treatments-xml/data/0B/19/12/0B1912EFBBAE50A18A5E28D041E4FEA9.xml
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.227.101907" ID-Pensoft-Pub="1314-2003-227-43" ID-Pensoft-UUID="AD08E6F7EEA15D67BB611F0B8B730623" ModsDocID="1314-2003-227-43" checkinTime="1685632887257" checkinUser="pensoft" docAuthor="Arista, Jessy Patricia, Hagsater, Eric, Santiago, Elizabeth, Edquen, Jose D., Pariente, Eli, Oliva, Manuel &amp; Salazar, Gerardo A." docDate="2023" docId="0B1912EFBBAE50A18A5E28D041E4FEA9" docLanguage="en" docName="PhytoKeys 227: 43-87" docOrigin="PhytoKeys 227" docPubDate="2023-06-01" docSource="http://dx.doi.org/10.3897/phytokeys.227.101907" docTitle="Epidendrum parvireflexilobum Hagsater, J. P. Arista &amp; Edquen 2023, sp. nov." docType="treatment" docVersion="1" id="AD08E6F7EEA15D67BB611F0B8B730623" lastPageNumber="43" masterDocId="AD08E6F7EEA15D67BB611F0B8B730623" masterDocTitle="New and noteworthy species of the genus Epidendrum (Orchidaceae, Laeliinae) from the Area de Conservacion Privada La Pampa del Burro, Amazonas, Peru" masterLastPageNumber="87" masterPageNumber="43" pageNumber="43" updateTime="1685632887257" updateUser="pensoft">
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<mods:title id="D52BFAED34C4CAEFA5C09D296F899920">New and noteworthy species of the genus Epidendrum (Orchidaceae, Laeliinae) from the Area de Conservacion Privada La Pampa del Burro, Amazonas, Peru</mods:title>
</mods:titleInfo>
<mods:name id="5B7A16EB503486F8C6E1EFDEF5FB206B" type="personal">
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<mods:namePart id="97E1E44FF24C6DA6B7C0A6B3905DDEA5">Arista, Jessy Patricia</mods:namePart>
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<mods:affiliation id="444FB212C98AD623F86191ED89910D74">Instituto de Investigacion, Innovacion y Desarrollo del Sector Agrario y Agroindustrial (IIDAA), Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
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<mods:namePart id="8680F0F08AE8122F3DD79D08347B7F9E">Hagsater, Eric</mods:namePart>
<mods:nameIdentifier id="0E58B28A163F606DC55C995C581C3C8A" type="ORCID">https://orcid.org/0000-0002-2371-9427</mods:nameIdentifier>
<mods:affiliation id="1550EFCF5EB8F6B21C648A0E403FF566">Escuela de Posgrado, Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
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<mods:namePart id="8F493D501BEF239C6F7AAEE785C9DB90">Santiago, Elizabeth</mods:namePart>
<mods:nameIdentifier id="CC71DDC1A7DF5BE7E062A9742E437E02" type="ORCID">https://orcid.org/0000-0002-1368-9025</mods:nameIdentifier>
<mods:affiliation id="4F49E051DABF877FF95A588B9AF743A4">Escuela de Posgrado, Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
</mods:name>
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<mods:namePart id="C54DA69A83A7739DEBA6F7E15B245DA6">Edquen, Jose D.</mods:namePart>
<mods:nameIdentifier id="161ED84A97DE3A9575F4C4F0F98AD99A" type="ORCID">https://orcid.org/0000-0002-7289-3974</mods:nameIdentifier>
<mods:affiliation id="5DF27206A291AD62F45385BABDEEB564">Instituto de Investigacion, Innovacion y Desarrollo del Sector Agrario y Agroindustrial (IIDAA), Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
</mods:name>
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<mods:namePart id="9E334EB2CC00F197946923E937939735">Pariente, Eli</mods:namePart>
<mods:nameIdentifier id="E87EDD98B9C4F7CDCC9E9F264739FD89" type="ORCID">https://orcid.org/0000-0002-9197-0218</mods:nameIdentifier>
<mods:affiliation id="FCF18B5D0DE9D53DCE7BE440EC9EADA8">Instituto de Investigacion, Innovacion y Desarrollo del Sector Agrario y Agroindustrial (IIDAA), Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="17F887104A1D5F90D76CF2E5736DE75B">Oliva, Manuel</mods:namePart>
<mods:nameIdentifier id="164D4788A8A6C0A637E21F62E136AD31" type="ORCID">https://orcid.org/0000-0002-9670-0970</mods:nameIdentifier>
<mods:affiliation id="155418A43AFB40D7E516DC6C0CC2C793">Instituto de Investigacion, Innovacion y Desarrollo del Sector Agrario y Agroindustrial (IIDAA), Universidad Nacional Toribio Rodriguez de Mendoza de Amazonas, Chachapoyas, Amazonas, Peru</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="FAE975B7572FC7E521F474A8C015ABA6">Salazar, Gerardo A.</mods:namePart>
<mods:nameIdentifier id="A75AF60184B3BBBD438C23E4334B8F32" type="ORCID">https://orcid.org/0000-0002-5203-5374</mods:nameIdentifier>
<mods:affiliation id="2C4BA51478610D78620450FF50F43AB2">Herbario AMO, Montanas Calizas 490, Lomas de Chapultepec. Miguel Hidalgo, Mexico City, 11000, Mexico</mods:affiliation>
<mods:nameIdentifier id="97B1D2B6F3669387F18877D608879F7D" type="email">gasc@ib.unam.mx</mods:nameIdentifier>
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<mods:typeOfResource id="E5DD1EE75597976EF9F1FDBBE0B3EA7C">text</mods:typeOfResource>
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<mods:titleInfo id="56CBA75759AA9EBA8F5260432D4994B9">
<mods:title id="25B2A438378CB143740AE8BF50871C10">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="420871F8BC735F4F2BA7404BF1D96C58">
<mods:date id="6DBF3644F21C0B311BD422895C31BB2D">2023</mods:date>
<mods:detail id="CC1A51B700C212926643BE174E874C76" type="pubDate">
<mods:number id="894E36C88111A09750818A601A92CFCA">2023-06-01</mods:number>
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<mods:detail id="9A3E018F15C5BA3B20CB29CD1708F9FF" type="volume">
<mods:number id="177EFE9B16801E4BCAAC159CDFB1E850">227</mods:number>
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<mods:extent id="3FE13EFFE0DF9C93A34D84BF1CDC48F1" unit="page">
<mods:start id="A047104E57DD9A309AC8FB7B72F53B04">43</mods:start>
<mods:end id="3768AF4234FD931C24E335C0CA4B563D">87</mods:end>
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<mods:location id="ADCFD42511E88B6410F4AA58565850A5">
<mods:url id="E34FC8537C578C970DF3B97DA3064758">http://dx.doi.org/10.3897/phytokeys.227.101907</mods:url>
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<mods:classification id="24928C5CF98A7E637CDEB56575C7C524">journal article</mods:classification>
<mods:identifier id="549D36A999A5E37480FF57D77F874064" type="DOI">http://dx.doi.org/10.3897/phytokeys.227.101907</mods:identifier>
<mods:identifier id="37601AA69EA34601F2BEE3E5B3B9EF84" type="Pensoft-Pub">1314-2003-227-43</mods:identifier>
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<treatment id="0B1912EFBBAE50A18A5E28D041E4FEA9" LSID="urn:lsid:plazi:treatment:0B1912EFBBAE50A18A5E28D041E4FEA9" httpUri="http://treatment.plazi.org/id/0B1912EFBBAE50A18A5E28D041E4FEA9" lastPageNumber="43" pageId="0" pageNumber="43">
<subSubSection id="C793A5FFF45ED5A54CC9503602EEF55D" pageId="0" pageNumber="43" type="nomenclature">
<paragraph id="82A108BE4ECDE21B71617DF66FB296C5" pageId="0" pageNumber="43">
<taxonomicName id="39A848ADE98B23469C55E9CE7D563F98" LSID="0B1912EF-BBAE-50A1-8A5E-28D041E4FEA9" authority="Hágsater, J. P. Arista &amp; Edquén" authorityName="Hagsater, J. P. Arista &amp; Edquen" authorityYear="2023" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum parvireflexilobum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="parvireflexilobum" status="sp. nov.">
Epidendrum parvireflexilobum
<normalizedToken id="0E71604BD0425830CF0F2E6C520ADDD9" originalValue="Hágsater">Hagsater</normalizedToken>
, J.P.Arista &amp;
<normalizedToken id="87E73898477FF38DD02A591293EF32D9" originalValue="Edquén">Edquen</normalizedToken>
</taxonomicName>
<taxonomicNameLabel id="29980EBF287BF3F85FCFB5EEFD7E97DA" pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="CD2B94830F11D60DC8E1B4869A11B064" pageId="0" pageNumber="43" type="description">
<paragraph id="B01A54FEED28A50F63476E018D785CE9" pageId="0" pageNumber="43">
<figureCitation id="0A1601673D0EB77AEB736EBBFCB2FE82" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Epidendrum parvireflexilobum from Arista et al. 49 A habit B flower C dissected perianth D inflorescence E longitudinal section of flower F column and ovary apex from below (left) and side (right) G anther from above (up) and below (down), and pollinarium (Photographs by J. D. Edquen; plate prepared by A. Cisneros)." figureDoi="10.3897/phytokeys.227.101907.figure12" httpUri="https://binary.pensoft.net/fig/857088" pageId="0" pageNumber="43">Fig. 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="341665D9ED424706E7B53A633A4ABA70" pageId="0" pageNumber="43" type="type material">
<paragraph id="6AB740576DD7CADB8A37E0E3970A5D3F" pageId="0" pageNumber="43">Type material.</paragraph>
<paragraph id="380985590289AE768FF2E46600EBCFAA" pageId="0" pageNumber="43">
<emphasis id="D19CA4A1FD6D8704B2B10B3E54CD93F5" bold="true" pageId="0" pageNumber="43">Peru. Amazonas</emphasis>
: Prov.
