1037 lines
130 KiB
XML
1037 lines
130 KiB
XML
<document id="FA0FE4FFCA03526D3781D0E913BA8443" ID-CLB-Dataset="9059" ID-DOI="10.11646/zootaxa.4857.1.4" ID-GBIF-Dataset="c4a1490d-15c3-446f-8676-e6c80096bcb6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4396474" ID-ZooBank="F0AF75AD-BAE2-4B7D-9CCB-3D9477F350BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609197892225" checkinUser="plazi" docAuthor="Lindquist, Evert E., Oconnor, Barry M., Shaw, Matthew D. & Sidorchuk, Ekaterina A." docDate="2020" docId="03E2879C4F48FF9EFF66A92BFE4ED269" docLanguage="en" docName="zootaxa.4857.1.4.pdf" docOrigin="Zootaxa 4857 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Berlesia multisetosa Lindquist & Oconnor & Shaw & Sidorchuk 2020, new species" docType="treatment" docVersion="6" lastPageNumber="26" masterDocId="FFDBFFE44F45FF8BFFF1AB6DFFACD13A" masterDocTitle="Review of the genera Berlesia Canestrini, 1884, and Katydiseius Fain & Lukoschus 1983, the subfamily Katydiseiinae Fain & Lukoschus, 1983, and their family group relationships (Acari: Mesostigmata: Gamasina), with description of three new species parasitic on gryllacridid crickets (Orthoptera)" masterLastPageNumber="70" masterPageNumber="5" pageNumber="18" updateTime="1698848037799" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="C2EEF806A4A22739840913ADA1BD836F">
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<mods:title id="5CB8D5C1F7A54E14A13BA42BCFBE16BC">Review of the genera Berlesia Canestrini, 1884, and Katydiseius Fain & Lukoschus 1983, the subfamily Katydiseiinae Fain & Lukoschus, 1983, and their family group relationships (Acari: Mesostigmata: Gamasina), with description of three new species parasitic on gryllacridid crickets (Orthoptera)</mods:title>
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<mods:name id="C0169B7CB62430481DFD968FA89E1C73" type="personal">
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<mods:roleTerm id="271E70039B699B3FC5E97F2DDC9E795C">Author</mods:roleTerm>
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<mods:namePart id="E650A169CAC1DF590322663BB9163C70">Lindquist, Evert E.</mods:namePart>
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<mods:name id="F5BD731A479CB95ADA96587FA2CFE77D" type="personal">
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<mods:roleTerm id="E464D2C9B3F81E6EF059A7DBD1C070B0">Author</mods:roleTerm>
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<mods:namePart id="27C397D5319C750049CFD248439EDEB7">Oconnor, Barry M.</mods:namePart>
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<mods:nameIdentifier id="FA77EFFD991D0F2D99D72DBB0DC653C1" type="ORCID">0000-0003-4588-0788</mods:nameIdentifier>
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<mods:affiliation id="BAA08AF67212E825E133F7FCE8E62532">Department of Ecology and Evolutionary Biology, Museum of Zoology, University of Michigan, Ann Arbor, Michigan, 48108, USA. bmoc @ umich. edu, https: // orcid. org / 0000 - 0003 - 4588 - 0788</mods:affiliation>
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</mods:name>
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<mods:name id="F0348B487AD5458F311A2E51F70AD009" type="personal">
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<mods:role id="7FFD7C157C6376D6013D434B59C951B2">
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<mods:roleTerm id="90011D035A215B0E1ADA1EEF0A61EF8E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7E71C7E6A951CE445E7C86B3C9314121">Shaw, Matthew D.</mods:namePart>
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<mods:nameIdentifier id="322FC5CB5B6C3186BF869354EAF8FB9C" type="ORCID">0000-0002-5037-3740</mods:nameIdentifier>
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<mods:affiliation id="03387586A5870EE8814AEDDDEF39F2A5">Terrestrial Invertebrates, South Australian Museum, North Tce, Adelaide 5000, South Australia, Australia matthew. shaw @ samuseum. sa. gov. au, https: // orcid. org / 0000 - 0002 - 5037 - 3740</mods:affiliation>
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<mods:namePart id="71500B3A776C3FB827333783F2B3BBE7">Sidorchuk, Ekaterina A.</mods:namePart>
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<mods:affiliation id="FB0BF993B0B29B82BC285E93F11FB119">Arthropoda Laboratory, Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya ulitsa 123, 117997 Moscow, Russia. International Complex Research Laboratory for the Study of Climate Change, Land Usage, and Biodiversity, Tyumen State University, 10 Semakova Str., 625003, Tyumen, Russia.</mods:affiliation>
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<mods:typeOfResource id="8DF2573A6550ED893D81FD2D6DAEB21C">text</mods:typeOfResource>
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<mods:title id="24642917473541DF204B444B53DF3BF2">Zootaxa</mods:title>
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<mods:part id="16218F4D6DBBE97A0AF419298F4C42E2">
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<mods:date id="552935F54F5697BE0996243E06F610E7">2020</mods:date>
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<mods:number id="6D605B113EAB5A8DCB704DA8C3A3D736">2020-09-28</mods:number>
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<mods:number id="41F103FF2156DA217D5F1FFEA55257C3">4857</mods:number>
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<mods:identifier id="EA96C09802AA09BFE64E54387571C0B8" type="DOI">10.11646/zootaxa.4857.1.4</mods:identifier>
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<mods:identifier id="D3762E0BF6D2D84F17839D3EF0FAA59D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E2879C4F48FF9EFF66A92BFE4ED269" ID-DOI="http://doi.org/10.5281/zenodo.4396472" ID-GBIF-Taxon="176438117" ID-Zenodo-Dep="4396472" LSID="urn:lsid:plazi:treatment:03E2879C4F48FF9EFF66A92BFE4ED269" httpUri="http://treatment.plazi.org/id/03E2879C4F48FF9EFF66A92BFE4ED269" lastPageId="21" lastPageNumber="26" pageId="13" pageNumber="18">
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<subSubSection id="C35165014F48FF86FF66A92BFD8ED35A" box="[151,546,582,609]" pageId="13" pageNumber="18" type="nomenclature">
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<paragraph id="8BF4368A4F48FF86FF66A92BFD8ED35A" blockId="13.[151,546,582,645]" box="[151,546,582,609]" pageId="13" pageNumber="18">
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<heading id="D0BC81E64F48FF86FF66A92BFD8ED35A" bold="true" box="[151,546,582,609]" fontSize="11" level="1" pageId="13" pageNumber="18" reason="1">
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<emphasis id="B93FEA984F48FF86FF66A92BFD8ED35A" bold="true" box="[151,546,582,609]" pageId="13" pageNumber="18">
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<taxonomicName id="4C4B4D094F48FF86FF66A92BFE27D35B" authority="Lindquist & Oconnor & Shaw & Sidorchuk, 2020" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[151,395,582,609]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="18" phylum="Arthropoda" rank="species" species="multisetosa" status="sp. nov.">
|
||
<emphasis id="B93FEA984F48FF86FF66A92BFE27D35B" bold="true" box="[151,395,582,609]" italics="true" pageId="13" pageNumber="18">Berlesia multisetosa</emphasis>
|
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</taxonomicName>
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||
<taxonomicNameLabel id="A20C57E34F48FF86FE63A92AFD8ED35A" box="[402,546,582,609]" pageId="13" pageNumber="18" rank="species">new species</taxonomicNameLabel>
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</emphasis>
|
||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C35165014F48FF86FF66A906FE91D3BF" box="[151,317,619,645]" pageId="13" pageNumber="18" type="description">
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<paragraph id="8BF4368A4F48FF86FF66A906FE91D3BF" blockId="13.[151,546,582,645]" box="[151,317,619,645]" pageId="13" pageNumber="18">
|
||
(
|
||
<figureCitation id="13702A0F4F48FF86FF6EA906FE99D3BF" box="[159,309,619,645]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStart-5="FIGURE 12" captionStartId-0="15.[151,250,1574,1598]" captionStartId-1="16.[151,250,1467,1491]" captionStartId-2="18.[151,250,1882,1906]" captionStartId-3="19.[151,250,1104,1128]" captionStartId-4="20.[151,250,1099,1123]" captionStartId-5="20.[151,250,1834,1858]" captionTargetBox-0="[151,1435,189,1544]" captionTargetBox-1="[152,1431,595,1440]" captionTargetBox-2="[167,1423,189,1855]" captionTargetBox-3="[155,1432,184,1077]" captionTargetBox-4="[155,1431,185,1068]" captionTargetBox-5="[157,1433,1222,1807]" captionTargetId-0="figure-17@15.[151,1435,181,1550]" captionTargetId-1="figure-267@16.[151,1436,591,1443]" captionTargetId-2="figure-21@18.[164,1423,181,1858]" captionTargetId-3="figure-17@19.[151,1436,181,1081]" captionTargetId-4="figure-21@20.[151,1436,181,1075]" captionTargetId-5="figure-67@20.[151,1437,1219,1810]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="20" captionText-0="FIGURE 7. Berlesia multisetosa n. sp., female. A, idiosoma dorsally; B, idiosoma ventrally; C, subcapitulum; D, E, chelicerae, dorsally, in working position, E, enlarged detail; F, peritreme; G, sperm access system; H, J, variability of gnathotectum outline. A, B, G to same scale, C, D, F, H, J to same scale. Abbreviations: co, corniculus; d lf, dorsal lyrifissure; d s, dorsal seta; id, lyrifissure; gp, glandular pore; lb, labrum; mi, internal mala; p, unidentified pore-like structure; p d, pilus dentilis; ss, salivary stylus; t, tooth; td, tendon; others denote setae." captionText-1="FIGURE 8. Berlesia multisetosa n. sp., female, legs.A, C, F, G, legs dorsally except separated coxae ventrally:A, leg I, subapical tarsal setae shown with circles; B, tarsus I dorsally; C, detail of sensory cluster on tarsus I; D–G, legs dorsally except separated coxae ventrally: D, leg II; E, detail of tarsus II tip ventrally; F, leg III, tarsus partly shown; G, leg IV, tarsus partly shown; H, tarsus IV ventrally. A, D–H to same scale." captionText-2="FIGURE 9. Berlesia multisetosa n. sp., male. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, D, chelicera and spermatodactyl, C, dorsal aspect, D, ventral; E, gnathotectum; F, subcapitulum and palpus; G, H, palp tarsus, G, ventral aspect, H, dorsal, enlarged; J, peritreme.A, B to same scale, C–G to same scale, H, J to same scale.Abbreviations: ar, arthrodial envelope; f d, fixed digit; m d, movable digit; others as in Fig. 7." captionText-3="FIGURE 10. Berlesia multisetosa n. sp., male, legs, coxae detached. A, leg I ventrally, most setae of tarsal subapex, beyond transverse dotted line, omitted; B, leg II ventrally; C, leg III ventrally; D, leg IV ventrolaterally. All to same scale." captionText-4="FIGURE 11. Berlesia multisetosa n. sp., deutonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, apical half of chelicera, ventrally, at resting position; D, variations of peritremes; E–G, variability of gnathotectum outline.A, B to same scale; C, E–G to same scale." captionText-5="FIGURE 12. Berlesia multisetosa n. sp., protonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, right chelicerae dorsally, at resting position; D, gnathotectum; E, subcapitulum. A, B to same scale, C–E to same scale." figureDoi-0="http://doi.org/10.5281/zenodo.4396490" figureDoi-1="http://doi.org/10.5281/zenodo.4396492" figureDoi-2="http://doi.org/10.5281/zenodo.4396494" figureDoi-3="http://doi.org/10.5281/zenodo.4396496" figureDoi-4="http://doi.org/10.5281/zenodo.4396498" figureDoi-5="http://doi.org/10.5281/zenodo.4396500" httpUri-0="https://zenodo.org/record/4396490/files/figure.png" httpUri-1="https://zenodo.org/record/4396492/files/figure.png" httpUri-2="https://zenodo.org/record/4396494/files/figure.png" httpUri-3="https://zenodo.org/record/4396496/files/figure.png" httpUri-4="https://zenodo.org/record/4396498/files/figure.png" httpUri-5="https://zenodo.org/record/4396500/files/figure.png" pageId="13" pageNumber="18">Figures 7–12</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSection id="E2C42F614F48FF86FF66A9DFFC6FD2F3" pageId="13" pageNumber="18" type="multiple">
|
||
<subSubSection id="C35165014F48FF86FF66A9DFFC6FD2F3" pageId="13" pageNumber="18" type="diagnosis">
|
||
<paragraph id="8BF4368A4F48FF86FF66A9DFFEA6D3F6" blockId="13.[151,1436,690,969]" box="[151,266,690,716]" pageId="13" pageNumber="18">
|
||
<heading id="D0BC81E64F48FF86FF66A9DFFEA6D3F6" bold="true" box="[151,266,690,716]" fontSize="11" level="1" pageId="13" pageNumber="18" reason="1">
|
||
<emphasis id="B93FEA984F48FF86FF66A9DFFEA6D3F6" bold="true" box="[151,266,690,716]" pageId="13" pageNumber="18">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F48FF86FF66A9BAFC6FD2F3" blockId="13.[151,1436,690,969]" pageId="13" pageNumber="18">
|
||
Adult: dorsal shield entirely smooth, female’s with irregular indication of lateral incisions at level of setae
|
||
<emphasis id="B93FEA984F48FF86FAC1A9BBFAE5D3CA" box="[1328,1353,726,752]" italics="true" pageId="13" pageNumber="18">s6</emphasis>
|
||
; vertex setae
|
||
<emphasis id="B93FEA984F48FF86FF27A996FF47D22F" box="[214,235,763,789]" italics="true" pageId="13" pageNumber="18">j1</emphasis>
|
||
, mid-dorsals (
|
||
<emphasis id="B93FEA984F48FF86FE7FA997FD93D22F" box="[398,575,762,789]" italics="true" pageId="13" pageNumber="18">j4–j6, z5, J1–J3</emphasis>
|
||
) and
|
||
<emphasis id="B93FEA984F48FF86FD8FA996FD35D22F" box="[638,665,763,789]" italics="true" pageId="13" pageNumber="18">J4</emphasis>
|
||
of moderate length (e.g.,
|
||
<emphasis id="B93FEA984F48FF86FC48A996FC63D22F" box="[953,975,763,789]" italics="true" pageId="13" pageNumber="18">j4</emphasis>
|
||
nearly as long as interval between
|
||
<emphasis id="B93FEA984F48FF86FAA6A996FAC1D22F" box="[1367,1389,763,789]" italics="true" pageId="13" pageNumber="18">j4</emphasis>
|
||
and
|
||
<emphasis id="B93FEA984F48FF86FF66A872FF01D203" box="[151,173,799,825]" italics="true" pageId="13" pageNumber="18">j5</emphasis>
|
||
); dorsal shield setae
|
||
<emphasis id="B93FEA984F48FF86FE6DA872FE1FD203" box="[412,435,799,825]" italics="true" pageId="13" pageNumber="18">j3</emphasis>
|
||
and
|
||
<emphasis id="B93FEA984F48FF86FE1BA872FDAFD203" box="[490,515,799,825]" italics="true" pageId="13" pageNumber="18">z4</emphasis>
|
||
slightly longer, but shorter than diagonal interval between them, and shorter than transverse interval between
|
||
<emphasis id="B93FEA984F48FF86FE3FA82EFE48D267" box="[462,484,835,861]" italics="true" pageId="13" pageNumber="18">j3</emphasis>
|
||
; setae
|
||
<emphasis id="B93FEA984F48FF86FDC1A82EFDE4D267" box="[560,584,835,861]" italics="true" pageId="13" pageNumber="18">z1</emphasis>
|
||
and
|
||
<emphasis id="B93FEA984F48FF86FD8CA82EFD3BD267" box="[637,663,835,861]" italics="true" pageId="13" pageNumber="18">J2</emphasis>
|
||
present. Dorsolateral areas of idiosoma with setae of
|
||
<emphasis id="B93FEA984F48FF86FB1BA82EFAB8D266" box="[1258,1300,835,860]" italics="true" pageId="13" pageNumber="18">r–R</emphasis>
|
||
and
|
||
<emphasis id="B93FEA984F48FF86FAB9A82EFAFAD267" box="[1352,1366,835,861]" italics="true" pageId="13" pageNumber="18">S</emphasis>
|
||
series hypertrichous. Soft ventral cuticle hypertrichous, with about 15 pairs of opisthogastric setae. Leg I genu and tibia each with 11 setae,
|
||
<emphasis id="B93FEA984F48FF86FE9CA8E6FE0CD29F" box="[365,416,907,933]" italics="true" pageId="13" pageNumber="18">pd-3</emphasis>
|
||
and
|
||
<emphasis id="B93FEA984F48FF86FE25A8E6FDA9D29F" box="[468,517,907,933]" italics="true" pageId="13" pageNumber="18">av-2</emphasis>
|
||
absent. Tarsus IV with seta
|
||
<emphasis id="B93FEA984F48FF86FCC9A8E6FCC6D29F" box="[824,874,907,933]" italics="true" pageId="13" pageNumber="18">pv-1</emphasis>
|
||
bluntly spine-like and asymmetrically shorter than attenuated
|
||
<emphasis id="B93FEA984F48FF86FEE0A8C2FEEFD2F3" box="[273,323,943,969]" italics="true" pageId="13" pageNumber="18">av-1</emphasis>
|
||
. Peritremes embellished with internal cellular structures.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
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||
<subSection id="E2C42F614F48FF9EFF66A89BFE4ED269" lastPageId="21" lastPageNumber="26" pageId="13" pageNumber="18" type="multiple">
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<subSubSection id="C35165014F48FF98FF66A89BFCC2D7C8" lastPageId="19" lastPageNumber="24" pageId="13" pageNumber="18" type="description">
|
||
<paragraph id="8BF4368A4F48FF86FF66A89BFE8DD52A" blockId="13.[151,1437,1014,2013]" box="[151,289,1014,1040]" pageId="13" pageNumber="18">
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<heading id="D0BC81E64F48FF86FF66A89BFE8DD52A" bold="true" box="[151,289,1014,1040]" fontSize="11" level="1" pageId="13" pageNumber="18" reason="1">
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<emphasis id="B93FEA984F48FF86FF66A89BFE8DD52A" bold="true" box="[151,289,1014,1040]" pageId="13" pageNumber="18">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F48FF86FF66AF77FDEDD4F8" blockId="13.[151,1437,1014,2013]" pageId="13" pageNumber="18">
|
||
<emphasis id="B93FEA984F48FF86FF66AF77FE2FD50F" bold="true" box="[151,387,1050,1077]" pageId="13" pageNumber="18">
|
||
<emphasis id="B93FEA984F48FF86FF66AF77FED0D50F" bold="true" box="[151,380,1050,1077]" italics="true" pageId="13" pageNumber="18">Idiosomatic dorsum</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B93FEA984F48FF86FE78AF76FDBAD50F" box="[393,534,1051,1077]" italics="true" pageId="13" pageNumber="18">Adult female</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F48FF86FDD5AF76FDD1D50F" box="[548,637,1051,1077]" captionStart="FIGURE 7" captionStartId="15.[151,250,1574,1598]" captionTargetBox="[151,1435,189,1544]" captionTargetId="figure-17@15.[151,1435,181,1550]" captionTargetPageId="15" captionText="FIGURE 7. Berlesia multisetosa n. sp., female. A, idiosoma dorsally; B, idiosoma ventrally; C, subcapitulum; D, E, chelicerae, dorsally, in working position, E, enlarged detail; F, peritreme; G, sperm access system; H, J, variability of gnathotectum outline. A, B, G to same scale, C, D, F, H, J to same scale. Abbreviations: co, corniculus; d lf, dorsal lyrifissure; d s, dorsal seta; id, lyrifissure; gp, glandular pore; lb, labrum; mi, internal mala; p, unidentified pore-like structure; p d, pilus dentilis; ss, salivary stylus; t, tooth; td, tendon; others denote setae." figureDoi="http://doi.org/10.5281/zenodo.4396490" httpUri="https://zenodo.org/record/4396490/files/figure.png" pageId="13" pageNumber="18">Fig. 7A</figureCitation>
|
||
). Idiosoma 625–700 long, 412–432 wide between insertions of setae
|
||
<emphasis id="B93FEA984F48FF86FA8DAF76FA39D50F" box="[1404,1429,1051,1077]" italics="true" pageId="13" pageNumber="18">r3</emphasis>
|
||
; dorsal shield 520–530 long, greatest width 290–320 at level between insertions of
|
||
<emphasis id="B93FEA984F48FF86FBCCAF52FBF9D563" box="[1085,1109,1087,1113]" italics="true" pageId="13" pageNumber="18">s4</emphasis>
|
||
(n=2), unornamented except for subdermal sigillar patterns; shield with irregularly formed lateral incisions at level of setae
|
||
<emphasis id="B93FEA984F48FF86FB2DAF0FFB50D546" box="[1244,1276,1122,1148]" italics="true" pageId="13" pageNumber="18">s6.</emphasis>
|
||
Dorsal shield with usually 10 to 14 pairs (5 to 8 podonotal, 4 to 6 opisthonotal) of smooth setae of which any of
|
||
<emphasis id="B93FEA984F48FF86FAE1AFEAFA84D59B" box="[1296,1320,1159,1185]" italics="true" pageId="13" pageNumber="18">z2</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F48FF86FAC8AFEAFAFDD59B" box="[1337,1361,1159,1185]" italics="true" pageId="13" pageNumber="18">s5</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F48FF86FA90AFEAFAD0D59B" box="[1377,1404,1159,1185]" italics="true" pageId="13" pageNumber="18">J4</emphasis>
|
||
or
|
||
<emphasis id="B93FEA984F48FF86FF66AFC6FF19D5FF" box="[151,181,1195,1221]" italics="true" pageId="13" pageNumber="18">Z4</emphasis>
|
||
asymmetrically on/off shield margin; of shield setae,
|
||
<emphasis id="B93FEA984F48FF86FCE9AFC6FC82D5FF" box="[792,814,1195,1221]" italics="true" pageId="13" pageNumber="18">j3</emphasis>
|
||
(60–65),
|
||
<emphasis id="B93FEA984F48FF86FC6CAFC6FC1AD5FF" box="[925,950,1195,1221]" italics="true" pageId="13" pageNumber="18">z4</emphasis>
|
||
(62–75),
|
||
<emphasis id="B93FEA984F48FF86FBD4AFC6FB92D5FF" box="[1061,1086,1195,1221]" italics="true" pageId="13" pageNumber="18">s4</emphasis>
|
||
(62–75),
|
||
<emphasis id="B93FEA984F48FF86FB5FAFC6FB6AD5FF" box="[1198,1222,1195,1221]" italics="true" pageId="13" pageNumber="18">s5</emphasis>
|
||
(60–70) of similar lengths;
|
||
<emphasis id="B93FEA984F48FF86FF09AFA2FEA2D5D3" box="[248,270,1231,1257]" italics="true" pageId="13" pageNumber="18">j4</emphasis>
|
||
(45–65),
|
||
<emphasis id="B93FEA984F48FF86FE8CAFA2FE3ED5D3" box="[381,402,1231,1257]" italics="true" pageId="13" pageNumber="18">j5</emphasis>
|
||
(40–60),
|
||
<emphasis id="B93FEA984F48FF86FDF0AFA3FDBBD5D2" box="[513,535,1230,1256]" italics="true" pageId="13" pageNumber="18">j6</emphasis>
|
||
(31–53),
|
||
<emphasis id="B93FEA984F48FF86FD77AFA2FD33D5D3" box="[646,671,1231,1257]" italics="true" pageId="13" pageNumber="18">z5</emphasis>
|
||
(33–52),
|
||
<emphasis id="B93FEA984F48FF86FCFFAFA2FCFED5D3" box="[782,850,1231,1257]" italics="true" pageId="13" pageNumber="18">J1–J4</emphasis>
|
||
somewhat shorter but not tiny (22–40), with
|
||
<emphasis id="B93FEA984F48FF86FAA2AFA2FAC0D5D3" box="[1363,1388,1231,1257]" italics="true" pageId="13" pageNumber="18">z4</emphasis>
|
||
and
|
||
<emphasis id="B93FEA984F48FF86FF66AF9EFF1CD437" box="[151,176,1267,1293]" italics="true" pageId="13" pageNumber="18">s4</emphasis>
|
||
(62–75),
|
||
<emphasis id="B93FEA984F48FF86FED2AF9EFE97D437" box="[291,315,1267,1293]" italics="true" pageId="13" pageNumber="18">s5</emphasis>
|
||
(60–67),
|
||
<emphasis id="B93FEA984F48FF86FE5FAF9EFE67D437" box="[430,459,1267,1293]" italics="true" pageId="13" pageNumber="18">Z1</emphasis>
|
||
(57–70) and
|
||
<emphasis id="B93FEA984F48FF86FD98AF9EFD2AD437" box="[617,646,1267,1293]" italics="true" pageId="13" pageNumber="18">Z4</emphasis>
|
||
(60–72) similarly longer. Soft dorsal peripheral cuticle with setae
|
||
<emphasis id="B93FEA984F48FF86FA77AF9EFA30D437" box="[1414,1436,1267,1293]" italics="true" pageId="13" pageNumber="18">j1</emphasis>
|
||
(47–52),
|
||
<emphasis id="B93FEA984F48FF86FF0FAE7AFEBBD40B" box="[254,279,1303,1329]" italics="true" pageId="13" pageNumber="18">z1</emphasis>
|
||
(40–43),
|
||
<emphasis id="B93FEA984F48FF86FE77AE7AFE33D40B" box="[390,415,1303,1329]" italics="true" pageId="13" pageNumber="18">z2</emphasis>
|
||
(54) anteriorly,
|
||
<emphasis id="B93FEA984F48FF86FDA6AE7BFDDCD40A" box="[599,624,1302,1328]" italics="true" pageId="13" pageNumber="18">s6</emphasis>
|
||
(60–65) and a hypertrichy of 12–15
|
||
<emphasis id="B93FEA984F48FF86FBE0AE7AFBB0D40A" box="[1041,1052,1303,1328]" italics="true" pageId="13" pageNumber="18">r</emphasis>
|
||
-setae (50–62) anterolaterally, and
|
||
<emphasis id="B93FEA984F48FF86FF66AE56FF18D46F" box="[151,180,1339,1365]" italics="true" pageId="13" pageNumber="18">Z2</emphasis>
|
||
(57–65),
|
||
<emphasis id="B93FEA984F48FF86FED5AE56FEC5D46F" box="[292,361,1339,1365]" italics="true" pageId="13" pageNumber="18">S1–S4</emphasis>
|
||
(55–70),
|
||
<emphasis id="B93FEA984F48FF86FE28AE56FE59D46F" box="[473,501,1339,1365]" italics="true" pageId="13" pageNumber="18">S5</emphasis>
|
||
(75) and hypertrichy of 10–15
|
||
<emphasis id="B93FEA984F48FF86FCA9AE56FCC5D46E" box="[856,873,1339,1364]" italics="true" pageId="13" pageNumber="18">R</emphasis>
|
||
-setae (52–65) posterolaterally. Dorsal shield with complement of nine pairs of discernible pore-like structures (four podonotal, five opisthonotal), of which three pairs (one podonotal, two opisthonotal) superficially appear secretory (gland pores) and six pairs (three podonotal, three opisthonotal) non-secretory (poroids).
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F48FF86FF36AEA6FA88D63C" blockId="13.[151,1437,1014,2013]" pageId="13" pageNumber="18">
|
||
<emphasis id="B93FEA984F48FF86FF36AEA6FEECD4DF" box="[199,320,1483,1509]" italics="true" pageId="13" pageNumber="18">Adult male</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F48FF86FEBEAEA6FE04D4DF" box="[335,424,1483,1509]" captionStart="FIGURE 9" captionStartId="18.[151,250,1882,1906]" captionTargetBox="[167,1423,189,1855]" captionTargetId="figure-21@18.[164,1423,181,1858]" captionTargetPageId="18" captionText="FIGURE 9. Berlesia multisetosa n. sp., male. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, D, chelicera and spermatodactyl, C, dorsal aspect, D, ventral; E, gnathotectum; F, subcapitulum and palpus; G, H, palp tarsus, G, ventral aspect, H, dorsal, enlarged; J, peritreme.A, B to same scale, C–G to same scale, H, J to same scale.Abbreviations: ar, arthrodial envelope; f d, fixed digit; m d, movable digit; others as in Fig. 7." figureDoi="http://doi.org/10.5281/zenodo.4396494" httpUri="https://zenodo.org/record/4396494/files/figure.png" pageId="13" pageNumber="18">Fig. 9A</figureCitation>
|
||
). Idiosoma 580–612 long, 370–437 wide between insertions of setae
|
||
<emphasis id="B93FEA984F48FF86FB58AEA6FB6FD4DF" box="[1193,1219,1483,1509]" italics="true" pageId="13" pageNumber="18">r3</emphasis>
|
||
; dorsal shield 520– 532 long, greatest width 337–365 at level between insertions of
|
||
<emphasis id="B93FEA984F48FF86FC81AE83FC26D732" box="[880,906,1518,1544]" italics="true" pageId="13" pageNumber="18">s6</emphasis>
|
||
(n=3), unornamented, with irregular but entire lateral margins, which widened to hold setae
|
||
<emphasis id="B93FEA984F48FF86FD60AD7EFD0AD717" box="[657,678,1555,1581]" italics="true" pageId="13" pageNumber="18">j1</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F48FF86FD44AD7EFD61D717" box="[693,717,1555,1581]" italics="true" pageId="13" pageNumber="18">z1</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F48FF86FD2AAD7EFD58D717" box="[731,756,1555,1581]" italics="true" pageId="13" pageNumber="18">s3</emphasis>
|
||
anteriorly and
|
||
<emphasis id="B93FEA984F48FF86FC6FAD7EFC49D717" box="[926,997,1555,1581]" italics="true" pageId="13" pageNumber="18">Z2, S2</emphasis>
|
||
laterally. Dorsal shield with usually 23 to 25 pairs (14 or 15 podonotal, 9 or 10 opisthonotal) of smooth setae of which
|
||
<emphasis id="B93FEA984F48FF86FBE0AD5BFB86D76A" box="[1041,1066,1590,1616]" italics="true" pageId="13" pageNumber="18">s6</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F48FF86FBC9AD5AFBF9D76B" box="[1080,1109,1591,1617]" italics="true" pageId="13" pageNumber="18">Z2</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F48FF86FB92AD5AFB2DD76B" box="[1123,1153,1591,1617]" italics="true" pageId="13" pageNumber="18">Z4</emphasis>
|
||
often asymmetrically off shield margin; relative setal lengths much as in female, except
|
||
<emphasis id="B93FEA984F48FF86FC92AD36FC0AD74F" box="[867,934,1627,1653]" italics="true" pageId="13" pageNumber="18">J1–J4</emphasis>
|
||
shorter (20–30) compared to
|
||
<emphasis id="B93FEA984F48FF86FB0BAD37FA9FD74E" box="[1274,1331,1626,1652]" italics="true" pageId="13" pageNumber="18">j4–j6</emphasis>
|
||
(37–57); other setal lengths
|
||
<emphasis id="B93FEA984F48FF86FE9FAD12FE28D7A3" box="[366,388,1663,1689]" italics="true" pageId="13" pageNumber="18">j1</emphasis>
|
||
(35–42),
|
||
<emphasis id="B93FEA984F48FF86FE04AD12FDA7D7A3" box="[501,523,1663,1689]" italics="true" pageId="13" pageNumber="18">j3</emphasis>
|
||
(58–68),
|
||
<emphasis id="B93FEA984F48FF86FD8AAD12FD38D7A3" box="[635,660,1663,1689]" italics="true" pageId="13" pageNumber="18">z1</emphasis>
|
||
(25–32),
|
||
<emphasis id="B93FEA984F48FF86FCF4AD12FCB2D7A3" box="[773,798,1663,1689]" italics="true" pageId="13" pageNumber="18">z2</emphasis>
|
||
(55–62),
|
||
<emphasis id="B93FEA984F48FF86FC7EAD12FC04D7A3" box="[911,936,1663,1689]" italics="true" pageId="13" pageNumber="18">z4</emphasis>
|
||
(60–65),
|
||
<emphasis id="B93FEA984F48FF86FBE9AD13FBF4D7A2" box="[1048,1112,1662,1688]" italics="true" pageId="13" pageNumber="18">s4–s6</emphasis>
|
||
(57–67), about eight
|
||
<emphasis id="B93FEA984F48FF86FAA1AD12FAF7D7A2" box="[1360,1371,1663,1688]" italics="true" pageId="13" pageNumber="18">r</emphasis>
|
||
-setae (40–52),
|
||
<emphasis id="B93FEA984F48FF86FEF1ADCEFEE5D787" box="[256,329,1699,1725]" italics="true" pageId="13" pageNumber="18">Z1–Z2</emphasis>
|
||
(60–65),
|
||
<emphasis id="B93FEA984F48FF86FE4AADCEFE75D787" box="[443,473,1699,1725]" italics="true" pageId="13" pageNumber="18">Z3</emphasis>
|
||
(60–72),
|
||
<emphasis id="B93FEA984F48FF86FDBBADCEFD3FD787" box="[586,659,1699,1725]" italics="true" pageId="13" pageNumber="18">Z4–Z5</emphasis>
|
||
(65–77),
|
||
<emphasis id="B93FEA984F48FF86FCF4ADCEFCE7D787" box="[773,843,1699,1725]" italics="true" pageId="13" pageNumber="18">S1–S3</emphasis>
|
||
(62–68),
|
||
<emphasis id="B93FEA984F48FF86FC4DADCEFC74D787" box="[956,984,1699,1725]" italics="true" pageId="13" pageNumber="18">S4</emphasis>
|
||
(65–72),
|
||
<emphasis id="B93FEA984F48FF86FBBBADCEFBCAD787" box="[1098,1126,1699,1725]" italics="true" pageId="13" pageNumber="18">S5</emphasis>
|
||
(68–85), about ten
|
||
<emphasis id="B93FEA984F48FF86FABBADCEFAF7D786" box="[1354,1371,1699,1724]" italics="true" pageId="13" pageNumber="18">R</emphasis>
|
||
-setae (52–68). Dorsal shield with complement of eleven pairs of discernible pore-like structures (4 podonotal, 7 opisthonotal), most of which are difficult to discern whether secretory (gland pores) or non-secretory (poroids).
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F48FF85FF36AC63FB2AD35B" blockId="13.[151,1437,1014,2013]" lastBlockId="14.[151,1437,150,1905]" lastPageId="14" lastPageNumber="19" pageId="13" pageNumber="18">
|
||
<emphasis id="B93FEA984F48FF86FF36AC63FE0CD613" bold="true" box="[199,416,1806,1833]" italics="true" pageId="13" pageNumber="18">Idiosomatic venter</emphasis>
|
||
.
|
||
<emphasis id="B93FEA984F48FF86FE41AC62FDEDD613" box="[432,577,1807,1833]" italics="true" pageId="13" pageNumber="18">Adult female</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F48FF86FDA2AC62FD02D613" box="[595,686,1807,1833]" captionStart="FIGURE 7" captionStartId="15.[151,250,1574,1598]" captionTargetBox="[151,1435,189,1544]" captionTargetId="figure-17@15.[151,1435,181,1550]" captionTargetPageId="15" captionText="FIGURE 7. Berlesia multisetosa n. sp., female. A, idiosoma dorsally; B, idiosoma ventrally; C, subcapitulum; D, E, chelicerae, dorsally, in working position, E, enlarged detail; F, peritreme; G, sperm access system; H, J, variability of gnathotectum outline. A, B, G to same scale, C, D, F, H, J to same scale. Abbreviations: co, corniculus; d lf, dorsal lyrifissure; d s, dorsal seta; id, lyrifissure; gp, glandular pore; lb, labrum; mi, internal mala; p, unidentified pore-like structure; p d, pilus dentilis; ss, salivary stylus; t, tooth; td, tendon; others denote setae." figureDoi="http://doi.org/10.5281/zenodo.4396490" httpUri="https://zenodo.org/record/4396490/files/figure.png" pageId="13" pageNumber="18">Fig. 7B</figureCitation>
|
||
). Tritosternum base 23–25 long and 23–26 wide at its base, with paired ciliate laciniae fused for about 5–7 of entire length (ca 100). Presternal area with a delineated, lightly sclerotized, streaky region without clear platelets. Sternal shield mid-length (98–100) much less than narrowest width (148–155) between legs II, faintly scabrous; sternal setae
|
||
<emphasis id="B93FEA984F48FF86FCC9AC16FCF4D6AF" box="[824,856,1915,1941]" italics="true" pageId="13" pageNumber="18">st1</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F48FF86FC9BAC16FC26D6AF" box="[874,906,1915,1941]" italics="true" pageId="13" pageNumber="18">st2</emphasis>
|
||
attenuated, of similar length (55–67). Sternal setae
|
||
<emphasis id="B93FEA984F48FF86FF26ACF2FF54D683" box="[215,248,1951,1977]" italics="true" pageId="13" pageNumber="18">st3</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F48FF86FEF7ACF2FE8AD683" box="[262,294,1951,1977]" italics="true" pageId="13" pageNumber="18">st4</emphasis>
|
||
similarly attenuated (50–60), with poroid
|
||
<emphasis id="B93FEA984F48FF86FCF5ACF2FC8BD683" box="[772,807,1951,1977]" italics="true" pageId="13" pageNumber="18">iv3</emphasis>
|
||
on soft cuticle. Endopodal strips rudimentary alongside coxae III–IV. Epigynal shield smooth, its rounded hyaline anterior margin reaching or slightly overlapping posterior edge of sternal shield, its lateral margins slightly widened at insertions of attenuated setae
|
||
<emphasis id="B93FEA984F4BFF85FB78ABFAFB06D18B" box="[1161,1194,151,177]" italics="true" pageId="14" pageNumber="19">st5</emphasis>
|
||
(57–65), its posterior margin slightly convex; shield 205–223 long, 60–63 at narrowest width between legs IV, 77–85 at widest level at setae
|
||
<emphasis id="B93FEA984F4BFF85FF26ABB2FF54D1C3" box="[215,248,223,249]" italics="true" pageId="14" pageNumber="19">st5</emphasis>
|
||
. Postgenital groove undeveloped, without elements of platelets. Opisthosomatic venter with two pairs of weak metapodal platelets, lateral pair 30–32 x 8–12, inner pair 17–20 x 7–10. Anal shield subrhomboidal, lightly lineated; shield greatest width (128) slightly less than length (132–147, including cribrum 20), cribrum width 75; postanal seta (60–70) about 1.5 longer than paranal setae (37–50) inserted in line with posterior margin of anus. Soft cuticle with ca 15 pairs of opisthogastric setae of similar lengths (43–57) flanking anal shield,
|
||
<emphasis id="B93FEA984F4BFF85FB4BAA02FB4AD0B3" box="[1210,1254,367,393]" italics="true" pageId="14" pageNumber="19">JV5</emphasis>
|
||
not distinguishable posterolaterally; of four or five pairs of pore-like structures on soft cuticle, paranal gland pores well separated from lateral margins of anal shield. Peritrematal shield and peritreme extending barely to mid-level of coxae III; peritreme with embellished internal chambers (
|
||
<figureCitation id="13702A0F4F4BFF85FD5DAAB6FCAED0CF" box="[684,770,475,501]" captionStart="FIGURE 7" captionStartId="15.[151,250,1574,1598]" captionTargetBox="[151,1435,189,1544]" captionTargetId="figure-17@15.[151,1435,181,1550]" captionTargetPageId="15" captionText="FIGURE 7. Berlesia multisetosa n. sp., female. A, idiosoma dorsally; B, idiosoma ventrally; C, subcapitulum; D, E, chelicerae, dorsally, in working position, E, enlarged detail; F, peritreme; G, sperm access system; H, J, variability of gnathotectum outline. A, B, G to same scale, C, D, F, H, J to same scale. Abbreviations: co, corniculus; d lf, dorsal lyrifissure; d s, dorsal seta; id, lyrifissure; gp, glandular pore; lb, labrum; mi, internal mala; p, unidentified pore-like structure; p d, pilus dentilis; ss, salivary stylus; t, tooth; td, tendon; others denote setae." figureDoi="http://doi.org/10.5281/zenodo.4396490" httpUri="https://zenodo.org/record/4396490/files/figure.png" pageId="14" pageNumber="19">Fig. 7F</figureCitation>
|
||
), its length 45, including stigmatal diameter of ca 22; peritrematal shield continuous posteriorly with short exopodal strip extending partly around posterior margin of coxa IV. Exopodal elements absent alongside coxae II–IV. Spermathecal apparatus with simple, elongate (90–95) tubulus leading from solenostome between bases of legs III–IV to an unpaired sacculus (
|
||
<figureCitation id="13702A0F4F4BFF85FBEEA92AFBD5D35B" box="[1055,1145,583,609]" captionStart="FIGURE 7" captionStartId="15.[151,250,1574,1598]" captionTargetBox="[151,1435,189,1544]" captionTargetId="figure-17@15.[151,1435,181,1550]" captionTargetPageId="15" captionText="FIGURE 7. Berlesia multisetosa n. sp., female. A, idiosoma dorsally; B, idiosoma ventrally; C, subcapitulum; D, E, chelicerae, dorsally, in working position, E, enlarged detail; F, peritreme; G, sperm access system; H, J, variability of gnathotectum outline. A, B, G to same scale, C, D, F, H, J to same scale. Abbreviations: co, corniculus; d lf, dorsal lyrifissure; d s, dorsal seta; id, lyrifissure; gp, glandular pore; lb, labrum; mi, internal mala; p, unidentified pore-like structure; p d, pilus dentilis; ss, salivary stylus; t, tooth; td, tendon; others denote setae." figureDoi="http://doi.org/10.5281/zenodo.4396490" httpUri="https://zenodo.org/record/4396490/files/figure.png" pageId="14" pageNumber="19">Fig. 7G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F4BFF85FF36A906FA87D547" blockId="14.[151,1437,150,1905]" pageId="14" pageNumber="19">
|
||
<emphasis id="B93FEA984F4BFF85FF36A906FEEDD3BF" box="[199,321,619,645]" italics="true" pageId="14" pageNumber="19">Adult male</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F4BFF85FEA3A906FE07D3BF" box="[338,427,619,645]" captionStart="FIGURE 9" captionStartId="18.[151,250,1882,1906]" captionTargetBox="[167,1423,189,1855]" captionTargetId="figure-21@18.[164,1423,181,1858]" captionTargetPageId="18" captionText="FIGURE 9. Berlesia multisetosa n. sp., male. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, D, chelicera and spermatodactyl, C, dorsal aspect, D, ventral; E, gnathotectum; F, subcapitulum and palpus; G, H, palp tarsus, G, ventral aspect, H, dorsal, enlarged; J, peritreme.A, B to same scale, C–G to same scale, H, J to same scale.Abbreviations: ar, arthrodial envelope; f d, fixed digit; m d, movable digit; others as in Fig. 7." figureDoi="http://doi.org/10.5281/zenodo.4396494" httpUri="https://zenodo.org/record/4396494/files/figure.png" pageId="14" pageNumber="19">Fig. 9B</figureCitation>
|
||
). Tritosternum formed as in female, with laciniae fused for only about one-tenth of entire length (ca 100). Presternal region with two pairs of small platelets anterior to genital opening. Sternitigenital shield length 245–255, narrowest widths 115–125 at level between coxae II, 100–107 between coxae III; posterior margin width 62–80; shield unornamented, narrowly separated from anterior edge of endopodal strips beside coxae IV; alveoli of setae
|
||
<emphasis id="B93FEA984F4BFF85FEBCA996FEC2D22F" box="[333,366,763,789]" italics="true" pageId="14" pageNumber="19">st5</emphasis>
|
||
on corners of irregularly shaped posterior margin of shield; setae
|
||
<emphasis id="B93FEA984F4BFF85FB94A996FB18D22F" box="[1125,1204,763,789]" italics="true" pageId="14" pageNumber="19">st1–st5</emphasis>
|
||
similarly attenuated as in female,
|
||
<emphasis id="B93FEA984F4BFF85FEDEA872FEFCD203" box="[303,336,799,825]" italics="true" pageId="14" pageNumber="19">st1</emphasis>
|
||
and
|
||
<emphasis id="B93FEA984F4BFF85FE79A872FE05D203" box="[392,425,799,825]" italics="true" pageId="14" pageNumber="19">st5</emphasis>
|
||
(53–67) slightly longer than
|
||
<emphasis id="B93FEA984F4BFF85FD04A872FCE9D203" box="[757,837,799,825]" italics="true" pageId="14" pageNumber="19">st2–st4</emphasis>
|
||
(50–65). Paragenital poroids on soft cuticle flanking posterolateral corners of sternitigenital shield. Postgenital groove not developed. Inguinal region with five or six pairs of sigilla indicating unusual muscular development connected with deeply retractable bases of chelicerae. Anal shield similar in rhomboid form, slight ornamentation as in female, its anterior and lateral apices more irregular; shield longer (130–135, including cribrum ca 15–20 long) than wide (103–112, cribrum width 60–62); postanal seta (57–62) ca 1.5 to nearly twice as long as paranal setae (30–40). Soft cuticle with about 15 pairs of opisthogastric setae flanking anal shield; these setae gradually slightly lengthening posteriorly, those in
|
||
<emphasis id="B93FEA984F4BFF85FB65A89AFAC9D52B" box="[1172,1381,1015,1041]" italics="true" pageId="14" pageNumber="19">JV1-JV2-ZV1-ZV2</emphasis>
|
||
area 43–48, those mid-posteriorly 45–62, and posterior-most
|
||
<emphasis id="B93FEA984F4BFF85FCFBAF76FC99D50F" box="[778,821,1051,1077]" italics="true" pageId="14" pageNumber="19">JV5</emphasis>
|
||
(62–75); of five pairs of pore-like structures evident on soft cuticle, paranal gland pores well separated from lateral margins of anal shield. Short but embellished peritremes (40–45), and peritrematal shield with brief poststigmatal exopodal extension as in female (
|
||
<figureCitation id="13702A0F4F4BFF85FB7CAF0EFB4CD547" box="[1165,1248,1123,1149]" captionStart="FIGURE 9" captionStartId="18.[151,250,1882,1906]" captionTargetBox="[167,1423,189,1855]" captionTargetId="figure-21@18.[164,1423,181,1858]" captionTargetPageId="18" captionText="FIGURE 9. Berlesia multisetosa n. sp., male. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, D, chelicera and spermatodactyl, C, dorsal aspect, D, ventral; E, gnathotectum; F, subcapitulum and palpus; G, H, palp tarsus, G, ventral aspect, H, dorsal, enlarged; J, peritreme.A, B to same scale, C–G to same scale, H, J to same scale.Abbreviations: ar, arthrodial envelope; f d, fixed digit; m d, movable digit; others as in Fig. 7." figureDoi="http://doi.org/10.5281/zenodo.4396494" httpUri="https://zenodo.org/record/4396494/files/figure.png" pageId="14" pageNumber="19">Figs 9J</figureCitation>
|
||
,
|
||
<figureCitation id="13702A0F4F4BFF85FB1DAF0EFAB3D547" box="[1260,1311,1123,1149]" captionStart="FIGURE 20" captionStartId="50.[151,250,1734,1758]" captionTargetBox="[151,1435,885,1711]" captionTargetId="figure-335@50.[151,1436,885,1711]" captionTargetPageId="50" captionText="FIGURE 20. Berlesia multisetosa n. sp., male. A, ventral habitus, showing chelicerae fully withdrawn into idiosoma, yet spermatodactyl apices still protruding from subcapitulum; B, detail of peritreme; C–E, details of appendage apices: C, palp genua, tibiae, tarsi flanking tips of salivary stylets and internal malae; D, E, pretarsus and adjacent tarsal structures, showing reduced claws enveloped by pulvillus, D, leg II, E, leg I. Abbreviations: lb, labrum; mi, internal mala; sd, spermatodactyl; ss, salivary stylus; pv, pulvillus; others denote setae." figureDoi="http://doi.org/10.5281/zenodo.4396516" httpUri="https://zenodo.org/record/4396516/files/figure.png" pageId="14" pageNumber="19">20B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F4BFF85FF36AFEBFC74D733" blockId="14.[151,1437,150,1905]" pageId="14" pageNumber="19">
|
||
<taxonomicName id="4C4B4D094F4BFF85FF36AFEBFEF7D59B" authorityName="Krefft" authorityYear="1871" box="[199,347,1158,1185]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="14" pageNumber="19" phylum="Nematoda" rank="genus">
|
||
<emphasis id="B93FEA984F4BFF85FF36AFEBFEF7D59B" bold="true" box="[199,347,1158,1185]" italics="true" pageId="14" pageNumber="19">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B93FEA984F4BFF85FE9BAFEAFE56D59B" box="[362,506,1159,1185]" italics="true" pageId="14" pageNumber="19">Adult female</emphasis>
|
||
. Gnathotectum anterior margin obtusely smoothly triangular, with a blunt, medial anterior projection (
|
||
<figureCitation id="13702A0F4F4BFF85FE85AFC6FE44D5FF" box="[372,488,1195,1221]" captionStart="FIGURE 7" captionStartId="15.[151,250,1574,1598]" captionTargetBox="[151,1435,189,1544]" captionTargetId="figure-17@15.[151,1435,181,1550]" captionTargetPageId="15" captionText="FIGURE 7. Berlesia multisetosa n. sp., female. A, idiosoma dorsally; B, idiosoma ventrally; C, subcapitulum; D, E, chelicerae, dorsally, in working position, E, enlarged detail; F, peritreme; G, sperm access system; H, J, variability of gnathotectum outline. A, B, G to same scale, C, D, F, H, J to same scale. Abbreviations: co, corniculus; d lf, dorsal lyrifissure; d s, dorsal seta; id, lyrifissure; gp, glandular pore; lb, labrum; mi, internal mala; p, unidentified pore-like structure; p d, pilus dentilis; ss, salivary stylus; t, tooth; td, tendon; others denote setae." figureDoi="http://doi.org/10.5281/zenodo.4396490" httpUri="https://zenodo.org/record/4396490/files/figure.png" pageId="14" pageNumber="19">Figs 7H, J</figureCitation>
|
||
). Cheliceral shaft, excluding basal section, 155–158 long; fixed digit with blunt apex, vestige of pilus dentilis, and a vestigial tooth midway on paraxial face (
|
||
<figureCitation id="13702A0F4F4BFF85FC29AFA2FB9ED5D3" box="[984,1074,1231,1257]" captionStart="FIGURE 7" captionStartId="15.[151,250,1574,1598]" captionTargetBox="[151,1435,189,1544]" captionTargetId="figure-17@15.[151,1435,181,1550]" captionTargetPageId="15" captionText="FIGURE 7. Berlesia multisetosa n. sp., female. A, idiosoma dorsally; B, idiosoma ventrally; C, subcapitulum; D, E, chelicerae, dorsally, in working position, E, enlarged detail; F, peritreme; G, sperm access system; H, J, variability of gnathotectum outline. A, B, G to same scale, C, D, F, H, J to same scale. Abbreviations: co, corniculus; d lf, dorsal lyrifissure; d s, dorsal seta; id, lyrifissure; gp, glandular pore; lb, labrum; mi, internal mala; p, unidentified pore-like structure; p d, pilus dentilis; ss, salivary stylus; t, tooth; td, tendon; others denote setae." figureDoi="http://doi.org/10.5281/zenodo.4396490" httpUri="https://zenodo.org/record/4396490/files/figure.png" pageId="14" pageNumber="19">Fig. 7E</figureCitation>
|
||
); movable digit (35) with three coarse teeth flared laterally (
|
||
<figureCitation id="13702A0F4F4BFF85FE24AF9EFDFFD437" box="[469,595,1267,1293]" captionStart="FIGURE 7" captionStartId="15.[151,250,1574,1598]" captionTargetBox="[151,1435,189,1544]" captionTargetId="figure-17@15.[151,1435,181,1550]" captionTargetPageId="15" captionText="FIGURE 7. Berlesia multisetosa n. sp., female. A, idiosoma dorsally; B, idiosoma ventrally; C, subcapitulum; D, E, chelicerae, dorsally, in working position, E, enlarged detail; F, peritreme; G, sperm access system; H, J, variability of gnathotectum outline. A, B, G to same scale, C, D, F, H, J to same scale. Abbreviations: co, corniculus; d lf, dorsal lyrifissure; d s, dorsal seta; id, lyrifissure; gp, glandular pore; lb, labrum; mi, internal mala; p, unidentified pore-like structure; p d, pilus dentilis; ss, salivary stylus; t, tooth; td, tendon; others denote setae." figureDoi="http://doi.org/10.5281/zenodo.4396490" httpUri="https://zenodo.org/record/4396490/files/figure.png" pageId="14" pageNumber="19">Figs 7D, E</figureCitation>
|
||
). Subcapitulum with 11 to 13 rows of deutosternal denticles, each similarly narrow, with a single denticle, or any row occasionally with two contiguous denticles (
|
||
<figureCitation id="13702A0F4F4BFF85FB88AE7AFB7FD40B" box="[1145,1235,1303,1329]" captionStart="FIGURE 7" captionStartId="15.[151,250,1574,1598]" captionTargetBox="[151,1435,189,1544]" captionTargetId="figure-17@15.[151,1435,181,1550]" captionTargetPageId="15" captionText="FIGURE 7. Berlesia multisetosa n. sp., female. A, idiosoma dorsally; B, idiosoma ventrally; C, subcapitulum; D, E, chelicerae, dorsally, in working position, E, enlarged detail; F, peritreme; G, sperm access system; H, J, variability of gnathotectum outline. A, B, G to same scale, C, D, F, H, J to same scale. Abbreviations: co, corniculus; d lf, dorsal lyrifissure; d s, dorsal seta; id, lyrifissure; gp, glandular pore; lb, labrum; mi, internal mala; p, unidentified pore-like structure; p d, pilus dentilis; ss, salivary stylus; t, tooth; td, tendon; others denote setae." figureDoi="http://doi.org/10.5281/zenodo.4396490" httpUri="https://zenodo.org/record/4396490/files/figure.png" pageId="14" pageNumber="19">Fig. 7C</figureCitation>
|
||
); subcapitular setae
|
||
<emphasis id="B93FEA984F4BFF85FF31AE56FF47D46F" box="[192,235,1339,1365]" italics="true" pageId="14" pageNumber="19">hp1</emphasis>
|
||
(60–63) and slightly longer
|
||
<emphasis id="B93FEA984F4BFF85FDC1AE56FDF7D46F" box="[560,603,1339,1365]" italics="true" pageId="14" pageNumber="19">hp3</emphasis>
|
||
(75–80) similarly attenuated,
|
||
<emphasis id="B93FEA984F4BFF85FC43AE56FC70D46F" box="[946,988,1339,1365]" italics="true" pageId="14" pageNumber="19">hp2</emphasis>
|
||
(20–25) and capitular setae
|
||
<emphasis id="B93FEA984F4BFF85FAD0AE56FA97D46E" box="[1313,1339,1339,1364]" italics="true" pageId="14" pageNumber="19">pc</emphasis>
|
||
(22–27) similarly short, but
|
||
<emphasis id="B93FEA984F4BFF85FE83AE32FE20D442" box="[370,396,1375,1400]" italics="true" pageId="14" pageNumber="19">pc</emphasis>
|
||
strongly bulbiform basally. Corniculi somewhat hyaline, blade-like, bluntly pointed apically, moderately short (35–38), closely adjacent to similarly short tubular salivary styli (ca 37–40) and longer, slender, finely fimbriated internal malae (87–90). Palp length 150–162; palptrochanter setae dissimilar in length,
|
||
<emphasis id="B93FEA984F4BFF85FADFAECAFAE4D4FB" box="[1326,1352,1447,1473]" italics="true" pageId="14" pageNumber="19">v1</emphasis>
|
||
22–25,
|
||
<emphasis id="B93FEA984F4BFF85FF66AEA6FF1ED4DF" box="[151,178,1483,1509]" italics="true" pageId="14" pageNumber="19">v2</emphasis>
|
||
38–52; palpus with femoral seta
|
||
<emphasis id="B93FEA984F4BFF85FDD6AEA6FDF8D4DF" box="[551,596,1483,1509]" italics="true" pageId="14" pageNumber="19">al-1</emphasis>
|
||
and genual setae
|
||
<emphasis id="B93FEA984F4BFF85FCEBAEA6FCE4D4DF" box="[794,840,1483,1509]" italics="true" pageId="14" pageNumber="19">al-1</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F4BFF85FCA4AEA6FC2FD4DF" box="[853,899,1483,1509]" italics="true" pageId="14" pageNumber="19">al-2</emphasis>
|
||
blunt, not spatulate; palpfemoral seta
|
||
<emphasis id="B93FEA984F4BFF85FADEAEA6FA91D4DF" box="[1327,1341,1483,1509]" italics="true" pageId="14" pageNumber="19">d</emphasis>
|
||
(30–32) slightly spine-like, shorter than femur length; palptarsal apotele two-tined.
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F4BFF85FF36AD7EFDD0D64B" blockId="14.[151,1437,150,1905]" pageId="14" pageNumber="19">
|
||
<emphasis id="B93FEA984F4BFF85FF36AD7EFEE4D717" box="[199,328,1555,1581]" italics="true" pageId="14" pageNumber="19">Adult male.</emphasis>
|
||
Gnathotectum with simple, triangular anterior margin much as in female, but sides acutely angular, tapering to single median projection (
|
||
<figureCitation id="13702A0F4F4BFF85FDCFAD5AFD3AD76B" box="[574,662,1591,1617]" captionStart="FIGURE 9" captionStartId="18.[151,250,1882,1906]" captionTargetBox="[167,1423,189,1855]" captionTargetId="figure-21@18.[164,1423,181,1858]" captionTargetPageId="18" captionText="FIGURE 9. Berlesia multisetosa n. sp., male. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, D, chelicera and spermatodactyl, C, dorsal aspect, D, ventral; E, gnathotectum; F, subcapitulum and palpus; G, H, palp tarsus, G, ventral aspect, H, dorsal, enlarged; J, peritreme.A, B to same scale, C–G to same scale, H, J to same scale.Abbreviations: ar, arthrodial envelope; f d, fixed digit; m d, movable digit; others as in Fig. 7." figureDoi="http://doi.org/10.5281/zenodo.4396494" httpUri="https://zenodo.org/record/4396494/files/figure.png" pageId="14" pageNumber="19">Fig. 9E</figureCitation>
|
||
). Cheliceral shaft, excluding short basal section 120–125 long; fixed digit straight, short (30–42) more slender and pointed than in female; movable digit bluntly pointed beyond apex of fixed digit, reduced, amalgamated into spermatodactyl base which extends anteroventrally; spermatodactyl greatly elongated, sinuous, gradually tapering, 425–465 long, without elaborations (
|
||
<figureCitation id="13702A0F4F4BFF85FC05ADCEFBD9D787" box="[1012,1141,1699,1725]" captionStart="FIGURE 9" captionStartId="18.[151,250,1882,1906]" captionTargetBox="[167,1423,189,1855]" captionTargetId="figure-21@18.[164,1423,181,1858]" captionTargetPageId="18" captionText="FIGURE 9. Berlesia multisetosa n. sp., male. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, D, chelicera and spermatodactyl, C, dorsal aspect, D, ventral; E, gnathotectum; F, subcapitulum and palpus; G, H, palp tarsus, G, ventral aspect, H, dorsal, enlarged; J, peritreme.A, B to same scale, C–G to same scale, H, J to same scale.Abbreviations: ar, arthrodial envelope; f d, fixed digit; m d, movable digit; others as in Fig. 7." figureDoi="http://doi.org/10.5281/zenodo.4396494" httpUri="https://zenodo.org/record/4396494/files/figure.png" pageId="14" pageNumber="19">Figs 9C, D</figureCitation>
|
||
). Unlike female, corniculi shorter (22–25), more truncated apically, greatly exceeded by elongated salivary styli (63–70), and capitular setae
|
||
<emphasis id="B93FEA984F4BFF85FF66AD86FF1DD63E" box="[151,177,1771,1796]" italics="true" pageId="14" pageNumber="19">pc</emphasis>
|
||
nearly setiform, slightly bulbous basally. Form and length of other gnathosomatic structures much as in female: internal malae (120–130); setae
|
||
<emphasis id="B93FEA984F4BFF85FDF0AC62FD80D613" box="[513,556,1807,1833]" italics="true" pageId="14" pageNumber="19">hp1</emphasis>
|
||
(62–63) and
|
||
<emphasis id="B93FEA984F4BFF85FD33AC62FD41D613" box="[706,749,1807,1833]" italics="true" pageId="14" pageNumber="19">hp3</emphasis>
|
||
(65–77) much longer than
|
||
<emphasis id="B93FEA984F4BFF85FBD1AC62FBE5D613" box="[1056,1097,1807,1833]" italics="true" pageId="14" pageNumber="19">hp2</emphasis>
|
||
(18–20) and
|
||
<emphasis id="B93FEA984F4BFF85FB10AC62FB57D612" box="[1249,1275,1807,1832]" italics="true" pageId="14" pageNumber="19">pc</emphasis>
|
||
(23–25); deutosternum with 11–12 rows of mostly contiguously doubled denticles (
|
||
<figureCitation id="13702A0F4F4BFF85FC43AC5EFBA4D677" box="[946,1032,1843,1869]" captionStart="FIGURE 9" captionStartId="18.[151,250,1882,1906]" captionTargetBox="[167,1423,189,1855]" captionTargetId="figure-21@18.[164,1423,181,1858]" captionTargetPageId="18" captionText="FIGURE 9. Berlesia multisetosa n. sp., male. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, D, chelicera and spermatodactyl, C, dorsal aspect, D, ventral; E, gnathotectum; F, subcapitulum and palpus; G, H, palp tarsus, G, ventral aspect, H, dorsal, enlarged; J, peritreme.A, B to same scale, C–G to same scale, H, J to same scale.Abbreviations: ar, arthrodial envelope; f d, fixed digit; m d, movable digit; others as in Fig. 7." figureDoi="http://doi.org/10.5281/zenodo.4396494" httpUri="https://zenodo.org/record/4396494/files/figure.png" pageId="14" pageNumber="19">Fig. 9F</figureCitation>
|
||
). Palp length 160–165; palpfemoral seta
|
||
<emphasis id="B93FEA984F4BFF85FF3BAC3AFF74D64B" box="[202,216,1879,1905]" italics="true" pageId="14" pageNumber="19">d</emphasis>
|
||
(25–30) similar in form as in female.
|
||
</paragraph>
|
||
<caption id="DF3466024F4AFF84FF66AD4BFDD8D7F7" ID-DOI="http://doi.org/10.5281/zenodo.4396490" ID-Zenodo-Dep="4396490" httpUri="https://zenodo.org/record/4396490/files/figure.png" pageId="15" pageNumber="20" startId="15.[151,250,1574,1598]" targetBox="[151,1435,189,1544]" targetPageId="15">
|
||
<paragraph id="8BF4368A4F4AFF84FF66AD4BFDD8D7F7" blockId="15.[151,1437,1574,1742]" pageId="15" pageNumber="20">
|
||
<emphasis id="B93FEA984F4AFF84FF66AD4BFEBED704" bold="true" box="[151,274,1574,1598]" pageId="15" pageNumber="20">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C4B4D094F4AFF84FEE9AD4BFE49D704" authority="Lindquist & Oconnor & Shaw & Sidorchuk, 2020" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[280,485,1574,1598]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="20" phylum="Arthropoda" rank="species" species="multisetosa" status="sp. nov.">
|
||
<emphasis id="B93FEA984F4AFF84FEE9AD4BFE49D704" box="[280,485,1574,1598]" italics="true" pageId="15" pageNumber="20">Berlesia multisetosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA984F4AFF84FE1AAD4BFD8FD704" bold="true" box="[491,547,1574,1598]" pageId="15" pageNumber="20">
|
||
<taxonomicNameLabel id="A20C57E34F4AFF84FE1AAD4BFD8FD704" box="[491,547,1574,1598]" pageId="15" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female. A, idiosoma dorsally; B, idiosoma ventrally; C, subcapitulum; D, E, chelicerae, dorsally, in working position, E, enlarged detail; F, peritreme; G, sperm access system; H, J, variability of gnathotectum outline. A, B, G to same scale, C, D, F, H, J to same scale. Abbreviations:
|
||
<emphasis id="B93FEA984F4AFF84FCBBAD03FCCED7BC" bold="true" box="[842,866,1646,1670]" pageId="15" pageNumber="20">co</emphasis>
|
||
, corniculus;
|
||
<emphasis id="B93FEA984F4AFF84FC19AD03FBA1D7BC" bold="true" box="[1000,1037,1646,1670]" pageId="15" pageNumber="20">d lf</emphasis>
|
||
, dorsal lyrifissure;
|
||
<emphasis id="B93FEA984F4AFF84FB26AD03FB5BD7BC" bold="true" box="[1239,1271,1646,1670]" pageId="15" pageNumber="20">d s</emphasis>
|
||
, dorsal seta;
|
||
<emphasis id="B93FEA984F4AFF84FA70AD03FA3AD7BC" bold="true" box="[1409,1430,1646,1670]" pageId="15" pageNumber="20">id</emphasis>
|
||
, lyrifissure;
|
||
<emphasis id="B93FEA984F4AFF84FEFDADFFFE8BD790" bold="true" box="[268,295,1682,1706]" pageId="15" pageNumber="20">gp</emphasis>
|
||
, glandular pore;
|
||
<emphasis id="B93FEA984F4AFF84FE25ADFFFE45D790" bold="true" box="[468,489,1682,1706]" pageId="15" pageNumber="20">lb</emphasis>
|
||
, labrum;
|
||
<emphasis id="B93FEA984F4AFF84FDBDADFFFDC4D790" bold="true" box="[588,616,1682,1706]" pageId="15" pageNumber="20">mi</emphasis>
|
||
, internal mala;
|
||
<emphasis id="B93FEA984F4AFF84FCF6ADFFFCB9D790" bold="true" box="[775,789,1682,1706]" pageId="15" pageNumber="20">p</emphasis>
|
||
, unidentified pore-like structure;
|
||
<emphasis id="B93FEA984F4AFF84FB9DADFFFB22D790" bold="true" box="[1132,1166,1682,1706]" pageId="15" pageNumber="20">p d</emphasis>
|
||
, pilus dentilis;
|
||
<emphasis id="B93FEA984F4AFF84FADAADFFFA93D790" bold="true" box="[1323,1343,1682,1706]" pageId="15" pageNumber="20">ss</emphasis>
|
||
, salivary stylus;
|
||
<emphasis id="B93FEA984F4AFF84FF11ADDBFF44D7F4" bold="true" box="[224,232,1718,1742]" pageId="15" pageNumber="20">t</emphasis>
|
||
, tooth;
|
||
<emphasis id="B93FEA984F4AFF84FEC7ADDBFEE1D7F4" bold="true" box="[310,333,1718,1742]" pageId="15" pageNumber="20">td</emphasis>
|
||
, tendon; others denote setae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF4368A4F4AFF9BFF36AC7EFA23D302" blockId="15.[151,1437,1811,2015]" lastBlockId="16.[151,1437,151,568]" lastPageId="16" lastPageNumber="21" pageId="15" pageNumber="20">
|
||
<emphasis id="B93FEA984F4AFF84FF36AC7EFE37D617" box="[199,411,1811,1837]" italics="true" pageId="15" pageNumber="20">
|
||
<emphasis id="B93FEA984F4AFF84FF36AC7EFEA9D617" bold="true" box="[199,261,1811,1837]" italics="true" pageId="15" pageNumber="20">Legs.</emphasis>
|
||
Adult female
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F4AFF84FE5BAC7EFE43D617" box="[426,495,1811,1837]" captionStart="FIGURE 8" captionStartId="16.[151,250,1467,1491]" captionTargetBox="[152,1431,595,1440]" captionTargetId="figure-267@16.[151,1436,591,1443]" captionTargetPageId="16" captionText="FIGURE 8. Berlesia multisetosa n. sp., female, legs.A, C, F, G, legs dorsally except separated coxae ventrally:A, leg I, subapical tarsal setae shown with circles; B, tarsus I dorsally; C, detail of sensory cluster on tarsus I; D–G, legs dorsally except separated coxae ventrally: D, leg II; E, detail of tarsus II tip ventrally; F, leg III, tarsus partly shown; G, leg IV, tarsus partly shown; H, tarsus IV ventrally. A, D–H to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396492" httpUri="https://zenodo.org/record/4396492/files/figure.png" pageId="15" pageNumber="20">Figs 8</figureCitation>
|
||
A–H). Legs I (425–450) nearly as long as legs IV (460–470), and about 0.8 as long as dorsal shield; other leg lengths (excluding pretarsi): II (375–380), III (385–400). Leg I pretarsus (37–38) to apex of pulvillus, tarsus (115–125) about 1.7–2.1 longer than each of the similarly long femur (67–70), genu (53–63), and tibia (60–65). Legs II–IV with tarsus/tibia length ratios about 1.8–2.3. Complement of setae on leg segments holotrichous for femora II-III-IV (11-6-6) as presented for the genus, but femur I with 11 setae, lacking
|
||
<emphasis id="B93FEA984F4AFF84FAEEACCCFAFDD681" box="[1311,1361,1953,1979]" italics="true" pageId="15" pageNumber="20">pv-2</emphasis>
|
||
; chaetotactic formulae for genua I-II-III-IV: 11 (2-3/1, 2/1-2), 11 (2-3/1, 2/1-2), 9 (2-2/1, 2/1-1), 10 (2-2/1, 3/1-1); tibiae: 11 (2-3/1, 2/1-2), 10 (2-2/1, 2/1-2), 8 (2-1/1, 2/1-1), 10 (2-1/1, 3/1-2); genu I and tibia I without
|
||
<emphasis id="B93FEA984F55FF9BFB33ABFAFB59D18B" box="[1218,1269,151,177]" italics="true" pageId="16" pageNumber="21">pd-3</emphasis>
|
||
and
|
||
<emphasis id="B93FEA984F55FF9BFADBABFAFAF0D18B" box="[1322,1372,151,177]" italics="true" pageId="16" pageNumber="21">av-2</emphasis>
|
||
; tibia III without
|
||
<emphasis id="B93FEA984F55FF9BFEEBABD7FEEBD1EE" box="[282,327,186,212]" italics="true" pageId="16" pageNumber="21">pl-2</emphasis>
|
||
. Tarsus I with seta
|
||
<emphasis id="B93FEA984F55FF9BFDD6ABD7FDE5D1EE" box="[551,585,186,212]" italics="true" pageId="16" pageNumber="21">md</emphasis>
|
||
(55–60) longer than other setae on basal half of segment (
|
||
<figureCitation id="13702A0F4F55FF9BFB19ABD7FAEDD1EE" box="[1256,1345,186,212]" captionStart="FIGURE 8" captionStartId="16.[151,250,1467,1491]" captionTargetBox="[152,1431,595,1440]" captionTargetId="figure-267@16.[151,1436,591,1443]" captionTargetPageId="16" captionText="FIGURE 8. Berlesia multisetosa n. sp., female, legs.A, C, F, G, legs dorsally except separated coxae ventrally:A, leg I, subapical tarsal setae shown with circles; B, tarsus I dorsally; C, detail of sensory cluster on tarsus I; D–G, legs dorsally except separated coxae ventrally: D, leg II; E, detail of tarsus II tip ventrally; F, leg III, tarsus partly shown; G, leg IV, tarsus partly shown; H, tarsus IV ventrally. A, D–H to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396492" httpUri="https://zenodo.org/record/4396492/files/figure.png" pageId="16" pageNumber="21">Fig. 8B</figureCitation>
|
||
); dorsal subapical cluster of sensory structures with a cluster of three, tightly adjacent, tiny, pointed setae surrounded by three solenidia, one of which biramous (
|
||
<figureCitation id="13702A0F4F55FF9BFD97AA6CFD6CD021" box="[614,704,257,283]" captionStart="FIGURE 8" captionStartId="16.[151,250,1467,1491]" captionTargetBox="[152,1431,595,1440]" captionTargetId="figure-267@16.[151,1436,591,1443]" captionTargetPageId="16" captionText="FIGURE 8. Berlesia multisetosa n. sp., female, legs.A, C, F, G, legs dorsally except separated coxae ventrally:A, leg I, subapical tarsal setae shown with circles; B, tarsus I dorsally; C, detail of sensory cluster on tarsus I; D–G, legs dorsally except separated coxae ventrally: D, leg II; E, detail of tarsus II tip ventrally; F, leg III, tarsus partly shown; G, leg IV, tarsus partly shown; H, tarsus IV ventrally. A, D–H to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396492" httpUri="https://zenodo.org/record/4396492/files/figure.png" pageId="16" pageNumber="21">Fig. 8C</figureCitation>
|
||
). Tarsi II–IV with a finely serrated ventroapical ridge basal to a short, rounded process (
|
||
<figureCitation id="13702A0F4F55FF9BFE59AA49FD82D005" box="[424,558,292,319]" captionStart="FIGURE 8" captionStartId="16.[151,250,1467,1491]" captionTargetBox="[152,1431,595,1440]" captionTargetId="figure-267@16.[151,1436,591,1443]" captionTargetPageId="16" captionText="FIGURE 8. Berlesia multisetosa n. sp., female, legs.A, C, F, G, legs dorsally except separated coxae ventrally:A, leg I, subapical tarsal setae shown with circles; B, tarsus I dorsally; C, detail of sensory cluster on tarsus I; D–G, legs dorsally except separated coxae ventrally: D, leg II; E, detail of tarsus II tip ventrally; F, leg III, tarsus partly shown; G, leg IV, tarsus partly shown; H, tarsus IV ventrally. A, D–H to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396492" httpUri="https://zenodo.org/record/4396492/files/figure.png" pageId="16" pageNumber="21">Figs. 8E, H</figureCitation>
|
||
), and with dorsoapical setae (
|
||
<emphasis id="B93FEA984F55FF9BFC72AA49FC05D004" box="[899,937,292,318]" italics="true" pageId="16" pageNumber="21">d-1</emphasis>
|
||
) (12–15) nearly half as long as pretarsus to base of claws (ca 25). Coxal seta
|
||
<emphasis id="B93FEA984F55FF9BFDE3AA24FD80D058" box="[530,556,329,354]" italics="true" pageId="16" pageNumber="21">av</emphasis>
|
||
on leg I and
|
||
<emphasis id="B93FEA984F55FF9BFD33AA24FD70D058" box="[706,732,329,354]" italics="true" pageId="16" pageNumber="21">pv</emphasis>
|
||
on legs II, III strongly bulbiform basally, like subcapitular
|
||
<emphasis id="B93FEA984F55FF9BFA8BAA24FA38D058" box="[1402,1428,329,354]" italics="true" pageId="16" pageNumber="21">pc</emphasis>
|
||
; coxa IV seta
|
||
<emphasis id="B93FEA984F55FF9BFEDBAA01FE9AD0BF" box="[298,310,364,389]" italics="true" pageId="16" pageNumber="21">v</emphasis>
|
||
setiform. Trochanters I–IV with basal seta
|
||
<emphasis id="B93FEA984F55FF9BFCEAAA01FCE1D0BC" box="[795,845,364,390]" italics="true" pageId="16" pageNumber="21">pv-2</emphasis>
|
||
attenuated (50–55), and III–IV
|
||
<emphasis id="B93FEA984F55FF9BFB43AA06FB62D0BF" box="[1202,1230,363,389]" italics="true" pageId="16" pageNumber="21">ad</emphasis>
|
||
slightly spinelike. Tarsus IV with apical seta
|
||
<emphasis id="B93FEA984F55FF9BFE4AAAE2FE41D093" box="[443,493,399,425]" italics="true" pageId="16" pageNumber="21">pv-1</emphasis>
|
||
asymmetrically shorter, bluntly spine-like, than attenuated
|
||
<emphasis id="B93FEA984F55FF9BFB8EAAE2FB1DD093" box="[1151,1201,399,425]" italics="true" pageId="16" pageNumber="21">av-1</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F55FF9BFB4EAAE2FAB5D093" box="[1215,1305,399,425]" captionStart="FIGURE 8" captionStartId="16.[151,250,1467,1491]" captionTargetBox="[152,1431,595,1440]" captionTargetId="figure-267@16.[151,1436,591,1443]" captionTargetPageId="16" captionText="FIGURE 8. Berlesia multisetosa n. sp., female, legs.A, C, F, G, legs dorsally except separated coxae ventrally:A, leg I, subapical tarsal setae shown with circles; B, tarsus I dorsally; C, detail of sensory cluster on tarsus I; D–G, legs dorsally except separated coxae ventrally: D, leg II; E, detail of tarsus II tip ventrally; F, leg III, tarsus partly shown; G, leg IV, tarsus partly shown; H, tarsus IV ventrally. A, D–H to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396492" httpUri="https://zenodo.org/record/4396492/files/figure.png" pageId="16" pageNumber="21">Fig. 8H</figureCitation>
|
||
). Legs I–IV each with particular setae enlarged, spine-like: I Fe
|
||
<emphasis id="B93FEA984F55FF9BFD20AADFFCA8D0F6" box="[721,772,434,460]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(65),
|
||
<emphasis id="B93FEA984F55FF9BFCB4AADFFCD5D0F6" box="[837,889,434,460]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(45); I Ge
|
||
<emphasis id="B93FEA984F55FF9BFC01AADFFB8FD0F6" box="[1008,1059,434,460]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(50–60),
|
||
<emphasis id="B93FEA984F55FF9BFB7EAADFFB6ED0F6" box="[1167,1218,434,460]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(55–58); II Fe
|
||
<emphasis id="B93FEA984F55FF9BFA98AADFFA30D0F6" box="[1385,1436,434,460]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(57–63),
|
||
<emphasis id="B93FEA984F55FF9BFF0FAABBFE9ED0CA" box="[254,306,470,496]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(47–48); II Ge
|
||
<emphasis id="B93FEA984F55FF9BFE15AABBFDBBD0CA" box="[484,535,470,496]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(50–52),
|
||
<emphasis id="B93FEA984F55FF9BFD77AABBFD07D0CA" box="[646,683,470,496]" italics="true" pageId="16" pageNumber="21">pd-</emphasis>
|
||
1 (47–48); II Ti
|
||
<emphasis id="B93FEA984F55FF9BFC93AABBFC39D0CA" box="[866,917,470,496]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
not differentiated; III Fe
|
||
<emphasis id="B93FEA984F55FF9BFB44AABBFB44D0CA" box="[1205,1256,470,496]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(52–54); III Ge
|
||
<emphasis id="B93FEA984F55FF9BFF66AA94FF66D329" box="[151,202,505,531]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(57–62),
|
||
<emphasis id="B93FEA984F55FF9BFEC7AA94FEC5D329" box="[310,361,505,531]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(50–52); III Ti
|
||
<emphasis id="B93FEA984F55FF9BFDE4AA94FD9DD329" box="[533,561,505,531]" italics="true" pageId="16" pageNumber="21">ad</emphasis>
|
||
(42),
|
||
<emphasis id="B93FEA984F55FF9BFD83AA94FD0AD329" box="[626,678,505,531]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(30); IV Fe
|
||
<emphasis id="B93FEA984F55FF9BFCDCAA94FCCCD329" box="[813,864,505,531]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(57–62),
|
||
<emphasis id="B93FEA984F55FF9BFC3DAA97FC4AD329" box="[972,998,506,531]" italics="true" pageId="16" pageNumber="21">pv</emphasis>
|
||
not differentiated; IV Ge
|
||
<emphasis id="B93FEA984F55FF9BFAF2AA94FA9AD329" box="[1283,1334,505,531]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(66–68),
|
||
<emphasis id="B93FEA984F55FF9BFF66A970FF67D30D" box="[151,203,541,567]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(62–65); IV Ti
|
||
<emphasis id="B93FEA984F55FF9BFE8AA970FE02D30D" box="[379,430,541,567]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(53–60),
|
||
<emphasis id="B93FEA984F55FF9BFDEDA970FDFCD30D" box="[540,592,541,567]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(55–60). Relative lengths of other leg setae as shown in
|
||
<figureCitation id="13702A0F4F55FF9BFB3DA970FAE6D302" box="[1228,1354,541,568]" captionStart="FIGURE 8" captionStartId="16.[151,250,1467,1491]" captionTargetBox="[152,1431,595,1440]" captionTargetId="figure-267@16.[151,1436,591,1443]" captionTargetPageId="16" captionText="FIGURE 8. Berlesia multisetosa n. sp., female, legs.A, C, F, G, legs dorsally except separated coxae ventrally:A, leg I, subapical tarsal setae shown with circles; B, tarsus I dorsally; C, detail of sensory cluster on tarsus I; D–G, legs dorsally except separated coxae ventrally: D, leg II; E, detail of tarsus II tip ventrally; F, leg III, tarsus partly shown; G, leg IV, tarsus partly shown; H, tarsus IV ventrally. A, D–H to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396492" httpUri="https://zenodo.org/record/4396492/files/figure.png" pageId="16" pageNumber="21">Figs 8A, D</figureCitation>
|
||
, F–H.
|
||
</paragraph>
|
||
<caption id="DF3466024F55FF9BFF66AED6FDCDD705" ID-DOI="http://doi.org/10.5281/zenodo.4396492" ID-Zenodo-Dep="4396492" httpUri="https://zenodo.org/record/4396492/files/figure.png" pageId="16" pageNumber="21" startId="16.[151,250,1467,1491]" targetBox="[152,1431,595,1440]" targetPageId="16">
|
||
<paragraph id="8BF4368A4F55FF9BFF66AED6FDCDD705" blockId="16.[151,1437,1467,1599]" pageId="16" pageNumber="21">
|
||
<emphasis id="B93FEA984F55FF9BFF66AED6FEBED4E9" bold="true" box="[151,274,1467,1491]" pageId="16" pageNumber="21">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C4B4D094F55FF9BFEE9AED6FE49D4E9" authority="Lindquist & Oconnor & Shaw & Sidorchuk, 2020" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[280,485,1467,1491]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="21" phylum="Arthropoda" rank="species" species="multisetosa" status="sp. nov.">
|
||
<emphasis id="B93FEA984F55FF9BFEE9AED6FE49D4E9" box="[280,485,1467,1491]" italics="true" pageId="16" pageNumber="21">Berlesia multisetosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA984F55FF9BFE1BAED6FD8FD4E9" bold="true" box="[490,547,1467,1491]" pageId="16" pageNumber="21">
|
||
<taxonomicNameLabel id="A20C57E34F55FF9BFE1BAED6FD8FD4E9" box="[490,547,1467,1491]" pageId="16" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female, legs. A, C, F, G, legs dorsally except separated coxae ventrally: A, leg I, subapical tarsal setae shown with circles; B, tarsus I dorsally; C, detail of sensory cluster on tarsus I; D–G, legs dorsally except separated coxae ventrally: D, leg II; E, detail of tarsus II tip ventrally; F, leg III, tarsus partly shown; G, leg IV, tarsus partly shown; H, tarsus IV ventrally. A, D–H to same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF4368A4F55FF9AFF36ADE9FCB1D1EF" blockId="16.[151,1437,1668,2012]" lastBlockId="17.[151,1437,150,2009]" lastPageId="17" lastPageNumber="22" pageId="16" pageNumber="21">
|
||
<emphasis id="B93FEA984F55FF9BFF36ADE9FEEED7A4" box="[199,322,1668,1694]" italics="true" pageId="16" pageNumber="21">Adult male</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F55FF9BFEA2ADE9FE04D7A4" box="[339,424,1668,1694]" captionStart="FIGURE 10" captionStartId="19.[151,250,1104,1128]" captionTargetBox="[155,1432,184,1077]" captionTargetId="figure-17@19.[151,1436,181,1081]" captionTargetPageId="19" captionText="FIGURE 10. Berlesia multisetosa n. sp., male, legs, coxae detached. A, leg I ventrally, most setae of tarsal subapex, beyond transverse dotted line, omitted; B, leg II ventrally; C, leg III ventrally; D, leg IV ventrolaterally. All to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396496" httpUri="https://zenodo.org/record/4396496/files/figure.png" pageId="16" pageNumber="21">Figs 10</figureCitation>
|
||
A–D). Legs similar in relative dimensions to those of female; lengths I 415–430, II 360– 378, III 375–400, IV 440–470. Legs I and IV without dimorphism in thickness of segments, legs II and III thickersegmented than in female. Leg I with form and relative sizes of setae similar to those of female, except femur I with 12 setae, including
|
||
<emphasis id="B93FEA984F55FF9BFE59AD83FE76D632" box="[424,474,1774,1800]" italics="true" pageId="16" pageNumber="21">pv-2</emphasis>
|
||
, and with
|
||
<emphasis id="B93FEA984F55FF9BFDBEAD83FDC5D63D" box="[591,617,1774,1799]" italics="true" pageId="16" pageNumber="21">av</emphasis>
|
||
slightly bulbiform. Legs I–IV retain attenuated form of basal seta
|
||
<emphasis id="B93FEA984F55FF9BFABCAD83FAD3D632" box="[1357,1407,1774,1800]" italics="true" pageId="16" pageNumber="21">pv-2</emphasis>
|
||
of trochanter. Legs II–IV with particular setae dimorphically modified: leg II Fe
|
||
<emphasis id="B93FEA984F55FF9BFC08AC7FFBBFD611" box="[1017,1043,1810,1835]" italics="true" pageId="16" pageNumber="21">av</emphasis>
|
||
spine-like; Ge
|
||
<emphasis id="B93FEA984F55FF9BFB4FAC7FFB74D611" box="[1214,1240,1810,1835]" italics="true" pageId="16" pageNumber="21">av</emphasis>
|
||
, Ti
|
||
<emphasis id="B93FEA984F55FF9BFAF5AC7FFAB2D611" box="[1284,1310,1810,1835]" italics="true" pageId="16" pageNumber="21">av</emphasis>
|
||
, and tarsus
|
||
<emphasis id="B93FEA984F55FF9BFF66AC58FF14D674" box="[151,184,1845,1870]" italics="true" pageId="16" pageNumber="21">mv</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFF37AC59FF5BD674" box="[198,247,1844,1870]" italics="true" pageId="16" pageNumber="21">av-2</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFEF4AC59FE9AD674" box="[261,310,1844,1870]" italics="true" pageId="16" pageNumber="21">av-1</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFEB4AC59FEDAD674" box="[325,374,1844,1870]" italics="true" pageId="16" pageNumber="21">pv-1</emphasis>
|
||
bulbiform; leg III coxal
|
||
<emphasis id="B93FEA984F55FF9BFD7AAC58FD09D674" box="[651,677,1845,1870]" italics="true" pageId="16" pageNumber="21">pv</emphasis>
|
||
simple; Tr
|
||
<emphasis id="B93FEA984F55FF9BFCD4AC59FCEDD674" box="[805,833,1844,1870]" italics="true" pageId="16" pageNumber="21">ad</emphasis>
|
||
markedly arched; Fe
|
||
<emphasis id="B93FEA984F55FF9BFBC5AC59FBE6D674" box="[1076,1098,1844,1870]" italics="true" pageId="16" pageNumber="21">al</emphasis>
|
||
less spine-like, and tarsus
|
||
<emphasis id="B93FEA984F55FF9BFA84AC58FA3AD674" box="[1397,1430,1845,1870]" italics="true" pageId="16" pageNumber="21">mv</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFF66AC3AFF65D64B" box="[151,201,1879,1905]" italics="true" pageId="16" pageNumber="21">av-2</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFF27AC3AFEA5D64B" box="[214,265,1879,1905]" italics="true" pageId="16" pageNumber="21">pv-2</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFEE7AC3AFEEBD64B" box="[278,327,1879,1905]" italics="true" pageId="16" pageNumber="21">av-1</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFEA4AC3AFE2BD64B" box="[341,391,1879,1905]" italics="true" pageId="16" pageNumber="21">pv-1</emphasis>
|
||
bulbiform; leg IV Tr
|
||
<emphasis id="B93FEA984F55FF9BFD89AC3AFD38D64B" box="[632,660,1879,1905]" italics="true" pageId="16" pageNumber="21">ad</emphasis>
|
||
markedly arched,
|
||
<emphasis id="B93FEA984F55FF9BFC92AC35FCD1D64B" box="[867,893,1880,1905]" italics="true" pageId="16" pageNumber="21">av</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFC7AAC3AFC10D64B" box="[907,956,1879,1905]" italics="true" pageId="16" pageNumber="21">pv-1</emphasis>
|
||
thickened, more spine-like; Fe
|
||
<emphasis id="B93FEA984F55FF9BFAECAC3AFA95D64B" box="[1309,1337,1879,1905]" italics="true" pageId="16" pageNumber="21">pd</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFAB7AC3AFAF0D64B" box="[1350,1372,1879,1905]" italics="true" pageId="16" pageNumber="21">pl</emphasis>
|
||
more spine-like; Ge
|
||
<emphasis id="B93FEA984F55FF9BFEB1AC17FEDFD6AE" box="[320,371,1914,1940]" italics="true" pageId="16" pageNumber="21">pd-2</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFE72AC17FE1AD6AE" box="[387,438,1914,1940]" italics="true" pageId="16" pageNumber="21">pd-3</emphasis>
|
||
and Ti
|
||
<emphasis id="B93FEA984F55FF9BFDE0AC17FD92D6AE" box="[529,574,1914,1940]" italics="true" pageId="16" pageNumber="21">pl-1</emphasis>
|
||
larger, more spine-like; tarsus
|
||
<emphasis id="B93FEA984F55FF9BFC53AC16FC78D6AF" box="[930,980,1915,1941]" italics="true" pageId="16" pageNumber="21">av-2</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFC15AC16FBBAD6AF" box="[996,1046,1915,1941]" italics="true" pageId="16" pageNumber="21">pv-2</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFBD7AC16FBFBD6AF" box="[1062,1111,1915,1941]" italics="true" pageId="16" pageNumber="21">av-1</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F55FF9BFB96AC16FB35D6AF" box="[1127,1177,1915,1941]" italics="true" pageId="16" pageNumber="21">pv-1</emphasis>
|
||
bulbiform. Lengths of spine-like setae similarly enlarged as in female: I Fe
|
||
<emphasis id="B93FEA984F55FF9BFD2BACF3FCA1D682" box="[730,781,1950,1976]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(48–62),
|
||
<emphasis id="B93FEA984F55FF9BFC89ACF3FC00D682" box="[888,940,1950,1976]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(37–45); I Ge
|
||
<emphasis id="B93FEA984F55FF9BFBBDACF3FBD3D682" box="[1100,1151,1950,1976]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(45–52),
|
||
<emphasis id="B93FEA984F55FF9BFB1AACF3FAB2D682" box="[1259,1310,1950,1976]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(52–60); II Fe
|
||
<emphasis id="B93FEA984F55FF9BFF4BACACFF41D6E1" box="[186,237,1985,2011]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(55–65),
|
||
<emphasis id="B93FEA984F55FF9BFEABACACFE21D6E1" box="[346,397,1985,2011]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(42–49); II Ge
|
||
<emphasis id="B93FEA984F55FF9BFDCAACACFDC2D6E1" box="[571,622,1985,2011]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(38–45),
|
||
<emphasis id="B93FEA984F55FF9BFD2DACACFCA3D6E1" box="[732,783,1985,2011]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
(50–57); II Ti
|
||
<emphasis id="B93FEA984F55FF9BFC45ACACFC4BD6E1" box="[948,999,1985,2011]" italics="true" pageId="16" pageNumber="21">pd-1</emphasis>
|
||
not differentiated; III Fe
|
||
<emphasis id="B93FEA984F55FF9BFAF3ACACFA99D6E1" box="[1282,1333,1985,2011]" italics="true" pageId="16" pageNumber="21">ad-1</emphasis>
|
||
(40–50); III Ge
|
||
<emphasis id="B93FEA984F54FF9AFF13ABFAFEB9D18B" box="[226,277,151,177]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(43–53),
|
||
<emphasis id="B93FEA984F54FF9AFE75ABFAFE1BD18B" box="[388,439,151,177]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(47–50); III Ti
|
||
<emphasis id="B93FEA984F54FF9AFD99ABFAFD28D18B" box="[616,644,151,177]" italics="true" pageId="17" pageNumber="22">ad</emphasis>
|
||
(35),
|
||
<emphasis id="B93FEA984F54FF9AFD38ABFAFD50D18B" box="[713,764,151,177]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(27); IV Fe
|
||
<emphasis id="B93FEA984F54FF9AFC78ABFAFC10D18B" box="[905,956,151,177]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(57–70),
|
||
<emphasis id="B93FEA984F54FF9AFBDBABFAFBE8D18A" box="[1066,1092,151,176]" italics="true" pageId="17" pageNumber="22">pv</emphasis>
|
||
not differentiated; IV Ge
|
||
<emphasis id="B93FEA984F54FF9AFA98ABFAFA30D18B" box="[1385,1436,151,177]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(67–70),
|
||
<emphasis id="B93FEA984F54FF9AFF0FABD7FE9DD1EE" box="[254,305,186,212]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(67–75); IV Ti
|
||
<emphasis id="B93FEA984F54FF9AFE13ABD7FDB9D1EE" box="[482,533,186,212]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(55–65),
|
||
<emphasis id="B93FEA984F54FF9AFD72ABD7FD1AD1EE" box="[643,694,186,212]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(60–67).
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F54FF9AFF36ABB0FA39D32B" blockId="17.[151,1437,150,2009]" pageId="17" pageNumber="22">
|
||
<emphasis id="B93FEA984F54FF9AFF36ABB0FDF4D1C2" bold="true" box="[199,600,221,248]" pageId="17" pageNumber="22">
|
||
Deutonymph.
|
||
<emphasis id="B93FEA984F54FF9AFE83ABB0FDF4D1C2" bold="true" box="[370,600,221,248]" italics="true" pageId="17" pageNumber="22">Idiosomatic dorsum</emphasis>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F54FF9AFD98ABB0FD7ED1CD" box="[617,722,221,247]" captionStart="FIGURE 11" captionStartId="20.[151,250,1099,1123]" captionTargetBox="[155,1431,185,1068]" captionTargetId="figure-21@20.[151,1436,181,1075]" captionTargetPageId="20" captionText="FIGURE 11. Berlesia multisetosa n. sp., deutonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, apical half of chelicera, ventrally, at resting position; D, variations of peritremes; E–G, variability of gnathotectum outline.A, B to same scale; C, E–G to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396498" httpUri="https://zenodo.org/record/4396498/files/figure.png" pageId="17" pageNumber="22">Fig. 11A</figureCitation>
|
||
). Idiosoma 475–500 long, 362–365 wide between insertions of setae
|
||
<emphasis id="B93FEA984F54FF9AFF26AA6DFF5DD020" box="[215,241,256,282]" italics="true" pageId="17" pageNumber="22">r3</emphasis>
|
||
(n=3); dorsal shields weakly sclerotized, unornamented, faintly divided by transverse strip of soft cuticle between setae
|
||
<emphasis id="B93FEA984F54FF9AFECAAA4EFEFDD007" box="[315,337,291,317]" italics="true" pageId="17" pageNumber="22">j6</emphasis>
|
||
and
|
||
<emphasis id="B93FEA984F54FF9AFE79AA49FE0ED004" box="[392,418,292,318]" italics="true" pageId="17" pageNumber="22">J1</emphasis>
|
||
. Podonotal shield 210–225 long, ca 215–225 wide at its greatest width at level of setae
|
||
<emphasis id="B93FEA984F54FF9AFA8DAA49FA39D004" box="[1404,1429,292,318]" italics="true" pageId="17" pageNumber="22">s5</emphasis>
|
||
, with ca 12 pairs of setae (
|
||
<emphasis id="B93FEA984F54FF9AFE49AA2AFD53D05A" box="[440,767,326,353]" italics="true" pageId="17" pageNumber="22">j1, j3–j6, z1, z2, z4. z5, s4–s6</emphasis>
|
||
); opisthonotal shield ca 175–195 long and 175–205 at widest level near setae
|
||
<emphasis id="B93FEA984F54FF9AFE85AA07FE3ED0BE" box="[372,402,362,388]" italics="true" pageId="17" pageNumber="22">Z2</emphasis>
|
||
, with nine pairs of setae (
|
||
<emphasis id="B93FEA984F54FF9AFD48AA07FCF9D0BE" box="[697,853,362,388]" italics="true" pageId="17" pageNumber="22">J1–J4, Z1–Z5</emphasis>
|
||
). Soft cuticle with ca 25 pairs of dorsolateral setae, including uncertain homologies of the
|
||
<emphasis id="B93FEA984F54FF9AFDB2AAE3FDE2D09D" box="[579,590,398,423]" italics="true" pageId="17" pageNumber="22">s</emphasis>
|
||
-and
|
||
<emphasis id="B93FEA984F54FF9AFD77AAE0FD38D09D" box="[646,660,397,423]" italics="true" pageId="17" pageNumber="22">S</emphasis>
|
||
-series among hypertrichous elements of the marginal
|
||
<emphasis id="B93FEA984F54FF9AFB1CAAE3FABBD09D" box="[1261,1303,398,423]" italics="true" pageId="17" pageNumber="22">r–R</emphasis>
|
||
-series (
|
||
<figureCitation id="13702A0F4F54FF9AFA9EAAE0FF66D0F0" captionStart="FIGURE 11" captionStartId="20.[151,250,1099,1123]" captionTargetBox="[155,1431,185,1068]" captionTargetId="figure-21@20.[151,1436,181,1075]" captionTargetPageId="20" captionText="FIGURE 11. Berlesia multisetosa n. sp., deutonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, apical half of chelicera, ventrally, at resting position; D, variations of peritremes; E–G, variability of gnathotectum outline.A, B to same scale; C, E–G to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396498" httpUri="https://zenodo.org/record/4396498/files/figure.png" pageId="17" pageNumber="22">Fig. 11A</figureCitation>
|
||
). Form and relative lengths of dorsal setae more homologous than in adult, many of them slightly barbed as shown in
|
||
<figureCitation id="13702A0F4F54FF9AFEF6AABEFEC2D0D4" box="[263,366,467,494]" captionStart="FIGURE 11" captionStartId="20.[151,250,1099,1123]" captionTargetBox="[155,1431,185,1068]" captionTargetId="figure-21@20.[151,1436,181,1075]" captionTargetPageId="20" captionText="FIGURE 11. Berlesia multisetosa n. sp., deutonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, apical half of chelicera, ventrally, at resting position; D, variations of peritremes; E–G, variability of gnathotectum outline.A, B to same scale; C, E–G to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396498" httpUri="https://zenodo.org/record/4396498/files/figure.png" pageId="17" pageNumber="22">Fig. 11A</figureCitation>
|
||
; other than
|
||
<emphasis id="B93FEA984F54FF9AFE05AAB9FDA1D0D4" box="[500,525,468,494]" italics="true" pageId="17" pageNumber="22">z1</emphasis>
|
||
clearly shortest (18–28), setal lengths range from
|
||
<emphasis id="B93FEA984F54FF9AFBBAAAB9FBCCD0D4" box="[1099,1120,468,494]" italics="true" pageId="17" pageNumber="22">j1</emphasis>
|
||
(35–40) to
|
||
<emphasis id="B93FEA984F54FF9AFB19AAB9FAA9D0D4" box="[1256,1285,468,494]" italics="true" pageId="17" pageNumber="22">Z5</emphasis>
|
||
longest (65). Peritreme similarly short (length 37–38, including stigma diameter 15–17) and embellished as in adult (
|
||
<figureCitation id="13702A0F4F54FF9AFAD0AA9AFA2BD32B" box="[1313,1415,503,529]" captionStart="FIGURE 11" captionStartId="20.[151,250,1099,1123]" captionTargetBox="[155,1431,185,1068]" captionTargetId="figure-21@20.[151,1436,181,1075]" captionTargetPageId="20" captionText="FIGURE 11. Berlesia multisetosa n. sp., deutonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, apical half of chelicera, ventrally, at resting position; D, variations of peritremes; E–G, variability of gnathotectum outline.A, B to same scale; C, E–G to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396498" httpUri="https://zenodo.org/record/4396498/files/figure.png" pageId="17" pageNumber="22">Fig. 11D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F54FF9AFF36A974FE09D277" blockId="17.[151,1437,150,2009]" pageId="17" pageNumber="22">
|
||
<emphasis id="B93FEA984F54FF9AFF36A974FE32D30E" bold="true" box="[199,414,537,564]" italics="true" pageId="17" pageNumber="22">Idiosomatic venter</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F54FF9AFE5CA977FDBFD30E" box="[429,531,538,564]" captionStart="FIGURE 11" captionStartId="20.[151,250,1099,1123]" captionTargetBox="[155,1431,185,1068]" captionTargetId="figure-21@20.[151,1436,181,1075]" captionTargetPageId="20" captionText="FIGURE 11. Berlesia multisetosa n. sp., deutonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, apical half of chelicera, ventrally, at resting position; D, variations of peritremes; E–G, variability of gnathotectum outline.A, B to same scale; C, E–G to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396498" httpUri="https://zenodo.org/record/4396498/files/figure.png" pageId="17" pageNumber="22">Fig. 11B</figureCitation>
|
||
). Tritosternum similar in form to that of adult, with laciniae fused for only about 5–10 of entire length (87–92 excluding base). Presternal area devoid of ornamentation. Intercoxal shield 182–200 long, weakly sclerotized, bearing five pairs of setae and three pairs of poroids, seta
|
||
<emphasis id="B93FEA984F54FF9AFBCFA90CFBF3D341" box="[1086,1119,609,635]" italics="true" pageId="17" pageNumber="22">st5</emphasis>
|
||
sometimes barely off shield edge; setae
|
||
<emphasis id="B93FEA984F54FF9AFEEAA9E9FEC7D3A4" box="[283,363,644,670]" italics="true" pageId="17" pageNumber="22">st1–st4</emphasis>
|
||
(43–50) slightly shorter than
|
||
<emphasis id="B93FEA984F54FF9AFD4DA9E9FD70D3A4" box="[700,732,644,670]" italics="true" pageId="17" pageNumber="22">st5</emphasis>
|
||
(52–57); paragenital poroids on soft cuticle flanking posterior edge of shield. Opisthogaster with oboval, smooth anal shield; length 95–100, greatest width 75–80; shield with postanal seta (45–50) about 1.3 longer and stouter than paranal setae (35–37). Soft cuticle with hypertrichous complement and relative lengths of setae flanking anal shield much as in adult, with ca 15 pairs of slender opisthogastric setae flanked by 5–10 pairs of thicker
|
||
<emphasis id="B93FEA984F54FF9AFDB1A87CFDFDD210" box="[576,593,785,810]" italics="true" pageId="17" pageNumber="22">R</emphasis>
|
||
-marginal setae of similar lengths (40–47), the more posterolateral elements of them thicker, blunter.
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F54FF9AFF36A83BFEC4D57E" blockId="17.[151,1437,150,2009]" pageId="17" pageNumber="22">
|
||
<emphasis id="B93FEA984F54FF9AFF36A83BFECED24B" bold="true" box="[199,354,854,881]" italics="true" pageId="17" pageNumber="22">
|
||
<taxonomicName id="4C4B4D094F54FF9AFF36A83BFEF1D24B" authorityName="Krefft" authorityYear="1871" box="[199,349,854,881]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="17" pageNumber="22" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Gnathotectum, chelicerae, palpal structures, and subcapitulum much as in female, except setal pair
|
||
<emphasis id="B93FEA984F54FF9AFF3AA816FF49D2AE" box="[203,229,891,916]" italics="true" pageId="17" pageNumber="22">pc</emphasis>
|
||
simple. Cheliceral shaft, excluding basal section, 132–138 long; movable digit 28–35 long. Subcapitulum with 11 to 13 rows of deutosternal denticles formed as in female; subcapitular setae
|
||
<emphasis id="B93FEA984F54FF9AFBCDA8F0FBCAD28D" box="[1084,1126,925,951]" italics="true" pageId="17" pageNumber="22">hp1</emphasis>
|
||
(45–50) and slightly longer
|
||
<emphasis id="B93FEA984F54FF9AFF66A8ADFF6ED2E0" box="[151,194,960,986]" italics="true" pageId="17" pageNumber="22">hp3</emphasis>
|
||
(52–58) similarly attenuated,
|
||
<emphasis id="B93FEA984F54FF9AFDEFA8ADFDE4D2E0" box="[542,584,960,986]" italics="true" pageId="17" pageNumber="22">hp2</emphasis>
|
||
(13–15) and capitular setae
|
||
<emphasis id="B93FEA984F54FF9AFC65A8ACFC02D2E0" box="[916,942,961,986]" italics="true" pageId="17" pageNumber="22">pc</emphasis>
|
||
(15–18) similarly short. Corniculi (25–30) formed as in female, exceeded by elongated tubular salivary styli (45–50) and even longer, slender, finely fimbriated internal malae (70–75). Palpus length 127–133; palptrochanter setae
|
||
<emphasis id="B93FEA984F54FF9AFC64AF6AFC02D51B" box="[917,942,1031,1057]" italics="true" pageId="17" pageNumber="22">v1</emphasis>
|
||
12–15,
|
||
<emphasis id="B93FEA984F54FF9AFBF9AF6AFB8FD51B" box="[1032,1059,1031,1057]" italics="true" pageId="17" pageNumber="22">v2</emphasis>
|
||
30–34; palpgenual seta
|
||
<emphasis id="B93FEA984F54FF9AFADEAF6AFA91D51B" box="[1327,1341,1031,1057]" italics="true" pageId="17" pageNumber="22">d</emphasis>
|
||
(22–23) slightly spine-like.
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F54FF9AFF36AF20FB0AD4BB" blockId="17.[151,1437,150,2009]" pageId="17" pageNumber="22">
|
||
<emphasis id="B93FEA984F54FF9AFF36AF20FEA9D55D" bold="true" box="[199,261,1101,1127]" italics="true" pageId="17" pageNumber="22">Legs.</emphasis>
|
||
Leg lengths, excluding pretarsi, I (325–350), II (278–312), III (287–315), IV (335–360). Leg setation with normal deutonymphal complement of setae as given for adult female, with occasional variabilities. Tarsi II to IV with ventroapical process and pretarsal structures formed as in female; apical dorsosetal processes (
|
||
<emphasis id="B93FEA984F54FF9AFAC0AFFEFAFBD597" box="[1329,1367,1171,1197]" italics="true" pageId="17" pageNumber="22">d-1</emphasis>
|
||
) relatively short (9–10, about 0.6 as long as pretarsus to base of claws 13–15) as in female; form and relative lengths of setae much as in female, with particular setae enlarged, spine-like: I Fe
|
||
<emphasis id="B93FEA984F54FF9AFC16AFB7FBB6D5CE" box="[999,1050,1242,1268]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(45–50),
|
||
<emphasis id="B93FEA984F54FF9AFB7BAFB7FB12D5CE" box="[1162,1214,1242,1268]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(30–33); I Ge
|
||
<emphasis id="B93FEA984F54FF9AFA98AFB7FA30D5CE" box="[1385,1436,1242,1268]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(38–45),
|
||
<emphasis id="B93FEA984F54FF9AFF0CAF90FE9DD42D" box="[253,305,1277,1303]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(37–43); II Fe
|
||
<emphasis id="B93FEA984F54FF9AFE28AF90FDA0D42D" box="[473,524,1277,1303]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(50–53),
|
||
<emphasis id="B93FEA984F54FF9AFD88AF90FD01D42D" box="[633,685,1277,1303]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(32–33); II Ge
|
||
<emphasis id="B93FEA984F54FF9AFCABAF90FC21D42D" box="[858,909,1277,1303]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(35–38),
|
||
<emphasis id="B93FEA984F54FF9AFC0BAF90FB81D42D" box="[1018,1069,1277,1303]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(34–35); III Fe
|
||
<emphasis id="B93FEA984F54FF9AFB11AF90FABFD42D" box="[1248,1299,1277,1303]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(45–50); III Ge
|
||
<emphasis id="B93FEA984F54FF9AFF4EAE4DFF5ED400" box="[191,242,1312,1338]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(42–43),
|
||
<emphasis id="B93FEA984F54FF9AFE93AE4DFE39D400" box="[354,405,1312,1338]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(37–40); IV Fe
|
||
<emphasis id="B93FEA984F54FF9AFDBCAE4DFD2CD400" box="[589,640,1312,1338]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(47–52); IV Ge
|
||
<emphasis id="B93FEA984F54FF9AFCCCAE4DFCDCD400" box="[829,880,1312,1338]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(45–20),
|
||
<emphasis id="B93FEA984F54FF9AFC11AE4DFBBFD400" box="[992,1043,1312,1338]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(40–48); IV Ti
|
||
<emphasis id="B93FEA984F54FF9AFB36AE4DFB55D400" box="[1223,1273,1312,1338]" italics="true" pageId="17" pageNumber="22">ad-1</emphasis>
|
||
(40–47),
|
||
<emphasis id="B93FEA984F54FF9AFA98AE4DFA30D400" box="[1385,1436,1312,1338]" italics="true" pageId="17" pageNumber="22">pd-1</emphasis>
|
||
(36–42). Tarsus IV with asymmetry in lengths of setae (
|
||
<emphasis id="B93FEA984F54FF9AFCF9AE29FC87D464" box="[776,811,1348,1374]" italics="true" pageId="17" pageNumber="22">v-1</emphasis>
|
||
) as in female. Claws of all legs strongly formed (length ca 13), projecting freely from pulvillus (as in
|
||
<figureCitation id="13702A0F4F54FF9AFD67AE0AFD56D4BB" box="[662,762,1383,1409]" captionStart="FIGURE 19" captionStartId="49.[151,250,1824,1848]" captionTargetBox="[151,1436,181,1799]" captionTargetId="figure-17@49.[151,1436,181,1800]" captionTargetPageId="49" captionText="FIGURE 19. Berlesia hospitabilis n. sp.,details. A–C, gnathosomas ventrally: A, female, asterisks showing lateral borders of cheliceral bases inside; B, male; C, deutonymph (same as in D, J–K) with arrowed pharate female structures inside; D, copula, slightly parted in lactic acid, male artificially shrunk; E, F, deutonymphal ambulacra: E, leg II; F, leg I; G, male ambulacrum of leg I (similar in all legs of adult male, female and of protonymph); H, spermatophore; J, same copula, when first found on host’s wing; K, detail of D, area of remaining contact. Arrowheads indicate structures of pharate female (black) and her deutonymphal skin (white). Abbreviations: cxIII, cxIV, leg coxae III and IV; sd, spermatodactyl; sf, spermatophore; st, solenostome." figureDoi="http://doi.org/10.5281/zenodo.4396514" httpUri="https://zenodo.org/record/4396514/files/figure.png" pageId="17" pageNumber="22">Fig. 19E</figureCitation>
|
||
), in contrast to adult and protonymph.
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F54FF9AFF36AEE7FC85D7DB" blockId="17.[151,1437,150,2009]" pageId="17" pageNumber="22">
|
||
<emphasis id="B93FEA984F54FF9AFF36AEE7FDFCD49E" bold="true" box="[199,592,1417,1444]" pageId="17" pageNumber="22">
|
||
Protonymph.
|
||
<emphasis id="B93FEA984F54FF9AFE9AAEE4FDFCD49E" bold="true" box="[363,592,1417,1444]" italics="true" pageId="17" pageNumber="22">Idiosomatic dorsum</emphasis>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F54FF9AFDAEAEE7FD6BD49E" box="[607,711,1418,1444]" captionStart="FIGURE 12" captionStartId="20.[151,250,1834,1858]" captionTargetBox="[157,1433,1222,1807]" captionTargetId="figure-67@20.[151,1437,1219,1810]" captionTargetPageId="20" captionText="FIGURE 12. Berlesia multisetosa n. sp., protonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, right chelicerae dorsally, at resting position; D, gnathotectum; E, subcapitulum. A, B to same scale, C–E to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396500" httpUri="https://zenodo.org/record/4396500/files/figure.png" pageId="17" pageNumber="22">Fig. 12A</figureCitation>
|
||
). Idiosoma 370 long, 268 wide at widest level of setae
|
||
<emphasis id="B93FEA984F54FF9AFAD8AEE7FAEED49E" box="[1321,1346,1418,1444]" italics="true" pageId="17" pageNumber="22">r3</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F54FF9AFABEAEE7FAC4D49E" box="[1359,1384,1418,1444]" italics="true" pageId="17" pageNumber="22">r5</emphasis>
|
||
(n = 1), with weakly sclerotized dorsal shields; podonotal shield 200 long and 178 at widest level between setae
|
||
<emphasis id="B93FEA984F54FF9AFAB5AEC0FAF1D4FD" box="[1348,1373,1453,1479]" italics="true" pageId="17" pageNumber="22">s4</emphasis>
|
||
, with ten pairs of setae (
|
||
<emphasis id="B93FEA984F54FF9AFE97AEBDFD2BD4D0" box="[358,647,1488,1515]" italics="true" pageId="17" pageNumber="22">j1, j3–j6, z1, z2, z4, z5, s4</emphasis>
|
||
); pygidial shield ca 80 long and 152 at widest level between setae
|
||
<emphasis id="B93FEA984F54FF9AFA89AEBDFA39D4D0" box="[1400,1429,1488,1514]" italics="true" pageId="17" pageNumber="22">Z4</emphasis>
|
||
, with five pairs of setae (
|
||
<emphasis id="B93FEA984F54FF9AFE57AE99FDF7D734" box="[422,603,1524,1550]" italics="true" pageId="17" pageNumber="22">J3, J4, Z4, Z5, S</emphasis>
|
||
5). Soft cuticle with ca 12 pairs of podonotal setae, indicating some hypertrichy among
|
||
<emphasis id="B93FEA984F54FF9AFEC7AD7AFEEDD70A" box="[310,321,1559,1584]" italics="true" pageId="17" pageNumber="22">s</emphasis>
|
||
-and
|
||
<emphasis id="B93FEA984F54FF9AFE8AAD7AFE2AD70A" box="[379,390,1559,1584]" italics="true" pageId="17" pageNumber="22">r</emphasis>
|
||
-series, and 12–14 pairs of opisthonotal setae including
|
||
<emphasis id="B93FEA984F54FF9AFBF0AD7AFB32D70B" box="[1025,1182,1559,1585]" italics="true" pageId="17" pageNumber="22">J1, J2, Z1–Z3</emphasis>
|
||
and some hypertrichy among
|
||
<emphasis id="B93FEA984F54FF9AFF1DAD57FF56D76E" box="[236,250,1594,1620]" italics="true" pageId="17" pageNumber="22">S</emphasis>
|
||
-and
|
||
<emphasis id="B93FEA984F54FF9AFEC4AD56FEEAD76E" box="[309,326,1595,1620]" italics="true" pageId="17" pageNumber="22">R</emphasis>
|
||
-series). Form and relative lengths of dorsal setae more homologous than in adult, many of them stout, blunt, but smooth as shown in
|
||
<figureCitation id="13702A0F4F54FF9AFDC0AD30FD34D74D" box="[561,664,1629,1655]" captionStart="FIGURE 12" captionStartId="20.[151,250,1834,1858]" captionTargetBox="[157,1433,1222,1807]" captionTargetId="figure-67@20.[151,1437,1219,1810]" captionTargetPageId="20" captionText="FIGURE 12. Berlesia multisetosa n. sp., protonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, right chelicerae dorsally, at resting position; D, gnathotectum; E, subcapitulum. A, B to same scale, C–E to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396500" httpUri="https://zenodo.org/record/4396500/files/figure.png" pageId="17" pageNumber="22">Fig. 12A</figureCitation>
|
||
; on podonotal shield, other than
|
||
<emphasis id="B93FEA984F54FF9AFBF5AD30FBB1D74D" box="[1028,1053,1629,1655]" italics="true" pageId="17" pageNumber="22">z1</emphasis>
|
||
clearly shortest (15), setal lengths range from
|
||
<emphasis id="B93FEA984F54FF9AFEEAADEDFE98D7A0" box="[283,308,1664,1690]" italics="true" pageId="17" pageNumber="22">z2</emphasis>
|
||
(20) to
|
||
<emphasis id="B93FEA984F54FF9AFE61ADEDFE09D7A0" box="[400,421,1664,1690]" italics="true" pageId="17" pageNumber="22">j6</emphasis>
|
||
slightly longest (28); on pygidial shield, setal lengths range from
|
||
<emphasis id="B93FEA984F54FF9AFB61ADEDFB06D7A0" box="[1168,1194,1664,1690]" italics="true" pageId="17" pageNumber="22">J4</emphasis>
|
||
(12) to
|
||
<emphasis id="B93FEA984F54FF9AFAF7ADECFA8FD7A1" box="[1286,1315,1665,1691]" italics="true" pageId="17" pageNumber="22">Z5</emphasis>
|
||
(23); setal lengths on soft cuticle of similar range length (12–20). Peritreme short (length 32, including stigma diameter 13), with slight indication of embellishment as formed in adult.
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F54FF9AFF36AD84FD02D6E3" blockId="17.[151,1437,150,2009]" pageId="17" pageNumber="22">
|
||
<emphasis id="B93FEA984F54FF9AFF36AD84FE33D63E" bold="true" box="[199,415,1769,1796]" italics="true" pageId="17" pageNumber="22">Idiosomatic venter</emphasis>
|
||
(
|
||
<figureCitation id="13702A0F4F54FF9AFE5EAD87FDBAD63E" box="[431,534,1770,1796]" captionStart="FIGURE 12" captionStartId="20.[151,250,1834,1858]" captionTargetBox="[157,1433,1222,1807]" captionTargetId="figure-67@20.[151,1437,1219,1810]" captionTargetPageId="20" captionText="FIGURE 12. Berlesia multisetosa n. sp., protonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, right chelicerae dorsally, at resting position; D, gnathotectum; E, subcapitulum. A, B to same scale, C–E to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396500" httpUri="https://zenodo.org/record/4396500/files/figure.png" pageId="17" pageNumber="22">Fig. 12B</figureCitation>
|
||
). Tritosternum formed much as in adult, with laciniae fused for only about 8 of total length (72). Presternal area devoid of ornamentation. Sternal shield not clearly delineated, distance between bases of setae
|
||
<emphasis id="B93FEA984F54FF9AFEC6AC5DFEFBD670" box="[311,343,1840,1866]" italics="true" pageId="17" pageNumber="22">st1</emphasis>
|
||
to
|
||
<emphasis id="B93FEA984F54FF9AFE8BAC5DFE37D670" box="[378,411,1840,1866]" italics="true" pageId="17" pageNumber="22">st3</emphasis>
|
||
ca 105, with setae
|
||
<emphasis id="B93FEA984F54FF9AFD9EAC5DFD13D670" box="[623,703,1840,1867]" italics="true" pageId="17" pageNumber="22">st1–st3</emphasis>
|
||
(35–40) and poroids
|
||
<emphasis id="B93FEA984F54FF9AFC5CAC5DFC63D670" box="[941,975,1840,1866]" italics="true" pageId="17" pageNumber="22">iv1</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F54FF9AFC2CAC5DFBACD670" box="[989,1024,1840,1866]" italics="true" pageId="17" pageNumber="22">iv2</emphasis>
|
||
; setae
|
||
<emphasis id="B93FEA984F54FF9AFBBDAC5CFBC1D671" box="[1100,1133,1841,1867]" italics="true" pageId="17" pageNumber="22">st5</emphasis>
|
||
short (20), inserted closely anterior to opisthogastric setae, and flanked by paragenital poroids (one of which asymmetrically formed into minute seta,
|
||
<figureCitation id="13702A0F4F54FF9AFF0CAC1AFEC8D6AB" box="[253,356,1911,1937]" captionStart="FIGURE 12" captionStartId="20.[151,250,1834,1858]" captionTargetBox="[157,1433,1222,1807]" captionTargetId="figure-67@20.[151,1437,1219,1810]" captionTargetPageId="20" captionText="FIGURE 12. Berlesia multisetosa n. sp., protonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, right chelicerae dorsally, at resting position; D, gnathotectum; E, subcapitulum. A, B to same scale, C–E to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396500" httpUri="https://zenodo.org/record/4396500/files/figure.png" pageId="17" pageNumber="22">Fig. 12B</figureCitation>
|
||
). Anal shield smooth, oval, length ca 58 including cribrum, width 55 (cribrum strip width 35); postanal seta (37) about 1.3 longer than paranal setae (28). Soft opisthogastric integument with four pairs of setae,
|
||
<emphasis id="B93FEA984F54FF9AFF66ACD3FF6ED6E2" box="[151,194,1982,2008]" italics="true" pageId="17" pageNumber="22">JV1</emphasis>
|
||
(37),
|
||
<emphasis id="B93FEA984F54FF9AFEF7ACD3FE9ED6E2" box="[262,306,1982,2008]" italics="true" pageId="17" pageNumber="22">JV2</emphasis>
|
||
(30),
|
||
<emphasis id="B93FEA984F54FF9AFE87ACD3FE08D6E2" box="[374,420,1982,2008]" italics="true" pageId="17" pageNumber="22">ZV2</emphasis>
|
||
(22),
|
||
<emphasis id="B93FEA984F54FF9AFE19ACD3FDB8D6E2" box="[488,532,1982,2008]" italics="true" pageId="17" pageNumber="22">JV5</emphasis>
|
||
shortest (15).
|
||
</paragraph>
|
||
<caption id="DF3466024F57FF99FF66AC37FD6ED6E4" ID-DOI="http://doi.org/10.5281/zenodo.4396494" ID-Zenodo-Dep="4396494" httpUri="https://zenodo.org/record/4396494/files/figure.png" pageId="18" pageNumber="23" startId="18.[151,250,1882,1906]" targetBox="[167,1423,189,1855]" targetPageId="18">
|
||
<paragraph id="8BF4368A4F57FF99FF66AC37FD6ED6E4" blockId="18.[151,1436,1882,2014]" pageId="18" pageNumber="23">
|
||
<emphasis id="B93FEA984F57FF99FF66AC37FEB8D648" bold="true" box="[151,276,1882,1906]" pageId="18" pageNumber="23">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C4B4D094F57FF99FEEDAC37FE40D648" authority="Lindquist & Oconnor & Shaw & Sidorchuk, 2020" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[284,492,1882,1906]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="23" phylum="Arthropoda" rank="species" species="multisetosa" status="sp. nov.">
|
||
<emphasis id="B93FEA984F57FF99FEEDAC37FE40D648" box="[284,492,1882,1906]" italics="true" pageId="18" pageNumber="23">Berlesia multisetosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA984F57FF99FE05AC37FD83D648" bold="true" box="[500,559,1882,1906]" pageId="18" pageNumber="23">
|
||
<taxonomicNameLabel id="A20C57E34F57FF99FE05AC37FD83D648" box="[500,559,1882,1906]" pageId="18" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, D, chelicera and spermatodactyl, C, dorsal aspect, D, ventral; E, gnathotectum; F, subcapitulum and palpus; G, H, palp tarsus, G, ventral aspect, H, dorsal, enlarged; J, peritreme. A, B to same scale, C–G to same scale, H, J to same scale. Abbreviations:
|
||
<emphasis id="B93FEA984F57FF99FB5FACCFFB6AD680" bold="true" box="[1198,1222,1954,1978]" pageId="18" pageNumber="23">ar</emphasis>
|
||
, arthrodial envelope;
|
||
<emphasis id="B93FEA984F57FF99FF66ACABFF18D6E4" bold="true" box="[151,180,1990,2014]" pageId="18" pageNumber="23">f d</emphasis>
|
||
, fixed digit;
|
||
<emphasis id="B93FEA984F57FF99FEC6ACABFECCD6E4" bold="true" box="[311,352,1990,2014]" pageId="18" pageNumber="23">m d</emphasis>
|
||
, movable digit; others as in Fig. 7.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3466024F56FF98FF66AF3DFAB6D5B7" ID-DOI="http://doi.org/10.5281/zenodo.4396496" ID-Zenodo-Dep="4396496" httpUri="https://zenodo.org/record/4396496/files/figure.png" pageId="19" pageNumber="24" startId="19.[151,250,1104,1128]" targetBox="[155,1432,184,1077]" targetPageId="19">
|
||
<paragraph id="8BF4368A4F56FF98FF66AF3DFAB6D5B7" blockId="19.[151,1437,1104,1165]" pageId="19" pageNumber="24">
|
||
<emphasis id="B93FEA984F56FF98FF66AF3DFE8DD552" bold="true" box="[151,289,1104,1128]" pageId="19" pageNumber="24">FIGURE 10.</emphasis>
|
||
<taxonomicName id="4C4B4D094F56FF98FED9AF3CFE5BD553" authority="Lindquist & Oconnor & Shaw & Sidorchuk, 2020" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[296,503,1105,1129]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="19" pageNumber="24" phylum="Arthropoda" rank="species" species="multisetosa" status="sp. nov.">
|
||
<emphasis id="B93FEA984F56FF98FED9AF3CFE5BD553" box="[296,503,1105,1129]" italics="true" pageId="19" pageNumber="24">Berlesia multisetosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA984F56FF98FE0EAF3CFD96D553" bold="true" box="[511,570,1105,1129]" pageId="19" pageNumber="24">
|
||
<taxonomicNameLabel id="A20C57E34F56FF98FE0EAF3CFD96D553" box="[511,570,1105,1129]" pageId="19" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male, legs, coxae detached. A, leg I ventrally, most setae of tarsal subapex, beyond transverse dotted line, omitted; B, leg II ventrally; C, leg III ventrally; D, leg IV ventrolaterally. All to same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF4368A4F56FF98FF36AFB3FCC0D4CF" blockId="19.[151,1437,1246,1958]" pageId="19" pageNumber="24">
|
||
<emphasis id="B93FEA984F56FF98FF36AFB3FECED5C3" bold="true" box="[199,354,1246,1273]" italics="true" pageId="19" pageNumber="24">
|
||
<taxonomicName id="4C4B4D094F56FF98FF36AFB3FEF1D5C3" authorityName="Krefft" authorityYear="1871" box="[199,349,1246,1273]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="19" pageNumber="24" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Gnathotectum with triangular smooth anterior margin, tapering to single blunt projection (
|
||
<figureCitation id="13702A0F4F56FF98FA9EAFB2FF60D427" captionStart="FIGURE 12" captionStartId="20.[151,250,1834,1858]" captionTargetBox="[157,1433,1222,1807]" captionTargetId="figure-67@20.[151,1437,1219,1810]" captionTargetPageId="20" captionText="FIGURE 12. Berlesia multisetosa n. sp., protonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, right chelicerae dorsally, at resting position; D, gnathotectum; E, subcapitulum. A, B to same scale, C–E to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396500" httpUri="https://zenodo.org/record/4396500/files/figure.png" pageId="19" pageNumber="24">Fig. 12D</figureCitation>
|
||
). Cheliceral shaft, excluding short basal section 120 long; fixed digit straight, bluntly bifid apically; movable digit (28) formed much as in female, with three teeth flared laterally (
|
||
<figureCitation id="13702A0F4F56FF98FC46AE4AFB8CD47B" box="[951,1056,1319,1345]" captionStart="FIGURE 12" captionStartId="20.[151,250,1834,1858]" captionTargetBox="[157,1433,1222,1807]" captionTargetId="figure-67@20.[151,1437,1219,1810]" captionTargetPageId="20" captionText="FIGURE 12. Berlesia multisetosa n. sp., protonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, right chelicerae dorsally, at resting position; D, gnathotectum; E, subcapitulum. A, B to same scale, C–E to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396500" httpUri="https://zenodo.org/record/4396500/files/figure.png" pageId="19" pageNumber="24">Fig. 12C</figureCitation>
|
||
). Subcapitulum with 12 similarly narrow rows of deutosternal denticles, most with a single denticle but basal and apical rows with two contiguous denticles (
|
||
<figureCitation id="13702A0F4F56FF98FEFBAE02FEC2D4B3" box="[266,366,1391,1417]" captionStart="FIGURE 12" captionStartId="20.[151,250,1834,1858]" captionTargetBox="[157,1433,1222,1807]" captionTargetId="figure-67@20.[151,1437,1219,1810]" captionTargetPageId="20" captionText="FIGURE 12. Berlesia multisetosa n. sp., protonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, right chelicerae dorsally, at resting position; D, gnathotectum; E, subcapitulum. A, B to same scale, C–E to same scale." figureDoi="http://doi.org/10.5281/zenodo.4396500" httpUri="https://zenodo.org/record/4396500/files/figure.png" pageId="19" pageNumber="24">Fig. 12E</figureCitation>
|
||
); subcapitular setae
|
||
<emphasis id="B93FEA984F56FF98FDA1AE02FDD6D4B3" box="[592,634,1391,1417]" italics="true" pageId="19" pageNumber="24">hp1</emphasis>
|
||
attenuated (30), longer than
|
||
<emphasis id="B93FEA984F56FF98FC4AAE02FC49D4B3" box="[955,997,1391,1417]" italics="true" pageId="19" pageNumber="24">hp3</emphasis>
|
||
(15),
|
||
<emphasis id="B93FEA984F56FF98FBD6AE02FBFDD4B3" box="[1063,1105,1391,1417]" italics="true" pageId="19" pageNumber="24">hp2</emphasis>
|
||
and capitular setae
|
||
<emphasis id="B93FEA984F56FF98FADFAE1DFAE4D4B3" box="[1326,1352,1392,1417]" italics="true" pageId="19" pageNumber="24">pc</emphasis>
|
||
similar in short (12) simple form. Corniculi hyaline, blade-like, narrowly rounded apically, moderately short (25), slightly exceeded by tubular salivary styli (40). Palpus length 115, with normal protonymphal complement of setae; single palptrochanter seta 5; palpfemoral seta
|
||
<emphasis id="B93FEA984F56FF98FDA1AEB6FDF2D4CF" box="[592,606,1499,1525]" italics="true" pageId="19" pageNumber="24">d</emphasis>
|
||
(18) slightly spine-like.
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F56FF98FF36AE92FA35D7F7" blockId="19.[151,1437,1246,1958]" pageId="19" pageNumber="24">
|
||
<emphasis id="B93FEA984F56FF98FF36AE92FEA9D723" bold="true" box="[199,261,1535,1561]" italics="true" pageId="19" pageNumber="24">Legs.</emphasis>
|
||
Leg lengths, excluding pretarsi, I 265, II 225, III 225, IV 255. Leg chaetotaxy of protonymph typical of general holotrichous pattern presented for free-living
|
||
<taxonomicName id="4C4B4D094F56FF98FD00AD4EFCCDD707" box="[753,865,1571,1597]" class="Arachnida" higherTaxonomySource="GBIF" infraOrder="Gamasina" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="24" phylum="Arthropoda" rank="infraOrder">Gamasina</taxonomicName>
|
||
by Evans (1963): coxae, 2-2-2-1; trochanters, 4-4- 4-4; femora, 10 (2 2/1 2/1 2)-8 (1 2/1 2/1 1)-5 (1 2/1 1/0 0)-4 (1 2/0 1/0 0); genua, 8 (1 2/1 2/1 1)-6 (1 2/0 2/0 1)-6 (1 2/0 2/0 1)-5 (1 2/0 2/0 0); tibiae, 8 (1 2/1 2/1 1)-7 (1 1/1 2/1 1)-7 (1 1/1 2/1 1)-7 (1 1/1 2/1 1). Legs I–IV each with particular setae enlarged, spinelike, much as noted for deutonymph. Tarsi II–IV with ventroapical process, pretarsal structures and weakly developed claws formed as in female. Tarsus IV with asymmetry of setae (
|
||
<emphasis id="B93FEA984F56FF98FB27ADDEFB56D7F7" box="[1238,1274,1715,1741]" italics="true" pageId="19" pageNumber="24">v-1</emphasis>
|
||
) as in female.
|
||
</paragraph>
|
||
<paragraph id="8BF4368A4F56FF98FF36ADBAFCC2D7C8" blockId="19.[151,1437,1246,1958]" box="[199,878,1751,1778]" pageId="19" pageNumber="24">
|
||
<emphasis id="B93FEA984F56FF98FF36ADBAFEBBD7CB" bold="true" box="[199,279,1751,1777]" pageId="19" pageNumber="24">Larva.</emphasis>
|
||
No material available (adult females nymphiparous).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165014F56FF9EFF36AD96FC4BD0BC" lastPageId="21" lastPageNumber="26" pageId="19" pageNumber="24" type="materials_examined">
|
||
<paragraph id="8BF4368A4F56FF98FF36AD96FC09D69F" blockId="19.[151,1437,1246,1958]" pageId="19" pageNumber="24">
|
||
<emphasis id="B93FEA984F56FF98FF36AD96FEDBD62E" bold="true" box="[199,375,1786,1813]" pageId="19" pageNumber="24">Type material.</emphasis>
|
||
All specimens were collected by E.E. Lindquist or E.A. Sidorchuk from four adult crickets, identified as “
|
||
<taxonomicName id="4C4B4D094F56FF98FEC2AC72FDA1D603" authorityName="Karney" authorityYear="1937" box="[307,525,1823,1849]" class="Insecta" family="Haglidae" genus="Craspedogryllacris" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEA984F56FF98FEC2AC72FDA1D603" box="[307,525,1823,1849]" italics="true" pageId="19" pageNumber="24">Craspedogryllacris</emphasis>
|
||
</taxonomicName>
|
||
sp. 21” by David Rentz, collected from low coastal heathland of the southwestern region of
|
||
<collectingCountry id="F35C761A4F56FF98FEF5AC2EFEC7D667" box="[260,363,1859,1885]" name="Australia" pageId="19" pageNumber="24">Australia</collectingCountry>
|
||
(
|
||
<collectingRegion id="498FF8684F56FF98FE8BAC2EFDE4D667" box="[378,584,1859,1885]" country="Australia" name="Western Australia" pageId="19" pageNumber="24">Western Australia</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE7F504D4F56FF98FDA2AC2EFD1ED667" box="[595,690,1859,1885]" degrees="33" direction="south" minutes="33" orientation="latitude" pageId="19" pageNumber="24" precision="925" value="-33.55">33°33’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7F504D4F56FF98FD4FAC2EFC87D664" box="[702,811,1859,1886]" degrees="115" direction="east" minutes="01" orientation="longitude" pageId="19" pageNumber="24" precision="925" value="115.01667">115°01’E</geoCoordinate>
|
||
, Cape Naturaliste, West Coast Road,
|
||
<date id="FFF5104A4F56FF98FB25AC2EFAC0D664" box="[1236,1388,1859,1886]" pageId="19" pageNumber="24" value="1995-12-10">10 Dec. 1995</date>
|
||
and
|
||
<date id="FFF5104A4F56FF98FF66AC0AFEE9D6BB" box="[151,325,1895,1921]" pageId="19" pageNumber="24" value="1999-03-13">13 March 1999</date>
|
||
, coll. R.Hale).
|
||
<materialsCitation id="3B233CD74F56FF98FE1CAC0AFC09D69F" ID-GBIF-Occurrence="3010620303" collectingDate="1995-12-10" location="Australian Natural Insect Collection" municipality="Craspedogryllacris" pageId="19" pageNumber="24" specimenCount="4" specimenCount-adult="1" specimenCount-female="1" typeStatus="holotype">
|
||
<typeStatus id="54F088284F56FF98FE1CAC0AFD29D6BB" box="[493,645,1895,1921]" pageId="19" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
|
||
: nymphiparous
|
||
<specimenCount id="9D4DFD034F56FF98FCC6AC0AFCC3D6BB" box="[823,879,1895,1921]" pageId="19" pageNumber="24" type="adult">adult</specimenCount>
|
||
<specimenCount id="9D4DFD034F56FF98FC87AC0AFC69D6BB" box="[886,965,1895,1921]" pageId="19" pageNumber="24" type="female">female</specimenCount>
|
||
,
|
||
<specimenCount id="9D4DFD034F56FF98FC21AC0AFC46D6BB" box="[976,1002,1895,1921]" pageId="19" pageNumber="24" type="generic">ex</specimenCount>
|
||
<taxonomicName id="4C4B4D094F56FF98FC00AC0AFB67D6BB" authorityName="Karney" authorityYear="1937" box="[1009,1227,1895,1921]" class="Insecta" family="Haglidae" genus="Craspedogryllacris" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEA984F56FF98FC00AC0AFB67D6BB" box="[1009,1227,1895,1921]" italics="true" pageId="19" pageNumber="24">
|
||
<collectingMunicipality id="6B90ACF04F56FF98FC00AC0AFB67D6BB" box="[1009,1227,1895,1921]" pageId="19" pageNumber="24">Craspedogryllacris</collectingMunicipality>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<specimenCount id="9D4DFD034F56FF98FB23AC0AFB5ED6BB" box="[1234,1266,1895,1921]" pageId="19" pageNumber="24" type="generic">sp.</specimenCount>
|
||
21 #3,
|
||
<date id="FFF5104A4F56FF98FAB4AC0AFF7ED69C" pageId="19" pageNumber="24" value="1995-12-10">
|
||
<collectingDate id="EFB1E9A24F56FF98FAB4AC0AFF7ED69C" pageId="19" pageNumber="24" value="1995-12-10">10 Dec. 1995</collectingDate>
|
||
</date>
|
||
, deposited in the
|
||
<location id="8E9460514F56FF98FE65ACE6FC80D69F" LSID="urn:lsid:plazi:treatment:03E2879C4F48FF9EFF66A92BFE4ED269:8E9460514F56FF98FE65ACE6FC80D69F" box="[404,812,1931,1957]" municipality="Craspedogryllacris" name="Australian Natural Insect Collection" pageId="19" pageNumber="24">Australian Natural Insect Collection</location>
|
||
,
|
||
<location id="8E9460514F56FF98FCC6ACE6FC0DD69F" LSID="urn:lsid:plazi:treatment:03E2879C4F48FF9EFF66A92BFE4ED269:8E9460514F56FF98FCC6ACE6FC0DD69F" box="[823,929,1931,1957]" municipality="Craspedogryllacris" name="Canberra" pageId="19" pageNumber="24">Canberra</location>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF3466024F51FF9FFF66AF26FED4D591" ID-DOI="http://doi.org/10.5281/zenodo.4396498" ID-Zenodo-Dep="4396498" httpUri="https://zenodo.org/record/4396498/files/figure.png" pageId="20" pageNumber="25" startId="20.[151,250,1099,1123]" targetBox="[155,1431,185,1068]" targetPageId="20">
|
||
<paragraph id="8BF4368A4F51FF9FFF66AF26FED4D591" blockId="20.[151,1436,1099,1195]" pageId="20" pageNumber="25">
|
||
<emphasis id="B93FEA984F51FF9FFF66AF26FE8CD559" bold="true" box="[151,288,1099,1123]" pageId="20" pageNumber="25">FIGURE 11.</emphasis>
|
||
<taxonomicName id="4C4B4D094F51FF9FFED9AF26FE5BD559" authority="Lindquist & Oconnor & Shaw & Sidorchuk, 2020" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[296,503,1099,1123]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="25" phylum="Arthropoda" rank="species" species="multisetosa" status="sp. nov.">
|
||
<emphasis id="B93FEA984F51FF9FFED9AF26FE5BD559" box="[296,503,1099,1123]" italics="true" pageId="20" pageNumber="25">Berlesia multisetosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA984F51FF9FFDF1AF26FD96D559" bold="true" box="[512,570,1099,1123]" pageId="20" pageNumber="25">
|
||
<taxonomicNameLabel id="A20C57E34F51FF9FFDF1AF26FD96D559" box="[512,570,1099,1123]" pageId="20" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, deutonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, apical half of chelicera, ventrally, at resting position; D, variations of peritremes; E–G, variability of gnathotectum outline. A, B to same scale; C, E–G to same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3466024F51FF9FFF66AC47FB04D65C" ID-DOI="http://doi.org/10.5281/zenodo.4396500" ID-Zenodo-Dep="4396500" httpUri="https://zenodo.org/record/4396500/files/figure.png" pageId="20" pageNumber="25" startId="20.[151,250,1834,1858]" targetBox="[157,1433,1222,1807]" targetPageId="20">
|
||
<paragraph id="8BF4368A4F51FF9FFF66AC47FB04D65C" blockId="20.[151,1437,1834,1894]" pageId="20" pageNumber="25">
|
||
<emphasis id="B93FEA984F51FF9FFF66AC47FE8CD678" bold="true" box="[151,288,1834,1858]" pageId="20" pageNumber="25">FIGURE 12.</emphasis>
|
||
<taxonomicName id="4C4B4D094F51FF9FFED6AC47FE59D678" authority="Lindquist & Oconnor & Shaw & Sidorchuk, 2020" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[295,501,1834,1858]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="25" phylum="Arthropoda" rank="species" species="multisetosa" status="sp. nov.">
|
||
<emphasis id="B93FEA984F51FF9FFED6AC47FE59D678" box="[295,501,1834,1858]" italics="true" pageId="20" pageNumber="25">Berlesia multisetosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEA984F51FF9FFE0DAC47FD99D678" bold="true" box="[508,565,1834,1858]" pageId="20" pageNumber="25">
|
||
<taxonomicNameLabel id="A20C57E34F51FF9FFE0DAC47FD99D678" box="[508,565,1834,1858]" pageId="20" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, protonymph. A, idiosoma dorsally; B, idiosoma ventrally, partial; C, right chelicerae dorsally, at resting position; D, gnathotectum; E, subcapitulum. A, B to same scale, C–E to same scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF4368A4F50FF9EFF36ABFAFC4BD0BC" blockId="21.[151,1437,151,851]" pageId="21" pageNumber="26">
|
||
<typeStatus id="54F088284F50FF9EFF36ABFAFECAD18B" box="[199,358,151,177]" pageId="21" pageNumber="26" type="paratype">PARATYPES</typeStatus>
|
||
: one female, one male, one deutonymph with same data as
|
||
<typeStatus id="54F088284F50FF9EFC1FABFAFBF8D18B" box="[1006,1108,151,177]" pageId="21" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
; one male, two deutonymphs, one protonymph, with same data as
|
||
<typeStatus id="54F088284F50FF9EFDD4ABD6FD2BD1EF" box="[549,647,187,213]" pageId="21" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
except ex
|
||
<emphasis id="B93FEA984F50FF9EFD0CABD6FCBBD1EF" box="[765,791,187,213]" italics="true" pageId="21" pageNumber="26">C.</emphasis>
|
||
sp. 21 #4; one male, with same data as
|
||
<typeStatus id="54F088284F50FF9EFB3AABD6FA81D1EF" box="[1227,1325,187,213]" pageId="21" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
except ex
|
||
<emphasis id="B93FEA984F50FF9EFF66ABB2FF1DD1C3" box="[151,177,223,249]" italics="true" pageId="21" pageNumber="26">C.</emphasis>
|
||
sp. 21 #2,
|
||
<date id="FFF5104A4F50FF9EFEDCABB3FE6AD1C2" box="[301,454,222,249]" pageId="21" pageNumber="26" value="1999-05-13">13 May 1999</date>
|
||
; seven deutonymphs, with same data as
|
||
<typeStatus id="54F088284F50FF9EFC7CABB3FC43D1C2" box="[909,1007,222,248]" pageId="21" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
except ex
|
||
<emphasis id="B93FEA984F50FF9EFB98ABB2FB2ED1C3" box="[1129,1154,223,249]" italics="true" pageId="21" pageNumber="26">C.</emphasis>
|
||
sp. 21 #1,
|
||
<date id="FFF5104A4F50FF9EFB0FABB3FA3BD1C2" box="[1278,1431,222,249]" pageId="21" pageNumber="26" value="1999-05-13">13 May 1999</date>
|
||
; of the seven deutonymphs ex cricket #1, five were on the wings, one on the abdominal venter, and one on the trochanter of leg III. One female, one male, three deutonymphs, one protonymph deposited in the CNC, Ottawa; one male, two deutonymphs deposited in the ZIN, St.-
|
||
<collectingRegion id="498FF8684F50FF9EFD4CAA25FC91D058" box="[701,829,328,354]" country="Russia" name="Saint Petersburg" pageId="21" pageNumber="26">Petersburg</collectingRegion>
|
||
; one male, two deutonymphs deposited in the UMMZ, Ann Arbor; one male, three deutonymphs deposited in the ANIC, Canberra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165014F50FF9EFF36AAFDFCFFD32E" pageId="21" pageNumber="26" type="etymology">
|
||
<paragraph id="8BF4368A4F50FF9EFF36AAFDFCFFD32E" blockId="21.[151,1437,151,851]" pageId="21" pageNumber="26">
|
||
<emphasis id="B93FEA984F50FF9EFF36AAFDFEE2D090" bold="true" box="[199,334,400,426]" pageId="21" pageNumber="26">Etymology.</emphasis>
|
||
The specific name is a compound adjective derived from the Latin words
|
||
<emphasis id="B93FEA984F50FF9EFB7CAAE2FB7CD093" box="[1165,1232,399,425]" italics="true" pageId="21" pageNumber="26">multi-</emphasis>
|
||
, meaning “many” or “multiple” and
|
||
<emphasis id="B93FEA984F50FF9EFE91AAD9FE1DD0F7" box="[352,433,436,461]" italics="true" pageId="21" pageNumber="26">setosus</emphasis>
|
||
, meaning “bristly” or “hairy”, and refers to the hypertrichous condition of the idiosoma of all instars of this species. As the gender of the generic name
|
||
<taxonomicName id="4C4B4D094F50FF9EFCB7AABBFC0FD0CA" authorityName="Canestrini" authorityYear="1884" box="[838,931,470,496]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEA984F50FF9EFCB7AABBFC0FD0CA" box="[838,931,470,496]" italics="true" pageId="21" pageNumber="26">Berlesia</emphasis>
|
||
</taxonomicName>
|
||
should be feminine (Sauro Simoni, personal communication), the specific name is accordingly
|
||
<taxonomicName id="4C4B4D094F50FF9EFD3DAA97FCE0D32E" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[716,844,506,532]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="26" phylum="Arthropoda" rank="species" species="multisetosa">
|
||
<emphasis id="B93FEA984F50FF9EFD3DAA97FCE0D32E" box="[716,844,506,532]" italics="true" pageId="21" pageNumber="26">multisetosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165014F50FF9EFF36A970FEB8D236" pageId="21" pageNumber="26" type="diagnosis">
|
||
<paragraph id="8BF4368A4F50FF9EFF36A970FEB8D236" blockId="21.[151,1437,151,851]" pageId="21" pageNumber="26">
|
||
<emphasis id="B93FEA984F50FF9EFF36A970FE60D30D" bold="true" box="[199,460,541,567]" pageId="21" pageNumber="26">Differential diagnosis.</emphasis>
|
||
Adults, deutonymphs and protonymphs of
|
||
<taxonomicName id="4C4B4D094F50FF9EFC5BA973FBE4D30D" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[938,1096,541,567]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="26" phylum="Arthropoda" rank="species" species="multisetosa">
|
||
<emphasis id="B93FEA984F50FF9EFC5BA973FBE4D30D" box="[938,1096,541,567]" italics="true" pageId="21" pageNumber="26">B. multisetosa</emphasis>
|
||
</taxonomicName>
|
||
are immediately distinguished from those of other known species in being hypertrichous on the lateral and opisthogastric regions of the idiosoma. Also, those setae are of similarly moderate lengths, none being little more than twice as long as the others. Unlike
|
||
<taxonomicName id="4C4B4D094F50FF9EFF66A9E5FE94D398" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[151,312,648,674]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="26" phylum="Arthropoda" rank="species" species="hospitabilis">
|
||
<emphasis id="B93FEA984F50FF9EFF66A9E5FE94D398" box="[151,312,648,674]" italics="true" pageId="21" pageNumber="26">B. hospitabilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4B4D094F50FF9EFE9CA9E5FDACD39B" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[365,512,648,673]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="26" phylum="Arthropoda" rank="species" species="vorontsovi">
|
||
<emphasis id="B93FEA984F50FF9EFE9CA9E5FDACD39B" box="[365,512,648,673]" italics="true" pageId="21" pageNumber="26">B. vorontsovi</emphasis>
|
||
</taxonomicName>
|
||
, paravertical setae
|
||
<emphasis id="B93FEA984F50FF9EFD27A9E5FD43D398" box="[726,751,648,674]" italics="true" pageId="21" pageNumber="26">z1</emphasis>
|
||
and setae
|
||
<emphasis id="B93FEA984F50FF9EFC93A9E5FCD0D398" box="[866,892,648,674]" italics="true" pageId="21" pageNumber="26">J2</emphasis>
|
||
are present. On adults, the genu and tibia of leg I each have 11 setae, with only two ventral setae (
|
||
<emphasis id="B93FEA984F50FF9EFD41A9C6FD4ED3FF" box="[688,738,683,709]" italics="true" pageId="21" pageNumber="26">av-1</emphasis>
|
||
,
|
||
<emphasis id="B93FEA984F50FF9EFD01A9C6FC8DD3FF" box="[752,801,683,709]" italics="true" pageId="21" pageNumber="26">pv-1</emphasis>
|
||
), whereas those of
|
||
<taxonomicName id="4C4B4D094F50FF9EFC0BA9C6FBD1D3FF" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[1018,1149,683,709]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="26" phylum="Arthropoda" rank="species" species="hospitabilis">
|
||
<emphasis id="B93FEA984F50FF9EFC0BA9C6FBD1D3FF" box="[1018,1149,683,709]" italics="true" pageId="21" pageNumber="26">hospitabilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4B4D094F50FF9EFB43A9C1FA8BD3FF" authorityName="Lindquist & Oconnor & Shaw & Sidorchuk" authorityYear="2020" box="[1202,1319,684,709]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="26" phylum="Arthropoda" rank="species" species="vorontsovi">
|
||
<emphasis id="B93FEA984F50FF9EFB43A9C1FA8BD3FF" box="[1202,1319,684,709]" italics="true" pageId="21" pageNumber="26">vorontsovi</emphasis>
|
||
</taxonomicName>
|
||
each have the third ventral seta (
|
||
<emphasis id="B93FEA984F50FF9EFE7FA9A2FE6CD3D3" box="[398,448,719,745]" italics="true" pageId="21" pageNumber="26">av-2</emphasis>
|
||
) present. The asymmetrically spine-like form of subapical seta
|
||
<emphasis id="B93FEA984F50FF9EFB7DA9A2FB12D3D3" box="[1164,1214,719,745]" italics="true" pageId="21" pageNumber="26">pv-1</emphasis>
|
||
on tarsus IV is also distinctive.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165014F50FF9EFF36A87BFE4ED269" pageId="21" pageNumber="26" type="discussion">
|
||
<paragraph id="8BF4368A4F50FF9EFF36A87BFE4ED269" blockId="21.[151,1437,151,851]" pageId="21" pageNumber="26">
|
||
<emphasis id="B93FEA984F50FF9EFF36A87BFE97D20A" bold="true" box="[199,315,790,816]" pageId="21" pageNumber="26">Remarks.</emphasis>
|
||
This is the first species of the genus
|
||
<taxonomicName id="4C4B4D094F50FF9EFD28A878FC9AD215" authorityName="Canestrini" authorityYear="1884" box="[729,822,789,815]" class="Arachnida" family="Lebertiidae" genus="Berlesia" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEA984F50FF9EFD28A878FC9AD215" box="[729,822,789,815]" italics="true" pageId="21" pageNumber="26">Berlesia</emphasis>
|
||
</taxonomicName>
|
||
to be described with all active instars having peripherally hypertrichous idiosomas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |