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<document id="06AF74FCECB6945DA586FFC78B80949D" ID-CLB-Dataset="20537" ID-DOI="10.15553/c2018v732a7" ID-GBIF-Dataset="4c48979e-961e-4f52-aea5-35622edab76f" ID-ISSN="2235-3658" ID-Zenodo-Dep="5724607" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637764901401" checkinUser="carolina" docAuthor="Applequist, Wendy L." docDate="2018" docId="0B2CC003FFE8146A6827D3B24A6042E5" docLanguage="en" docName="Candollea.73.2.221-244.pdf" docOrigin="Candollea 73 (2)" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Homalium fortunatii Appleq. 2018, spec. nova" docType="treatment" docVersion="12" lastPageNumber="232" masterDocId="F715B87BFFE314616856D2104F69413B" masterDocTitle="A revision of the Malagasy species of Homalium sect. Blackwellia (Salicaceae)" masterLastPageNumber="244" masterPageNumber="221" pageNumber="232" updateTime="1699470032420" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="9E0656B773656237791FC104241A87AC">A revision of the Malagasy species of Homalium sect. Blackwellia (Salicaceae)</mods:title>
</mods:titleInfo>
<mods:name id="739556CBAF7F285E593854DB6E73B4A5" type="personal">
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<mods:namePart id="39F45993FBFD31F6166DA09932980734">Applequist, Wendy L.</mods:namePart>
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<mods:typeOfResource id="8A889196F6E52FA95C74C91E6313479B">text</mods:typeOfResource>
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<mods:title id="E2CBF1CE595E6A530035B34E038483BD">Candollea</mods:title>
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<mods:part id="D5831978787649E3BE46EEF88F873CB7">
<mods:date id="869FFB406DCC4F60B0A583CF60E683B0">2018</mods:date>
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<mods:number id="85E3A941D3A410EB126DB851192589B0">2018-11-05</mods:number>
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<mods:number id="942014F3BD3CB0C62AD0B03071724C5A">73</mods:number>
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<treatment id="0B2CC003FFE8146A6827D3B24A6042E5" ID-DOI="http://doi.org/10.5281/zenodo.6314437" ID-GBIF-Taxon="193438785" ID-Zenodo-Dep="6314437" LSID="urn:lsid:plazi:treatment:0B2CC003FFE8146A6827D3B24A6042E5" httpUri="http://treatment.plazi.org/id/0B2CC003FFE8146A6827D3B24A6042E5" lastPageNumber="232" pageId="11" pageNumber="232">
<subSubSection id="CB9F229EFFE8146A6827D3B24DC54082" box="[113,684,416,442]" pageId="11" pageNumber="232" type="nomenclature">
<paragraph id="833A7115FFE8146A6827D3B24DC54082" blockId="11.[113,684,416,442]" box="[113,684,416,442]" pageId="11" pageNumber="232">
<heading id="D872C679FFE8146A6827D3B24DC54082" box="[113,684,416,442]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="232" reason="2">
7.
<taxonomicName id="44850A96FFE8146A68C1D3B14EBC4081" ID-CoL="3MCL4" authority="Appleq." authorityName="Appleq." authorityYear="2018" box="[151,469,417,442]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="fortunatii" status="sp. nov.">
<emphasis id="B1F1AD07FFE8146A68C1D3B14E104082" bold="true" box="[151,377,417,441]" italics="true" pageId="11" pageNumber="232">Homalium fortunatii</emphasis>
Appleq.
</taxonomicName>
,
<emphasis id="B1F1AD07FFE8146A6989D3B24D274081" bold="true" box="[479,590,418,442]" pageId="11" pageNumber="232">
<taxonomicNameLabel id="AAC2107CFFE8146A6989D3B24D274081" box="[479,590,418,442]" pageId="11" pageNumber="232" rank="species">spec. nova</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1BBE6D90FFE8146A6A0BD3B24DF54081" box="[605,668,418,442]" captionStart="Fig" captionStartId="12.[136,164,1914,1933]" captionTargetBox="[174,1416,236,1835]" captionTargetId="figure-37@12.[253,1357,298,1819]" captionTargetPageId="12" captionText="Fig. 2. Homalium fortunatii Appleq. A. Flowering branch; B. Flowers. [Randrianasolo et al. 1358, TAN] [Drawing: R.L. Andriamiarisoa]" figureDoi="http://doi.org/10.5281/zenodo.6238376" httpUri="https://zenodo.org/record/6238376/files/figure.png" pageId="11" pageNumber="232">Fig. 2</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CB9F229EFFE8146A68C1D3DC4E09435F" pageId="11" pageNumber="232" type="materials_examined">
<paragraph id="833A7115FFE8146A68C1D3DC4E09435F" blockId="11.[151,764,460,612]" pageId="11" pageNumber="232">
<materialsCitation id="33ED7B48FFE8146A68C1D3DC4D95437F" ID-GBIF-Occurrence="3470313553" collectingDate="2011-02-04" collectionCode="G, P, TAN" collectorName="Randrianasolo et al" country="Madagascar" county="Ambalabe-Sahanionaka" elevation="302" latitude="-19.304722" location="Vatomandry" longLatPrecision="21" longitude="48.622776" municipality="Toamasina" pageId="11" pageNumber="232" specimenCode="MO-6703445!" specimenCount="1" stateProvince="Holotypus" typeStatus="holotype">
<emphasis id="B1F1AD07FFE8146A68C1D3DC4DEF40DF" bold="true" box="[151,646,460,484]" pageId="11" pageNumber="232">
<typeStatus id="5C3ECFB7FFE8146A68C1D3DC4E7940DF" box="[151,272,460,484]" pageId="11" pageNumber="232" type="holotype">Holotypus</typeStatus>
:
<collectingCountry id="FB923185FFE8146A694AD3DC4ED140DF" box="[284,440,460,484]" name="Madagascar" pageId="11" pageNumber="232">MADAGASCAR</collectingCountry>
. Prov.
<collectingMunicipality id="635EEB6FFFE8146A6A55D3DC4DE940DF" box="[515,640,460,484]" pageId="11" pageNumber="232">Toamasina</collectingMunicipality>
:
</emphasis>
<location id="865A27CEFFE8146A6ADBD3DD4FD3433F" LSID="urn:lsid:plazi:treatment:0B2CC003FFE8146A6827D3B24A6042E5:865A27CEFFE8146A6ADBD3DD4FD3433F" country="Madagascar" county="Ambalabe-Sahanionaka" latitude="-19.304722" longLatPrecision="21" longitude="48.622776" municipality="Toamasina" name="Vatomandry" pageId="11" pageNumber="232" stateProvince="Holotypus">Vatomandry</location>
,
<location id="865A27CEFFE8146A6893D3FC4E5A433F" LSID="urn:lsid:plazi:treatment:0B2CC003FFE8146A6827D3B24A6042E5:865A27CEFFE8146A6893D3FC4E5A433F" box="[197,307,492,516]" country="Madagascar" county="Ambalabe-Sahanionaka" latitude="-19.304722" longLatPrecision="21" longitude="48.622776" municipality="Toamasina" name="Ambalabe" pageId="11" pageNumber="232" stateProvince="Holotypus">Ambalabe</location>
,
<location id="865A27CEFFE8146A6968D3FC4D28433F" LSID="urn:lsid:plazi:treatment:0B2CC003FFE8146A6827D3B24A6042E5:865A27CEFFE8146A6968D3FC4D28433F" box="[318,577,492,516]" country="Madagascar" county="Ambalabe-Sahanionaka" latitude="-19.304722" longLatPrecision="21" longitude="48.622776" municipality="Toamasina" name="along trail to Ambatobe" pageId="11" pageNumber="232" stateProvince="Holotypus">along trail to Ambatobe</location>
,
<location id="865A27CEFFE8146A6A1DD3FD4EAA431F" LSID="urn:lsid:plazi:treatment:0B2CC003FFE8146A6827D3B24A6042E5:865A27CEFFE8146A6A1DD3FD4EAA431F" country="Madagascar" county="Ambalabe-Sahanionaka" latitude="-19.304722" longLatPrecision="21" longitude="48.622776" municipality="Toamasina" name="a deviation from Ambalabe-Sahanionaka trail" pageId="11" pageNumber="232" stateProvince="Holotypus">a deviation from Ambalabe-Sahanionaka trail</location>
,
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<geoCoordinate id="E6B117D2FFE8146A6A6CD01C4DC2431F" box="[570,683,524,548]" degrees="48" direction="east" minutes="37" orientation="longitude" pageId="11" pageNumber="232" precision="15" seconds="22" value="48.622776">48°37'22&quot;E</geoCoordinate>
,
<quantity id="447DDCF0FFE8146A6AE3D01C4D9E431F" box="[693,759,524,548]" metricMagnitude="2" metricUnit="m" metricValue="3.02" pageId="11" pageNumber="232" unit="m" value="302.0">
<elevation id="08A89626FFE8146A6AE3D01C4D9E431F" box="[693,759,524,548]" metricMagnitude="2" metricUnit="m" metricValue="3.02" pageId="11" pageNumber="232" unit="m" value="302.0">302 m</elevation>
</quantity>
,
<date id="F73B57D5FFE8146A68C1D03C4F9C437F" box="[151,245,556,580]" pageId="11" pageNumber="232" value="2011-02-04">
<collectingDate id="E77FAE3DFFE8146A68C1D03C4F9C437F" box="[151,245,556,580]" pageId="11" pageNumber="232" value="2011-02-04">4.II.2011</collectingDate>
</date>
, fl. &amp; fr.,
<emphasis id="B1F1AD07FFE8146A6903D03C4D38437F" box="[341,593,556,580]" italics="true" pageId="11" pageNumber="232">
<collectorName id="2E7014C3FFE8146A6903D03C4D70437F" box="[341,537,556,580]" pageId="11" pageNumber="232">Randrianasolo et al</collectorName>
. 1358
</emphasis>
(
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;
</materialsCitation>
<materialsCitation id="33ED7B48FFE8146A68C1D05C4E304358" ID-GBIF-Occurrence="3470313563" box="[151,345,587,612]" collectingDate="2011-02-04" collectionCode="G, P, TAN" country="Madagascar" county="Ambalabe-Sahanionaka" elevation="302" latitude="-19.304722" location="Prov" longLatPrecision="21" longitude="48.622776" municipality="Ambatobe" pageId="11" pageNumber="232" specimenCount="1" stateProvince="Holotypus" typeStatus="isotype">
<typeStatus id="5C3ECFB7FFE8146A68C1D05C4FAB435F" box="[151,194,588,612]" pageId="11" pageNumber="232" type="isotype">isotype</typeStatus>
:
<collectionCode id="E594E9D0FFE8146A689BD05C4F8F435F" box="[205,230,588,612]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="11" pageNumber="232" type="Herbarium">G</collectionCode>
!,
<collectionCode id="E594E9D0FFE8146A68A3D05C4E60435F" box="[245,265,588,612]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="11" pageNumber="232" type="Herbarium">P</collectionCode>
!;
<collectionCode id="E594E9D0FFE8146A6941D05C4E39435F" box="[279,336,588,612]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="11" pageNumber="232" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CB9F229EFFE8146A68C1D0674E084274" pageId="11" pageNumber="232" type="discussion">
<paragraph id="833A7115FFE8146A68C1D0674E084274" blockId="11.[151,764,631,847]" pageId="11" pageNumber="232">
<emphasis id="B1F1AD07FFE8146A68C1D0674D2543B4" box="[151,588,631,655]" italics="true" pageId="11" pageNumber="232">
<taxonomicName id="44850A96FFE8146A68C1D0674EA543B4" authority="Appleq." authorityName="Appleq." box="[151,460,631,655]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="fortunatii">Homalium fortunatii Appleq.</taxonomicName>
differs from
</emphasis>
<taxonomicName id="44850A96FFE8146A6A05D0684E774394" authority="(Tul.) Baill." authorityName="Baill." baseAuthorityName="Tul." class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="thuarsianum">
<emphasis id="B1F1AD07FFE8146A6A05D0684D9243B4" box="[595,763,631,655]" italics="true" pageId="11" pageNumber="232">H. thuarsianum</emphasis>
(
<emphasis id="B1F1AD07FFE8146A68F7D0874E774394" box="[161,286,663,687]" italics="true" pageId="11" pageNumber="232">Tul.) Baill.</emphasis>
</taxonomicName>
<emphasis id="B1F1AD07FFE8146A6971D0884E0B43D4" italics="true" pageId="11" pageNumber="232">in having minutely pubescent young twigs; elliptical to narrowly elliptical (to broadly elliptical or obovate) leaves, (3.4)7.513</emphasis>
×
<emphasis id="B1F1AD07FFE8146A692AD0C64E084274" italics="true" pageId="11" pageNumber="232">
(1.4)
<quantity id="447DDCF0FFE8146A69EDD0C74D4343D4" box="[443,554,726,751]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="5.3" metricValueMin="3.5" pageId="11" pageNumber="232" unit="cm" value="4.4" valueMax="5.3" valueMin="3.5">3.55.3 cm</quantity>
, with base convex (to rounded) and apex cuspidate with rounded tip (to rounded); and sepals and petals slightly shorter than, to slightly longer than, the calyx tube.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CB9F229EFFE8146A68C1D1684EF54734" pageId="11" pageNumber="232" type="description">
<paragraph id="833A7115FFE8146A68C1D1684EF54734" blockId="11.[113,766,886,1551]" pageId="11" pageNumber="232">
<emphasis id="B1F1AD07FFE8146A68C1D1684FAD42B4" box="[151,196,888,911]" italics="true" pageId="11" pageNumber="232">Tree</emphasis>
to
<quantity id="447DDCF0FFE8146A68BED1674E2842B5" box="[232,321,886,911]" metricMagnitude="1" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="11" pageNumber="232" unit="m" value="13.5" valueMax="15.0" valueMin="12.0">1215 m</quantity>
; young twigs minutely pubescent becoming glabrous.
<emphasis id="B1F1AD07FFE8146A6952D1884E234294" box="[260,330,920,943]" italics="true" pageId="11" pageNumber="232">Leaves</emphasis>
elliptical to narrowly elliptical (broadly elliptical to obovate in abnormally small leaves), (3.4)7.513 × (1.4 )
<quantity id="447DDCF0FFE8146A689BD1C74E3B42D5" box="[205,338,982,1007]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="5.3" metricValueMin="3.5" pageId="11" pageNumber="232" unit="cm" value="4.4" valueMax="5.3" valueMin="3.5">3.5 5.3 cm</quantity>
; petiole
<quantity id="447DDCF0FFE8146A69EDD1C74D5E42D5" box="[443,567,982,1007]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="6.5" metricValueMin="2.0" pageId="11" pageNumber="232" unit="mm" value="4.25" valueMax="6.5" valueMin="2.0">2 6.5 mm</quantity>
; base convex (to rounded), often attenuate at extreme base; apex cuspidate with rounded tip (rounded); margins shallowly repand, with glands small, often slightly protruding; adaxial surface drying dark brown, sometimes gray-tinged, abaxial surface brown.
<emphasis id="B1F1AD07FFE8146A6827D6674F9245B4" box="[113,251,1143,1167]" italics="true" pageId="11" pageNumber="232">Inflorescences</emphasis>
paniculate, lateral, sometimes cauliflorous on small twigs,
<quantity id="447DDCF0FFE8146A68AAD6874E024595" box="[252,363,1174,1199]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" metricValueMax="11.5" metricValueMin="5.0" pageId="11" pageNumber="232" unit="cm" value="8.25" valueMax="11.5" valueMin="5.0">511.5 cm</quantity>
, pubescent; flowers pedicellate with pedicels
<quantity id="447DDCF0FFE8146A6884D6A74E2E45F5" box="[210,327,1206,1231]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="2.0" metricValueMin="0.7" pageId="11" pageNumber="232" unit="mm" value="1.35" valueMax="2.0" valueMin="0.7">0.7 2 mm</quantity>
; bracts lanceolate to ovate,
<quantity id="447DDCF0FFE8146A6A2FD6A74D9245F5" box="[633,763,1206,1231]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.5" metricValueMin="1.8" pageId="11" pageNumber="232" unit="mm" value="2.65" valueMax="3.5" valueMin="1.8">1.83.5 mm</quantity>
(&lt;
<quantity id="447DDCF0FFE8146A68D8D6C74FA345D5" box="[142,202,1239,1263]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="232" unit="mm" value="1.0">1 mm</quantity>
at apex).
<emphasis id="B1F1AD07FFE8146A6978D6C74E1645D4" box="[302,383,1239,1263]" italics="true" pageId="11" pageNumber="232">Flowers</emphasis>
79-merous; calyx tube funnelform with rounded base, erect-pubescent; sepals ligulate (to narrowly lanceolate) with acute (to rounded) apex,
<quantity id="447DDCF0FFE8146A6A20D7074D9E4415" box="[630,759,1302,1327]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="11" pageNumber="232" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.12.5 mm</quantity>
; petals ligulate with acute (to rounded-acute) apex,
<quantity id="447DDCF0FFE8146A6A2AD7274D914475" box="[636,760,1334,1359]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.7" metricValueMin="1.8" pageId="11" pageNumber="232" unit="mm" value="2.25" valueMax="2.7" valueMin="1.8">1.82.7 mm</quantity>
; sepals and petals post-flowering pinkish white or reddish, slightly shorter than calyx tube to slightly longer, ascending, ciliate with long soft hairs, abaxial surface sparsely pubescent, adaxial surface sparsely villous; sepal glands pubescent; filaments
<quantity id="447DDCF0FFE8146A68EDD7C74E2B44D5" box="[187,322,1494,1519]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="11" pageNumber="232" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
, basally villous; ovary villous; styles 45,
<quantity id="447DDCF0FFE8146A6827D7E74F9E4735" box="[113,247,1526,1551]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="11" pageNumber="232" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.60.8 mm</quantity>
, basally villous.
</paragraph>
</subSubSection>
<subSubSection id="CB9F229EFFE8146A68C1D4274D1B4794" pageId="11" pageNumber="232" type="etymology">
<paragraph id="833A7115FFE8146A68C1D4274D1B4794" blockId="11.[113,763,1591,1711]" pageId="11" pageNumber="232">
<emphasis id="B1F1AD07FFE8146A68C1D4274D6E4774" box="[151,519,1591,1615]" italics="true" pageId="11" pageNumber="232">
Etymology.
<taxonomicName id="44850A96FFE8146A697ED4274D6E4774" box="[296,519,1591,1615]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="fortunatii">Homalium fortunatii</taxonomicName>
</emphasis>
is named for botanist Fortunat Rakotoarivony; he was among the collectors of the
<typeStatus id="5C3ECFB7FFE8146A6827D4674FC947B4" box="[113,160,1655,1679]" pageId="11" pageNumber="232">type</typeStatus>
specimen and is a strong advocate of conservation and botanical research in the region of its collection.
</paragraph>
</subSubSection>
<subSubSection id="CB9F229EFFE8146A68C1D4C74FB94614" pageId="11" pageNumber="232" type="vernacular_names">
<paragraph id="833A7115FFE8146A68C1D4C74FB94614" blockId="11.[113,763,1751,1903]" pageId="11" pageNumber="232">
<emphasis id="B1F1AD07FFE8146A68C1D4C74E8A47D4" box="[151,483,1751,1775]" italics="true" pageId="11" pageNumber="232">Vernacular names and use.</emphasis>
“ Hazombatofotsy ”
<emphasis id="B1F1AD07FFE8146A6827D4E64EFC4634" box="[113,405,1782,1807]" italics="true" pageId="11" pageNumber="232">(Randrianasolo et al. 1358);</emphasis>
“Vazanaomby”
<emphasis id="B1F1AD07FFE8146A6A17D4E64FB94614" italics="true" pageId="11" pageNumber="232">(Randrianasolo et al. 1358).</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CB9F229EFFE8146A68C1D5274EE54654" pageId="11" pageNumber="232" type="discussion">
<paragraph id="833A7115FFE8146A68C1D5274EE54654" blockId="11.[113,763,1751,1903]" pageId="11" pageNumber="232">
Wood of
<emphasis id="B1F1AD07FFE8146A6943D5284EDC4674" box="[277,437,1847,1871]" italics="true" pageId="11" pageNumber="232">H. fortunatii</emphasis>
is used for construction
<emphasis id="B1F1AD07FFE8146A6827D5464EE54654" box="[113,396,1878,1903]" italics="true" pageId="11" pageNumber="232">(Randrianasolo et al. 1358).</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CB9F229EFFE8146A6B11D3074B5B4095" pageId="11" pageNumber="232" type="distribution">
<paragraph id="833A7115FFE8146A6B11D3074B5B4095" blockId="11.[801,1452,279,431]" pageId="11" pageNumber="232">
<emphasis id="B1F1AD07FFE8146A6B11D3074CEC4074" italics="true" pageId="11" pageNumber="232">
Distribution, ecology and conservation status.
<taxonomicName id="44850A96FFE8146A6D6AD3074CEC4074" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="fortunatii">Homalium fortunatii</taxonomicName>
</emphasis>
is known from two low-elevation humid forests in
<collectingRegion id="4141BFF7FFE8146A6B77D3474CF34054" box="[801,922,343,367]" country="Madagascar" name="Toamasina" pageId="11" pageNumber="232">Toamasina</collectingRegion>
. Because of its very small known Area of Occupancy (only
<quantity id="447DDCF0FFE8146A6BF1D3674CB740B4" box="[935,990,375,399]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="232" unit="km" value="8.0">8 km</quantity>
²) its conservation status may be estimated as “Vulnerable” [VU D2].
</paragraph>
</subSubSection>
<subSubSection id="CB9F229EFFE8146A6B11D3C84BCF43F4" pageId="11" pageNumber="232" type="discussion">
<paragraph id="833A7115FFE8146A6B11D3C84BCF43F4" blockId="11.[801,1452,471,879]" pageId="11" pageNumber="232">
<emphasis id="B1F1AD07FFE8146A6B11D3C84B1C40D4" box="[839,1141,471,495]" italics="true" pageId="11" pageNumber="232">
Notes.
<taxonomicName id="44850A96FFE8146A6BC9D3C74B1C40D4" box="[927,1141,471,495]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="fortunatii">Homalium fortunatii</taxonomicName>
</emphasis>
is among the
<taxonomicName id="44850A96FFE8146A6D5CD3C84CF04334" authority="(Tul.) Baill." authorityName="Baill." baseAuthorityName="Tul." class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="thuarsianum">
<emphasis id="B1F1AD07FFE8146A6D5CD3C84AC540D4" box="[1290,1452,471,495]" italics="true" pageId="11" pageNumber="232">H. thuarsianum</emphasis>
(Tul.) Baill.
</taxonomicName>
group of species with paniculate inflorescences and pubescent sepal glands.
<taxonomicName id="44850A96FFE8146A6C72D0074A744314" box="[1060,1309,535,559]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="thuarsianum">
<emphasis id="B1F1AD07FFE8146A6C72D0074A744314" box="[1060,1309,535,559]" italics="true" pageId="11" pageNumber="232">Homalium thuarsianum</emphasis>
</taxonomicName>
has glabrous young twigs, usually oblanceolate to obovate (to oblongoblanceolate or elliptical leaves) with cuneate to convex bases and variable (often rounded, but rarely rounded-cuspidate) apices, and sepals and petals that are consistently longer than the calyx tube when fully developed.
</paragraph>
</subSubSection>
<subSubSection id="CB9F229EFFE8146A6B11D0C74A6042E5" pageId="11" pageNumber="232" type="materials_examined">
<paragraph id="833A7115FFE8146A6B11D0C74CC34254" blockId="11.[801,1452,471,879]" pageId="11" pageNumber="232">
Material of
<taxonomicName id="44850A96FFE8146A6B8CD0C84B0743D4" box="[986,1134,727,751]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="fortunatii">
<emphasis id="B1F1AD07FFE8146A6B8CD0C84B0743D4" box="[986,1134,727,751]" italics="true" pageId="11" pageNumber="232">H. fortunatii</emphasis>
</taxonomicName>
has been misidentified as
<taxonomicName id="44850A96FFE8146A6B77D0E84CD84234" box="[801,945,759,783]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="erianthum">
<emphasis id="B1F1AD07FFE8146A6B77D0E84CD84234" box="[801,945,759,783]" italics="true" pageId="11" pageNumber="232">H. erianthum</emphasis>
</taxonomicName>
, which likewise has relatively large leaves and paniculate inflorescences.
<taxonomicName id="44850A96FFE8146A6C6DD1074A744214" box="[1083,1309,791,815]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="232" phylum="Tracheophyta" rank="species" species="erianthum">
<emphasis id="B1F1AD07FFE8146A6C6DD1074A744214" box="[1083,1309,791,815]" italics="true" pageId="11" pageNumber="232">Homalium erianthum</emphasis>
</taxonomicName>
has glabrous sepal glands and much longer stamens, and is confined to Anstiranana.
</paragraph>
<paragraph id="833A7115FFE8146A6B11D1864A6042E5" blockId="11.[801,1452,917,990]" pageId="11" pageNumber="232">
<materialsCitation id="33ED7B48FFE8146A6B11D1864A6D42E7" ID-GBIF-Occurrence="3470313341" collectingDate="1988-02-19" collectionCode="MO, P" collectorName="Schatz" country="Madagascar" elevation="75" latitude="-15.75" location="Masoala Peninsula" longLatPrecision="1284" longitude="50.033333" municipality="Masoala Peninsula" pageId="11" pageNumber="232" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
<emphasis id="B1F1AD07FFE8146A6B11D1864CC94293" box="[839,928,918,936]" italics="true" pageId="11" pageNumber="232">
<typeStatus id="5C3ECFB7FFE8146A6B11D1864CF54293" box="[839,924,918,936]" pageId="11" pageNumber="232" type="paratype">Paratypus</typeStatus>
.
</emphasis>
<emphasis id="B1F1AD07FFE8146A6BE0D1854BBD4293" bold="true" box="[950,1236,917,936]" pageId="11" pageNumber="232">
<collectingCountry id="FB923185FFE8146A6BE0D1854B5B4293" box="[950,1074,917,936]" name="Madagascar" pageId="11" pageNumber="232">MADAGASCAR</collectingCountry>
. Prov. Toamasina:
</emphasis>
<location id="865A27CEFFE8146A6C8CD1854A174293" LSID="urn:lsid:plazi:treatment:0B2CC003FFE8146A6827D3B24A6042E5:865A27CEFFE8146A6C8CD1854A174293" box="[1242,1406,917,936]" country="Madagascar" latitude="-15.75" longLatPrecision="1284" longitude="50.033333" municipality="Masoala Peninsula" name="Masoala Peninsula" pageId="11" pageNumber="232" stateProvince="Toamasina">Masoala Peninsula</location>
,
<location id="865A27CEFFE8146A6DDED1854B0142F8" LSID="urn:lsid:plazi:treatment:0B2CC003FFE8146A6827D3B24A6042E5:865A27CEFFE8146A6DDED1854B0142F8" country="Madagascar" latitude="-15.75" longLatPrecision="1284" longitude="50.033333" municipality="Masoala Peninsula" name="trail just above Antalavia River on S bank" pageId="11" pageNumber="232" stateProvince="Toamasina">trail just above Antalavia River on S bank</location>
,
<locationDeviation id="92886837FFE8146A6C27D1A04A7E42F8" box="[1137,1303,943,963]" pageId="11" pageNumber="232">
<quantity id="447DDCF0FFE8146A6C27D1A04BDA42F8" box="[1137,1203,943,963]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="11" pageNumber="232" unit="km" value="1.5" valueMax="2.0" valueMin="1.0">12 km</quantity>
from coast
</locationDeviation>
,
<geoCoordinate id="E6B117D2FFE8146A6D76D1A04A3442F8" box="[1312,1373,944,963]" degrees="15" direction="south" minutes="45" orientation="latitude" pageId="11" pageNumber="232" precision="925" value="-15.75">15°45'S</geoCoordinate>
<geoCoordinate id="E6B117D2FFE8146A6D34D1A04ACE42F8" box="[1378,1447,944,963]" degrees="50" direction="east" minutes="02" orientation="longitude" pageId="11" pageNumber="232" precision="925" value="50.033333">50°02'E</geoCoordinate>
,
<quantity id="447DDCF0FFE8146A6B77D1DA4C1142E5" box="[801,888,970,990]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="11" pageNumber="232" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
<elevation id="08A89626FFE8146A6B77D1DA4C1142E5" box="[801,888,970,990]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="11" pageNumber="232" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50100 m</elevation>
</quantity>
,
<date id="F73B57D5FFE8146A6BD7D1DB4CBF42E5" box="[897,982,971,990]" pageId="11" pageNumber="232" value="1988-02-19">
<collectingDate id="E77FAE3DFFE8146A6BD7D1DB4CBF42E5" box="[897,982,971,990]" pageId="11" pageNumber="232" value="1988-02-19">19.II.1988</collectingDate>
</date>
, fl. &amp; fr.,
<emphasis id="B1F1AD07FFE8146A6C70D1DA4BDD42E5" box="[1062,1204,970,990]" italics="true" pageId="11" pageNumber="232">
<collectorName id="2E7014C3FFE8146A6C70D1DA4B3542E6" box="[1062,1116,970,989]" pageId="11" pageNumber="232">Schatz</collectorName>
et al. 1928
</emphasis>
(
<collectionCode id="E594E9D0FFE8146A6C97D1DB4B8042E5" box="[1217,1257,971,990]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="11" pageNumber="232" type="Herbarium">MO</collectionCode>
,
<collectionCode id="E594E9D0FFE8146A6CA6D1DB4B9442E5" box="[1264,1277,971,990]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="11" pageNumber="232" type="Herbarium">P</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>