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<mods:title id="096C2C4267BC6015DAA60053DC85313D">The Ants (Hymenoptera: Formicidae) of Bolshaya Svetlovodnaya (Late Eocene of Sikhote-Alin, Russian Far East)</mods:title>
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<mods:namePart id="89D93D6A3E7772D1421E07957E06BA65">Dlussky, G. M.</mods:namePart>
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<paragraph id="8312D848FFF66640FC6FFD65FB36FD06" box="[936,1275,680,701]" pageId="3" pageNumber="134">
<heading id="D85A6F24FFF66640FC6FFD65FB36FD06" bold="true" box="[936,1275,680,701]" centered="true" fontSize="9" level="3" pageId="3" pageNumber="134" reason="0">
<emphasis id="6E7CBBB8A44B0AA11C83361F96F94EE2" bold="true" pageId="3" pageNumber="134">Genus</emphasis>
<taxonomicName id="44ADA3CBFFF66640FC36FD65FB36FD06" ID-CoL="62R6N" ID-ENA="121500" authority="Lund, 1831" authorityName="Lund" authorityYear="1831" box="[1009,1275,680,701]" class="Insecta" family="Formicidae" genus="Dolichoderus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B1D9045AFFF66640FC36FD65FBB3FD06" bold="true" box="[1009,1150,680,701]" italics="true" pageId="3" pageNumber="134">Dolichoderus</emphasis>
Lund, 1831
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8312D848FFF66640FC9BFD2FFC55FCAF" pageId="3" pageNumber="134">
Type species
<taxonomicName id="44ADA3CBFFF66640FC37FD2CFA83FD4C" authority="Lund, 1831" authorityName="Lund" authorityYear="1831" box="[1008,1358,737,759]" class="Insecta" family="Formicidae" genus="Formica" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="attelaboides">
<emphasis id="B1D9045AFFF66640FC37FD2CFB05FD4C" bold="true" box="[1008,1224,737,759]" italics="true" pageId="3" pageNumber="134">Formica attelaboides</emphasis>
Lund, 1831
</taxonomicName>
, by monotypy.
</paragraph>
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<subSubSection id="CBB78BC3FFF66647FC9BFCD0FD80FCEC" lastPageId="4" lastPageNumber="135" pageId="3" pageNumber="134" type="diagnosis">
<paragraph id="8312D848FFF66640FC9BFCD0FAB5FBA7" pageId="3" pageNumber="134">
<emphasis id="B1D9045AFFF66640FC9BFCD0FC0BFC89" bold="true" box="[860,966,797,818]" pageId="3" pageNumber="134">Diagnosis</emphasis>
(for fossil imprints). Waist consisting of 1 segment (petiole); gaster without constriction between 1st and 2nd gastral (abdominal III and IV) segments. Head elliptical or oval, widest behind midlength of head sides; sometimes retort-shaped or subrectangular with rounded occipital corners, but never rectangular and longer than wide as in
<taxonomicName id="44ADA3CBFFF66640FC58FC01FB74FC5A" authority="Carpenter, 1930" authorityName="Carpenter" authorityYear="1930" box="[927,1209,972,993]" class="Insecta" family="Formicidae" genus="Protazteca" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B1D9045AFFF66640FC58FC01FBC7FC5A" bold="true" box="[927,1034,972,993]" italics="true" pageId="3" pageNumber="134">Protazteca</emphasis>
<bibRefCitation id="04641A4B2445A283EB051BD542464E68" author="Carpenter F. M." journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pagination="1 - 66" part="70" refId="ref19512" refString="Carpenter F. M. 1930. The fossil ants of North America. Bulletin of the Museum of Comparative Zoology. 70 (1): 1 66." title="The fossil ants of North America" type="journal article" year="1930">Carpenter, 1930</bibRefCitation>
</taxonomicName>
. Eyes of moderate size. Antennae of gynes and workers 12-segmented; antennae of males 13-segmented, with very short scape.
</paragraph>
<caption id="D7D288C0FFF66640FF35F893FBC2F864" pageId="3" pageNumber="134" targetBox="[193,1393,1086,1842]" targetPageId="3">
<caption id="D7D288C0FFF66640FF35F893FD65F8D4" box="[242,680,1886,1903]" pageId="3" pageNumber="134" targetBox="[193,1393,1086,1842]" targetPageId="3">
<paragraph id="8312D848FFF66640FF35F893FD65F8D4" box="[242,680,1886,1903]" pageId="3" pageNumber="134">
Figs 58. Species of genera
<taxonomicName id="44ADA3CBFFF66640FE01F892FDE0F8D4" authorityName="Lund" authorityYear="1831" box="[454,557,1887,1903]" class="Insecta" family="Formicidae" genus="Dolichoderus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B1D9045AFFF66640FE01F892FDE0F8D4" bold="true" box="[454,557,1887,1903]" italics="true" pageId="3" pageNumber="134">Dolichoderus</emphasis>
</taxonomicName>
and
<taxonomicName id="44ADA3CBFFF66640FD95F892FD69F8D4" authorityName="Donisthorpe" authorityYear="1920" box="[594,676,1887,1903]" class="Insecta" family="Formicidae" genus="Emplastus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B1D9045AFFF66640FD95F892FD69F8D4" bold="true" box="[594,676,1887,1903]" italics="true" pageId="3" pageNumber="134">Emplastus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8312D848FFF66640FF35F8B8FC31F827" pageId="3" pageNumber="134">
5
<emphasis id="B1D9045AFFF66640FED4F8B8FE3FF83D" bold="true" box="[275,498,1909,1926]" pageId="3" pageNumber="134">
<taxonomicName id="44ADA3CBFFF66640FED4F8B8FE72F83D" authority="Dlussky &amp; Rasnitsyn &amp; Perfilieva, 2015" authorityName="Dlussky &amp; Rasnitsyn &amp; Perfilieva" authorityYear="2015" box="[275,447,1909,1926]" class="Insecta" family="Formicidae" genus="Dolichoderus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="pinguis" status="sp. n.">
<emphasis id="B1D9045AFFF66640FED4F8B8FE72F83D" bold="true" box="[275,447,1909,1926]" italics="true" pageId="3" pageNumber="134">Dolichoderus pinguis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAEAB921FFF66640FE02F8BBFE3FF83D" box="[453,498,1910,1926]" pageId="3" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype PIN 3429/1134, male; 6
<emphasis id="B1D9045AFFF66640FCE8F8B8FC31F83D" bold="true" box="[815,1020,1909,1926]" pageId="3" pageNumber="134">
<taxonomicName id="44ADA3CBFFF66640FCE8F8B8FC05F83D" authority="Dlussky &amp; Rasnitsyn &amp; Perfilieva, 2015" authorityName="Dlussky &amp; Rasnitsyn &amp; Perfilieva" authorityYear="2015" box="[815,968,1909,1926]" class="Insecta" family="Formicidae" genus="Emplastus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="macrops" status="sp. n.">
<emphasis id="B1D9045AFFF66640FCE8F8B8FC05F83D" bold="true" box="[815,968,1909,1926]" italics="true" pageId="3" pageNumber="134">Emplastus macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAEAB921FFF66640FC08F8BBFC31F83D" box="[975,1020,1910,1926]" pageId="3" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype PIN 3429/1187, male; 7
<emphasis id="B1D9045AFFF66640FAE2F8B8FEFBF826" bold="true" pageId="3" pageNumber="134">
<taxonomicName id="44ADA3CBFFF66640FAE2F8B8FECBF827" authority="Dlussky &amp; Rasnitsyn &amp; Perfilieva, 2015" authorityName="Dlussky &amp; Rasnitsyn &amp; Perfilieva" authorityYear="2015" class="Insecta" family="Formicidae" genus="Emplastus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="elongatus" status="sp. n.">
<emphasis id="B1D9045AFFF66640FAE2F8B8FECBF827" bold="true" italics="true" pageId="3" pageNumber="134">Emplastus elongatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAEAB921FFF66640FECCF841FEFBF826" box="[267,310,1932,1949]" pageId="3" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype PIN 3429/1115, male; 8
<emphasis id="B1D9045AFFF66640FD96F841FCC6F826" bold="true" box="[593,779,1932,1949]" pageId="3" pageNumber="134">
<taxonomicName id="44ADA3CBFFF66640FD96F841FD11F827" authority="Dlussky &amp; Rasnitsyn &amp; Perfilieva, 2015" authorityName="Dlussky &amp; Rasnitsyn &amp; Perfilieva" authorityYear="2015" box="[593,732,1932,1948]" class="Insecta" family="Formicidae" genus="Emplastus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="dubius" status="sp. n.">
<emphasis id="B1D9045AFFF66640FD96F841FD11F827" bold="true" box="[593,732,1932,1948]" italics="true" pageId="3" pageNumber="134">Emplastus dubius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAEAB921FFF66640FD27F841FCC6F826" box="[736,779,1932,1949]" pageId="3" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype PIN 3429/109, gyne.
</paragraph>
<paragraph id="8312D848FFF66640FF35F86FFDB9F808" box="[242,628,1954,1971]" pageId="3" pageNumber="134">
Рис.58. Виды родов
<taxonomicName id="44ADA3CBFFF66640FE62F86FFDC6F808" authorityName="Lund" authorityYear="1831" box="[421,523,1954,1971]" class="Insecta" family="Formicidae" genus="Dolichoderus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B1D9045AFFF66640FE62F86FFDC6F808" bold="true" box="[421,523,1954,1971]" italics="true" pageId="3" pageNumber="134">Dolichoderus</emphasis>
</taxonomicName>
и
<taxonomicName id="44ADA3CBFFF66640FDD8F86FFDBDF808" authorityName="Donisthorpe" authorityYear="1920" box="[543,624,1954,1971]" class="Insecta" family="Formicidae" genus="Emplastus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B1D9045AFFF66640FDD8F86FFDBDF808" bold="true" box="[543,624,1954,1971]" italics="true" pageId="3" pageNumber="134">Emplastus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8312D848FFF66640FF35F874FBC2F864" pageId="3" pageNumber="134">
5
<emphasis id="B1D9045AFFF66640FED3F875FE23F871" bold="true" box="[276,494,1976,1994]" pageId="3" pageNumber="134">
<taxonomicName id="44ADA3CBFFF66640FED3F875FE74F872" authority="Dlussky &amp; Rasnitsyn &amp; Perfilieva, 2015" authorityName="Dlussky &amp; Rasnitsyn &amp; Perfilieva" authorityYear="2015" box="[276,441,1976,1993]" class="Insecta" family="Formicidae" genus="Dolichoderus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="pinguis" status="sp. n.">
<emphasis id="B1D9045AFFF66640FED3F875FE74F872" bold="true" box="[276,441,1976,1993]" italics="true" pageId="3" pageNumber="134">Dolichoderus pinguis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAEAB921FFF66640FE07F874FE23F871" box="[448,494,1977,1994]" pageId="3" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, голотип PIN 3429/1134, самец; 6
<emphasis id="B1D9045AFFF66640FCE5F875FC3DF871" bold="true" box="[802,1008,1976,1994]" pageId="3" pageNumber="134">
<taxonomicName id="44ADA3CBFFF66640FCE5F875FC71F872" authority="Dlussky &amp; Rasnitsyn &amp; Perfilieva, 2015" authorityName="Dlussky &amp; Rasnitsyn &amp; Perfilieva" authorityYear="2015" box="[802,956,1976,1993]" class="Insecta" family="Formicidae" genus="Emplastus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="macrops" status="sp. n.">
<emphasis id="B1D9045AFFF66640FCE5F875FC71F872" bold="true" box="[802,956,1976,1993]" italics="true" pageId="3" pageNumber="134">Emplastus macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAEAB921FFF66640FC04F874FC3DF871" box="[963,1008,1977,1994]" pageId="3" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, голотип PIN 3429/1187, самец; 7
<emphasis id="B1D9045AFFF66640FAE2F875FEFBF85B" bold="true" pageId="3" pageNumber="134">
<taxonomicName id="44ADA3CBFFF66640FAE2F875FECBF864" authority="Dlussky &amp; Rasnitsyn &amp; Perfilieva, 2015" authorityName="Dlussky &amp; Rasnitsyn &amp; Perfilieva" authorityYear="2015" class="Insecta" family="Formicidae" genus="Emplastus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="elongatus" status="sp. n.">
<emphasis id="B1D9045AFFF66640FAE2F875FECBF864" bold="true" italics="true" pageId="3" pageNumber="134">Emplastus elongatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAEAB921FFF66640FECCF81DFEFBF85B" box="[267,310,2000,2016]" pageId="3" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, голотип PIN 3429/1115, самец; 8
<emphasis id="B1D9045AFFF66640FD9CF802FCD8F85B" bold="true" box="[603,789,1999,2016]" pageId="3" pageNumber="134">
<taxonomicName id="44ADA3CBFFF66640FD9CF802FD28F864" authority="Dlussky &amp; Rasnitsyn &amp; Perfilieva, 2015" authorityName="Dlussky &amp; Rasnitsyn &amp; Perfilieva" authorityYear="2015" box="[603,741,1999,2015]" class="Insecta" family="Formicidae" genus="Emplastus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="dubius" status="sp. n.">
<emphasis id="B1D9045AFFF66640FD9CF802FD28F864" bold="true" box="[603,741,1999,2015]" italics="true" pageId="3" pageNumber="134">Emplastus dubius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAEAB921FFF66640FD2EF81DFCD8F85B" box="[745,789,2000,2016]" pageId="3" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, голотип PIN 3429/109, самка.
</paragraph>
</caption>
<paragraph id="8312D848FFF16647FF7AFED9FE54FE62" pageId="4" pageNumber="135">Mandibles triangular, dentate. Propodeum bispinate, bidentate, or distinctly angular in side view, always with concave declivity. Petiole of various form. Forewing with cells 1+2r, 3r, rm and mcu closed; rm usually triangular. Cell 3r touching wing margin. Icu&gt; 1.45. Integument thick, often coarsely sculptured. Male genitalia small, not permanently external.</paragraph>
<paragraph id="8312D848FFF16647FF2AFE2FFD80FCEC" pageId="4" pageNumber="135">
<emphasis id="B1D9045AFFF16647FF2AFE2FFE68FE4C" bold="true" box="[237,421,482,503]" pageId="4" pageNumber="135">Species included.</emphasis>
The genus
<taxonomicName id="44ADA3CBFFF16647FDD0FE2CFD56FE4C" authorityName="Lund" authorityYear="1831" box="[535,667,481,503]" class="Insecta" family="Formicidae" genus="Dolichoderus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B1D9045AFFF16647FDD0FE2CFD56FE4C" bold="true" box="[535,667,481,503]" italics="true" pageId="4" pageNumber="135">Dolichoderus</emphasis>
</taxonomicName>
comprises 132species and 20subspecies in the modern fauna [
<bibRefCitation id="8BF2C26C6C1D9CB1D5E6ECE965612C8D" author="Bolton B." journalOrPublisher="Memoirs of the American Entomological Institute" publicationUrl="http://antcat.org" refId="ref19444" refString="Bolton B. 2015. An online catalog of the ants of the world. Available at: http: // antcat. org (accessed May, 2015)." title="An online catalog of the ants of the world" type="url" year="2015">Bolton, 2015</bibRefCitation>
]. The vast majority of them are distributed in the Neotropical, Indo-Australian, and Australian Regions. Only 9species are known from the Oriental Region, 4from the Nearctic Region and 2 from the Palearctic Region, and none have been recorded from the Afrotropical Region. A total of 47 extinct species have been described. The oldest species
<taxonomicName id="44ADA3CBFFF16647FEB9FD01FDE0FD5A" authority="Dlussky" authorityName="Dlussky" box="[382,557,716,737]" class="Insecta" family="Formicidae" genus="Dolichoderus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="kohlsi">
<emphasis id="B1D9045AFFF16647FEB9FD01FE1BFD5A" bold="true" box="[382,470,716,737]" italics="true" pageId="4" pageNumber="135">D.kohlsi</emphasis>
<bibRefCitation id="53DB632C025BA93FC719B5331749A3D1" author="Dlussky G. M." journalOrPublisher="Paleontological Journal" pagination="50 - 63" part="36" refId="ref19859" refString="Dlussky G. M. 2002. Ants of the genus Dolichoderus Lund (Hymenoptera: Formicidae) from the Baltic and Rovno Ambers. Paleontological Journal. 36 (1): 50 63." title="Ants of the genus Dolichoderus Lund (Hymenoptera: Formicidae) from the Baltic and Rovno Ambers" type="journal article" year="2002">Dlussky et Rasnitsyn, 2002</bibRefCitation>
</taxonomicName>
is recorded from the Middle Eocene Green River Formation, USA [
<bibRefCitation id="7AB6C47711FA773F94F02562EB6CF4A6" author="Dlussky G. M. &amp; Rasnitsyn A. P." journalOrPublisher="Russian Entomological Journal. 2002" pagination="411 - 436" part="11" refId="ref20187" refString="Dlussky G. M., Rasnitsyn A. P. 2003. Ants (Hymenoptera: Formicidae) of Formation Green River and some other Middle Eocene deposits of North America. Russian Entomological Journal. 2002. 11: 411 436." title="Ants (Hymenoptera: Formicidae) of Formation Green River and some other Middle Eocene deposits of North America" type="journal article" year="2003">Dlussky, Rasnitsyn, 2003</bibRefCitation>
]. Nearly half of the fossil species (24) have been described from Late Eocene European ambers [
<bibRefCitation id="2667BB6BBD9EAE081A3BBAEF88DF9A93" author="Dlussky G. M." journalOrPublisher="Paleontological Journal" pagination="50 - 63" part="36" refId="ref19859" refString="Dlussky G. M. 2002. Ants of the genus Dolichoderus Lund (Hymenoptera: Formicidae) from the Baltic and Rovno Ambers. Paleontological Journal. 36 (1): 50 63." title="Ants of the genus Dolichoderus Lund (Hymenoptera: Formicidae) from the Baltic and Rovno Ambers" type="journal article" year="2002">Dlussky, 2002</bibRefCitation>
,
<bibRefCitation id="682F5A7676A1A80D67A11C9B53D23A03" author="Dlussky G. M." journalOrPublisher="Vestnik Zoologii" pagination="497 - 514" part="42" refId="ref19988" refString="Dlussky G. M. 2008. New species of ants of the genus Dolichoderus (Hymenoptera, Formicidae) from Late Eocene ambers of Europe. Vestnik Zoologii. 42 (6): 497 514 (in Russian)." title="New species of ants of the genus Dolichoderus (Hymenoptera, Formicidae) from Late Eocene ambers of Europe" type="journal article" year="2008">2008</bibRefCitation>
].
</paragraph>
</subSubSection>
</treatment>
</document>