearctic species of
<taxonomicName id="47EFDAE883356D4AFDB9FEB4FD4AFE9C" ID-CoL="8VS4C" box="[591,694,281,307]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B29B7D7983356D4AFDB9FEB4FD4AFE9C" bold="true" box="[591,694,281,307]" italics="true" pageId="11" pageNumber="12">Stethusa</emphasis>
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</subSubSection>
<subSubSection id="C8F5F2E083356D4AFEFEFEC9FAD6FC79" pageId="11" pageNumber="12" type="key">
<keyStep id="3B1BB9CE83356D4AFEFEFEC9FAD7FD49" pageId="11" pageNumber="12">
<paragraph id="8050A16B83356D4AFEFEFEC9FAD6FDE9" blockId="11.[264,1324,356,982]" pageId="11" pageNumber="12">
<keyLead id="3B1E025E83356D4AFEFEFEC9FAD6FDE9" pageId="11" pageNumber="12">
1 Antennal article 11 with two subbasal impressions densely covered with microsetae (
<figureCitation id="18D4BDEE83356D4AFEB7FE21FE30FE09" box="[321,460,396,422]" captionStart="FIGURES 14 ­ 17" captionStartId="9.[264,382,1243,1267]" captionTargetBox="[347,1240,317,1218]" captionTargetId="figure@9.[337,1249,307,1227]" captionTargetPageId="9" captionText="FIGURES 14 ­ 17. Details of Stethusa dichroa (Gravenhorst) (male (14 ­ 16) and female (17), Eddyville, Illinois). 14 right mesotibia; 15 antennal article 11; 16 right antenna; 17 right metatarsus. Scale bar 0.1 mm (15, 17), 0.2 mm (14), 0.4 mm (16)." httpUri="https://zenodo.org/record/156497/files/figure.png" pageId="11" pageNumber="12">Figs. 15­16</figureCitation>
). Internal sac of aedeagus with two strong distal spines (
<figureCitation id="18D4BDEE83356D4AFB9AFE21FEA5FE61" captionStart="FIGURES 34 ­ 39" captionStartId="18.[264,382,1757,1781]" captionTargetBox="[284,1296,654,1714]" captionTargetId="figure@18.[274,1313,646,1730]" captionTargetPageId="18" captionText="FIGURES 34 ­ 39. Aedeagus of Stethusa dichroa (Gravenhorst) (males, 12 km NNE Lawrence, Kansas (35 ­ 36, 38 ­ 39); 18 km SSE Lawrence (34, 37 )). 34 everted internal sac, parameral view; 35 ­ 36 details of retracted internal sac, abparameral view; 37 everted internal sac, lateral view; 38 ­ 39 details of retracted internal sac, lateral view. SB ­ sclerotized band. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156501/files/figure.png" pageId="11" pageNumber="12">Figs. 34­35, 37­ 38</figureCitation>
). Female with accessory sclerite (
<figureCitation id="18D4BDEE83356D4AFD1CFE19FCBEFE61" box="[746,834,436,462]" captionStart="FIGURES 40 ­ 42" captionStartId="19.[264,382,1234,1258]" captionTargetBox="[344,1251,575,1191]" captionTargetId="figure@19.[335,1251,566,1207]" captionTargetPageId="19" captionText="FIGURES 40 ­ 42. Details of Stethusa dichroa (Gravenhorst) (female, 12 km NNE Lawrence, Kansas). 40 female abdominal segments 9 ­ 10, ventral view; 41 ­ 42 spermatheca of the same specimen illustrated at different angles. The arrow indicates the point where the spermathecal gland is attached. Scale bar 0.2 mm (40), 0.1 mm (41 ­ 42)." httpUri="https://zenodo.org/record/156502/files/figure.png" pageId="11" pageNumber="12">Fig. 40</figureCitation>
). Distal end of spermatheca bent towards spermathecal gland (
<figureCitation id="18D4BDEE83356D4AFDDBFE71FD45FE59" box="[557,697,476,502]" captionStart="FIGURES 40 ­ 42" captionStartId="19.[264,382,1234,1258]" captionTargetBox="[344,1251,575,1191]" captionTargetId="figure@19.[335,1251,566,1207]" captionTargetPageId="19" captionText="FIGURES 40 ­ 42. Details of Stethusa dichroa (Gravenhorst) (female, 12 km NNE Lawrence, Kansas). 40 female abdominal segments 9 ­ 10, ventral view; 41 ­ 42 spermatheca of the same specimen illustrated at different angles. The arrow indicates the point where the spermathecal gland is attached. Scale bar 0.2 mm (40), 0.1 mm (41 ­ 42)." httpUri="https://zenodo.org/record/156502/files/figure.png" pageId="11" pageNumber="12">Figs. 41­42</figureCitation>
). Body length 2.0­3.0 mm, pronotal length
<quantity id="47170C8E83356D4AFB4CFE71FE96FDB1" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.3" metricValueMin="4.1" pageId="11" pageNumber="12" unit="mm" value="0.47" valueMax="0.53" valueMin="0.41">0.41­0.53 mm</quantity>
. Widely distributed in the eastern
<collectingCountry id="F8F8E1FB83356D4AFCEBFDA9FC41FDB1" box="[797,957,516,542]" name="United States of America" pageId="11" pageNumber="12">United States</collectingCountry>
(
<figureCitation id="18D4BDEE83356D4AFC27FDA9FBC7FDB1" box="[977,1083,516,542]" captionStart="FIGURE 110" captionStartId="39.[264,368,1695,1719]" captionTargetBox="[349,1238,846,1668]" captionTargetId="figure@39.[349,1239,846,1668]" captionTargetPageId="39" captionText="FIGURE 110. Geographical distribution of Stethusa dichroa (Gravenhorst) in North America. The exact localities in Vermont and Maine are unknown." httpUri="https://zenodo.org/record/156516/files/figure.png" pageId="11" pageNumber="12">Fig. 110</figureCitation>
). Also known from
<collectingCountry id="F8F8E1FB83356D4AFECEFD81FE6DFDE9" box="[312,401,556,582]" name="Bolivia" pageId="11" pageNumber="12">Bolivia</collectingCountry>
,
<collectingCountry id="F8F8E1FB83356D4AFE6BFD81FDF7FDE9" box="[413,523,556,582]" name="Paraguay" pageId="11" pageNumber="12">Paraguay</collectingCountry>
, the Galápagos Islands and the Antilles..1.
<taxonomicName id="47EFDAE883356D4AFBF1FD81FAD6FDE9" ID-CoL="52GKB" authority="Gravenhorst" authorityName="Gravenhorst" box="[1031,1322,556,582]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="B29B7D7983356D4AFBF1FD81FB83FDEA" box="[1031,1151,556,581]" italics="true" pageId="11" pageNumber="12">S. dichroa</emphasis>
(Gravenhorst)
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</paragraph>
<paragraph id="8050A16B83356D4AFEFEFDF9FAD7FD49" blockId="11.[264,1324,356,982]" pageId="11" pageNumber="12">
<keyLead id="3B1E025E83356D4AFEFEFDF9FAD7FD49" pageId="11" pageNumber="12">
Antennal article 11 without subbasal impressions, microsetae evenly distributed. Internal sac of aedeagus without strong distal spines (
<figureCitation id="18D4BDEE83356D4AFC3DFDD1FBA4FD39" box="[971,1112,636,662]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 details of retracted internal sac, abparameral view; 54 details of retracted internal sac, lateral view; 55 copulatory piece, lateral view; 56 copulatory piece, abparameral view; 57 spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="11" pageNumber="12">Figs. 53­54</figureCitation>
,
<figureCitation id="18D4BDEE83356D4AFB9EFDD1FB4DFD39" box="[1128,1201,636,662]" captionStart="FIGURES 63 ­ 70" captionStartId="27.[264,382,1254,1278]" captionTargetBox="[306,1276,315,1222]" captionTargetId="figure@27.[298,1289,306,1231]" captionTargetPageId="27" captionText="FIGURES 63 ­ 70. Aedeagus and spermatheca of Stethusa spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (63 ­ 69); and female 9 km W Buffalo Gap, Virginia (70 )). 63 median lobe, parameral view; 64 apex of median lobe, parameral view; 65 median lobe, lateral view; 66 apex of median lobe, lateral view; 67 details of retracted internal sac, abparameral view; 68 details of retracted internal sac, lateral view; 69 apex of left paramere; 70 spermatheca. Scale bar 0.1 mm (64, 66, 69 ­ 70), 0.2 mm (63, 65, 67 ­ 68)." httpUri="https://zenodo.org/record/156507/files/figure.png" pageId="11" pageNumber="12">67­68</figureCitation>
). Female without accessory sclerite. Distal end of spermatheca bent away from spermathecal gland (
<figureCitation id="18D4BDEE83356D4AFE7FFD61FE17FD49" box="[393,491,716,742]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 details of retracted internal sac, abparameral view; 54 details of retracted internal sac, lateral view; 55 copulatory piece, lateral view; 56 copulatory piece, abparameral view; 57 spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="11" pageNumber="12">Figs. 57</figureCitation>
,
<figureCitation id="18D4BDEE83356D4AFE01FD61FDEAFD49" box="[503,534,716,742]" captionStart="FIGURES 63 ­ 70" captionStartId="27.[264,382,1254,1278]" captionTargetBox="[306,1276,315,1222]" captionTargetId="figure@27.[298,1289,306,1231]" captionTargetPageId="27" captionText="FIGURES 63 ­ 70. Aedeagus and spermatheca of Stethusa spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (63 ­ 69); and female 9 km W Buffalo Gap, Virginia (70 )). 63 median lobe, parameral view; 64 apex of median lobe, parameral view; 65 median lobe, lateral view; 66 apex of median lobe, lateral view; 67 details of retracted internal sac, abparameral view; 68 details of retracted internal sac, lateral view; 69 apex of left paramere; 70 spermatheca. Scale bar 0.1 mm (64, 66, 69 ­ 70), 0.2 mm (63, 65, 67 ­ 68)." httpUri="https://zenodo.org/record/156507/files/figure.png" pageId="11" pageNumber="12">70</figureCitation>
) ....................................................................................................... 2
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</keyStep>
<keyStep id="3B1BB9CE83356D4AFEFEFD59FAD6FC79" pageId="11" pageNumber="12">
<paragraph id="8050A16B83356D4AFEFEFD59FAD6FCF1" blockId="11.[264,1324,356,982]" pageId="11" pageNumber="12">
<keyLead id="3B1E025E83356D4AFEFEFD59FAD6FCF1" pageId="11" pageNumber="12">
2 Body larger, length
<quantity id="47170C8E83356D4AFDD4FD59FD52FCA1" box="[546,686,756,782]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="11" pageNumber="12" unit="mm" value="3.35" valueMax="3.5" valueMin="3.2">3.2­3.5 mm</quantity>
, pronotal length
<quantity id="47170C8E83356D4AFC83FD59FBE3FCA1" box="[885,1055,756,782]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="6.1" metricValueMin="5.4" pageId="11" pageNumber="12" unit="mm" value="0.575" valueMax="0.61" valueMin="0.54">0.54­0.61 mm</quantity>
. Apex of median lobe in lateral view straight (
<figureCitation id="18D4BDEE83356D4AFDABFCB1FD17FC99" box="[605,747,796,822]" captionStart="FIGURES 48 ­ 52" captionStartId="22.[264,382,1402,1426]" captionTargetBox="[288,1293,492,1373]" captionTargetId="figure@22.[278,1309,478,1384]" captionTargetPageId="22" captionText="FIGURES 48 ­ 52. Aedeagus of Stethusa klimschi (Bernhauer) (lectotype (48 ­ 51); and male from Vicksburg, Mississippi (52 )). 48 median lobe, parameral view; 49 apex of median lobe, parameral view; 50 median lobe, lateral view; 51 apex of median lobe, lateral view; 52 apex of left paramere. Scale bar 0.1 mm (49, 51 ­ 52), 0.2 mm (48, 50)." httpUri="https://zenodo.org/record/156504/files/figure.png" pageId="11" pageNumber="12">Figs. 50­51</figureCitation>
). Spermatheca longer (
<figureCitation id="18D4BDEE83356D4AFBF9FCB1FB97FC99" box="[1039,1131,796,822]" captionStart="FIGURES 53 ­ 57" captionStartId="23.[264,382,1144,1168]" captionTargetBox="[284,1298,315,1116]" captionTargetId="figure@23.[281,1306,308,1121]" captionTargetPageId="23" captionText="FIGURES 53 ­ 57. Aedeagus and spermatheca of Stethusa klimschi (Bernhauer) (lectotype (53 ­ 54); male (55 ­ 56) and female (57) from Vicksburg, Mississippi). 53 details of retracted internal sac, abparameral view; 54 details of retracted internal sac, lateral view; 55 copulatory piece, lateral view; 56 copulatory piece, abparameral view; 57 spermatheca. Scale bar 0.2 mm (53 ­ 54), 0.1 mm (55 ­ 57)." httpUri="https://zenodo.org/record/156505/files/figure.png" pageId="11" pageNumber="12">Fig. 57</figureCitation>
). Known from Louisiana, Mississippi and southern Indiana (
<figureCitation id="18D4BDEE83356D4AFCBAFCE9FC51FCF1" box="[844,941,836,862]" captionStart="FIGURE 111" captionStartId="40.[264,367,1214,1238]" captionTargetBox="[349,1238,316,1190]" captionTargetId="figure@40.[349,1239,316,1190]" captionTargetPageId="40" captionText="FIGURE 111. Geographical distribution of Stethusa klimschi (Bernhauer) and S. spuriella (Casey). The exact locality in Ohio is unknown." httpUri="https://zenodo.org/record/156517/files/figure.png" pageId="11" pageNumber="12">Fig. 111</figureCitation>
)........ 2.
<taxonomicName id="47EFDAE883356D4AFBE1FCE9FAD6FCF1" ID-CoL="52GKD" authority="Bernhauer" authorityName="Bernhauer" box="[1047,1322,836,862]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="klimschi">
<emphasis id="B29B7D7983356D4AFBE1FCE9FB6AFCF2" box="[1047,1174,836,861]" italics="true" pageId="11" pageNumber="12">S. klimschi</emphasis>
(Bernhauer)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8050A16B83356D4AFEFEFCC1FAD6FC79" blockId="11.[264,1324,356,982]" pageId="11" pageNumber="12">
<keyLead id="3B1E025E83356D4AFEFEFCC1FAD6FC79" pageId="11" pageNumber="12">
Body smaller, length
<quantity id="47170C8E83356D4AFDC8FCC1FD31FC29" box="[574,717,876,902]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="11" pageNumber="12" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.1­2.5 mm</quantity>
, pronotal length
<quantity id="47170C8E83356D4AFC68FCC1FBB7FC29" box="[926,1099,876,902]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="4.4" metricValueMin="3.5999999999999996" pageId="11" pageNumber="12" unit="mm" value="0.4" valueMax="0.44" valueMin="0.36">0.36­0.44 mm</quantity>
. Apex of median lobe in lateral view bent paramerally (
<figureCitation id="18D4BDEE83356D4AFCFCFC39FC64FC01" box="[778,920,916,942]" captionStart="FIGURES 63 ­ 70" captionStartId="27.[264,382,1254,1278]" captionTargetBox="[306,1276,315,1222]" captionTargetId="figure@27.[298,1289,306,1231]" captionTargetPageId="27" captionText="FIGURES 63 ­ 70. Aedeagus and spermatheca of Stethusa spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (63 ­ 69); and female 9 km W Buffalo Gap, Virginia (70 )). 63 median lobe, parameral view; 64 apex of median lobe, parameral view; 65 median lobe, lateral view; 66 apex of median lobe, lateral view; 67 details of retracted internal sac, abparameral view; 68 details of retracted internal sac, lateral view; 69 apex of left paramere; 70 spermatheca. Scale bar 0.1 mm (64, 66, 69 ­ 70), 0.2 mm (63, 65, 67 ­ 68)." httpUri="https://zenodo.org/record/156507/files/figure.png" pageId="11" pageNumber="12">Figs. 65­66</figureCitation>
). Spermatheca shorter (
<figureCitation id="18D4BDEE83356D4AFB36FC39FAE0FC01" box="[1216,1308,916,942]" captionStart="FIGURES 63 ­ 70" captionStartId="27.[264,382,1254,1278]" captionTargetBox="[306,1276,315,1222]" captionTargetId="figure@27.[298,1289,306,1231]" captionTargetPageId="27" captionText="FIGURES 63 ­ 70. Aedeagus and spermatheca of Stethusa spuriella (Casey) (male, 18 km SSE Lawrence, Kansas (63 ­ 69); and female 9 km W Buffalo Gap, Virginia (70 )). 63 median lobe, parameral view; 64 apex of median lobe, parameral view; 65 median lobe, lateral view; 66 apex of median lobe, lateral view; 67 details of retracted internal sac, abparameral view; 68 details of retracted internal sac, lateral view; 69 apex of left paramere; 70 spermatheca. Scale bar 0.1 mm (64, 66, 69 ­ 70), 0.2 mm (63, 65, 67 ­ 68)." httpUri="https://zenodo.org/record/156507/files/figure.png" pageId="11" pageNumber="12">Fig. 70</figureCitation>
). Widely distributed in the eastern
<collectingCountry id="F8F8E1FB83356D4AFD4AFC11FCABFC79" box="[700,855,956,982]" name="United States of America" pageId="11" pageNumber="12">United States</collectingCountry>
(
<figureCitation id="18D4BDEE83356D4AFC9EFC11FC34FC79" box="[872,968,956,982]" captionStart="FIGURE 111" captionStartId="40.[264,367,1214,1238]" captionTargetBox="[349,1238,316,1190]" captionTargetId="figure@40.[349,1239,316,1190]" captionTargetPageId="40" captionText="FIGURE 111. Geographical distribution of Stethusa klimschi (Bernhauer) and S. spuriella (Casey). The exact locality in Ohio is unknown." httpUri="https://zenodo.org/record/156517/files/figure.png" pageId="11" pageNumber="12">Fig. 111</figureCitation>
).......... 3.
<taxonomicName id="47EFDAE883356D4AFBB5FC11FAD6FC79" ID-CoL="52GKH" authority="Casey" authorityName="Casey" box="[1091,1322,956,982]" class="Insecta" family="Staphylinidae" genus="Stethusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="spuriella">
<emphasis id="B29B7D7983356D4AFBB5FC11FB34FC7A" box="[1091,1224,956,981]" italics="true" pageId="11" pageNumber="12">S. spuriella</emphasis>
(Casey)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>