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<mods:title id="41B7E721D056963A1474C620BFAEDEE7">The ammonoids from the Gattendorfia Limestone of Oberrödinghausen (Early Carboniferous; Rhenish Mountains, Germany)</mods:title>
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<mods:date id="EC4AC916E58B962BBE5ED8D8597F5C35">2023</mods:date>
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<paragraph id="8BFCED02CA558513FD98FE99FC6C854E" blockId="92.[598,990,265,291]" box="[598,990,265,291]" pageId="92" pageNumber="93">
<heading id="D0B45A6ECA558513FD98FE99FC6C854E" box="[598,990,265,291]" centered="true" fontSize="11" level="2" pageId="92" pageNumber="93" reason="2">
Genus
<taxonomicName id="4C439681CA558513FD66FE99FC6C854E" ID-CoL="63XZ" authority="Korn, 1993" authorityName="Korn" authorityYear="1993" box="[680,990,265,291]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FD66FE99FCE2854E" bold="true" box="[680,848,265,291]" italics="true" pageId="92" pageNumber="93">Nicimitoceras</emphasis>
<bibRefCitation id="EFD290F3CA558513FC96FE99FC6C854E" author="Korn D." box="[856,990,265,291]" pageId="92" pageNumber="93" pagination="581 - 593" refId="ref114371" refString="Korn D. 1993. The ammonoid faunal change near the Devonian - Carboniferous boundary. Annales de la Societe geologique de Belgique 115: 581 - 593." type="journal article" year="1993">Korn, 1993</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C359BE89CA558513FF73FEDDFEEA850B" box="[189,344,332,359]" pageId="92" pageNumber="93" type="type_taxon">
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<heading id="D0B45A6ECA558513FF73FEDDFEEA850B" bold="true" box="[189,344,332,359]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="B9373110CA558513FF73FEDDFEEA850B" bold="true" box="[189,344,332,359]" pageId="92" pageNumber="93">
<typeStatus id="54F853A0CA558513FF73FEDDFF48850A" box="[189,250,333,359]" pageId="92" pageNumber="93">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CA558513FF73FEEEFC1085FA" box="[189,930,381,407]" pageId="92" pageNumber="93" type="reference_group">
<paragraph id="8BFCED02CA558513FF73FEEEFC1085FA" blockId="92.[189,930,381,407]" box="[189,930,381,407]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="AB53CA2CCA558513FF73FEEEFC1085FA" box="[189,930,381,407]" pageId="92" pageNumber="93">
<treatmentCitation id="0AE2CB13CA558513FF73FEEEFD1685FA" author="Vohringer E." box="[189,676,381,407]" page="120" pageId="92" pageNumber="93" year="1960">
<taxonomicName id="4C439681CA558513FF73FEEEFD1685FA" authority="Vohringer, 1960: 120" authorityName="Vohringer" authorityPageNumber="120" authorityYear="1960" box="[189,676,381,407]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="subacre">
<emphasis id="B9373110CA558513FF73FEEEFE2C85FA" box="[189,414,381,407]" italics="true" pageId="92" pageNumber="93">Imitoceras subacre</emphasis>
<bibRefCitation id="EFD290F3CA558513FE6BFEEDFD1685FA" author="Vohringer E." box="[421,676,381,407]" pageId="92" pageNumber="93" pagination="107 - 196" refId="ref118120" refString="Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196." type="journal article" year="1960">Vöhringer, 1960: 120</bibRefCitation>
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; original designation.
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</paragraph>
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<subSubSection id="C359BE89CA558513FF73FE50FF4D87CF" pageId="92" pageNumber="93" type="diagnosis">
<paragraph id="8BFCED02CA558513FF73FE50FE3A85B7" blockId="92.[189,392,448,474]" box="[189,392,448,474]" pageId="92" pageNumber="93">
<heading id="D0B45A6ECA558513FF73FE50FE3A85B7" bold="true" box="[189,392,448,474]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="B9373110CA558513FF73FE50FE3A85B7" bold="true" box="[189,392,448,474]" pageId="92" pageNumber="93">Genus diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA558513FF73FE61FDBF8616" blockId="92.[188,1399,497,635]" pageId="92" pageNumber="93">
Genus of the
<taxonomicName id="4C439681CA558513FEA6FE61FDFB8666" authorityName="Korn" authorityYear="1994" box="[360,585,497,523]" class="Cephalopoda" family="Prionoceratidae" kingdom="Animalia" order="Goniatitina" pageId="92" pageNumber="116" phylum="Mollusca" rank="subFamily" subFamily="Acutimitoceratinae">Acutimitoceratinae</taxonomicName>
with a discoidal conch with moderately high to high coiling rate (WER = 1.902.20); inner whorls subinvolute to evolute to variable degree. Ornament with convex or slightly biconvex growth lines. Suture line with short, V-shaped or lanceolate external lobe (0.50 to 0.75 depth of the adventive lobe).
</paragraph>
<paragraph id="8BFCED02CA558513FF73FD34FE1E86D3" blockId="92.[189,428,676,702]" box="[189,428,676,702]" pageId="92" pageNumber="93">
<heading id="D0B45A6ECA558513FF73FD34FE1E86D3" bold="true" box="[189,428,676,702]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="B9373110CA558513FF73FD34FE1E86D3" bold="true" box="[189,428,676,702]" pageId="92" pageNumber="93">Genus composition</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA558513FF73FD45FA858757" blockId="92.[189,1399,725,826]" pageId="92" pageNumber="93">
Central Europe (
<bibRefCitation id="EFD290F3CA558513FE45FD45FD888682" author="Schmidt H." box="[395,570,725,751]" pageId="92" pageNumber="93" pagination="98 - 171" refId="ref117457" refString="Schmidt H. 1924. Zwei Cephalopodenfaunen an der Devon-Carbongrenze im Sauerland. Jahrbuch der Preussischen Geologischen Landesanstalt 44: 98 - 171." type="journal article" year="1924">Schmidt 1924</bibRefCitation>
;
<bibRefCitation id="EFD290F3CA558513FD82FD45FCA28682" author="Vohringer E." box="[588,784,725,751]" pageId="92" pageNumber="93" pagination="107 - 196" refId="ref118120" refString="Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196." type="journal article" year="1960">Vöhringer 1960</bibRefCitation>
;
<bibRefCitation id="EFD290F3CA558513FCECFD46FC1A8682" author="Korn D." box="[802,936,725,752]" pageId="92" pageNumber="93" pagination="71 - 89" refId="ref113987" refString="Korn D. 1984. Die Goniatiten der Stockumer Imitoceras - Kalklinsen (Ammonoidea; Devon / Karbon-Grenze). Courier Forschungsinstitut Senckenberg 67: 71 - 89. https: // doi. org / 10.1127 / njgpm / 1984 / 1984 / 66" type="journal article" year="1984">Korn 1984</bibRefCitation>
):
<taxonomicName id="4C439681CA558513FC08FD45FAC08682" authority="Schmidt, 1924" authorityName="Schmidt" authorityYear="1924" box="[966,1394,725,751]" class="Cephalopoda" family="Prionoceratidae" genus="Aganides" kingdom="Animalia" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="carinatus">
<emphasis id="B9373110CA558513FC08FD45FB028682" box="[966,1200,725,751]" italics="true" pageId="92" pageNumber="93">Aganides carinatus</emphasis>
<bibRefCitation id="EFD290F3CA558513FB73FD45FAC08682" author="Schmidt H." box="[1213,1394,725,751]" pageId="92" pageNumber="93" pagination="98 - 171" refId="ref117457" refString="Schmidt H. 1924. Zwei Cephalopodenfaunen an der Devon-Carbongrenze im Sauerland. Jahrbuch der Preussischen Geologischen Landesanstalt 44: 98 - 171." type="journal article" year="1924">Schmidt, 1924</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C439681CA558513FF73FD6BFD0A8779" authority="Vohringer, 1960" authorityName="Vohringer" authorityYear="1960" box="[189,696,762,788]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="trochiforme">
<emphasis id="B9373110CA558513FF73FD6BFE648779" box="[189,470,762,788]" italics="true" pageId="92" pageNumber="93">Imitoceras trochiforme</emphasis>
<bibRefCitation id="EFD290F3CA558513FE27FD6AFD0A8779" author="Vohringer E." box="[489,696,762,788]" pageId="92" pageNumber="93" pagination="107 - 196" refId="ref118120" refString="Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196." type="journal article" year="1960">Vöhringer, 1960</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C439681CA558513FD01FD6BFB2E8779" authority="Vohringer, 1960" authorityName="Vohringer" authorityYear="1960" box="[719,1180,762,788]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="subacre">
<emphasis id="B9373110CA558513FD01FD6BFC098779" box="[719,955,762,788]" italics="true" pageId="92" pageNumber="93">Imitoceras subacre</emphasis>
<bibRefCitation id="EFD290F3CA558513FC03FD6AFB2E8779" author="Vohringer E." box="[973,1180,762,788]" pageId="92" pageNumber="93" pagination="107 - 196" refId="ref118120" refString="Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196." type="journal article" year="1960">Vöhringer, 1960</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C439681CA558513FB7AFD6BFE338754" authority="Vohringer, 1960" authorityName="Vohringer" authorityYear="1960" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="acre">
<emphasis id="B9373110CA558513FB7AFD6BFAC48779" box="[1204,1398,763,788]" italics="true" pageId="92" pageNumber="93">Imitoceras acre</emphasis>
<bibRefCitation id="EFD290F3CA558513FF73FC8FFE338754" author="Vohringer E." box="[189,385,799,825]" pageId="92" pageNumber="93" pagination="107 - 196" refId="ref118120" refString="Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196." type="journal article" year="1960">Vöhringer, 1960</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C439681CA558513FE43FCB0FC358754" authority="Vohringer, 1960" authorityName="Vohringer" authorityYear="1960" box="[397,903,799,825]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="heterolobatum">
<emphasis id="B9373110CA558513FE43FCB0FD0E8754" box="[397,700,799,825]" italics="true" pageId="92" pageNumber="93">Imitoceras heterolobatum</emphasis>
<bibRefCitation id="EFD290F3CA558513FD0DFC8FFC358754" author="Vohringer E." box="[707,903,799,825]" pageId="92" pageNumber="93" pagination="107 - 196" refId="ref118120" refString="Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196." type="journal article" year="1960">Vöhringer, 1960</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C439681CA558513FC5DFCB0FA818757" authority="Korn, 1984" authorityName="Korn" authorityYear="1984" box="[915,1331,800,826]" class="Cephalopoda" family="Gattendorfiidae" genus="Acutimitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="caesari">
<emphasis id="B9373110CA558513FC5DFCB0FB108754" box="[915,1186,800,826]" italics="true" pageId="92" pageNumber="93">Acutimitoceras caesari</emphasis>
<bibRefCitation id="EFD290F3CA558513FB64FCB0FA818757" author="Korn D." box="[1194,1331,800,826]" pageId="92" pageNumber="93" pagination="71 - 89" refId="ref113987" refString="Korn D. 1984. Die Goniatiten der Stockumer Imitoceras - Kalklinsen (Ammonoidea; Devon / Karbon-Grenze). Courier Forschungsinstitut Senckenberg 67: 71 - 89. https: // doi. org / 10.1127 / njgpm / 1984 / 1984 / 66" type="journal article" year="1984">Korn, 1984</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFCED02CA558513FF73FCF3FF4D87CF" blockId="92.[188,1398,867,930]" pageId="92" pageNumber="93">
North Africa (
<bibRefCitation id="EFD290F3CA558513FE93FCF3FD598710" author="Bockwinkel J. &amp; Ebbighausen V." box="[349,747,867,893]" pageId="92" pageNumber="93" pagination="87 - 129" refId="ref112039" refString="Bockwinkel J. &amp; Ebbighausen V. 2006. A new ammonoid fauna from the Gattendorfia - Eocanites Genozone of the Anti-Atlas (Early Carboniferous; Morocco). Fossil Record 9: 87 - 129. https: // doi. org / 10.5194 / fr- 9 - 87 - 2006" type="journal article" year="2006">Bockwinkel &amp; Ebbighausen 2006</bibRefCitation>
):
<taxonomicName id="4C439681CA558513FCCAFCF3FF4887CF" authority="Bockwinkel &amp; Ebbighausen, 2006" authorityName="Bockwinkel &amp; Ebbighausen" authorityYear="2006" class="Cephalopoda" family="Gattendorfiidae" genus="Acutimitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="mfisense">
<emphasis id="B9373110CA558513FCCAFCF3FBAD8710" box="[772,1055,867,893]" italics="true" pageId="92" pageNumber="93">Acutimitoceras mfisense</emphasis>
<bibRefCitation id="EFD290F3CA558513FBE8FCF3FF4887CF" author="Bockwinkel J. &amp; Ebbighausen V." pageId="92" pageNumber="93" pagination="87 - 129" refId="ref112039" refString="Bockwinkel J. &amp; Ebbighausen V. 2006. A new ammonoid fauna from the Gattendorfia - Eocanites Genozone of the Anti-Atlas (Early Carboniferous; Morocco). Fossil Record 9: 87 - 129. https: // doi. org / 10.5194 / fr- 9 - 87 - 2006" type="journal article" year="2006">Bockwinkel &amp; Ebbighausen, 2006</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C359BE89CA558512FF73FC5CFE64802F" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93" type="discussion">
<paragraph id="8BFCED02CA558513FF73FC5CFE9D878B" blockId="92.[189,303,972,998]" box="[189,303,972,998]" pageId="92" pageNumber="93">
<heading id="D0B45A6ECA558513FF73FC5CFE9D878B" bold="true" box="[189,303,972,998]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="B9373110CA558513FF73FC5CFE9D878B" bold="true" box="[189,303,972,998]" pageId="92" pageNumber="93">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFCED02CA558513FF73FC6CFE1180EA" blockId="92.[188,1399,1020,1159]" pageId="92" pageNumber="93">
<bibRefCitation id="EFD290F3CA558513FF73FC6CFE39807B" author="Vohringer E." box="[189,395,1020,1046]" pageId="92" pageNumber="93" pagination="107 - 196" refId="ref118120" refString="Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196." type="journal article" year="1960">Vöhringer (1960)</bibRefCitation>
placed his three new species “
<emphasis id="B9373110CA558513FD39FC6DFBBE807B" box="[759,1036,1020,1046]" italics="true" pageId="92" pageNumber="93">
<taxonomicName id="4C439681CA558513FD39FC6DFCCA807B" box="[759,888,1021,1046]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">Imitoceras</taxonomicName>
.
<taxonomicName id="4C439681CA558513FC4DFC6CFBBE807B" authorityName="Vohringer" authorityYear="1960" box="[899,1036,1020,1046]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="trochiforme">trochiforme</taxonomicName>
</emphasis>
”, “
<taxonomicName id="4C439681CA558513FBF8FC6DFB19807B" authorityName="Vohringer" authorityYear="1960" box="[1078,1195,1020,1046]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="subacre">
<emphasis id="B9373110CA558513FBF8FC6DFB19807B" box="[1078,1195,1020,1046]" italics="true" pageId="92" pageNumber="93">I. subacre</emphasis>
</taxonomicName>
” and “
<taxonomicName id="4C439681CA558513FB31FC6DFAF9807B" authorityName="Vohringer" authorityYear="1960" box="[1279,1355,1021,1046]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="acre">
<emphasis id="B9373110CA558513FB31FC6DFAF9807B" box="[1279,1355,1021,1046]" italics="true" pageId="92" pageNumber="93">I. acre</emphasis>
</taxonomicName>
” in the species group of “
<taxonomicName id="4C439681CA558513FE76FBB2FD3C8051" box="[440,654,1058,1084]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="lineare">
<emphasis id="B9373110CA558513FE76FBB2FD3C8051" box="[440,654,1058,1084]" italics="true" pageId="92" pageNumber="93">Imitoceras lineare</emphasis>
</taxonomicName>
”, which was putatively characterised by a conch that is involute in all growth stages and by the absence of shell constrictions. This concept can no longer be followed for several reasons:
</paragraph>
<paragraph id="8BFCED02CA558513FF73FB20FC6081E9" blockId="92.[189,1400,1200,1413]" pageId="92" pageNumber="93">
(1) “
<taxonomicName id="4C439681CA558513FF39FB20FE7F80A7" box="[247,461,1200,1226]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="lineare">
<emphasis id="B9373110CA558513FF39FB20FE7F80A7" box="[247,461,1200,1226]" italics="true" pageId="92" pageNumber="93">Imitoceras lineare</emphasis>
</taxonomicName>
” from the Late Devonian is a species of the genus
<taxonomicName id="4C439681CA558513FBD2FB20FB7C80A4" authorityName="Korn" authorityYear="1988" box="[1052,1230,1200,1225]" class="Cephalopoda" family="Prionoceratidae" genus="Mimimitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FBD2FB20FB7C80A4" box="[1052,1230,1200,1225]" italics="true" pageId="92" pageNumber="93">Mimimitoceras</emphasis>
</taxonomicName>
and possesses shell constrictions in the juvenile and middle growth stage (Korn 1994). The species has hardly any features in common with the three Early Carboniferous species and therefore cannot be considered closely related.
<taxonomicName id="4C439681CA558513FE66FAB0FD068154" baseAuthorityName="Munster" baseAuthorityYear="1839" box="[424,692,1311,1337]" class="Cephalopoda" family="Prionoceratidae" genus="Mimimitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="lineare">
<emphasis id="B9373110CA558513FE66FAB0FD068154" box="[424,692,1311,1337]" italics="true" pageId="92" pageNumber="93">Mimimitoceras lineare</emphasis>
</taxonomicName>
differs markedly in the low coiling rate (WER = 1.50) and the deep external lobe, which has the same depth when compared with the adventive lobe. Species of
<taxonomicName id="4C439681CA558513FEC4FAFBFE1D81E9" authorityName="Korn" authorityYear="1993" box="[266,431,1387,1412]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FEC4FAFBFE1D81E9" box="[266,431,1387,1412]" italics="true" pageId="92" pageNumber="93">Nicimitoceras</emphasis>
</taxonomicName>
have a very characteristic, short external lobe.
</paragraph>
<paragraph id="8BFCED02CA558513FF73FA3EFD938231" blockId="92.[189,1399,1453,1629]" pageId="92" pageNumber="93">
(2) Vöhrigers assumption that the three species have a conch that is involute at all growth stages is apparently based on the cross section of specimen GPIT-PV-638645 (
<taxonomicName id="4C439681CA558513FBDCFA44FB0E8180" authorityName="Vohringer" authorityYear="1960" box="[1042,1212,1491,1517]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="trochiforme">
<emphasis id="B9373110CA558513FBDCFA44FB0E8180" box="[1042,1212,1491,1517]" italics="true" pageId="92" pageNumber="93">N. trochiforme</emphasis>
</taxonomicName>
). However, this cross section is off-centre and does not allow the study of the innermost whorls; there is no evidence for the assumption of involute whorls. In fact, other preparations show that the inner whorls are subinvolute or subevolute.
</paragraph>
<paragraph id="8BFCED02CA558513FF73F916FBBD82CC" blockId="92.[189,1039,1670,1697]" box="[189,1039,1670,1697]" pageId="92" pageNumber="93">
The following genera must be discussed in the context of
<taxonomicName id="4C439681CA558513FCADF917FBBA82CD" authorityName="Korn" authorityYear="1993" box="[867,1032,1671,1696]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FCADF917FBBA82CD" box="[867,1032,1671,1696]" italics="true" pageId="92" pageNumber="93">Nicimitoceras</emphasis>
</taxonomicName>
:
</paragraph>
<paragraph id="8BFCED02CA558513FF73F959FB168384" blockId="92.[189,1399,1737,2025]" pageId="92" pageNumber="93">
<taxonomicName id="4C439681CA558513FF73F959FED8828E" authorityName="Ruan" authorityYear="1995" box="[189,362,1737,1763]" class="Cephalopoda" genus="Follimitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FF73F959FED8828E" box="[189,362,1737,1763]" italics="true" pageId="92" pageNumber="93">Follimitoceras</emphasis>
</taxonomicName>
, introduced by
<bibRefCitation id="EFD290F3CA558513FDFFF95AFD788289" author="Ruan Y." box="[561,714,1737,1764]" pageId="92" pageNumber="93" pagination="407 - 430" refId="ref117020" refString="Ruan Y. 1995. Tournaisian Ammonoids of Northern Xinjiang, China. Palaeontologia Cathayana 6: 407 - 430." type="journal article" year="1995">Ruan (1995)</bibRefCitation>
as a subgenus of
<taxonomicName id="4C439681CA558513FC7AF95AFBD48289" baseAuthorityName="Kullmann" baseAuthorityYear="2009" box="[948,1126,1738,1764]" class="Cephalopoda" family="Gattendorfiidae" genus="Acutimitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FC7AF95AFBD48289" box="[948,1126,1738,1764]" italics="true" pageId="92" pageNumber="93">Acutimitoceras</emphasis>
</taxonomicName>
with the
<typeStatus id="54F853A0CA558513FB2DF95AFAA48289" box="[1251,1302,1738,1764]" pageId="92" pageNumber="93">type</typeStatus>
species
<taxonomicName id="4C439681CA558513FF73F960FCBE8364" authority="Ruan, 1981" authorityName="Ruan" authorityYear="1981" box="[189,780,1775,1801]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="folliforme" subGenus="Imitoceras">
<emphasis id="B9373110CA558513FF73F960FE888364" box="[189,314,1776,1801]" italics="true" pageId="92" pageNumber="93">Imitoceras</emphasis>
(
<emphasis id="B9373110CA558513FE99F960FE668364" box="[343,468,1776,1801]" italics="true" pageId="92" pageNumber="93">Imitoceras</emphasis>
)
<emphasis id="B9373110CA558513FE3EF97FFDD68364" box="[496,612,1775,1801]" italics="true" pageId="92" pageNumber="93">folliforme</emphasis>
<bibRefCitation id="EFD290F3CA558513FDB8F97FFCBE8364" author="Ruan Y." box="[630,780,1775,1801]" pageId="92" pageNumber="93" pagination="1 - 152" refId="ref116988" refString="Ruan Y. 1981. Devonian and earliest Carboniferous Ammonoids from Guangxi and Guizhou. Memoires of Nanjing Institute of Geology and Paleontology, Academia Sinica 15: 1 - 152." type="journal article" year="1981">Ruan, 1981</bibRefCitation>
</taxonomicName>
. The subgenus was established for discoidal prionoceratid ammonoids with a deep, slightly pouched external lobe. The conch characters are very similar to
<taxonomicName id="4C439681CA558513FEF2F8AAFE53833E" authorityName="Korn" authorityYear="1993" box="[316,481,1850,1875]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FEF2F8AAFE53833E" box="[316,481,1850,1875]" italics="true" pageId="92" pageNumber="93">Nicimitoceras</emphasis>
</taxonomicName>
, with which it shares the lenticular conch shape and the high coiling rate. However,
<taxonomicName id="4C439681CA558513FEF9F8CEFE568315" authorityName="Ruan" authorityYear="1995" box="[311,484,1886,1912]" class="Cephalopoda" genus="Follimitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FEF9F8CEFE568315" box="[311,484,1886,1912]" italics="true" pageId="92" pageNumber="93">Follimitoceras</emphasis>
</taxonomicName>
differs in the deeper, pouched external lobe.
<taxonomicName id="4C439681CA558513FBCCF8CEFB1D8315" authorityName="Ruan" authorityYear="1995" box="[1026,1199,1886,1912]" class="Cephalopoda" genus="Follimitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FBCCF8CEFB1D8315" box="[1026,1199,1886,1912]" italics="true" pageId="92" pageNumber="93">Follimitoceras</emphasis>
</taxonomicName>
should therefore be regarded as a valid, independent genus.
<bibRefCitation id="EFD290F3CA558513FD0FF814FC3F83F3" author="Kullmann J." box="[705,909,1924,1950]" pageId="92" pageNumber="93" pagination="2 - 10" refId="ref115865" refString="Kullmann J. 2009. Prionoceratoidea, In: Seldon P. A. (Ed.), Treatise on Invertebrate Paleontology. Part L. Mollusca 4 Revised. Volume 2: Carboniferous and Permian Ammonoidea (Goniatitida and Prolecanitida): 2 - 10. The University of Kansas Paleontological Institute, Lawrence, Kansas." type="book chapter" year="2009">Kullmann (2009)</bibRefCitation>
regarded
<taxonomicName id="4C439681CA558513FBCBF814FB0283F3" authorityName="Ruan" authorityYear="1995" box="[1029,1200,1924,1950]" class="Cephalopoda" genus="Follimitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FBCBF814FB0283F3" box="[1029,1200,1924,1950]" italics="true" pageId="92" pageNumber="93">Follimitoceras</emphasis>
</taxonomicName>
, with a question mark, as a synonym of
<emphasis id="B9373110CA558513FE2EF839FDCE83AE" box="[480,636,1961,1987]" italics="true" pageId="92" pageNumber="93">
<taxonomicName id="4C439681CA558513FE2EF839FDCA83AE" authorityName="Weyer" authorityYear="1972" box="[480,632,1961,1987]" class="Cephalopoda" family="Gattendorfiidae" genus="Zadelsdorfia" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">Zadelsdorfia</taxonomicName>
.
</emphasis>
Such an attribution as proposed by Kullmann is very unlikely because of the conch shape and the open umbilicus in the adult stage in
<taxonomicName id="4C439681CA558513FBC7F85EFB2F8385" authorityName="Weyer" authorityYear="1972" box="[1033,1181,1998,2024]" class="Cephalopoda" family="Gattendorfiidae" genus="Zadelsdorfia" kingdom="Animalia" order="Goniatitida" pageId="92" pageNumber="93" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA558513FBC7F85EFB2F8385" box="[1033,1181,1998,2024]" italics="true" pageId="92" pageNumber="93">Zadelsdorfia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFCED02CA548512FF73FE99FEDB85DC" blockId="93.[189,1399,265,433]" pageId="93" pageNumber="94">
<taxonomicName id="4C439681CA548512FF73FE99FED1854E" authorityName="Ruan" authorityYear="1995" box="[189,355,265,291]" class="Cephalopoda" genus="Acrimitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FF73FE99FED1854E" box="[189,355,265,291]" italics="true" pageId="93" pageNumber="94">Acrimitoceras</emphasis>
</taxonomicName>
, also introduced by
<bibRefCitation id="EFD290F3CA548512FD97FE9AFD5C8549" author="Ruan Y." box="[601,750,265,292]" pageId="93" pageNumber="94" pagination="407 - 430" refId="ref117020" refString="Ruan Y. 1995. Tournaisian Ammonoids of Northern Xinjiang, China. Palaeontologia Cathayana 6: 407 - 430." type="journal article" year="1995">Ruan (1995)</bibRefCitation>
as a subgenus of
<taxonomicName id="4C439681CA548512FC0AFE99FBC4854E" baseAuthorityName="Kullmann" baseAuthorityYear="2009" box="[964,1142,265,291]" class="Cephalopoda" family="Gattendorfiidae" genus="Acutimitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FC0AFE99FBC4854E" box="[964,1142,265,291]" italics="true" pageId="93" pageNumber="94">Acutimitoceras</emphasis>
</taxonomicName>
with the
<typeStatus id="54F853A0CA548512FB29FE9AFAA88549" box="[1255,1306,266,292]" pageId="93" pageNumber="94">type</typeStatus>
species
<taxonomicName id="4C439681CA548512FF70FEBDFDF7852B" authority="Vohringer, 1960" authorityName="Vohringer" authorityYear="1960" box="[190,581,300,326]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="species" species="acre">
<emphasis id="B9373110CA548512FF70FEBDFEC5852B" box="[190,375,301,326]" italics="true" pageId="93" pageNumber="94">Imitoceras acre</emphasis>
<bibRefCitation id="EFD290F3CA548512FEB1FEBCFDF7852B" author="Vohringer E." box="[383,581,300,326]" pageId="93" pageNumber="94" pagination="107 - 196" refId="ref118120" refString="Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196." type="journal article" year="1960">Vöhringer, 1960</bibRefCitation>
</taxonomicName>
. The subgenus separates the acute end-member of an evolutionary lineage, not paying attention to the evolutionary lineage leading to it, i.e., possibly through
<taxonomicName id="4C439681CA548512FB1EFEC1FEAB85E0" baseAuthorityName="Vohringer" baseAuthorityYear="1960" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="species" species="subacre">
<emphasis id="B9373110CA548512FB1EFEC1FEAB85E0" italics="true" pageId="93" pageNumber="94">Nicimitoceras subacre</emphasis>
</taxonomicName>
, the
<typeStatus id="54F853A0CA548512FE9AFEE4FE3585E3" box="[340,391,372,398]" pageId="93" pageNumber="94">type</typeStatus>
species of
<taxonomicName id="4C439681CA548512FDC3FEE4FD0385E0" authorityName="Korn" authorityYear="1993" box="[525,689,372,397]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FDC3FEE4FD0385E0" box="[525,689,372,397]" italics="true" pageId="93" pageNumber="94">Nicimitoceras</emphasis>
</taxonomicName>
.
<taxonomicName id="4C439681CA548512FD0EFEE4FCD485E3" authorityName="Ruan" authorityYear="1995" box="[704,870,372,398]" class="Cephalopoda" genus="Acrimitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FD0EFEE4FCD485E3" box="[704,870,372,398]" italics="true" pageId="93" pageNumber="94">Acrimitoceras</emphasis>
</taxonomicName>
should therefore be regarded as synonym of
<taxonomicName id="4C439681CA548512FF70FE07FED085DD" authorityName="Korn" authorityYear="1993" box="[190,354,407,432]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FF70FE07FED085DD" box="[190,354,407,432]" italics="true" pageId="93" pageNumber="94">Nicimitoceras</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFCED02CA548512FF73FE48FCE38760" blockId="93.[188,1399,472,782]" pageId="93" pageNumber="94">
<emphasis id="B9373110CA548512FF73FE48FEFA859F" box="[189,328,472,498]" italics="true" pageId="93" pageNumber="94">Streeliceras</emphasis>
, introduced by
<bibRefCitation id="EFD290F3CA548512FDDDFE48FD73859F" author="Becker R. T." box="[531,705,472,498]" pageId="93" pageNumber="94" pagination="19 - 45" refId="ref111308" refString="Becker R. T. 1996. New faunal records and holostratigraphic correlation of the Hasselbachtal D / C- Boundary Auxiliary Stratotype (Germany). Annales de la Societe geologique de Belgique 117: 19 - 45." type="journal article" year="1996">Becker (1996)</bibRefCitation>
as a subgenus of
<taxonomicName id="4C439681CA548512FC7EFE49FBD0859E" baseAuthorityName="Kullmann" baseAuthorityYear="2009" box="[944,1122,473,499]" class="Cephalopoda" family="Gattendorfiidae" genus="Acutimitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FC7EFE49FBD0859E" box="[944,1122,473,499]" italics="true" pageId="93" pageNumber="94">Acutimitoceras</emphasis>
</taxonomicName>
with the
<typeStatus id="54F853A0CA548512FB2CFE49FAA7859E" box="[1250,1301,473,499]" pageId="93" pageNumber="94">type</typeStatus>
species
<taxonomicName id="4C439681CA548512FF73FE6CFD738678" authority="Vohringer, 1960" authorityName="Vohringer" authorityYear="1960" box="[189,705,507,534]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="species" species="heterolobatum">
<emphasis id="B9373110CA548512FF73FE6CFE5D867B" box="[189,495,508,534]" italics="true" pageId="93" pageNumber="94">Imitoceras heterolobatum</emphasis>
<bibRefCitation id="EFD290F3CA548512FE34FE6CFD738678" author="Vohringer E." box="[506,705,507,534]" pageId="93" pageNumber="94" pagination="107 - 196" refId="ref118120" refString="Vohringer E. 1960. Die Goniatiten der unterkarbonischen Gattendorfia - Stufe im Honnetal (Sauerland). Fortschritte in der Geologie von Rheinland und Westfalen 3: 107 - 196." type="journal article" year="1960">Vöhringer, 1960</bibRefCitation>
</taxonomicName>
. The subgenus was introduced for species that possess evolute inner whorls in combination with a short external lobe. Separation from
<taxonomicName id="4C439681CA548512FB57FDB0FA8C8654" authorityName="Korn" authorityYear="1993" box="[1177,1342,544,569]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FB57FDB0FA8C8654" box="[1177,1342,544,569]" italics="true" pageId="93" pageNumber="94">Nicimitoceras</emphasis>
</taxonomicName>
was only justified by the higher number of evolute juvenile whorls. However, numerous studies of cross-sections of acutimitoceratin ammonoids have shown that the juvenile umbilical width changes gradually between species of the different groups. In contrast, the short external lobe is a very constant character of the acutimitoceratin ammonoids of the “
<taxonomicName id="4C439681CA548512FD7FFD3CFC3D86AB" authority="Stufe" authorityName="Stufe" box="[689,911,684,710]" class="Cephalopoda" family="Gattendorfiidae" genus="Gattendorfia" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FD7FFD3CFCF586AB" box="[689,839,684,710]" italics="true" pageId="93" pageNumber="94">Gattendorfia</emphasis>
Stufe
</taxonomicName>
” and is always accompanied by lenticular conchs and high coiling rates. Therefore, “
<taxonomicName id="4C439681CA548512FD78FD41FC578687" authorityName="Vohringer" authorityYear="1960" box="[694,997,720,746]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="species" species="heterolobatum">
<emphasis id="B9373110CA548512FD78FD41FC578687" box="[694,997,720,746]" italics="true" pageId="93" pageNumber="94">Imitoceras heterolobatum</emphasis>
</taxonomicName>
” does hardly fulfil the criteria for an own subgenus and is thus kept in
<taxonomicName id="4C439681CA548512FDA5FD64FCA28760" authorityName="Korn" authorityYear="1993" box="[619,784,756,781]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FDA5FD64FCA28760" box="[619,784,756,781]" italics="true" pageId="93" pageNumber="94">Nicimitoceras</emphasis>
</taxonomicName>
here.
</paragraph>
<paragraph id="8BFCED02CA548512FF73FCA6FB20871E" blockId="93.[189,1399,821,883]" pageId="93" pageNumber="94">
<taxonomicName id="4C439681CA548512FF73FCA6FED08722" authorityName="Korn" authorityYear="1993" box="[189,354,822,847]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FF73FCA6FED08722" box="[189,354,822,847]" italics="true" pageId="93" pageNumber="94">Nicimitoceras</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C439681CA548512FDC9FCA5FD218722" baseAuthorityName="Korn &amp; Weyer" baseAuthorityYear="2023" box="[519,659,821,847]" class="Cephalopoda" family="Prionoceratidae" genus="Stockumites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FDC9FCA5FD218722" box="[519,659,821,847]" italics="true" pageId="93" pageNumber="94">Stockumites</emphasis>
</taxonomicName>
by the short external lobe and, in most cases, by the higher coiling rate (WER usually above 2.00, but usually below this value in
<taxonomicName id="4C439681CA548512FC3BFCC8FB33871F" baseAuthorityName="Korn &amp; Weyer" baseAuthorityYear="2023" box="[1013,1153,856,882]" class="Cephalopoda" family="Prionoceratidae" genus="Stockumites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FC3BFCC8FB33871F" box="[1013,1153,856,882]" italics="true" pageId="93" pageNumber="94">Stockumites</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BFCED02CA548512FF73FC0AFE64802F" blockId="93.[189,1399,922,1090]" pageId="93" pageNumber="94">
A genus with similar species is
<taxonomicName id="4C439681CA548512FDFEFC0BFD1F87D9" box="[560,685,923,948]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FDFEFC0BFD1F87D9" box="[560,685,923,948]" italics="true" pageId="93" pageNumber="94">Imitoceras</emphasis>
</taxonomicName>
, but in the species of this genus the external lobe is pouched in contrast to
<taxonomicName id="4C439681CA548512FE93FC2EFDB087BA" authorityName="Korn" authorityYear="1993" box="[349,514,958,983]" class="Cephalopoda" family="Gattendorfiidae" genus="Nicimitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FE93FC2EFDB087BA" box="[349,514,958,983]" italics="true" pageId="93" pageNumber="94">Nicimitoceras</emphasis>
</taxonomicName>
with a lanceolate or V-shaped external lobe. In the stratigraphically earliest members of the genus this feature may still be weakly developed (
<bibRefCitation id="EFD290F3CA548512FC1AFC71FAD58796" author="Bockwinkel J. &amp; Ebbighausen V." box="[980,1383,993,1019]" pageId="93" pageNumber="94" pagination="87 - 129" refId="ref112039" refString="Bockwinkel J. &amp; Ebbighausen V. 2006. A new ammonoid fauna from the Gattendorfia - Eocanites Genozone of the Anti-Atlas (Early Carboniferous; Morocco). Fossil Record 9: 87 - 129. https: // doi. org / 10.5194 / fr- 9 - 87 - 2006" type="journal article" year="2006">Bockwinkel &amp; Ebbighausen 2006</bibRefCitation>
), but the shape of the adventive lobe, the dorsal of which is curved inwards in
<taxonomicName id="4C439681CA548512FBBFFB95FB5C8073" box="[1137,1262,1029,1054]" class="Cephalopoda" family="Prionoceratidae" genus="Imitoceras" kingdom="Animalia" order="Goniatitida" pageId="93" pageNumber="94" phylum="Mollusca" rank="genus">
<emphasis id="B9373110CA548512FBBFFB95FB5C8073" box="[1137,1262,1029,1054]" italics="true" pageId="93" pageNumber="94">Imitoceras</emphasis>
</taxonomicName>
, is another distinguishing criterion.
</paragraph>
</subSubSection>
</treatment>
</document>