<normalizedToken id="A17469E1964E891FB4AA70A71A6C31A2" originalValue="Bongará">Bongara</normalizedToken>
: Yambrasbamba: Perla del Imaza, La Perla Vieja, Pampa del Burro, 1871 m, 22 Aug. 2021,
<emphasis id="DA2BAE68BF041CA8918FB2B1291688FE" italics="true" pageId="0" pageNumber="43">J. P. Arista</emphasis>
,
<emphasis id="1EB9C23CE49680A5A94AA58C9305B217" italics="true" pageId="0" pageNumber="43">
J. D.
<normalizedToken id="988E84E59D25E30FFFB37AABB7D93A04" originalValue="Edquén">Edquen</normalizedToken>
</emphasis>
,
<emphasis id="AE01A0960A299966788096321E03A444" italics="true" pageId="0" pageNumber="43">
E.
<normalizedToken id="9924E53F32B2744FC57366E1AEAC508B" originalValue="Yrigoín">Yrigoin</normalizedToken>
</emphasis>
,
<emphasis id="D5748F762B81D6DA9C29E703F6AAD6EB" italics="true" pageId="0" pageNumber="43">L. Iliquin 49</emphasis>
(holotype: KUELAP! [LCDP voucher]).
</paragraph>
</subSubSection>
<subSubSection id="0335C188DDFAF2A928E21F521D4D1B2E" pageId="0" pageNumber="43" type="diagnosis">
<paragraph id="48C716A15332B928C5D9464D4E3BEA5E" pageId="0" pageNumber="43">Diagnosis.</paragraph>
<paragraph id="3E565BA052D7D921166760853010114D" pageId="0" pageNumber="43">
Similar to
<taxonomicName id="299E0E568CFC67897008E9DEFBFB8623" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum reflexilobum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="reflexilobum">
<emphasis id="52E226F32003A1CACFBA77FA42B57B73" italics="true" pageId="0" pageNumber="43">Epidendrum reflexilobum</emphasis>
</taxonomicName>
C.Schweinf., but overall smaller, with plants 22-46 cm tall including the inflorescence (vs. 40-120 cm tall), sepals 9.5-10.8 mm long, elliptic (vs. sepals 12.5-13.5 mm long, obovate-elliptic), the lateral lobes of the lip semi-ovate, erect (vs. lateral lobes spreading, twisted 90° in natural position, narrowly obovate), the callus 3-ribbed, forming a cuneate platform only reaching middle of mid-lobe (vs. callus 3-ribbed, ribs not forming a platform, with the mid-rib longer and reaching apical sinus).
</paragraph>
</subSubSection>
<subSubSection id="DED0FFE33A5558A968AE6E2D045E7B91" pageId="0" pageNumber="43" type="description">
<paragraph id="6ABBCAD8A1A5393B87D0855A7A5666CD" pageId="0" pageNumber="43">Description.</paragraph>
<paragraph id="C32A4FFBE409FE499B4251C04C38EFED" pageId="0" pageNumber="43">
Epiphytic, sympodial, caespitose, erect
<emphasis id="2CA1CBEB442661F7E52D050082CF5D41" bold="true" italics="true" pageId="0" pageNumber="43">herb</emphasis>
, 22-46 cm tall including inflorescence.
<emphasis id="F07EE0C5D2574805024F1D9C09D8F863" bold="true" italics="true" pageId="0" pageNumber="43">Roots</emphasis>
2 mm in diameter, basal, terete, fleshy, white.
<emphasis id="972AF10CD54B0F94812B17F44DC30F76" bold="true" italics="true" pageId="0" pageNumber="43">Stems</emphasis>
8-22
<normalizedToken id="24FB4C02CE2FDBF7DC27A4049F6B7A90" originalValue="×">x</normalizedToken>
0.3 cm, simple, cane-like, terete to slightly compressed toward apex, thin, basal half covered by non-foliar sheaths.
<emphasis id="1C2D07502D3D0E59C6B543BC7D69EFD9" bold="true" italics="true" pageId="0" pageNumber="43">Leaves</emphasis>
7-12, distichous, distributed along upper 1/4 of stem; sheaths 4.4-6.0
<normalizedToken id="4C0CDF7114FFECB430541E84FA043F15" originalValue="×">x</normalizedToken>
0.4 cm, tubular, smooth, vinaceous, papyraceous when dry; blade 4.0-7.8
<normalizedToken id="AC5BEA5DD450BEF8E7C5E25605656299" originalValue="×">x</normalizedToken>
0.8-2.8 cm, oblong, apex unequally bilobed, articulate, coriaceous, smooth, medium green on both sides, margins entire.
<emphasis id="AA9E694F86896AB49069ADD78C7157A3" bold="true" italics="true" pageId="0" pageNumber="43">Spathe</emphasis>
lacking.
<emphasis id="F20F89F038490DAA461C177AE77C9C5F" bold="true" italics="true" pageId="0" pageNumber="43">Inflorescence</emphasis>
12-33 cm long, racemose to pluri-racemose, laxly flowered, cylindric; peduncle 10-28 cm long, elongate, covered by numerous tubular, imbricate bracts 4.5
<normalizedToken id="CBF6BCF285D6CDB41F26D45F4A80C168" originalValue="×">x</normalizedToken>
0.4 cm, acute, scarious when dry, striated, papyraceous; rachis ca. 4.8 cm long.
<emphasis id="C477A9EA3D8C44B0A1145355C9322AD2" bold="true" italics="true" pageId="0" pageNumber="43">Floral bracts</emphasis>
1-3
<normalizedToken id="440BBFFB9CD37ECBF6475B08F217611D" originalValue="×">x</normalizedToken>
0.7-3.0 mm, much shorter than ovary, decreasing in size toward apex, triangular, acuminate to acute, embracing.
<emphasis id="46213C2C3A3EB04C815B6DA73135CCD9" bold="true" italics="true" pageId="0" pageNumber="43">Flowers</emphasis>
ca. 6-12, per raceme, successive, 3-6 open at a time, non-resupinate, pale to bright red, callus yellow; fragrance none.
<emphasis id="389F62E9BCAB5C8B0A604442BDFB625D" bold="true" italics="true" pageId="0" pageNumber="43">Sepals</emphasis>
spreading, apex obliquely rounded, short apiculate, 7-veined, margin entire, spreading; dorsal sepal 9.5-10.2
<normalizedToken id="604B22E77EEAC1844F5A58F86EF16F52" originalValue="×">x</normalizedToken>
4.5-4.9 mm, elliptic; lateral sepals 9.9-10.8
<normalizedToken id="4A128C253A949EA11DC89017A062E8AE" originalValue="×">x</normalizedToken>
5.0-5.5 mm, elliptic, oblique, sub-obtuse.
<emphasis id="29C41A652CAA204B5FD77392EA85FA57" bold="true" italics="true" pageId="0" pageNumber="43">Petals</emphasis>
10.6-13.5
<normalizedToken id="92F0DA87C31FAEEC615F5E46733E10E2" originalValue="×">x</normalizedToken>
4.0-5.0 mm, extended, oblanceolate or cuneate-spathulate, apex acute, 3-5-veined, margin entire, spreading.
<emphasis id="8F9FE673196DA0519BDABD83710378EA" bold="true" italics="true" pageId="0" pageNumber="43">Lip</emphasis>
5.7-7.8
<normalizedToken id="F6A755D3B8906E65F2F7C870E9A0B930" originalValue="×">x</normalizedToken>
8.2-9.0 mm, united to column, deeply 3-lobed, in natural position mid-lobe flat, extended, lateral lobes sub-erect, base cordate, distal margins irregularly laciniate; callus low, rectangular, truncate to rounded, with mid-rib formed by 3 straight, parallel ribs on mid-lobe, mid-rib longer, with two divergent bifid calli at base of lateral lobes; lateral lobes 3.0-3.5
<normalizedToken id="78ADA3B887D64B4F17D7F3E85C229095" originalValue="×">x</normalizedToken>
4.2-5.9 mm, semi-obovate, erect in natural position; mid-lobe 4.6
<normalizedToken id="95BC75BDFE2221D6697E8139B96E5DB3" originalValue="×">x</normalizedToken>
8.2 mm, obcuneate, bifid, slightly divergent, deeply and narrowly emarginate.
<emphasis id="1A7617A0D6EE9345D7584F71D3533792" bold="true" italics="true" pageId="0" pageNumber="43">Column</emphasis>
4.0 mm long, straight, wider at apex, with a pair of long, apical recurved fleshy wings with distal margin erose.
<emphasis id="C5D1FD1B9C0B236B5DB98BF84A684C64" bold="true" italics="true" pageId="0" pageNumber="43">Clinandrium hood</emphasis>
very short, margin entire, leaving anther totally exposed.
<emphasis id="717C984E923B8B4564D73DCA9D996566" bold="true" italics="true" pageId="0" pageNumber="43">Anther</emphasis>
ovoid, apiculate, surface rugose, 4-celled.
<emphasis id="0B94A72687F049FBE22E5868A2CA64F5" bold="true" italics="true" pageId="0" pageNumber="43">Pollinia</emphasis>
4, narrowly obovoid, laterally compressed, caudicles formed by a pile of elongate pollen tetrads like a pile of tiles.
<emphasis id="94991A1836988C9E5655ABDE68872C48" bold="true" italics="true" pageId="0" pageNumber="43">Rostellum</emphasis>
apical, split; viscarium semi-liquid.
<emphasis id="867D4AD34D3FA939A7BA30684AF240FE" bold="true" italics="true" pageId="0" pageNumber="43">Lateral lobes of stigma</emphasis>
short, occupying
<normalizedToken id="1E182BE640588E10FF84B6CB83441012" originalValue="¼">1/4</normalizedToken>
length of stigmatic cavity.
<emphasis id="67BF88A8B0E1C8C165A6CF1374A73AC5" bold="true" italics="true" pageId="0" pageNumber="43">Cuniculus</emphasis>
penetrating half pedicellate ovary, minutely papillose.
<emphasis id="9387CFA9A3501CC90C7ECC498C732615" bold="true" italics="true" pageId="0" pageNumber="43">Ovary</emphasis>
20 mm long, terete, thin, not inflated, green to red, furrowed.
<emphasis id="7FC2BF76C1CC3E1B244B395D90FD48FA" bold="true" italics="true" pageId="0" pageNumber="43">Capsule</emphasis>
not seen.
</paragraph>
<caption id="4424B8D929636DB630292FAFB7B9B1F7" doi="10.3897/phytokeys.227.101907.figure12" httpUri="https://binary.pensoft.net/fig/857088" pageId="0" pageNumber="43" start="Figure 12" startId="F12">
<paragraph id="0A4926C10C359117528A11FE234BF6FB" pageId="0" pageNumber="43">
<emphasis id="9606B88B65E8CCEEB83CEAA9C12B610C" bold="true" pageId="0" pageNumber="43">Figure 12.</emphasis>
<taxonomicName id="0124C00C413C02B035375508B0858E9A" authorityName="Hagsater, J. P. Arista &amp; Edquen" authorityYear="2023" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum parvireflexilobum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="parvireflexilobum">
<emphasis id="A93419EF88860B4D9726607CEE73A023" italics="true" pageId="0" pageNumber="43">Epidendrum parvireflexilobum</emphasis>
</taxonomicName>
from
<emphasis id="49CF1BC024BAB36E2D89EB3BD3BCA73B" italics="true" pageId="0" pageNumber="43">Arista et al. 49</emphasis>
<emphasis id="92776CA0086F329D8F70D36025DE1FFA" bold="true" pageId="0" pageNumber="43">A</emphasis>
habit
<emphasis id="D7737B24580BFA27E7F92D99F9740198" bold="true" pageId="0" pageNumber="43">B</emphasis>
flower
<emphasis id="D6F446639443514682421B54E669076E" bold="true" pageId="0" pageNumber="43">C</emphasis>
dissected perianth
<emphasis id="7A6E0029DA7B1130C4DBE051898422A3" bold="true" pageId="0" pageNumber="43">D</emphasis>
inflorescence
<emphasis id="22D12E02F3198FBD930AEF61AA631B41" bold="true" pageId="0" pageNumber="43">E</emphasis>
longitudinal section of flower
<emphasis id="2ECC17C9A77070CEC2C96808459E3B36" bold="true" pageId="0" pageNumber="43">F</emphasis>
column and ovary apex from below (left) and side (right)
<emphasis id="CA87834228F7921FFC133323DEFD9685" bold="true" pageId="0" pageNumber="43">G</emphasis>
anther from above (up) and below (down), and pollinarium (Photographs by J. D.
<normalizedToken id="7182972E75E11F78F1C10D174A5B0B58" originalValue="Edquén">Edquen</normalizedToken>
; plate prepared by A. Cisneros).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D49D638C09E25E499657C80459DAF10B" pageId="0" pageNumber="43" type="materials_examined">
<paragraph id="A82572708546D51D1112A6B0781317A7" pageId="0" pageNumber="43">Additional specimens examined.</paragraph>
<paragraph id="E81DA0E49B6580B79500BF1D0A34BFAF" pageId="0" pageNumber="43">
<materialsCitation id="6EBB7517EC5E1CE07D3B35BCC8444CF8" collectingDate="2021-08-22" collectorName="Prov. Bongara, Yambrasbamba, La Perla Vieja" country="Peru" elevation="1871" location="Pampa del Burro" municipality="Perla del Imaza" specimenCount="1" stateProvince="Amazonas">
<emphasis id="FC97EB36D4CB5FCC835150A7CDB1A4FC" bold="true" pageId="0" pageNumber="43">
<collectingCountry id="27A1BDD412A93A85E3677FB8477CF334" name="Peru">Peru</collectingCountry>
.
<collectingRegion id="27A9A1C6AB5AE253D402008F8613713B" country="Peru" name="Amazonas">Amazonas</collectingRegion>
</emphasis>
:
<collectorName id="FF75FF5400F0FD022DDF0904599B71B8">
Prov.
<normalizedToken id="42029E055C65024A3901794374BCEC81" originalValue="Bongará">Bongara</normalizedToken>
</collectorName>
:
<collectorName id="735448537CE3E1E6E34857BD86531A3D">Yambrasbamba</collectorName>
:
<collectingMunicipality id="2F31B2D12F94B4160EF4494DFC77CD03">Perla del Imaza</collectingMunicipality>
,
<collectorName id="6CF27AA1B89F42F2A297E1E99B5150D3">La Perla Vieja</collectorName>
,
<location id="DE48A565E6F9A7B8D7D8413C450186C5" LSID="urn:lsid:plazi:treatment:0B1912EFBBAE50A18A5E28D041E4FEA9:DE48A565E6F9A7B8D7D8413C450186C5" country="Peru" municipality="Perla del Imaza" name="Pampa del Burro" stateProvince="Amazonas">Pampa del Burro</location>
,
<quantity id="C6F727A5D790E08BF4B3700830BA4A96" metricMagnitude="3" metricUnit="m" metricValue="1.871" unit="m" value="1871.0">
<elevation id="A0A90916D2B6F15BC0306A6BD3D2331F" metricMagnitude="3" metricUnit="m" metricValue="1.871" unit="m" value="1871.0">1871 m</elevation>
</quantity>
,
<collectingDate id="8EBDBC287D362A74238FD14C65DECF5B" value="2021-08-22">22 Aug. 2021</collectingDate>
,
<emphasis id="CE5A64C25552872678D64821F062ABFA" italics="true" pageId="0" pageNumber="43">Arista et al. 25</emphasis>
(KUELAP!)
</materialsCitation>
;
<materialsCitation id="EA41873582BB351A3132D37C446ED827" collectingDate="1990-10-01" country="Peru" location="Piura" specimenCount="1" stateProvince="Piura">
<emphasis id="71F53E294645DD575324C62B32864274" bold="true" pageId="0" pageNumber="43">
<collectingRegion id="2466C60A63AF79B920F51D547DED3546" country="Peru" name="Piura">Piura</collectingRegion>
</emphasis>
: Huancabamba,
<collectingDate id="B24531C458A2EB5EF22A641A5E2787BF" value="1990-10-01">1 Oct. 1990</collectingDate>
,
<emphasis id="DF66205D851CDD09B5FF42B5AB1A172A" italics="true" pageId="0" pageNumber="43">Castillo 5</emphasis>
(AMO!)
</materialsCitation>
;
<materialsCitation id="5607EFB2F0219E468A3FFAA56272ACEC" collectingDate="2015-12-06" collectorName="Rioja, Pardo Miguel Naranjos, Mirador Venceremos" country="Peru" elevation="1867" location="San Martin" specimenCount="1" stateProvince="San Martin">
<emphasis id="87D2760BF2B0BB7C47A9B5DF44358550" bold="true" pageId="0" pageNumber="43">
<collectingRegion id="86C39673CAF26647A8C970AAFC71B1F7" country="Peru" name="San Martin">
San
<normalizedToken id="B701C47E6820244BACCFC65F2C7C4E53" originalValue="Martín">Martin</normalizedToken>
</collectingRegion>
</emphasis>
:
<collectorName id="F1FF038C21A6F356DB36BA3A7DED6BAA">Rioja</collectorName>
:
<collectorName id="4DAC210D39475F55ECDAD857221CE17A">Pardo Miguel Naranjos</collectorName>
,
<collectorName id="84B9C7E5249BB4E1F8D0F09119E71409">Mirador Venceremos</collectorName>
,
<quantity id="FAF4DC8E53616474237197667D6A8A55" metricMagnitude="3" metricUnit="m" metricValue="1.8669999999999998" unit="m" value="1867.0">
<elevation id="714FA8EC349D736EE78103F74577A02B" metricMagnitude="3" metricUnit="m" metricValue="1.8669999999999998" unit="m" value="1867.0">1867 m</elevation>
</quantity>
,
<collectingDate id="6F18A86F0531CE1638035A6073D358E8" value="2015-12-06">6 Dec. 2015</collectingDate>
,
<emphasis id="97032429B4BEAA0C5AE5499F1FB6BE87" italics="true" pageId="0" pageNumber="43">
<normalizedToken id="C66780CD0AE833DB5D3C689F6EF6BF73" originalValue="Edquén">Edquen</normalizedToken>
1979
</emphasis>
(HURP!; color plate published in
<bibRefCitation id="F28DC1BA3A2A2AE505B95ACD368AD2FA" author="Hagsater, E" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="43" publicationUrl="https://herbarioamo.org/index_archivos/Fascicle18(1).pdf" refId="B34" refString="Hagsater, E, Santiago, E [Eds], 2020b. The Genus Epidendrum, Part 14, Icones Orchidacearum 18(1): t. 1801-1848. https://herbarioamo.org/index_archivos/Fascicle18(1).pdf" title="The Genus Epidendrum, Part 14, Icones Orchidacearum 18 (1): t. 1801 - 1848." url="https://herbarioamo.org/index_archivos/Fascicle18(1).pdf" year="2020 b">
<normalizedToken id="DDA5D44B3A16196CCFA0794556E231FB" originalValue="Hágsater">Hagsater</normalizedToken>
and Santiago 2020b
</bibRefCitation>
: t. 1835, as
<taxonomicName id="B6AA83DF667E1E4D1F32BC409A9D37E5" lsidName="E. reflexilobum" pageId="0" pageNumber="43" rank="species" species="reflexilobum">
<emphasis id="94E09607D8579A2CB440FB8EAF3B34A1" italics="true" pageId="0" pageNumber="43">E. reflexilobum</emphasis>
</taxonomicName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CB71DA229C257C6428EC2C723B6DD414" pageId="0" pageNumber="43" type="other records">
<paragraph id="3E00C6ECA3F541287A3B5119248C967C" pageId="0" pageNumber="43">Other records.</paragraph>
<paragraph id="4626FBBDAD97DDC8C1C0F17BE54FFA1E" pageId="0" pageNumber="43">
<emphasis id="4D4710CDF2DF0C77518AA88DD95F0F7A" bold="true" pageId="0" pageNumber="43">Peru. Amazonas</emphasis>
: Chachapoyas: Balsas-Leimebamba, Cordillera Calla Calla, ca. 2400 m, 4 Feb. 2022,
<emphasis id="481E8CEC64A79B6CBC677062A6527DC8" italics="true" pageId="0" pageNumber="43">Deza s.n.</emphasis>
, digital images (AMO!);
<emphasis id="B8D7DA515EC1D99CA81FEF079F4FC4B3" italics="true" pageId="0" pageNumber="43">ibid. loc.</emphasis>
2363 m,
<emphasis id="3CC409868DF8C23A12BCA2A4B21063BE" italics="true" pageId="0" pageNumber="43">Harding s.n.</emphasis>
, digital images (AMO!);
<emphasis id="DA9C598033469E0F5CF10A7C3A94B60C" bold="true" pageId="0" pageNumber="43">
San
<normalizedToken id="80E6D82751D4B4BFB5B5F528E5FBA95B" originalValue="Martín">Martin</normalizedToken>
</emphasis>
: Rioja: Nuevo Cajamarca, Sector Yuracyacu,
<normalizedToken id="5AD8CC6102A7B8738E98EAF4C4F7D0EF" originalValue="Río">Rio</normalizedToken>
Yuracyacu, 1905 m, 15 Jul. 2018,
<emphasis id="327C932755D4DB0EDBE935BEA538AE80" italics="true" pageId="0" pageNumber="43">
<normalizedToken id="53B9BED20C1877C94181019BAB1F729F" originalValue="Edquén">Edquen</normalizedToken>
1815
</emphasis>
, digital images (AMO!).
</paragraph>
</subSubSection>
<subSubSection id="E50805DD6378CBB5ED1494575904657A" pageId="0" pageNumber="43" type="distribution">
<paragraph id="037C59CC35C1D91F86CB811AFE5896A3" pageId="0" pageNumber="43">Distribution.</paragraph>
<paragraph id="1E857BFF6FC373D408D0C005D867C27A" pageId="0" pageNumber="43">
Known presently from northern Peru in the Regions of Amazonas and San
<normalizedToken id="AA9F0F5225D55BB72A4BA29F60BB7A54" originalValue="Martín">Martin</normalizedToken>
. Growing at 1850-1905 m elevation.
</paragraph>
</subSubSection>
<subSubSection id="96209FA2AF38B8BFA00916739201A4B4" pageId="0" pageNumber="43" type="habitat">
<paragraph id="20F1AED719A150D9E902367290B61590" pageId="0" pageNumber="43">Habitat and ecology.</paragraph>
<paragraph id="1DC18D0964818B6C9CFE71AC9B5539A6" pageId="0" pageNumber="43">
Epiphytic and terrestrial in humid montane forest, secondary forest and abandoned coffee plantations, on
<taxonomicName id="851649C860B6EE3764568D1E49FD52D1" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" lsidName="Coffea arabica" order="Gentianales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="arabica">
<emphasis id="26163BAC95E413FAFDC30B79633918E7" italics="true" pageId="0" pageNumber="43">Coffea arabica</emphasis>
</taxonomicName>
L. and
<taxonomicName id="ED56FEF08608B7B22A3015C48F71C85B" class="Magnoliopsida" family="Rutaceae" genus="Citrus" isHybrid="true" kingdom="Plantae" lsidName="Citrus aurantium" order="Sapindales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="aurantium">
Citrus
<normalizedToken id="B831ED379EC47DA340101F139A14503D" originalValue="×">x</normalizedToken>
aurantium
</taxonomicName>
L.
</paragraph>
</subSubSection>
<subSubSection id="2F7160846BE0C68B239C0A57341AA1BD" pageId="0" pageNumber="43" type="phenology">
<paragraph id="A315ADF4B8D117520EDC54578A60D47E" pageId="0" pageNumber="43">Phenology.</paragraph>
<paragraph id="B63775DEAA188C8B10143E11DF2BDC2A" pageId="0" pageNumber="43">Flowering from July to December.</paragraph>
</subSubSection>
<subSubSection id="E1CA25C8E8285379D33D3D8F45825344" pageId="0" pageNumber="43" type="taxonomic notes">
<paragraph id="37E62DC27FF406C398F570146287D3CB" pageId="0" pageNumber="43">Taxonomic notes.</paragraph>
<paragraph id="1A7C8EA469C00924FD19722292989AA5" pageId="0" pageNumber="43">
<taxonomicName id="9EF4FD2A702F87400504A9DE0E0D95D2" authorityName="Hagsater, J. P. Arista &amp; Edquen" authorityYear="2023" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum parvireflexilobum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="parvireflexilobum">
<emphasis id="D2989A7E0A212E861742D73DF4CE43AC" italics="true" pageId="0" pageNumber="43">Epidendrum parvireflexilobum</emphasis>
</taxonomicName>
belongs to the Schistochilum Group,
<taxonomicName id="33C8AAB03174D08588DFA8749C7009F2" genus="Secundum" lsidName="Secundum" pageId="0" pageNumber="43" rank="genus">Secundum</taxonomicName>
Subgroup, which is characterized by the caespitose habit, the erect, simple, cane-like stems, the normally elongate peduncle of the inflorescence, the erect raceme of generally non-resupinate, yellow, orange, red or purple flowers, and the lip adorned by a complex callus. The novelty is recognized by the shorter plants, up to 22-46 cm including the inflorescence, thus being one of the &quot;dwarf species&quot; in the
<taxonomicName id="9B1AD0B89F54BEEDDB7851A4329D6B13" genus="Secundum" lsidName="Secundum" pageId="0" pageNumber="43" rank="genus">
<emphasis id="89BD2E64BD2822EF5746343B222E9D06" italics="true" pageId="0" pageNumber="43">Secundum</emphasis>
</taxonomicName>
Subgroup, as most species can be over 150 cm tall; the floral segments are short and proportionately wide, the lateral lobes of the lip are flat and not twisted and the callus is formed by a single platform with 3 short ribs, that only reaches the middle of the mid-lobe.
<taxonomicName id="A48EE0202314FE107027DFCF6C2628B7" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum reflexilobum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="reflexilobum">
<emphasis id="6AA3545BE764285DD2329ED29F213CF0" italics="true" pageId="0" pageNumber="43">Epidendrum reflexilobum</emphasis>
</taxonomicName>
occurs in the regions of
<normalizedToken id="E1BB5498289F7837748CB70B51F5CA6A" originalValue="Huánuco">Huanuco</normalizedToken>
and
<normalizedToken id="F06AA4999CF61031C5FB6007941CF858" originalValue="Junín">Junin</normalizedToken>
, and is overall larger, has the same red flowers with a yellow callus, and the lateral lobes of the lip spreading, twisted 90° in natural position, narrowly obovate. The novelty is also similar to
<taxonomicName id="D1FEF1C992D85C9B006CF3B8AFA45113" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum macrocyphum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="macrocyphum">
<emphasis id="11C22F30A53D1D00FFC3D4168E348678" italics="true" pageId="0" pageNumber="43">Epidendrum macrocyphum</emphasis>
</taxonomicName>
Kraenzl. which has pink-purple flowers with a white callus formed by 3-5 straight parallel ribs on the mid-lobe, has the mid-rib longer nearly reaching the apical sinus, with two divergent bifid calli at the base of the lateral lobes, the longer segment projecting on the junction of the mid-lobe with the lateral lobes, and the lip in natural position has the mid-lobe flat, extended, the lateral lobes erect, embracing the entire column with outer margin strongly revolute. In a previous publication of
<taxonomicName id="9428FB1B36C6729A3BFB4CBB8EDEECBC" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum reflexilobum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="reflexilobum">
<emphasis id="23482CDAADB17F15080C5D892EFA8781" italics="true" pageId="0" pageNumber="43">Epidendrum reflexilobum</emphasis>
</taxonomicName>
(
<bibRefCitation id="C1280DEB3C77383066B2B3541AC5636F" author="Hagsater, E" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="43" publicationUrl="https://herbarioamo.org/index_archivos/Fascicle17(2).pdf" refId="B33" refString="Hagsater, E, Santiago, E [Eds], 2020a. The Genus Epidendrum, Part 13, Icones Orchidacearum 17(2): t. 1757-1800. https://herbarioamo.org/index_archivos/Fascicle17(2).pdf" title="The Genus Epidendrum, Part 13, Icones Orchidacearum 17 (2): t. 1757 - 1800." url="https://herbarioamo.org/index_archivos/Fascicle17(2).pdf" year="2020 a">
<normalizedToken id="49C90058B99C8588D9CD3D5C690DD010" originalValue="Hágsater">Hagsater</normalizedToken>
and Santiago 2020a
</bibRefCitation>
: t. 1835), the description is a mixture of the new species and
<taxonomicName id="CB2D65BEA0AC046A95B1CFBD0EB63C91" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum reflexilobum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="reflexilobum">
<emphasis id="F9AAAD2B6C0FCDE383113989E1214108" italics="true" pageId="0" pageNumber="43">Epidendrum reflexilobum</emphasis>
</taxonomicName>
; the latter species corresponds to the photograph on the text page, but the color plate corresponds to the new entity here described.
</paragraph>
</subSubSection>
<subSubSection id="75D4CBB0634979784B9A7EAA996CC0FD" pageId="0" pageNumber="43" type="etymology">
<paragraph id="5E0651213473B6FB97FD5B6DBCA4340D" pageId="0" pageNumber="43">Etymology.</paragraph>
<paragraph id="9DD96800B73C6FC1F420CA66B5779857" pageId="0" pageNumber="43">
From the Latin
<emphasis id="C87B830BE72E92AEB168EAFBBC95189D" italics="true" pageId="0" pageNumber="43">parvi</emphasis>
-, small, and
<taxonomicName id="71004E53C7814ADDBAAE3A4C5D357C0B" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Epidendrum reflexilobum" order="Asparagales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="species" species="reflexilobum">
<emphasis id="B9DD934D29EFD5DD1B7E9765A619D149" italics="true" pageId="0" pageNumber="43">Epidendrum reflexilobum</emphasis>
</taxonomicName>
, reflexed lobes, in reference to the smaller flowers and lobes of the lip relative to those of closely allied
<taxonomicName id="4E5485F72CF433E4D7435E38EBF15FF1" lsidName="E. reflexilobum" pageId="0" pageNumber="43" rank="species" species="reflexilobum">
<emphasis id="0F130CDD9CE2A67513F5B3A94978113D" italics="true" pageId="0" pageNumber="43">E. reflexilobum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>