289 lines
39 KiB
XML
289 lines
39 KiB
XML
<document id="3E892DD97FF4003D786C021BE5947FDB" ID-DOI="10.11646/zootaxa.5343.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8334181" ID-ZooBank="BB852CA9-6C6E-4016-9972-AAF2C378E41F" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1694429699348" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2023" docId="03A487CEFFFBFFD785A550F96A07F81E" docLanguage="en" docName="zootaxa.5343.3.2.pdf" docOrigin="Zootaxa 5343 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5343.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procloeon (Securiops) mutadens Kluge 2023, comb. n." docType="treatment" docVersion="3" lastPageNumber="262" masterDocId="FF9DFFB6FFE8FFC4853251076B7DFFDC" masterDocTitle="Redescription of the subgenus Securiops Jacobus, McCafferty & Gattolliat 2006 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)" masterLastPageNumber="272" masterPageNumber="243" pageNumber="262" updateTime="1694453124034" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo id="3FF066B16D70AA45FD34AE9D587AE22D">
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<mods:title id="630867F628DE8A768ED6F9F4F940DB92">Redescription of the subgenus Securiops Jacobus, McCafferty & Gattolliat 2006 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)</mods:title>
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<mods:namePart id="01B03D173844A7ABC95C97461B7FE49D">Kluge, Nikita J.</mods:namePart>
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<mods:title id="49EB7CDDE34C3A5265A4DE490CD7D7FB">Zootaxa</mods:title>
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<mods:part id="16D9BC03037F33C2BEF2D20BAAB74C62">
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<mods:date id="4A9D74BDD4EEAF6BBE43FF0085063986">2023</mods:date>
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<mods:number id="D455B7C1E2AC295A1BA9EB288C80A7D0">2023-09-08</mods:number>
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<mods:number id="C75A6D9CDD1AAAABE6A6B2852AE05FF8">5343</mods:number>
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<subSubSection id="C3176553FFFBFFD785A550F968E5FDC5" box="[151,920,510,537]" pageId="19" pageNumber="262" type="nomenclature">
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<paragraph id="8BB236D8FFFBFFD785A550F968E5FDC5" blockId="19.[151,920,510,573]" box="[151,920,510,537]" pageId="19" pageNumber="262">
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<heading id="D0FA81B4FFFBFFD785A550F968E5FDC5" bold="true" box="[151,920,510,537]" fontSize="11" level="1" pageId="19" pageNumber="262" reason="1">
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<emphasis id="B979EACAFFFBFFD785A550F968E5FDC5" bold="true" box="[151,920,510,537]" pageId="19" pageNumber="262">
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<taxonomicName id="4C0D4D5BFFFBFFD785A550F96858FDC4" authority="(Jacobus et al. 2006)" authorityName="Kluge" authorityYear="2023" baseAuthorityName="Jacobus, McCafferty & Gattolliat" baseAuthorityYear="2006" box="[151,805,510,537]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="262" phylum="Arthropoda" rank="species" species="mutadens" status="comb. n." subGenus="Securiops">
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<emphasis id="B979EACAFFFBFFD785A550F96A6EFDC5" bold="true" box="[151,275,510,537]" italics="true" pageId="19" pageNumber="262">Procloeon</emphasis>
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(
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<emphasis id="B979EACAFFFBFFD7841650F86AE6FDC5" bold="true" box="[292,411,511,537]" italics="true" pageId="19" pageNumber="262">Securiops</emphasis>
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)
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<emphasis id="B979EACAFFFBFFD7849E50F9695CFDC5" bold="true" box="[428,545,510,537]" italics="true" pageId="19" pageNumber="262">mutadens</emphasis>
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(
|
||
<bibRefCitation id="EF9C4B29FFFBFFD7870350F96860FDC4" author="Jacobus, L. M. & McCafferty, W. P. & Gattolliat, J. - L." box="[561,797,510,537]" pageId="19" pageNumber="262" pagination="129 - 140" refId="ref12262" refString="Jacobus, L. M., McCafferty, W. P. & Gattolliat, J. - L. (2006) Taxonomy of Afrotropical Securiops, new genus, and Cloeodes Traver (Ephemeroptera: Baetidae). African Entomology, 14 (1), 129 - 140." type="journal article" year="2006">
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||
Jacobus
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<emphasis id="B979EACAFFFBFFD787AD50F869AEFDC5" bold="true" box="[671,723,510,537]" italics="true" pageId="19" pageNumber="262">et al</emphasis>
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||
. 2006
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||
</bibRefCitation>
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||
)
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||
</taxonomicName>
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<taxonomicNameLabel id="A24A57B1FFFBFFD7861F50F868E5FDC5" box="[813,920,511,537]" pageId="19" pageNumber="262" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3176553FFFBFFD785A553246A48FDE1" box="[151,309,546,573]" pageId="19" pageNumber="262" type="description">
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<paragraph id="8BB236D8FFFBFFD785A553246A48FDE1" blockId="19.[151,920,510,573]" box="[151,309,546,573]" pageId="19" pageNumber="262">
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||
(
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||
<figureCitation id="13362A5DFFFBFFD785AD53246A50FDE1" box="[159,301,546,573]" captionStart="FIGURES 98–105" captionStartId="20.[151,264,1857,1881]" captionTargetBox="[151,1435,201,1834]" captionTargetId="figure-18@20.[151,1435,201,1834]" captionTargetPageId="20" captionText="FIGURES 98–105. Procloeon (Securiops) mutadens. 98–99, left and right mandibles; 100, labrum; 101, maxilla; 102, larval cuticle of metanotum with vestige of hind protopteron; 103, abdominal tergum V; 104, abdominal sternum V; 105, caudalii. Abbreviation: sc, scales and scale sockets." figureDoi="http://doi.org/10.5281/zenodo.8334229" httpUri="https://zenodo.org/record/8334229/files/figure.png" pageId="19" pageNumber="262">Figs 98–105</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3176553FFFBFFD785A553656804FD6B" pageId="19" pageNumber="262" type="reference_group">
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||
<paragraph id="8BB236D8FFFBFFD785A5536568F0FDA6" blockId="19.[151,1431,610,695]" box="[151,909,610,634]" pageId="19" pageNumber="262">
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||
<treatmentCitationGroup id="AB1D11F6FFFBFFD785A5536568F0FDA6" box="[151,909,610,634]" pageId="19" pageNumber="262">
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<treatmentCitation id="0AAC10C9FFFBFFD785A553656A88FDA6" author="Gillies, M. T." box="[151,501,610,634]" page="53" pageId="19" pageNumber="262" year="1988">
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||
<taxonomicName id="4C0D4D5BFFFBFFD785A553656A88FDA6" authority=": Gillies 1988: 53" authorityName="Gillies" authorityPageNumber="53" authorityYear="1988" box="[151,501,610,634]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="262" phylum="Arthropoda" rank="species" species="dentatum">
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||
<emphasis id="B979EACAFFFBFFD785A553656A38FDA6" box="[151,325,610,634]" italics="true" pageId="19" pageNumber="262">Cloeon dentatum</emphasis>
|
||
:
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<bibRefCitation id="EF9C4B29FFFBFFD7846053656A88FDA6" author="Gillies, M. T." box="[338,501,610,634]" pageId="19" pageNumber="262" pagination="49 - 59" refId="ref12030" refString="Gillies, M. T. (1988) Descriptions of the nymphs of some Afrotropical Baetidae (Ephemeroptera). I. Cloeon Leach and Rhithrocloeon Gillies. Aquatic Insects, 10 (1), 49 - 59. https: // doi. org / 10.1080 / 01650428809361310" type="journal article" year="1988">Gillies 1988: 53</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
(larva; non imago, nec Kimmins 1956).
|
||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BB236D8FFFBFFD785A553876EEAFD44" blockId="19.[151,1431,610,695]" box="[151,1431,640,665]" pageId="19" pageNumber="262">
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<treatmentCitationGroup id="AB1D11F6FFFBFFD785A553876EEAFD44" box="[151,1431,640,665]" pageId="19" pageNumber="262">
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<treatmentCitation id="0AAC10C9FFFBFFD785A55387692AFD45" author="Gillies, M. T." box="[151,599,640,665]" page="208" pageId="19" pageNumber="262" year="1990">
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||
<taxonomicName id="4C0D4D5BFFFBFFD785A55387692AFD45" authority=": Gillies 1990 a: 208" authorityName="Gillies" authorityPageNumber="208" authorityYear="1990" box="[151,599,640,665]" class="Insecta" family="Baetidae" genus="Potamocloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="262" phylum="Arthropoda" rank="species" species="dentatum">
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||
<emphasis id="B979EACAFFFBFFD785A553876AF3FD44" box="[151,398,640,664]" italics="true" pageId="19" pageNumber="262">Potamocloeon dentatum</emphasis>
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||
:
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||
<bibRefCitation id="EF9C4B29FFFBFFD784AE5387692AFD45" author="Gillies, M. T." box="[412,599,640,665]" pageId="19" pageNumber="262" pagination="207 - 208" refId="ref12081" refString="Gillies, M. T. (1990 a) A new genus for the Afrotropical mayfly, Cloeon dentatum Kimmins (Ephem., Baetidae). Entomologist's Monthly Magazine, 126, 207 - 208." type="journal article" year="1990">Gillies 1990a: 208</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
(larva, non imago, nec Kimmins 1956);
|
||
<treatmentCitation id="0AAC10C9FFFBFFD786C653876E3BFD45" author="Jacobus, L. M. & McCafferty, W. P." box="[1012,1350,640,665]" page="474" pageId="19" pageNumber="262" year="2005">
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||
<bibRefCitation id="EF9C4B29FFFBFFD786C653876E3BFD45" author="Jacobus, L. M. & McCafferty, W. P." box="[1012,1350,640,665]" pageId="19" pageNumber="262" pagination="474 - 475" refId="ref12218" refString="Jacobus, L. M. & McCafferty, W. P. (2005) Validation of Potamocloeon macaferiorum Lugo-Ortiz (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 107 (2), 474 - 475." type="journal article" year="2005">Jacobus & McCafferty 2005: 474</bibRefCitation>
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</treatmentCitation>
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(larva).
|
||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BB236D8FFFBFFD785A553986804FD6B" blockId="19.[151,1431,610,695]" box="[151,889,671,695]" pageId="19" pageNumber="262">
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<treatmentCitationGroup id="AB1D11F6FFFBFFD785A553986804FD6B" box="[151,889,671,695]" pageId="19" pageNumber="262">
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<treatmentCitation id="0AAC10C9FFFBFFD785A553986855FD6B" author="Jacobus, L. M. & McCafferty, W. P. & Gattolliat, J. - L." box="[151,808,671,695]" page="137" pageId="19" pageNumber="262" year="2006">
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||
<taxonomicName id="4C0D4D5BFFFBFFD785A553986855FD6B" ID-CoL="6XZCG" authority="Jacobus, McCafferty & Gattolliat 2006: 137" authorityName="Jacobus, McCafferty & Gattolliat" authorityPageNumber="137" authorityYear="2006" box="[151,808,671,695]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="262" phylum="Arthropoda" rank="species" species="mutadens">
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||
<emphasis id="B979EACAFFFBFFD785A553986A1FFD6B" box="[151,354,671,695]" italics="true" pageId="19" pageNumber="262">Securiops mutadens</emphasis>
|
||
<bibRefCitation id="EF9C4B29FFFBFFD7845B53986855FD6B" author="Jacobus, L. M. & McCafferty, W. P. & Gattolliat, J. - L." box="[361,808,671,695]" pageId="19" pageNumber="262" pagination="129 - 140" refId="ref12262" refString="Jacobus, L. M., McCafferty, W. P. & Gattolliat, J. - L. (2006) Taxonomy of Afrotropical Securiops, new genus, and Cloeodes Traver (Ephemeroptera: Baetidae). African Entomology, 14 (1), 129 - 140." type="journal article" year="2006">Jacobus, McCafferty & Gattolliat 2006: 137</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(larva).
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3176553FFFBFFD785A553E76FA4FD27" box="[151,1241,736,763]" pageId="19" pageNumber="262" type="materials_examined">
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<paragraph id="8BB236D8FFFBFFD785A553E76FA4FD27" blockId="19.[151,1437,736,1986]" box="[151,1241,736,763]" pageId="19" pageNumber="262">
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<emphasis id="B979EACAFFFBFFD785A553E76AFCFD26" bold="true" box="[151,385,736,762]" pageId="19" pageNumber="262">Material examined.</emphasis>
|
||
<materialsCitation id="3B653C85FFFBFFD784B553E76FA8FD27" box="[391,1237,736,763]" collectingDate="1993-02-19" collectionCode="R, T" collectorName="Gillies, Y" country="Gambia" location="Fatoto" municipality="Gambia" pageId="19" pageNumber="262" specimenCount="1" specimenCount-female="1">
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<collectingCountry id="F31A7648FFFBFFD784B553E76A86FD26" box="[391,507,736,762]" name="Gambia" pageId="19" pageNumber="262">GAMBIA</collectingCountry>
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,
|
||
<location id="8ED26003FFFBFFD7873553E6692CFD27" LSID="urn:lsid:plazi:treatment:03A487CEFFFBFFD785A550F96A07F81E:8ED26003FFFBFFD7873553E6692CFD27" box="[519,593,737,763]" country="Gambia" municipality="Gambia" name="Fatoto" pageId="19" pageNumber="262">Fatoto</location>
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,
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||
<collectionCode id="ED1CAE1DFFFBFFD7876F53E6690EFD27" box="[605,627,737,763]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="262">R</collectionCode>
|
||
.
|
||
<collectingMunicipality id="6BD6ACA2FFFBFFD7874F53E769A4FD26" box="[637,729,736,762]" pageId="19" pageNumber="262">Gambia</collectingMunicipality>
|
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,
|
||
<date id="FFB31018FFFBFFD787D653E76826FD26" box="[740,859,736,762]" pageId="19" pageNumber="262" value="1993-02-19">
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||
<collectingDate id="EFF7E9F0FFFBFFD787D653E76826FD26" box="[740,859,736,762]" pageId="19" pageNumber="262" value="1993-02-19">19.II.1993</collectingDate>
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||
</date>
|
||
, coll. M.
|
||
<collectionCode id="ED1CAE1DFFFBFFD7868C53E668ACFD27" box="[958,977,737,763]" name="Tavera, Department of Geology and Geophysics" pageId="19" pageNumber="262">T</collectionCode>
|
||
.
|
||
<collectorName id="26F8530EFFFBFFD786E953E76F37FD26" box="[987,1098,736,762]" pageId="19" pageNumber="262">Gillies, Y</collectorName>
|
||
127-8: 1 L
|
||
<specimenCount id="9D0BFD51FFFBFFD781F653E66FA8FD27" box="[1220,1237,737,763]" pageId="19" pageNumber="262" type="female">♀</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3176553FFFBFFD785F552036A07F81E" pageId="19" pageNumber="262" type="description">
|
||
<paragraph id="8BB236D8FFFBFFD785F552036988FCC2" blockId="19.[151,1437,736,1986]" box="[199,757,772,798]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F552036988FCC2" bold="true" box="[199,757,772,798]" pageId="19" pageNumber="262">Additions and corrections to larval description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFFBFFD785F552206834FC0E" blockId="19.[151,1437,736,1986]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F552206A0BFC9E" bold="true" box="[199,374,807,834]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F552206A12FC9E" bold="true" box="[199,367,807,834]" italics="true" pageId="19" pageNumber="262">Maxillary palp</emphasis>
|
||
.
|
||
</emphasis>
|
||
Length of 2nd (last) segment of maxillary palp was accepted as a key character separating species of
|
||
<taxonomicName id="4C0D4D5BFFFBFFD78587524B6A5EFCBA" authorityName=": Kluge" authorityYear="2020" box="[181,291,844,870]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACAFFFBFFD78587524B6A5EFCBA" box="[181,291,844,870]" italics="true" pageId="19" pageNumber="262">Securiops</emphasis>
|
||
</taxonomicName>
|
||
: in
|
||
<taxonomicName id="4C0D4D5BFFFBFFD7847F524B6AAEFCBA" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[333,467,844,870]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="262" phylum="Arthropoda" rank="species" species="mutadens">
|
||
<emphasis id="B979EACAFFFBFFD7847F524B6AAEFCBA" box="[333,467,844,870]" italics="true" pageId="19" pageNumber="262">S. mutadens</emphasis>
|
||
</taxonomicName>
|
||
it was said to be 0.55 of 1st segment length, in contrast to
|
||
<quantity id="4CF59B3DFFFBFFD7816B524B6FDBFCBA" box="[1113,1190,844,871]" metricMagnitude="-2" metricUnit="m" metricValue="2.159" pageId="19" pageNumber="262" unit="in" value="0.85">0.85 in</quantity>
|
||
<taxonomicName id="4C0D4D5BFFFBFFD7819E524B6E10FCBA" authorityName=": Kluge" authorityYear="2022" box="[1196,1389,844,870]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="262" phylum="Arthropoda" rank="species" species="macafertiorum">
|
||
<emphasis id="B979EACAFFFBFFD7819E524B6E10FCBA" box="[1196,1389,844,870]" italics="true" pageId="19" pageNumber="262">S. macafertiorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0D4D5BFFFBFFD785A552776A49FC56" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[151,308,880,906]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="262" phylum="Arthropoda" rank="species" species="mandradae">
|
||
<emphasis id="B979EACAFFFBFFD785A552776A49FC56" box="[151,308,880,906]" italics="true" pageId="19" pageNumber="262">S. mandradae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9C4B29FFFBFFD78470527769FBFC57" author="Jacobus, L. M. & McCafferty, W. P." box="[322,646,880,907]" pageId="19" pageNumber="262" pagination="474 - 475" refId="ref12218" refString="Jacobus, L. M. & McCafferty, W. P. (2005) Validation of Potamocloeon macaferiorum Lugo-Ortiz (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 107 (2), 474 - 475." type="journal article" year="2005">Jacobus & McCafferty 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9C4B29FFFBFFD787A052776812FC56" author="Jacobus, L. M. & McCafferty, W. P. & Gattolliat, J. - L." box="[658,879,880,907]" pageId="19" pageNumber="262" pagination="129 - 140" refId="ref12262" refString="Jacobus, L. M., McCafferty, W. P. & Gattolliat, J. - L. (2006) Taxonomy of Afrotropical Securiops, new genus, and Cloeodes Traver (Ephemeroptera: Baetidae). African Entomology, 14 (1), 129 - 140." type="journal article" year="2006">
|
||
Jacobus
|
||
<emphasis id="B979EACAFFFBFFD787C052766859FC56" box="[754,804,880,906]" italics="true" pageId="19" pageNumber="262">et al</emphasis>
|
||
. 2006
|
||
</bibRefCitation>
|
||
). This proportion (0.55) corresponds to the figure in the description (
|
||
<bibRefCitation id="EF9C4B29FFFBFFD7845A52936A85FC72" author="Gillies, M. T." box="[360,504,916,942]" pageId="19" pageNumber="262" pagination="49 - 59" refId="ref12030" refString="Gillies, M. T. (1988) Descriptions of the nymphs of some Afrotropical Baetidae (Ephemeroptera). I. Cloeon Leach and Rhithrocloeon Gillies. Aquatic Insects, 10 (1), 49 - 59. https: // doi. org / 10.1080 / 01650428809361310" type="journal article" year="1988">Gillies 1988</bibRefCitation>
|
||
: fig. 30), but in another larva from the same locality in
|
||
<collectingCountry id="F31A7648FFFBFFD7815652936F90FC72" box="[1124,1261,916,942]" name="Gambia" pageId="19" pageNumber="262">Gambia, the</collectingCountry>
|
||
2nd segment is longer, and its ratio to the 1st segment is about 0.7 (
|
||
<figureCitation id="13362A5DFFFBFFD787E852BF6846FC0E" box="[730,827,952,978]" captionStart="FIGURES 98–105" captionStartId="20.[151,264,1857,1881]" captionTargetBox="[151,1435,201,1834]" captionTargetId="figure-18@20.[151,1435,201,1834]" captionTargetPageId="20" captionText="FIGURES 98–105. Procloeon (Securiops) mutadens. 98–99, left and right mandibles; 100, labrum; 101, maxilla; 102, larval cuticle of metanotum with vestige of hind protopteron; 103, abdominal tergum V; 104, abdominal sternum V; 105, caudalii. Abbreviation: sc, scales and scale sockets." figureDoi="http://doi.org/10.5281/zenodo.8334229" httpUri="https://zenodo.org/record/8334229/files/figure.png" pageId="19" pageNumber="262">Fig. 101</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFFBFFD785F552DC68A7FBBE" blockId="19.[151,1437,736,1986]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F552DC6A2DFC2A" bold="true" box="[199,336,987,1014]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F552DC6A34FC2A" bold="true" box="[199,329,987,1014]" italics="true" pageId="19" pageNumber="262">Labial palp</emphasis>
|
||
.
|
||
</emphasis>
|
||
<bibRefCitation id="EF9C4B29FFFBFFD7846452DB6A8CFC2A" author="Gillies, M. T." box="[342,497,988,1014]" pageId="19" pageNumber="262" pagination="49 - 59" refId="ref12030" refString="Gillies, M. T. (1988) Descriptions of the nymphs of some Afrotropical Baetidae (Ephemeroptera). I. Cloeon Leach and Rhithrocloeon Gillies. Aquatic Insects, 10 (1), 49 - 59. https: // doi. org / 10.1080 / 01650428809361310" type="journal article" year="1988">Gillies (1988)</bibRefCitation>
|
||
described the labial palp as having ́two segments», taking the 2nd and 3dr segments fore one; however, the border separating these segments and the muscle going from the 2nd segment to the 3rd one are retained, as in other
|
||
<taxonomicName id="4C0D4D5BFFFBFFD7849D55236960FBE2" authorityName=": Kluge" authorityYear="2020" box="[431,541,1060,1086]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EACAFFFBFFD7849D55236960FBE2" box="[431,541,1060,1086]" italics="true" pageId="19" pageNumber="262">Securiops</emphasis>
|
||
</taxonomicName>
|
||
(as in
|
||
<figureCitation id="13362A5DFFFBFFD78742552369B8FBE2" box="[624,709,1060,1086]" captionStart="FIGURES 40–46" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-16@7.[151,1436,181,1817]" captionTargetPageId="7" captionText="FIGURES 40–46. Procloeon (Securiops) falcatum, labium. 40, integral labium; 41–46, parts of exuviae: 41, glossae and paraglossae, ventral view; 42, half of labium of another individual (ventral setae of glossae directed distally); 43, the same individual, with ventral setae of glossae directed proximally; 44, distal segments of labial palp; 45–46, enlarged glossae in ventral and dorsal view (ventral setae directed distally). Abbreviations: m2-3, muscle going from 2nd palpomere to base of 3rd palpomere; pm1, pm2, pm3, 1st–3rd palpomeres; s.d, dorsal setae of glossae; s.v, ventral setae of glossa." figureDoi="http://doi.org/10.5281/zenodo.8334193" httpUri="https://zenodo.org/record/8334193/files/figure.png" pageId="19" pageNumber="262">Fig. 40</figureCitation>
|
||
). Distal segment of labial palp without row of stout setae, with several stout setae near outer-distal and inner-distal corners (as in
|
||
<figureCitation id="13362A5DFFFBFFD78648554F68B3FBBE" box="[890,974,1096,1122]" captionStart="FIGURES 40–46" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-16@7.[151,1436,181,1817]" captionTargetPageId="7" captionText="FIGURES 40–46. Procloeon (Securiops) falcatum, labium. 40, integral labium; 41–46, parts of exuviae: 41, glossae and paraglossae, ventral view; 42, half of labium of another individual (ventral setae of glossae directed distally); 43, the same individual, with ventral setae of glossae directed proximally; 44, distal segments of labial palp; 45–46, enlarged glossae in ventral and dorsal view (ventral setae directed distally). Abbreviations: m2-3, muscle going from 2nd palpomere to base of 3rd palpomere; pm1, pm2, pm3, 1st–3rd palpomeres; s.d, dorsal setae of glossae; s.v, ventral setae of glossa." figureDoi="http://doi.org/10.5281/zenodo.8334193" httpUri="https://zenodo.org/record/8334193/files/figure.png" pageId="19" pageNumber="262">Fig. 44</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFFBFFD785F5556C6A04FB13" blockId="19.[151,1437,736,1986]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F5556C696AFB5A" bold="true" box="[199,535,1131,1158]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F5556C696DFB5A" bold="true" box="[199,528,1131,1158]" italics="true" pageId="19" pageNumber="262">Abdominal terga and sterna</emphasis>
|
||
.
|
||
</emphasis>
|
||
Lateral spines are present on abdominal segments
|
||
<collectionCode id="ED1CAE1DFFFBFFD78153556A6F02FB5B" box="[1121,1151,1133,1159]" pageId="19" pageNumber="262">IV</collectionCode>
|
||
–IX, varying from 2 to 8 spines on one side; their number is not as constant as reported by
|
||
<bibRefCitation id="EF9C4B29FFFBFFD786B655976F58FB76" author="Gillies, M. T." box="[900,1061,1168,1194]" pageId="19" pageNumber="262" pagination="49 - 59" refId="ref12030" refString="Gillies, M. T. (1988) Descriptions of the nymphs of some Afrotropical Baetidae (Ephemeroptera). I. Cloeon Leach and Rhithrocloeon Gillies. Aquatic Insects, 10 (1), 49 - 59. https: // doi. org / 10.1080 / 01650428809361310" type="journal article" year="1988">Gillies (1988)</bibRefCitation>
|
||
. Posterolateral spines are present on segments II–
|
||
<collectionCode id="ED1CAE1DFFFBFFD7847E55B26A08FB13" box="[332,373,1205,1231]" pageId="19" pageNumber="262">VII</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFFBFFD785F555DF6E21FA83" blockId="19.[151,1437,736,1986]" pageId="19" pageNumber="262">
|
||
Posterior margin of abdominal tergum I is smooth, without denticles; posterior margins of terga II–X have heavily sclerotized, brown, conic, sharply pointed, spine-like denticles separated by spaces wider than denticle width (
|
||
<figureCitation id="13362A5DFFFBFFD7800055FB6EF2FACA" box="[1330,1423,1276,1302]" captionStart="FIGURES 98–105" captionStartId="20.[151,264,1857,1881]" captionTargetBox="[151,1435,201,1834]" captionTargetId="figure-18@20.[151,1435,201,1834]" captionTargetPageId="20" captionText="FIGURES 98–105. Procloeon (Securiops) mutadens. 98–99, left and right mandibles; 100, labrum; 101, maxilla; 102, larval cuticle of metanotum with vestige of hind protopteron; 103, abdominal tergum V; 104, abdominal sternum V; 105, caudalii. Abbreviation: sc, scales and scale sockets." figureDoi="http://doi.org/10.5281/zenodo.8334229" httpUri="https://zenodo.org/record/8334229/files/figure.png" pageId="19" pageNumber="262">Fig. 103</figureCitation>
|
||
); on tergum IX row of denticles is interrupted medially, behind pair of submedian setae (as in
|
||
<figureCitation id="13362A5DFFFBFFD7814C54276FB3FAE7" box="[1150,1230,1312,1339]" captionStart="FIGURES 47–52" captionStartId="8.[151,264,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-18@8.[151,1435,181,1646]" captionTargetPageId="8" captionText="FIGURES 47–52. Procloeon (Securiops) falcatum, larva. 47, articulation of femur and tibia of fore leg, dorsal view; 48, the same, compressed to show junction of patella-tibial suture with row of fine hairs; 49, fragment of tarsus; 50, claw; 51, abdominal tergum X and margin of tergum IX; 52, abdominal tergum X and paraprocts." figureDoi="http://doi.org/10.5281/zenodo.8334199" httpUri="https://zenodo.org/record/8334199/files/figure.png" pageId="19" pageNumber="262">Fig. 51</figureCitation>
|
||
); projected portion of tergum X row with few larger denticles by sides, with few smaller denticles between them (as in
|
||
<figureCitation id="13362A5DFFFBFFD781E054436E2DFA83" box="[1234,1360,1348,1375]" captionStart="FIGURES 47–52" captionStartId="8.[151,264,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-18@8.[151,1435,181,1646]" captionTargetPageId="8" captionText="FIGURES 47–52. Procloeon (Securiops) falcatum, larva. 47, articulation of femur and tibia of fore leg, dorsal view; 48, the same, compressed to show junction of patella-tibial suture with row of fine hairs; 49, fragment of tarsus; 50, claw; 51, abdominal tergum X and margin of tergum IX; 52, abdominal tergum X and paraprocts." figureDoi="http://doi.org/10.5281/zenodo.8334199" httpUri="https://zenodo.org/record/8334199/files/figure.png" pageId="19" pageNumber="262">Figs 51–52</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFFBFFD785F5546F680FFA17" blockId="19.[151,1437,736,1986]" pageId="19" pageNumber="262">
|
||
Posterior margins of abdominal sterna I–
|
||
<collectionCode id="ED1CAE1DFFFBFFD787A5546E69C9FA5F" box="[663,692,1385,1411]" pageId="19" pageNumber="262">IV</collectionCode>
|
||
are smooth, without denticles; posterior margins of sterna
|
||
<collectionCode id="ED1CAE1DFFFBFFD78061546E6E0DFA5F" box="[1363,1392,1385,1411]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="19" pageNumber="262">VI</collectionCode>
|
||
–IX have colorless, conic, sharply pointed, spine-like denticles, denser than denticles on terga (
|
||
<figureCitation id="13362A5DFFFBFFD781B7548B6F9BFA7A" box="[1157,1254,1420,1446]" captionStart="FIGURES 98–105" captionStartId="20.[151,264,1857,1881]" captionTargetBox="[151,1435,201,1834]" captionTargetId="figure-18@20.[151,1435,201,1834]" captionTargetPageId="20" captionText="FIGURES 98–105. Procloeon (Securiops) mutadens. 98–99, left and right mandibles; 100, labrum; 101, maxilla; 102, larval cuticle of metanotum with vestige of hind protopteron; 103, abdominal tergum V; 104, abdominal sternum V; 105, caudalii. Abbreviation: sc, scales and scale sockets." figureDoi="http://doi.org/10.5281/zenodo.8334229" httpUri="https://zenodo.org/record/8334229/files/figure.png" pageId="19" pageNumber="262">Fig. 104</figureCitation>
|
||
). Paraproct with spine-like denticles larger than denticles on sterna (as in
|
||
<figureCitation id="13362A5DFFFBFFD7862054B76818FA17" box="[786,869,1456,1483]" captionStart="FIGURES 47–52" captionStartId="8.[151,264,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-18@8.[151,1435,181,1646]" captionTargetPageId="8" captionText="FIGURES 47–52. Procloeon (Securiops) falcatum, larva. 47, articulation of femur and tibia of fore leg, dorsal view; 48, the same, compressed to show junction of patella-tibial suture with row of fine hairs; 49, fragment of tarsus; 50, claw; 51, abdominal tergum X and margin of tergum IX; 52, abdominal tergum X and paraprocts." figureDoi="http://doi.org/10.5281/zenodo.8334199" httpUri="https://zenodo.org/record/8334199/files/figure.png" pageId="19" pageNumber="262">Fig. 52</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFFBFFD785F554D36F06F9EB" blockId="19.[151,1437,736,1986]" pageId="19" pageNumber="262">
|
||
Abdominal terga with sparse, small protuberances; terga III–X also with few, sparsely situated, oval scales in wide semilunar non-opercula-bearing sockets (
|
||
<figureCitation id="13362A5DFFFBFFD7879554FF6877F9CE" box="[679,778,1528,1554]" captionStart="FIGURES 98–105" captionStartId="20.[151,264,1857,1881]" captionTargetBox="[151,1435,201,1834]" captionTargetId="figure-18@20.[151,1435,201,1834]" captionTargetPageId="20" captionText="FIGURES 98–105. Procloeon (Securiops) mutadens. 98–99, left and right mandibles; 100, labrum; 101, maxilla; 102, larval cuticle of metanotum with vestige of hind protopteron; 103, abdominal tergum V; 104, abdominal sternum V; 105, caudalii. Abbreviation: sc, scales and scale sockets." figureDoi="http://doi.org/10.5281/zenodo.8334229" httpUri="https://zenodo.org/record/8334229/files/figure.png" pageId="19" pageNumber="262">Fig. 103</figureCitation>
|
||
). Sterna
|
||
<collectionCode id="ED1CAE1DFFFBFFD7865F54FE68F7F9CF" box="[877,906,1529,1555]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="19" pageNumber="262">VI</collectionCode>
|
||
–VIII also with such scales (
|
||
<figureCitation id="13362A5DFFFBFFD781FC54FF6E4CF9CE" box="[1230,1329,1528,1554]" captionStart="FIGURES 98–105" captionStartId="20.[151,264,1857,1881]" captionTargetBox="[151,1435,201,1834]" captionTargetId="figure-18@20.[151,1435,201,1834]" captionTargetPageId="20" captionText="FIGURES 98–105. Procloeon (Securiops) mutadens. 98–99, left and right mandibles; 100, labrum; 101, maxilla; 102, larval cuticle of metanotum with vestige of hind protopteron; 103, abdominal tergum V; 104, abdominal sternum V; 105, caudalii. Abbreviation: sc, scales and scale sockets." figureDoi="http://doi.org/10.5281/zenodo.8334229" httpUri="https://zenodo.org/record/8334229/files/figure.png" pageId="19" pageNumber="262">Fig. 104</figureCitation>
|
||
). Besides this, abdominal terga and sterna with ring-like sensilla and irregularly situated fine setae.
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFFBFFD785F557386E60F91A" blockId="19.[151,1437,736,1986]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F557386A5AF986" bold="true" box="[199,295,1599,1626]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F557386A5DF986" bold="true" box="[199,288,1599,1626]" italics="true" pageId="19" pageNumber="262">Tergalii</emphasis>
|
||
.
|
||
</emphasis>
|
||
Tergalii I–
|
||
<collectionCode id="ED1CAE1DFFFBFFD7849257466AC0F987" box="[416,445,1601,1627]" pageId="19" pageNumber="262">IV</collectionCode>
|
||
with dorsal lamella, tergalii
|
||
<collectionCode id="ED1CAE1DFFFBFFD787C857466873F987" box="[762,782,1601,1627]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="19" pageNumber="262" type="Museum">V</collectionCode>
|
||
–
|
||
<collectionCode id="ED1CAE1DFFFBFFD7862E5746683EF987" box="[796,835,1601,1627]" pageId="19" pageNumber="262">VII</collectionCode>
|
||
without dorsal lamella; tergalius
|
||
<collectionCode id="ED1CAE1DFFFBFFD7818757466FA1F987" box="[1205,1244,1601,1627]" pageId="19" pageNumber="262">VII</collectionCode>
|
||
enlarged (
|
||
<bibRefCitation id="EF9C4B29FFFBFFD7806257476BAEF9A2" author="Gillies, M. T." pageId="19" pageNumber="262" pagination="49 - 59" refId="ref12030" refString="Gillies, M. T. (1988) Descriptions of the nymphs of some Afrotropical Baetidae (Ephemeroptera). I. Cloeon Leach and Rhithrocloeon Gillies. Aquatic Insects, 10 (1), 49 - 59. https: // doi. org / 10.1080 / 01650428809361310" type="journal article" year="1988">Gillies 1988</bibRefCitation>
|
||
: figs 14–19). Tergalius
|
||
<collectionCode id="ED1CAE1DFFFBFFD784EF57626979F9A3" box="[477,516,1637,1663]" pageId="19" pageNumber="262">VII</collectionCode>
|
||
was figured with its anal margin directed up, like costal margins of other tergalii (
|
||
<bibRefCitation id="EF9C4B29FFFBFFD785AD578F6A53F97E" author="Gillies, M. T." box="[159,302,1672,1698]" pageId="19" pageNumber="262" pagination="49 - 59" refId="ref12030" refString="Gillies, M. T. (1988) Descriptions of the nymphs of some Afrotropical Baetidae (Ephemeroptera). I. Cloeon Leach and Rhithrocloeon Gillies. Aquatic Insects, 10 (1), 49 - 59. https: // doi. org / 10.1080 / 01650428809361310" type="journal article" year="1988">Gillies 1988</bibRefCitation>
|
||
: fig. 19); probably, the author confused the anal margin with the costal one. Actually, tergalius
|
||
<collectionCode id="ED1CAE1DFFFBFFD78078578E6E0CF97F" box="[1354,1393,1673,1699]" pageId="19" pageNumber="262">VII</collectionCode>
|
||
has costal margin sharply convex, and anal margin straight, that is typical for
|
||
<taxonomicName id="4C0D4D5BFFFBFFD786E257AB6F39F91A" authorityName="sensu Kluge" authorityYear="2016" box="[976,1092,1708,1734]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="262" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B979EACAFFFBFFD786E257AB6F39F91A" box="[976,1092,1708,1734]" italics="true" pageId="19" pageNumber="262">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24A57B1FFFBFFD7817957AA6F0EF91A" box="[1099,1139,1708,1735]" pageId="19" pageNumber="262" sensu="lato">s. l.</taxonomicNameLabel>
|
||
(as in
|
||
<figureCitation id="13362A5DFFFBFFD7818C57AB6E6DF91A" box="[1214,1296,1708,1734]" captionStart="FIGURES 1–24" captionStartId="2.[151,264,1672,1696]" captionTargetBox="[155,1427,188,1633]" captionTargetId="figure-18@2.[151,1436,179,1646]" captionTargetPageId="2" captionText="FIGURES 1–24. Right tergalii of Securiops with dorsal lamella turned out. 1–7, Procloeon (Securiops) falcatum, tergalii I–VII; 8–17, two individuals of P. (S.) macafertiorum: 8–11, tergalii I–II and V–VII; 12–16, tergalii I–V; 17, the same individual, margins of abdominal segments VII–IX with tergalius VII in natural position; 18–24, P. (S.) primasia, tergalii I–VII." figureDoi="http://doi.org/10.5281/zenodo.8334183" httpUri="https://zenodo.org/record/8334183/files/figure.png" pageId="19" pageNumber="262">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFFBFFD785F557C869D7F8A6" blockId="19.[151,1437,736,1986]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F557C86A4FF936" bold="true" box="[199,306,1743,1770]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F557C86A56F936" bold="true" box="[199,299,1743,1770]" italics="true" pageId="19" pageNumber="262">Caudalii</emphasis>
|
||
.
|
||
</emphasis>
|
||
On most part of cercus, lateral side of each segment with one long spine on posterior margin; these spines are gradually changed from stout and moderately long (about as long as segment lengths) in proximal part of cercus to delicate and very long (up to 3 times exceeding segment length) in middle and distal part of cercus (
|
||
<figureCitation id="13362A5DFFFBFFD7805D561F6BB8F88B" captionStart="FIGURES 98–105" captionStartId="20.[151,264,1857,1881]" captionTargetBox="[151,1435,201,1834]" captionTargetId="figure-18@20.[151,1435,201,1834]" captionTargetPageId="20" captionText="FIGURES 98–105. Procloeon (Securiops) mutadens. 98–99, left and right mandibles; 100, labrum; 101, maxilla; 102, larval cuticle of metanotum with vestige of hind protopteron; 103, abdominal tergum V; 104, abdominal sternum V; 105, caudalii. Abbreviation: sc, scales and scale sockets." figureDoi="http://doi.org/10.5281/zenodo.8334229" httpUri="https://zenodo.org/record/8334229/files/figure.png" pageId="19" pageNumber="262">Fig. 105</figureCitation>
|
||
). Posterior margin of each 4th segment of cercus and paracercus, followed by bark brown ring, with several pointed spines both on dorsal and ventral sides.
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFFBFFD785F556836A8AF842" blockId="19.[151,1437,736,1986]" box="[199,503,1924,1950]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F556836A07F842" bold="true" box="[199,378,1924,1950]" pageId="19" pageNumber="262">Winged stages.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8BB236D8FFFBFFD785F556AF6A07F81E" blockId="19.[151,1437,736,1986]" box="[199,378,1960,1986]" pageId="19" pageNumber="262">
|
||
<emphasis id="B979EACAFFFBFFD785F556AF6B80F81E" bold="true" box="[199,253,1960,1986]" pageId="19" pageNumber="262">Egg.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |