347 lines
42 KiB
XML
347 lines
42 KiB
XML
<document id="3B83DE19F9A9486F071F37766052724F" ID="10.11646/zootaxa.4205.3.6" ID-DOI="10.11646/zootaxa.4205.3.6" ID-GBIF-Dataset="d8d24aba-900e-4dcc-97cd-adb094ebe3ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="194329" ID-ZooBank="66FB30C3-66BC-4D79-9361-03F5C9F22498" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481096845972" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2016" docId="03D7CE3F3449F32DA8CF6001FF01FD94" docLanguage="en" docName="zootaxa.4205.3.6.pdf" docOrigin="Zootaxa 4205 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scydmaenus (Geoscydmaenus) madagassicus Franz" docType="treatment" docVersion="8" lastPageNumber="274" masterDocId="FFEEB647344FF324A8586722FFEEFFA9" masterDocTitle="Review of Scydmaenus (Geoscydmaenus) of Madagascar (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="274" masterPageNumber="265" pageNumber="271" updateTime="1698396176682" updateUser="plazi">
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<mods:titleInfo id="5977098F3FB20874191FF86D539BEEBA">
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<mods:title id="A8F4893FF5A3599A5D5FDB1F20FE5476">Review of Scydmaenus (Geoscydmaenus) of Madagascar (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="5B9A50C04389E450C6FD25CE3B8F2203" type="personal">
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<mods:namePart id="18AE93A95FDEC174A727EBA9E877EC23">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="C85A3827993EE8DF3CB1CD256C9280CC">text</mods:typeOfResource>
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<mods:title id="E783E509C4E5F0A34AE6030529BE492D">Zootaxa</mods:title>
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<mods:part id="3764EF0F72310C227A178859AC79CFAB">
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<mods:date id="0AE5CE46545F4BFD25FE57C47777CC4A">2016</mods:date>
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<mods:number id="2B5ECF350867FF3F0613FFCC412F8B90">4205</mods:number>
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<mods:detail id="8D7B9CE5B7073B74C9DFAF7A7E19DC2A" type="issue">
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<mods:number id="26E678AB968067BD89409FB0ADDAFB7C">3</mods:number>
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<mods:identifier id="005AFDCAF4E0B5AED09210CBA1ACDAB2" type="DOI">10.11646/zootaxa.4205.3.6</mods:identifier>
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<treatment id="03D7CE3F3449F32DA8CF6001FF01FD94" ID-DOI="http://doi.org/10.5281/zenodo.6053367" ID-GBIF-Taxon="124956958" ID-Zenodo-Dep="6053367" LSID="urn:lsid:plazi:treatment:03D7CE3F3449F32DA8CF6001FF01FD94" httpUri="http://treatment.plazi.org/id/03D7CE3F3449F32DA8CF6001FF01FD94" lastPageId="9" lastPageNumber="274" pageId="6" pageNumber="271">
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<subSubSection id="C3642CA23449F322A8CF6001FDDFF8F7" pageId="6" pageNumber="271" type="nomenclature">
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<paragraph id="8BC17F293449F322A8CF6001FCE0F894" blockId="6.[151,782,1827,1886]" box="[151,782,1827,1853]" pageId="6" pageNumber="271">
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<heading id="D089C8453449F322A8CF6001FCE0F894" bold="true" box="[151,782,1827,1853]" fontSize="11" level="1" pageId="6" pageNumber="271" reason="1">
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<taxonomicName id="4C7E04AA3449F322A8CF6001FCE0F894" ID-CoL="4W3VG" authority="Franz" authorityName="Franz" box="[151,782,1827,1853]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="madagassicus" subGenus="Geoscydmaenus">
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||
<emphasis id="B90AA33B3449F322A8CF6001FCE0F894" bold="true" box="[151,782,1827,1853]" pageId="6" pageNumber="271">
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||
<emphasis id="B90AA33B3449F322A8CF6001FEC0F894" bold="true" box="[151,302,1827,1853]" italics="true" pageId="6" pageNumber="271">Scydmaenus</emphasis>
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||
(
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||
<emphasis id="B90AA33B3449F322A9676001FDECF894" bold="true" box="[319,514,1827,1853]" italics="true" pageId="6" pageNumber="271">Geoscydmaenus</emphasis>
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||
)
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||
<emphasis id="B90AA33B3449F322AA4C6001FD52F894" bold="true" box="[532,700,1827,1853]" italics="true" pageId="6" pageNumber="271">madagassicus</emphasis>
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Franz
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</emphasis>
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</taxonomicName>
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||
</heading>
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</paragraph>
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<paragraph id="8BC17F293449F322A8CF6067FDDFF8F7" blockId="6.[151,782,1827,1886]" box="[151,561,1861,1886]" pageId="6" pageNumber="271">
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||
(
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<figureCitation id="134563AC3449F322A8C76067FEC0F8F7" box="[159,302,1861,1886]" captionStart-0="FIGURES 1 – 8" captionStart-1="FIGURES 9 – 11" captionStartId-0="4.[151,264,1881,1904]" captionStartId-1="5.[151,264,1613,1636]" captionTargetBox-0="[168,1436,295,1860]" captionTargetBox-1="[151,1436,193,1584]" captionTargetId-0="figure@4.[151,1436,273,1860]" captionTargetId-1="figure@5.[151,1436,193,1592]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1 – 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 – 4) and Sc. madagassicus Franz (2, 5 – 8). Set of original labels of holotype (1 – 2); dorsal habitus of male (3); head and prothorax in dorsal (4 – 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." captionText-1="FIGURES 9 – 11. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus madagassicus Franz (9 – 10) and Sc. isaloensis Franz (11). Pterothorax and abdomen in ventral view (9); pterothorax, transparent mount in ventral view (10); profemoral ventral tooth of male in lateral view (11). Abbreviations: aest 3, metanepisternum; ar, anterior ridge; lf, lateral fovea; lmfa, lateral metafurcal arm; mscc, mesocoxal cavity; mscs, mesocoxal socket; msvp, mesoventral intercoxal process; mtvc, metaventral carina; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus." httpUri-0="https://zenodo.org/record/194330/files/figure.png" httpUri-1="https://zenodo.org/record/267455/files/figure.png" pageId="6" pageNumber="271">Figs 2, 5–10</figureCitation>
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,
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<figureCitation id="134563AC3449F322A9616064FE6FF8F7" box="[313,385,1861,1886]" captionStart="FIGURES 12 – 15" captionStartId="6.[151,264,1482,1505]" captionTargetBox="[151,1427,210,1461]" captionTargetId="figure@6.[151,1436,193,1461]" captionTargetPageId="6" captionText="FIGURES 12 – 15. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (12, 14) and Sc. madagassicus Franz (13, 15). Right antenna of male in dorsal view (12 – 13); right fore leg of male in anterior view." httpUri="https://zenodo.org/record/194332/files/figure.png" pageId="6" pageNumber="271">13, 15</figureCitation>
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,
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<figureCitation id="134563AC3449F322A9D56064FE38F8F7" box="[397,470,1862,1886]" captionStart="FIGURES 16 – 21" captionStartId="7.[151,264,1906,1929]" captionTargetBox="[164,1415,224,1854]" captionTargetId="figure@7.[151,1436,193,1885]" captionTargetPageId="7" captionText="FIGURES 16 – 21. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (16 – 18) and Sc. madagassicus Franz (19 – 21). Aedeagus in ventral (16, 19) and lateral (17 – 18, 20) view (with partly exserted flagellum in Fig. 18); spermatheca (21)." httpUri="https://zenodo.org/record/194333/files/figure.png" pageId="6" pageNumber="271">19–21</figureCitation>
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,
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<figureCitation id="134563AC3449F322A9B96067FDC7F8F7" box="[481,553,1861,1886]" captionStart="FIGURES 22 – 25" captionStartId="8.[151,264,804,827]" captionTargetBox="[163,1428,208,766]" captionTargetId="figure@8.[151,1436,193,784]" captionTargetPageId="8" captionText="FIGURES 22 – 25. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (22 – 23) and Sc. madagassicus Franz (24 – 25). Aedeagus in ventral (22, 24) and lateral (23, 25) views." httpUri="https://zenodo.org/record/194334/files/figure.png" pageId="6" pageNumber="271">24–25</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3642CA23449F322A8CF60ACFC97F80A" box="[151,889,1934,1956]" pageId="6" pageNumber="271" type="reference_group">
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<paragraph id="8BC17F293449F322A8CF60ACFC97F80A" blockId="6.[151,889,1934,1956]" box="[151,889,1934,1956]" pageId="6" pageNumber="271">
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<treatmentCitationGroup id="AB6E58073449F322A8CF60ACFC97F80A" box="[151,889,1934,1956]" pageId="6" pageNumber="271">
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<taxonomicName id="4C7E04AA3449F322A8CF60ACFCF7F80A" ID-CoL="4W3VG" authority="Franz, 1986: 374" authorityName="Franz" authorityPageNumber="374" authorityYear="1986" box="[151,793,1934,1956]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="madagassicus" subGenus="Geoscydmaenus">
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<emphasis id="B90AA33B3449F322A8CF60ACFEFAF80A" box="[151,276,1934,1955]" italics="true" pageId="6" pageNumber="271">Scydmaenus</emphasis>
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(
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<emphasis id="B90AA33B3449F322A97B60ACFE29F80A" box="[291,455,1934,1955]" italics="true" pageId="6" pageNumber="271">Geoscydmaenus</emphasis>
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)
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<emphasis id="B90AA33B3449F322A98E60ACFD8AF80A" box="[470,612,1934,1955]" italics="true" pageId="6" pageNumber="271">madagassicus</emphasis>
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<treatmentCitation id="0ADF59383449F322AA3260ACFCF7F80A" author="Franz" box="[618,793,1934,1956]" page="374" pageId="6" pageNumber="271" year="1986">
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||
<bibRefCitation id="EFEF02D83449F322AA3260ACFD09F80D" author="Franz" box="[618,743,1934,1956]" pageId="6" pageNumber="271" refString="Franz, H. (1986) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986</bibRefCitation>
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: 374
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</treatmentCitation>
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||
</taxonomicName>
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, fig. 342.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF012FA13448F323A8CF6050FEA2F86F" httpUri="https://zenodo.org/record/194333/files/figure.png" pageId="7" pageNumber="272" targetBox="[164,1415,224,1854]" targetPageId="7">
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||
<paragraph id="8BC17F293448F323A8CF6050FEA2F86F" blockId="7.[151,1436,1906,1990]" pageId="7" pageNumber="272">
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||
<emphasis id="B90AA33B3448F323A8CF6050FEBAF821" bold="true" box="[151,340,1906,1929]" pageId="7" pageNumber="272">FIGURES 16–21.</emphasis>
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<taxonomicName id="4C7E04AA3448F323A9026051FD7CF821" authority="(Geoscydmaenus)" authorityName="Geoscydmaenus" baseAuthorityName="Geoscydmaenus" box="[346,658,1907,1928]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="272" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B90AA33B3448F323A9026051FE39F821" box="[346,471,1907,1928]" italics="true" pageId="7" pageNumber="272">Scydmaenus</emphasis>
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||
(
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<emphasis id="B90AA33B3448F323A9BD6051FD67F821" box="[485,649,1907,1928]" italics="true" pageId="7" pageNumber="272">Geoscydmaenus</emphasis>
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)
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</taxonomicName>
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of Madagascar.
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<taxonomicName id="4C7E04AA3448F323AB626051FB8CF821" authority="Franz" authorityName="Franz" box="[826,1122,1907,1928]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="272" phylum="Arthropoda" rank="species" species="isaloensis">
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<emphasis id="B90AA33B3448F323AB626051FBCFF821" box="[826,1057,1907,1928]" italics="true" pageId="7" pageNumber="272">Scydmaenus isaloensis</emphasis>
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Franz
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</taxonomicName>
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(16–18) and
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<taxonomicName id="4C7E04AA3448F323ACB16051FF3FF80E" authority="Franz" authorityName="Franz" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="272" phylum="Arthropoda" rank="species" species="madagassicus">
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<emphasis id="B90AA33B3448F323ACB16051FA72F821" box="[1257,1436,1907,1928]" italics="true" pageId="7" pageNumber="272">Sc. madagassicus</emphasis>
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Franz
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</taxonomicName>
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(19–21). Aedeagus in ventral (16, 19) and lateral (17–18, 20) view (with partly exserted flagellum in Fig. 18); spermatheca (21).
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</paragraph>
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</caption>
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<caption id="DF012FA13447F32CA8CF6406FC8AFCF3" httpUri="https://zenodo.org/record/194334/files/figure.png" pageId="8" pageNumber="273" targetBox="[163,1428,208,766]" targetPageId="8">
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||
<paragraph id="8BC17F293447F32CA8CF6406FC8AFCF3" blockId="8.[151,1436,804,858]" pageId="8" pageNumber="273">
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||
<emphasis id="B90AA33B3447F32CA8CF6406FEBAFC93" bold="true" box="[151,340,804,827]" pageId="8" pageNumber="273">FIGURES 22–25.</emphasis>
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<taxonomicName id="4C7E04AA3447F32CA9026407FD7CFC92" authority="Geoscydmaenus" authorityName="Geoscydmaenus" box="[346,658,805,827]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AA33B3447F32CA9026407FE39FC93" box="[346,471,805,826]" italics="true" pageId="8" pageNumber="273">Scydmaenus</emphasis>
|
||
(
|
||
<emphasis id="B90AA33B3447F32CA9BD6407FD67FC93" box="[485,649,805,826]" italics="true" pageId="8" pageNumber="273">Geoscydmaenus</emphasis>
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||
)
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||
</taxonomicName>
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||
of Madagascar.
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||
<taxonomicName id="4C7E04AA3447F32CAB626407FB8CFC92" authority="Franz" authorityName="Franz" box="[826,1122,805,827]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="isaloensis">
|
||
<emphasis id="B90AA33B3447F32CAB626407FBCFFC93" box="[826,1057,805,826]" italics="true" pageId="8" pageNumber="273">Scydmaenus isaloensis</emphasis>
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||
Franz
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||
</taxonomicName>
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||
(22–23) and
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<taxonomicName id="4C7E04AA3447F32CACB16404FF3FFCF0" authority="Franz" authorityName="Franz" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="madagassicus">
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||
<emphasis id="B90AA33B3447F32CACB16404FAE6FC92" box="[1257,1288,806,827]" italics="true" pageId="8" pageNumber="273">Sc.</emphasis>
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||
<emphasis id="B90AA33B3447F32CAD566407FA72FC93" box="[1294,1436,805,826]" italics="true" pageId="8" pageNumber="273">madagassicus</emphasis>
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||
Franz
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||
</taxonomicName>
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||
(24–25). Aedeagus in ventral (22, 24) and lateral (23, 25) views.
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||
</paragraph>
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||
</caption>
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||
<subSubSection id="C3642CA23447F32CA8CF64A4FB86FBA2" pageId="8" pageNumber="273" type="materials_examined">
|
||
<paragraph id="8BC17F293447F32CA8CF64A4FB86FBA2" blockId="8.[151,1437,902,2007]" pageId="8" pageNumber="273">
|
||
<materialsCitation id="3B1675743447F32CA8CF64A4FE34FBA2" ID-GBIF-Occurrence="1413803495" collectionCode="NHMW" collectorName="Foret de Sakahara & H. Franz & Mg" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="8" pageNumber="273" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
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||
<emphasis id="B90AA33B3447F32CA8CF64A4FD5DFC36" bold="true" box="[151,691,902,927]" pageId="8" pageNumber="273">
|
||
<typeStatus id="54C5C18B3447F32CA8CF64A4FF3CFC36" box="[151,210,902,927]" pageId="8" pageNumber="273">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54C5C18B3447F32CA90164A4FE28FC36" box="[345,454,902,927]" pageId="8" pageNumber="273" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<collectingCountry id="F3693FB93447F32CA98464A4FD43FC36" box="[476,685,902,927]" name="Madagascar" pageId="8" pageNumber="273">MADAGASCAR</collectingCountry>
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||
)
|
||
</emphasis>
|
||
:
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||
<specimenCount id="9D78B4A03447F32CAA9164A5FD0EFC37" box="[713,736,903,926]" pageId="8" pageNumber="273" type="male">♂</specimenCount>
|
||
, two labels (Fig.): "S-Madagaskar /
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||
<collectorName id="268B1AFF3447F32CACF664A5FA6BFC36" box="[1198,1413,902,927]" pageId="8" pageNumber="273">Forêt de Sakahara</collectorName>
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||
/ lg.
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||
<collectorName id="268B1AFF3447F32CA8E86488FEE0FC6A" box="[176,270,938,963]" pageId="8" pageNumber="273">H.Franz</collectorName>
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||
1969" [white, printed] with "
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||
<collectorName id="268B1AFF3447F32CAADD6489FD41FC6A" box="[645,687,939,963]" pageId="8" pageNumber="273">Mg</collectorName>
|
||
18" handwritten in blue on the reverse side;
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||
<taxonomicName id="4C7E04AA3447F32CACAF6488FA6AFC6A" box="[1271,1412,938,963]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="274" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
|
||
/ (
|
||
<taxonomicName id="4C7E04AA3447F32CA8C664ECFEB7FC4E" box="[158,345,974,999]" class="Insecta" family="Staphylinidae" genus="Geoscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="267" phylum="Arthropoda" rank="genus">Geoscydmaenus</taxonomicName>
|
||
) /
|
||
<taxonomicName id="4C7E04AA3447F32CA92D64ECFDE1FC4E" box="[373,527,974,999]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="madagassicus">madagassicus</taxonomicName>
|
||
m. / det.
|
||
<determinerName id="7E1D93ED3447F32CAA2864ECFD23FC4E" box="[624,717,974,999]" pageId="8" pageNumber="273">H.Franz</determinerName>
|
||
" [white, handwritten and printed]; "
|
||
<specimenCount id="9D78B4A03447F32CAC0664EDFB9BFC4E" box="[1118,1141,975,999]" pageId="8" pageNumber="273" type="male">♂</specimenCount>
|
||
" [white, printed]; "
|
||
<typeStatus id="54C5C18B3447F32CAD1264EDFA7DFC4E" box="[1354,1427,975,999]" pageId="8" pageNumber="273">Typus</typeStatus>
|
||
" [red, handwritten] (
|
||
<collectionCode id="ED6FE7EC3447F32CA92864D1FE3DFBA2" box="[368,467,1011,1035]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="273">NHMW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B1675743447F32CA9BC64D0FB8AFBA2" ID-GBIF-Occurrence="1413803493" box="[484,1124,1010,1035]" collectionCode="NHMW" collectorName="Foret de Sakahara & H. Franz & Mg" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="8" pageNumber="273" specimenCount="2" typeStatus="paratype">
|
||
<emphasis id="B90AA33B3447F32CA9BC64D0FD88FBA2" bold="true" box="[484,614,1010,1035]" pageId="8" pageNumber="273">
|
||
<typeStatus id="54C5C18B3447F32CA9BC64D0FD8FFBA2" box="[484,609,1010,1035]" pageId="8" pageNumber="273" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
23 exx., same data as for
|
||
<typeStatus id="54C5C18B3447F32CABD164D0FC05FBA2" box="[905,1003,1010,1035]" pageId="8" pageNumber="273" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED6FE7EC3447F32CABA264D1FBB2FBA2" box="[1018,1116,1011,1035]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="273">NHMW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3642CA23447F32CA89F6334FC24FB32" pageId="8" pageNumber="273" type="diagnosis">
|
||
<paragraph id="8BC17F293447F32CA89F6334FC24FB32" blockId="8.[151,1437,902,2007]" pageId="8" pageNumber="273">
|
||
<emphasis id="B90AA33B3447F32CA89F6334FE4FFB86" bold="true" box="[199,417,1046,1071]" pageId="8" pageNumber="273">Revised diagnosis.</emphasis>
|
||
Head broadest near posterior third of vertex, moderately narrowing anteriorly; profemur of male with clavate part strongly convex dorsally but weakly ventrally, with short and straight ventral projection located between proximal third and middle; protibia of male gradually broadened distally and nearly straight; aedeagus with indistinct median apical notch not separating lateral lobes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3642CA23447F32CA89F6384FD3AF8C2" pageId="8" pageNumber="273" type="description">
|
||
<paragraph id="8BC17F293447F32CA89F6384FDECFB4A" blockId="8.[151,1437,902,2007]" pageId="8" pageNumber="273">
|
||
<emphasis id="B90AA33B3447F32CA89F6384FE9AFB16" bold="true" box="[199,372,1190,1215]" pageId="8" pageNumber="273">Redescription.</emphasis>
|
||
Male very similar to
|
||
<taxonomicName id="4C7E04AA3447F32CAA286385FCE7FB17" box="[624,777,1190,1214]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="isaloensis">
|
||
<emphasis id="B90AA33B3447F32CAA286385FCE7FB17" box="[624,777,1190,1214]" italics="true" pageId="8" pageNumber="273">Sc. isaloensis</emphasis>
|
||
</taxonomicName>
|
||
, only differences and measurements are given below. BL
|
||
<quantity id="4C86D2CC3447F32CA8CF63E9FED4FB4B" box="[151,314,1227,1251]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.299999999999999" metricValueMin="8.0" pageId="8" pageNumber="273" unit="mm" value="0.815" valueMax="0.83" valueMin="0.8">0.80–0.83 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CA9D663E9FE18FB4A" box="[398,502,1227,1251]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="8" pageNumber="273" unit="mm" value="0.81">0.81 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC17F293447F32CA89F63CCFBB9FA3F" blockId="8.[151,1437,902,2007]" pageId="8" pageNumber="273">
|
||
Head (
|
||
<figureCitation id="134563AC3447F32CA94B63CCFE97FAAE" box="[275,377,1262,1287]" captionStart="FIGURES 1 – 8" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[168,1436,295,1860]" captionTargetId="figure@4.[151,1436,273,1860]" captionTargetPageId="4" captionText="FIGURES 1 – 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 – 4) and Sc. madagassicus Franz (2, 5 – 8). Set of original labels of holotype (1 – 2); dorsal habitus of male (3); head and prothorax in dorsal (4 – 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="8" pageNumber="273">Figs 5–8</figureCitation>
|
||
) broadest near posterior third of vertex and weakly narrowing anteriorly, in most specimens large punctures covering nearly entire surface of frons and vertex, but often small median area remains almost impunctate; HL
|
||
<quantity id="4C86D2CC3447F32CA90A6215FE19FAE7" box="[338,503,1335,1359]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="8" pageNumber="273" unit="mm" value="0.135" valueMax="0.14" valueMin="0.13">0.13–0.14 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CAA176215FD57FAE6" box="[591,697,1335,1359]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="273" unit="mm" value="0.13">0.13 mm</quantity>
|
||
), HW
|
||
<quantity id="4C86D2CC3447F32CAB5D6214FC80FAE7" box="[773,878,1334,1359]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="273" unit="mm" value="0.15">0.15 mm</quantity>
|
||
; AnL 0.3 9–0.43 (mean
|
||
<quantity id="4C86D2CC3447F32CACD86215FB04FAE6" box="[1152,1258,1335,1359]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="8" pageNumber="273" unit="mm" value="0.41">0.41 mm</quantity>
|
||
). Antenna (
|
||
<figureCitation id="134563AC3447F32CAD376214FF5AFADA" captionStart="FIGURES 12 – 15" captionStartId="6.[151,264,1482,1505]" captionTargetBox="[151,1427,210,1461]" captionTargetId="figure@6.[151,1436,193,1461]" captionTargetPageId="6" captionText="FIGURES 12 – 15. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (12, 14) and Sc. madagassicus Franz (13, 15). Right antenna of male in dorsal view (12 – 13); right fore leg of male in anterior view." httpUri="https://zenodo.org/record/194332/files/figure.png" pageId="8" pageNumber="273">Fig. 13</figureCitation>
|
||
) slightly longer in relation to body length than that of
|
||
<taxonomicName id="4C7E04AA3447F32CAB7C6279FC52FADA" box="[804,956,1371,1395]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="isaloensis">
|
||
<emphasis id="B90AA33B3447F32CAB7C6279FC52FADA" box="[804,956,1371,1395]" italics="true" pageId="8" pageNumber="273">Sc. isaloensis</emphasis>
|
||
</taxonomicName>
|
||
, with distinctly elongate antennomere VI, slightly elongate antennomere IX and distinctly asymmetrical antennomeres VI–VIII.
|
||
</paragraph>
|
||
<paragraph id="8BC17F293447F32CA89F6281FD98FA76" blockId="8.[151,1437,902,2007]" pageId="8" pageNumber="273">
|
||
Pronotum (
|
||
<figureCitation id="134563AC3447F32CA91E6280FE64FA12" box="[326,394,1442,1467]" captionStart="FIGURES 1 – 8" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[168,1436,295,1860]" captionTargetId="figure@4.[151,1436,273,1860]" captionTargetPageId="4" captionText="FIGURES 1 – 8. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (1, 3 – 4) and Sc. madagassicus Franz (2, 5 – 8). Set of original labels of holotype (1 – 2); dorsal habitus of male (3); head and prothorax in dorsal (4 – 5) and ventral (6) views; head in ventral (7) and lateral (8) view. Abbreviations: bst, basisternal part of prosternum; fr, frons; gp, gular plate; hr, hypostomal ridge; hy, hypomeron; occ, occipital constriction; ptp, posterior tentorial pit; vt, vertex." httpUri="https://zenodo.org/record/194330/files/figure.png" pageId="8" pageNumber="273">Fig. 5</figureCitation>
|
||
) of the same shape and proportions as that in
|
||
<taxonomicName id="4C7E04AA3447F32CABC56281FBDBFA12" box="[925,1077,1443,1467]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="isaloensis">
|
||
<emphasis id="B90AA33B3447F32CABC56281FBDBFA12" box="[925,1077,1443,1467]" italics="true" pageId="8" pageNumber="273">Sc. isaloensis</emphasis>
|
||
</taxonomicName>
|
||
; PL
|
||
<quantity id="4C86D2CC3447F32CAC376281FAFAFA13" box="[1135,1300,1443,1467]" metricMagnitude="-4" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="8" pageNumber="273" unit="mm" value="0.205" valueMax="0.21" valueMin="0.2">0.20–0.21 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CAD336281FF29FA76" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="273" unit="mm" value="0.21">0.21 mm</quantity>
|
||
), PW
|
||
<quantity id="4C86D2CC3447F32CA95362E5FE40FA77" box="[267,430,1479,1503]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.7999999999999998" pageId="8" pageNumber="273" unit="mm" value="0.19" valueMax="0.2" valueMin="0.18">0.18–0.20 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CAA5962E5FD87FA76" box="[513,617,1479,1503]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="273" unit="mm" value="0.19">0.19 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC17F293447F32CA89F62C8FB25F9AA" blockId="8.[151,1437,902,2007]" box="[199,1227,1514,1539]" pageId="8" pageNumber="273">
|
||
Elytra as in
|
||
<taxonomicName id="4C7E04AA3447F32CA91562C9FE0DF9AA" box="[333,483,1515,1539]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="isaloensis">
|
||
<emphasis id="B90AA33B3447F32CA91562C9FE0DF9AA" box="[333,483,1515,1539]" italics="true" pageId="8" pageNumber="273">Sc. isaloensis</emphasis>
|
||
</taxonomicName>
|
||
; EL
|
||
<quantity id="4C86D2CC3447F32CAA4462C9FD6CF9AB" box="[540,642,1515,1539]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="273" unit="mm" value="0.48">0.48 mm</quantity>
|
||
, EW
|
||
<quantity id="4C86D2CC3447F32CAA9862C9FC8DF9AB" box="[704,867,1515,1539]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="8" pageNumber="273" unit="mm" value="0.29" valueMax="0.3" valueMin="0.28">0.28–0.30 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CABEF62C9FBF1F9AA" box="[951,1055,1515,1539]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="273" unit="mm" value="0.28">0.28 mm</quantity>
|
||
), EI 1.58–1.73.
|
||
</paragraph>
|
||
<paragraph id="8BC17F293447F32CA89F612DFF1DF93A" blockId="8.[151,1437,902,2007]" pageId="8" pageNumber="273">
|
||
Legs (
|
||
<figureCitation id="134563AC3447F32CA956612CFE8FF98E" box="[270,353,1550,1575]" captionStart="FIGURES 12 – 15" captionStartId="6.[151,264,1482,1505]" captionTargetBox="[151,1427,210,1461]" captionTargetId="figure@6.[151,1436,193,1461]" captionTargetPageId="6" captionText="FIGURES 12 – 15. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (12, 14) and Sc. madagassicus Franz (13, 15). Right antenna of male in dorsal view (12 – 13); right fore leg of male in anterior view." httpUri="https://zenodo.org/record/194332/files/figure.png" pageId="8" pageNumber="273">Fig. 15</figureCitation>
|
||
) with profemur less thickened in distal third, its clavate part strongly convex dorsally but only weakly so ventrally; ventral femoral projection short, straight and perpendicular to the long axis of femur, not accompanied by long seta and with evenly rounded apex; protibia (
|
||
<figureCitation id="134563AC3447F32CABD96174FC3CF9C6" box="[897,978,1622,1647]" captionStart="FIGURES 12 – 15" captionStartId="6.[151,264,1482,1505]" captionTargetBox="[151,1427,210,1461]" captionTargetId="figure@6.[151,1436,193,1461]" captionTargetPageId="6" captionText="FIGURES 12 – 15. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (12, 14) and Sc. madagassicus Franz (13, 15). Right antenna of male in dorsal view (12 – 13); right fore leg of male in anterior view." httpUri="https://zenodo.org/record/194332/files/figure.png" pageId="8" pageNumber="273">Fig. 15</figureCitation>
|
||
) weakly and gradually thickened, almost straight.
|
||
</paragraph>
|
||
<paragraph id="8BC17F293447F32CA89F61BCFB91F972" blockId="8.[151,1437,902,2007]" pageId="8" pageNumber="273">
|
||
Aedeagus (
|
||
<figureCitation id="134563AC3447F32CA91361BCFE3AF91E" box="[331,468,1694,1719]" captionStart="FIGURES 16 – 21" captionStartId="7.[151,264,1906,1929]" captionTargetBox="[164,1415,224,1854]" captionTargetId="figure@7.[151,1436,193,1885]" captionTargetPageId="7" captionText="FIGURES 16 – 21. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (16 – 18) and Sc. madagassicus Franz (19 – 21). Aedeagus in ventral (16, 19) and lateral (17 – 18, 20) view (with partly exserted flagellum in Fig. 18); spermatheca (21)." httpUri="https://zenodo.org/record/194333/files/figure.png" pageId="8" pageNumber="273">Figs 19–20</figureCitation>
|
||
,
|
||
<figureCitation id="134563AC3447F32CA9BF61BCFDC1F91E" box="[487,559,1694,1719]" captionStart="FIGURES 22 – 25" captionStartId="8.[151,264,804,827]" captionTargetBox="[163,1428,208,766]" captionTargetId="figure@8.[151,1436,193,784]" captionTargetPageId="8" captionText="FIGURES 22 – 25. Scydmaenus (Geoscydmaenus) of Madagascar. Scydmaenus isaloensis Franz (22 – 23) and Sc. madagassicus Franz (24 – 25). Aedeagus in ventral (22, 24) and lateral (23, 25) views." httpUri="https://zenodo.org/record/194334/files/figure.png" pageId="8" pageNumber="273">24–25</figureCitation>
|
||
) elongate, AeL
|
||
<quantity id="4C86D2CC3447F32CAAA161BCFC89F91F" box="[761,871,1694,1719]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="273" unit="mm" value="0.25">0.25 mm</quantity>
|
||
; median lobe in ventral view with indistinctly demarcated apical portion, apex with indistinct median notch not separating lateral lobes.
|
||
</paragraph>
|
||
<paragraph id="8BC17F293447F32CA89F61C4FD3AF8C2" blockId="8.[151,1437,902,2007]" pageId="8" pageNumber="273">
|
||
<emphasis id="B90AA33B3447F32CA89F61C4FECBF956" bold="true" box="[199,293,1766,1791]" pageId="8" pageNumber="273">Female.</emphasis>
|
||
Similar to male, except for distinctly more slender profemora lacking ventral projections. BL
|
||
<quantity id="4C86D2CC3447F32CAD0561C5FF12F88B" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.4" metricValueMin="7.9" pageId="8" pageNumber="273" unit="mm" value="0.815" valueMax="0.84" valueMin="0.79">0.79– 0.84 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CA90A6029FE55F88A" box="[338,443,1803,1827]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="273" unit="mm" value="0.82">0.82 mm</quantity>
|
||
); HL
|
||
<quantity id="4C86D2CC3447F32CA9A56029FD4FF88B" box="[509,673,1803,1827]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="8" pageNumber="273" unit="mm" value="0.135" valueMax="0.14" valueMin="0.13">0.13–0.14 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CAAAF6029FC8EF88A" box="[759,864,1803,1827]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="273" unit="mm" value="0.13">0.13 mm</quantity>
|
||
), HW
|
||
<quantity id="4C86D2CC3447F32CABF36028FBA1F88B" box="[939,1103,1802,1827]" metricMagnitude="-4" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="8" pageNumber="273" unit="mm" value="0.155" valueMax="0.16" valueMin="0.15">0.15–0.16 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CACFD6028FAE0F88A" box="[1189,1294,1802,1827]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="273" unit="mm" value="0.16">0.16 mm</quantity>
|
||
), AnL
|
||
<quantity id="4C86D2CC3447F32CAD056029FF15F8EF" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="8" pageNumber="273" unit="mm" value="0.405" valueMax="0.43" valueMin="0.38">0.38– 0.43 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CA917600DFE59F8EE" box="[335,439,1839,1863]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="8" pageNumber="273" unit="mm" value="0.41">0.41 mm</quantity>
|
||
); PL
|
||
<quantity id="4C86D2CC3447F32CA9AA600DFD7BF8EF" box="[498,661,1839,1863]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="8" pageNumber="273" unit="mm" value="0.22" valueMax="0.23" valueMin="0.21">0.21–0.23 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CAAB1600DFCBFF8EE" box="[745,849,1839,1863]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="273" unit="mm" value="0.22">0.22 mm</quantity>
|
||
), PW
|
||
<quantity id="4C86D2CC3447F32CABCD600DFC13F8EF" box="[917,1021,1839,1863]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="273" unit="mm" value="0.2">0.20 mm</quantity>
|
||
; EL
|
||
<quantity id="4C86D2CC3447F32CAC69600CFB3BF8EF" box="[1073,1237,1838,1863]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="8" pageNumber="273" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CAD70600CFA7EF8EE" box="[1320,1424,1838,1863]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="273" unit="mm" value="0.46">0.46 mm</quantity>
|
||
), EW
|
||
<quantity id="4C86D2CC3447F32CA8916071FE82F8C3" box="[201,364,1875,1899]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="8" pageNumber="273" unit="mm" value="0.29" valueMax="0.3" valueMin="0.28">0.28–0.30 mm</quantity>
|
||
(mean
|
||
<quantity id="4C86D2CC3447F32CA9986071FDC6F8C2" box="[448,552,1875,1899]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="273" unit="mm" value="0.28">0.28 mm</quantity>
|
||
), EI 1.58–1.73.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3642CA23447F32CA89F6054FC38F826" box="[199,982,1910,1935]" pageId="8" pageNumber="273" type="distribution">
|
||
<paragraph id="8BC17F293447F32CA89F6054FC38F826" blockId="8.[151,1437,902,2007]" box="[199,982,1910,1935]" pageId="8" pageNumber="273">
|
||
<emphasis id="B90AA33B3447F32CA89F6054FE8EF826" bold="true" box="[199,352,1910,1935]" pageId="8" pageNumber="273">Distribution.</emphasis>
|
||
South-western
|
||
<collectingCountry id="F3693FB93447F32CAA486054FD77F826" box="[528,665,1910,1935]" name="Madagascar" pageId="8" pageNumber="273">Madagascar</collectingCountry>
|
||
, Atsimo-Andrefana Region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3642CA23447F32DA89F60B8FF01FD94" lastPageId="9" lastPageNumber="274" pageId="8" pageNumber="273" type="discussion">
|
||
<paragraph id="8BC17F293447F32CA89F60B8FA73F87E" blockId="8.[151,1437,902,2007]" pageId="8" pageNumber="273">
|
||
<emphasis id="B90AA33B3447F32CA89F60B8FED5F81A" bold="true" box="[199,315,1946,1971]" pageId="8" pageNumber="273">Remarks.</emphasis>
|
||
<bibRefCitation id="EFEF02D83447F32CA91E60B9FE32F81A" author="Franz" box="[326,476,1946,1971]" pageId="8" pageNumber="273" refString="Franz, H. (1986) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mit Ausschluss der Cephenniini). Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986)</bibRefCitation>
|
||
stated that
|
||
<taxonomicName id="4C7E04AA3447F32CAA3160B9FCDDF81A" box="[617,819,1947,1971]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="madagassicus">
|
||
<emphasis id="B90AA33B3447F32CAA3160B9FCDDF81A" box="[617,819,1947,1971]" italics="true" pageId="8" pageNumber="273">Sc. madagassicus</emphasis>
|
||
</taxonomicName>
|
||
had the head more elongate than
|
||
<taxonomicName id="4C7E04AA3447F32CAC9360B9FA88F81A" box="[1227,1382,1947,1971]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="isaloensis">
|
||
<emphasis id="B90AA33B3447F32CAC9360B9FA88F81A" box="[1227,1382,1947,1971]" italics="true" pageId="8" pageNumber="273">Sc. isaloensis</emphasis>
|
||
</taxonomicName>
|
||
; the pronotum in relation to the head broader than that in
|
||
<taxonomicName id="4C7E04AA3447F32CAB4C609DFC5EF87E" box="[788,944,1983,2007]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="isaloensis">
|
||
<emphasis id="B90AA33B3447F32CAB4C609DFC5EF87E" box="[788,944,1983,2007]" italics="true" pageId="8" pageNumber="273">Sc. isaloensis</emphasis>
|
||
</taxonomicName>
|
||
; and the profemur of male lacking tooth.
|
||
</paragraph>
|
||
<paragraph id="8BC17F293446F32DA8CF67BAFF01FD94" blockId="9.[151,1437,151,573]" pageId="9" pageNumber="274">
|
||
Moreover, the aedeagus of this species was described as "völlig anders gebaut" ("entirely differently built"), but the aedeagus of
|
||
<taxonomicName id="4C7E04AA3446F32DA970679EFE2FFF7D" box="[296,449,188,212]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="274" phylum="Arthropoda" rank="species" species="isaloensis">
|
||
<emphasis id="B90AA33B3446F32DA970679EFE2FFF7D" box="[296,449,188,212]" italics="true" pageId="9" pageNumber="274">Sc. isaloensis</emphasis>
|
||
</taxonomicName>
|
||
was illustrated only in the lateral view, whereas that of
|
||
<taxonomicName id="4C7E04AA3446F32DAC15679EFAF8FF7D" box="[1101,1302,188,212]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="274" phylum="Arthropoda" rank="species" species="madagassicus">
|
||
<emphasis id="B90AA33B3446F32DAC15679EFAF8FF7D" box="[1101,1302,188,212]" italics="true" pageId="9" pageNumber="274">Sc. madagassicus</emphasis>
|
||
</taxonomicName>
|
||
only in the dorsal view, so it is not possible to see any fundamental differences (especially knowing that Franz frequently illustrated aedeagi in a rather schematic and inaccurate way). Examination of the
|
||
<typeStatus id="54C5C18B3446F32DAC3A6627FB7CFEB4" box="[1122,1170,261,285]" pageId="9" pageNumber="274">type</typeStatus>
|
||
series of both species revealed that the head is equally long and broad in
|
||
<taxonomicName id="4C7E04AA3446F32DAA89660AFC86FEE9" box="[721,872,296,320]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="274" phylum="Arthropoda" rank="species" species="isaloensis">
|
||
<emphasis id="B90AA33B3446F32DAA89660AFC86FEE9" box="[721,872,296,320]" italics="true" pageId="9" pageNumber="274">Sc. isaloensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E04AA3446F32DABC6660AFB8AFEE9" box="[926,1124,296,320]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="274" phylum="Arthropoda" rank="species" species="madagassicus">
|
||
<emphasis id="B90AA33B3446F32DABC6660AFB8AFEE9" box="[926,1124,296,320]" italics="true" pageId="9" pageNumber="274">Sc. madagassicus</emphasis>
|
||
</taxonomicName>
|
||
; the species do not differ in the width of pronotum in relation to the width of head; and males of both species have the ventral femoral tooth. Consequently, all external morphological characters used by Franz to define these species have been misinterpreted, and his illustrations of aedeagi cannot be used to find any differences. Useful differences in the shape of profemora and tibiae, in the shape of the head; in the punctures on frons and vertex; and in proportions and shapes of antennomeres, were found during the present study; also the aedeagi truly differ in the general shape, although not so much in internal endophallic structures. Clearly these are closely related, but easily identifiable species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |