499 lines
62 KiB
XML
499 lines
62 KiB
XML
<document id="181DDE03C93D4DDCEFBD64172693C1F6" ID-CLB-Dataset="46719" ID-DOI="10.5281/zenodo.205110" ID-GBIF-Dataset="c8bab4f7-3ca7-429b-a361-7e0a06eb5c29" ID-ISSN="1175-5326" ID-Zenodo-Dep="205110" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460365820333" checkinUser="plazi" docAuthor="Fery, Hans & Hendrich, Lars" docDate="2011" docId="03B7A125221CFF90FF3AFC790F1AFB82" docLanguage="en" docName="zt02909p046.pdf" docOrigin="Zootaxa 2909" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hydroporus esseri Fery & Hendrich, 2011, sp. n." docType="treatment" docVersion="10" lastPageNumber="43" masterDocId="FF8ED95D221DFF95FFADFF880E01FFB6" masterDocTitle="Hydroporus esseri sp. n., a new diving beetle from southern Turkey (Coleoptera, Dytiscidae, Hydroporinae)" masterLastPageNumber="46" masterPageNumber="38" pageNumber="39" updateTime="1698295185274" updateUser="ExternalLinkService">
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<mods:mods id="732BFF02AF7692C43F61F9ACCCFD22D5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="359EC304521D33E3B357DCB3FDAD6DC1">
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<mods:title id="5DA458C18175087219E072496514D55C">Hydroporus esseri sp. n., a new diving beetle from southern Turkey (Coleoptera, Dytiscidae, Hydroporinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A26A38A67F82633DABD03B6BC01473FF" type="personal">
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<mods:role id="F5AC20AA4C1C4429F064CA313709F0AD">
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<mods:roleTerm id="EED06A502C457289960DA94292796F30">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="67ACC9EE036D627FAFE1E39E60C531DC">Fery, Hans</mods:namePart>
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</mods:name>
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<mods:name id="BC4896878090F23544B4AF51035EE1C5" type="personal">
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<mods:role id="303991D0B4553DB0020ED99D2147885D">
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<mods:roleTerm id="F56EC82A6096026C3279F7A62BF3E3C9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="319088AB35024E24CBE30ACC7AC1E563">Hendrich, Lars</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="2843A57E37BD7E6588073C1F00A9D836">text</mods:typeOfResource>
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<mods:relatedItem id="61EC253BCC27E8CC7E258EA0C6DDE36A" type="host">
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<mods:titleInfo id="3FCD4DF6253DDE18A273EC50266F5AEE">
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<mods:title id="EFA27B2974321C69E50BBE24323D96D9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1A007FC1CF352F58674C536CE1B3D84D">
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<mods:date id="F8246DDD9F80AEBC63EDB947542E18D1">2011</mods:date>
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<mods:detail id="B7067E1F6130C752F335472CF4087F6F" type="volume">
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<mods:number id="2EC67890A276FA91CA1611A757909D1A">2909</mods:number>
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</mods:detail>
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<mods:extent id="FED371767B238D697021A8C57D8A800E" unit="page">
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<mods:start id="1061E4A628A0AFB12D9B03E469EC243E">38</mods:start>
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<mods:end id="5A0F3A2B15DA0080201340D436F0C509">46</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="80CA874C59B0B5BE7951E87765C41AB8">journal article</mods:classification>
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<mods:identifier id="9B58EF46A1DE57A1E97EBF5D99569C8D" type="CLB-Dataset">46719</mods:identifier>
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<mods:identifier id="EF185F559385B2E99AD2C507ED187658" type="DOI">10.5281/zenodo.205110</mods:identifier>
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<mods:identifier id="F5D381CCE5C85F09F235E8C1C8B7FEE6" type="GBIF-Dataset">c8bab4f7-3ca7-429b-a361-7e0a06eb5c29</mods:identifier>
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<mods:identifier id="5DC2AC62DE14F7947D95274C691822BD" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="FE133DEF6110F29C15EB38C1562164EE" type="Zenodo-Dep">205110</mods:identifier>
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</mods:mods>
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<treatment id="03B7A125221CFF90FF3AFC790F1AFB82" ID-DOI="http://doi.org/10.5281/zenodo.6184646" ID-GBIF-Taxon="119410765" ID-Zenodo-Dep="6184646" LSID="urn:lsid:plazi:treatment:03B7A125221CFF90FF3AFC790F1AFB82" httpUri="http://treatment.plazi.org/id/03B7A125221CFF90FF3AFC790F1AFB82" lastPageId="5" lastPageNumber="43" pageId="1" pageNumber="39">
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<subSubSection id="C30443B8221CFF94FF3AFC790FBCFBBC" box="[151,445,1008,1035]" pageId="1" pageNumber="39" type="nomenclature">
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<paragraph id="8BA11033221CFF94FF3AFC790FBCFBBC" blockId="1.[151,445,1008,1035]" box="[151,445,1008,1035]" pageId="1" pageNumber="39">
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<heading id="D0E9A75F221CFF94FF3AFC790FBCFBBC" bold="true" box="[151,445,1008,1035]" fontSize="11" level="1" pageId="1" pageNumber="39" reason="1">
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<emphasis id="B96ACC21221CFF94FF3AFC790FBCFBBC" bold="true" box="[151,445,1008,1035]" pageId="1" pageNumber="39">
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<taxonomicName id="4C1E6BB0221CFF94FF3AFC790F72FBBD" ID-CoL="8LZLV" box="[151,371,1009,1035]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="39" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
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<emphasis id="B96ACC21221CFF94FF3AFC790F72FBBD" bold="true" box="[151,371,1009,1035]" italics="true" pageId="1" pageNumber="39">Hydroporus esseri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A259715A221CFF94FED6FC780FBCFBBC" box="[379,445,1008,1034]" pageId="1" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30443B8221CFF97FF3AFBBB0B97FA76" lastPageId="2" lastPageNumber="40" pageId="1" pageNumber="39" type="materials_examined">
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<paragraph id="8BA11033221CFF94FF3AFBBB0CA9FBD8" blockId="1.[151,1437,1074,2034]" pageId="1" pageNumber="39">
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<emphasis id="B96ACC21221CFF94FF3AFBBB0F37FBFD" bold="true" box="[151,310,1074,1099]" pageId="1" pageNumber="39">
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<typeStatus id="54A5AE91221CFF94FF3AFBBB0ED1FBFD" box="[151,208,1075,1099]" pageId="1" pageNumber="39">Type</typeStatus>
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locality.
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</emphasis>
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Southern
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<collectingCountry id="F30950A3221CFF94FE1DFBBA0C00FBFC" box="[432,513,1074,1098]" name="Turkey" pageId="1" pageNumber="39">Turkey</collectingCountry>
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, Antalya province, Manavgat district, road Sagirin–Beskonak, Köprülü Canyon;
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<geoCoordinate id="EE2A76F4221CFF94FF3AFBDE0F2EFBD8" box="[151,303,1110,1134]" direction="north" orientation="latitude" pageId="1" pageNumber="39" precision="1" value="37.117783">37.1177816N</geoCoordinate>
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<geoCoordinate id="EE2A76F4221CFF94FE9BFBDE0FCCFBD8" box="[310,461,1110,1134]" direction="east" orientation="longitude" pageId="1" pageNumber="39" precision="1" value="31.211155">31.2111551E</geoCoordinate>
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, altitude ca.
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<specimenCount id="9D18DBBA221CFF94FDF7FBDE0CA9FBD8" box="[602,680,1110,1134]" pageId="1" pageNumber="39" type="generic">
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||
<quantity id="4CE6BDD6221CFF94FDF7FBDE0CA5FBD8" box="[602,676,1110,1134]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="39" unit="m" value="150.0">150 m</quantity>
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.
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</specimenCount>
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</paragraph>
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<paragraph id="8BA11033221CFF94FF6AFBF30FEDFB48" blockId="1.[151,1437,1074,2034]" pageId="1" pageNumber="39">
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<emphasis id="B96ACC21221CFF94FF6AFBF30FD8FB25" bold="true" box="[199,473,1146,1171]" pageId="1" pageNumber="39">
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<typeStatus id="54A5AE91221CFF94FF6AFBF30F03FB25" box="[199,258,1147,1171]" pageId="1" pageNumber="39">Type</typeStatus>
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material studied.
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</emphasis>
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<typeStatus id="54A5AE91221CFF94FE49FBF20C4DFB24" box="[484,588,1146,1170]" pageId="1" pageNumber="39" type="holotype">Holotype</typeStatus>
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(male): "
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<date id="FFA036F3221CFF94FD17FBF20D2CFB24" box="[698,813,1146,1170]" pageId="1" pageNumber="39" value="2002-03-17">17.3.2002</date>
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<collectingCountry id="F30950A3221CFF94FC9AFBF20D86FB24" box="[823,903,1146,1170]" name="Turkey" pageId="1" pageNumber="39">Turkey</collectingCountry>
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Antalya prov., Manvagat distr., road Sagirin– Beskonak", "Köprülü Canyon, Köprülü River, backwater pool,
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<geoCoordinate id="EE2A76F4221CFF94FCC4FB160A03FB00" box="[873,1026,1182,1206]" direction="north" orientation="latitude" pageId="1" pageNumber="39" precision="1" value="37.117783">37.1177816N</geoCoordinate>
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<geoCoordinate id="EE2A76F4221CFF94FBA7FB160AA0FB00" box="[1034,1185,1182,1206]" direction="east" orientation="longitude" pageId="1" pageNumber="39" precision="1" value="31.211155">31.2111551E</geoCoordinate>
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,
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<quantity id="4CE6BDD6221CFF94FB03FB160AF8FB00" box="[1198,1273,1182,1206]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="39" unit="m" value="150.0">150 m</quantity>
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, J. Esser leg". [white printed labels]. "
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<typeStatus id="54A5AE91221CFF94FE0EFB4A0C10FB6C" box="[419,529,1218,1242]" pageId="1" pageNumber="39" type="holotype">Holotype</typeStatus>
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,
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<taxonomicName id="4C1E6BB0221CFF94FDB3FB4A0CEDFB6C" box="[542,748,1218,1242]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="39" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">Hydroporus esseri</taxonomicName>
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<emphasis id="B96ACC21221CFF94FD58FB4B0D37FB6D" bold="true" box="[757,822,1219,1243]" pageId="1" pageNumber="39">
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<taxonomicNameLabel id="A259715A221CFF94FD58FB4B0D37FB6D" box="[757,822,1219,1243]" pageId="1" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, Fery & Hendrich des. 2011" [red printed label]. The
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<typeStatus id="54A5AE91221CFF94FF3AFB6E0EF8FB48" box="[151,249,1254,1278]" pageId="1" pageNumber="39" type="holotype">holotype</typeStatus>
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is stored in the
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<collectionCode id="ED0F88F6221CFF94FE06FB6E0FE9FB48" box="[427,488,1254,1278]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="39">ZSM</collectionCode>
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.
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</paragraph>
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<paragraph id="8BA11033221CFF94FF6AFA820CA2FADC" blockId="1.[151,1437,1074,2034]" pageId="1" pageNumber="39">
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<emphasis id="B96ACC21221CFF94FF6AFA820F27FA95" bold="true" box="[199,294,1290,1315]" pageId="1" pageNumber="39">Habitus</emphasis>
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elongate oval (
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<figureCitation id="13250CB6221CFF94FE7DFA820C10FA94" box="[464,529,1290,1314]" captionStart="FIGURES 1 – 2" captionStartId="3.[151,264,1354,1377]" captionTargetBox="[276,1307,483,1331]" captionTargetId="figure@3.[276,1310,483,1331]" captionTargetPageId="3" captionText="FIGURES 1 – 2. Habitus of: (1) Hydroporus esseri sp. n., (2) Hydroporus umbrosus (specimen from Berlin, Germany)." httpUri="https://zenodo.org/record/205111/files/figure.png" pageId="1" pageNumber="39">Fig. 1</figureCitation>
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), maximum width behind middle of total length, in second third of elytral length. Pronotum largely black, head and elytra dark brown, sides of latter—in particular near shoulders—somewhat lighter. Entire surface reticulated, but not matt.
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</paragraph>
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<paragraph id="8BA11033221CFF94FF6AFAFF0A59F919" blockId="1.[151,1437,1074,2034]" pageId="1" pageNumber="39">
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||
<emphasis id="B96ACC21221CFF94FF6AFAFF0F06FA39" bold="true" box="[199,263,1399,1423]" pageId="1" pageNumber="39">Head</emphasis>
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dark brown with anterior part of clypeus (in particular above antennal cavity) and a transverse broad marking on vertex reddish brown; these lighter parts only diffusely delimited. Interocular distance about same as half of pronotal width at posterior angles. Clypeus with two longitudinal grooves behind anterior margin; these grooves rather flat and mainly apparent by distinct and dense punctation. Entire surface microreticulate with polygonal meshes, these more transverse near anterior margin of clypeus and on vertex. Punctures behind anterior margin of clypeus rather small and sparse; between clypeal grooves and on frons larger and denser, here their diameter equalling that of meshes; before vertex with some smaller punctures; vertex impunctate; beside inner margin of eyes with impressed puncture lines and meshes of reticulation replaced by small longitudinal wrinkles. Setae on head very indistinct, with only a very few in clypeal grooves and near margin of eyes.
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</paragraph>
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<paragraph id="8BA11033221CFF97FF6AF9330C09FF41" blockId="1.[151,1437,1074,2034]" lastBlockId="2.[151,1437,151,2012]" lastPageId="2" lastPageNumber="40" pageId="1" pageNumber="39">
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<emphasis id="B96ACC21221CFF94FF6AF9330F41F965" bold="true" box="[199,320,1723,1747]" pageId="1" pageNumber="39">Pronotum</emphasis>
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black; only laterally in anterior half narrowly and very diffusely brownish; lateral bead black. Bead thin, in lateral view distinctly narrower than half of diameter of antenna. Maximum width of pronotum between posterior angles; sides converging from posterior towards anterior angles, more or less evenly and slightly curved over entire length. Between disc and posterior margin distinctly depressed over about half of pronotal width, thus, disc appearing rather vaulted. Surface rather uniformly microreticulated, meshes more or less of same diameter as on head; only near anterior margin meshes more transverse. Near anterior margin unpunctured, behind without punctured line but with irregularly placed rather large punctures (larger than on clypeus); diameter larger than that of meshes, distance between puncture equalling one or at most two diameters of punctures; before posterior margin punctation similar. On disc and also more laterally with similar punctures, but punctation sparser, distance between punctures three to four times that of their diameter. Sublaterally punctures smaller, denser and near posterior angles somewhat longitudinally deformed. Except on disc most punctures with a transparent yellowish, very distinct and rather long (up to
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<quantity id="4CE6BDD6221FFF97FECCFF570FC7FF41" box="[353,454,223,247]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="40" unit="mm" value="0.05">0.05 mm</quantity>
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) seta.
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</paragraph>
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<paragraph id="8BA11033221FFF97FF6AFE8C0F0AFD8A" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="40">
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<emphasis id="B96ACC21221FFF97FF6AFE8C0F15FEAA" bold="true" box="[199,276,260,284]" pageId="2" pageNumber="40">Elytra</emphasis>
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dark brown, near sides a very little lighter, near shoulders still a little lighter; these areas very diffusely delimited; centrally elytra behind mid length without large blackish spot, here only almost imperceptibly darkened. In dorsal view sides of elytra almost evenly curved, only short before apex stronger curved, here not pointed. In lateral view elytral margin slightly ascending towards humeral angle; epipleuron visible until humeral angle. Elytral bead more or less as thin as pronotal bead, in lateral view both forming an angle at bases of pronotum and elytra. Reticulation on elytra more or less uniform, similar to that on pronotum. Punctures on elytra rather evenly distributed, somewhat larger and denser than on disc of pronotum. Elytral puncture lines absent. Setation in anterior central half less distinct (possibly rubbed off), but on rest of elytral surface very conspicuous, similar to that on pronotum.
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</paragraph>
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<paragraph id="8BA11033221FFF97FF6AFDC00F95FD7A" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="40">
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<emphasis id="B96ACC21221FFF97FF6AFDC00F81FDD6" bold="true" box="[199,384,583,608]" pageId="2" pageNumber="40">Ventral surface</emphasis>
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with head reddish brown, genae of same colour as gula. Prothoracic epipleuron (subnotum) and proepisternum dark brownish, prosternal process blackish brown; rest of ventral surface black, except dark brownish posterior part of metacoxal processes and last abdominal ventrite. Elytral epipleura brownish, near shoulders somewhat lighter.
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</paragraph>
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<paragraph id="8BA11033221FFF97FF68FD5F0D66FBEE" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="40">Prosternal process with blade lanceolate; in cross-section flatly tectiform, sides not flattened; margin only weakly beaded; apex not distinctly pointed; between procoxae without transverse ridge, but a weak protuberance; before with some extremely fine transverse carinae; anteriorly process not prolonged as narrow convexity onto prosternum; prosternum anteromedially flattened and very rugosely sculptured, here not particularly impressed. Vaulted part of lanceolate blade with several coarse punctures, near sides not flattened, but with coarse punctures and long setae. Posterior margins of metacoxal processes more or less straight and forming one line together; next to end of the midline of processes without distinct pointed prolongation. Midline of metacoxal processes very deeply impressed over its entire length. Lateral lines of metacoxal processes very distinct, carinate, weakly diverging anteriorly, reaching posterior margin of metaventrite; inside metacoxal lines with strongly impressed grooves. Second and third abdominal ventrite rigidly connected, without traces of a suture; however, when adequately illuminated a very flat impression perceptible instead of that suture.</paragraph>
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<paragraph id="8BA11033221FFF97FF68FBEC0DCBFA86" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="40">Almost entire ventral surface with reticulation, but not appearing matt; gula without reticulation. Sides of metaventrite, metacoxal plates, and first two abdominal ventrites with very coarse punctures; on centre of metaventrite and third to fifth abdominal ventrites less coarsely punctured; last abdominal ventrite very coarsely and densely punctured, particularly at apex; intralinear space of metacoxal processes only with a few small punctures, but impressed grooves beside metacoxal lines coarsely and densely punctured. Distinct and rather long setae present on metaventrite, metacoxal plates, abdominal ventrites and epipleura.</paragraph>
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<paragraph id="8BA11033221FFF97FF68FAB40B97FA76" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="40">Palpi, mouthparts and legs (including coxae and trochanters) brownish; pro- and mesotarsomeres slightly darkened; first five antennomeres lighter brownish, rest rather dark brownish. Third to sixth antennomeres rather short, fourth especially short, eleventh almost two times as long as tenth. First three pro- and mesotarsomeres with sucker hairs, but without sucker cups. Protarsal claws small and simple, without apparent secondary sexual modifications.</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30443B8221FFF90FF68FA440A9FFE42" lastPageId="5" lastPageNumber="43" pageId="2" pageNumber="40" type="description">
|
||
<paragraph id="8BA11033221FFF97FF68FA440C17F9BE" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="40">
|
||
<emphasis id="B96ACC21221FFF97FF68FA440F7AFA52" bold="true" box="[197,379,1484,1508]" pageId="2" pageNumber="40">Measurements.</emphasis>
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<typeStatus id="54A5AE91221FFF97FE2EFA440FF1FA52" box="[387,496,1484,1508]" pageId="2" pageNumber="40" type="holotype">Holotype</typeStatus>
|
||
: total length
|
||
<quantity id="4CE6BDD6221FFF97FD2FFA440CDDFA52" box="[642,732,1484,1508]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="40" unit="mm" value="2.4">2.4 mm</quantity>
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||
, total length without head
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||
<quantity id="4CE6BDD6221FFF97FBA7FA440A65FA52" box="[1034,1124,1484,1508]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="40" unit="mm" value="2.2">2.2 mm</quantity>
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||
, maximum width
|
||
<quantity id="4CE6BDD6221FFF97FA9DFA440B99FA52" box="[1328,1432,1484,1508]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="40" unit="mm" value="1.15">1.15 mm</quantity>
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, total length/maximum width 2.09.
|
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</paragraph>
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||
<paragraph id="8BA11033221FFF97FF6AF99C0D0EF9E6" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="40">
|
||
<emphasis id="B96ACC21221FFF97FF6AF99C0F0CF99A" bold="true" box="[199,269,1556,1580]" pageId="2" pageNumber="40">Male:</emphasis>
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Median lobe of aedeagus in ventral and lateral view as in
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<figureCitation id="13250CB6221FFF97FC36F99C0DDFF99A" box="[923,990,1556,1580]" captionStart="FIGURES 3 – 6" captionStartId="4.[151,264,1902,1925]" captionTargetBox="[253,1338,558,1857]" captionTargetId="figure@4.[239,1338,553,1865]" captionTargetPageId="4" captionText="FIGURES 3 – 6. Median lobe of aedeagus in ventral and lateral view and left paramere of: (3) and (4) Hydroporus esseri sp. n.; (5) and (6) Hydroporus umbrosus (specimen from Berlin, Germany)." httpUri="https://zenodo.org/record/205112/files/figure.png" pageId="2" pageNumber="40">Fig. 3</figureCitation>
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; left paramere as in
|
||
<figureCitation id="13250CB6221FFF97FB6FF99C0B07F99A" box="[1218,1286,1556,1580]" captionStart="FIGURES 3 – 6" captionStartId="4.[151,264,1902,1925]" captionTargetBox="[253,1338,558,1857]" captionTargetId="figure@4.[239,1338,553,1865]" captionTargetPageId="4" captionText="FIGURES 3 – 6. Median lobe of aedeagus in ventral and lateral view and left paramere of: (3) and (4) Hydroporus esseri sp. n.; (5) and (6) Hydroporus umbrosus (specimen from Berlin, Germany)." httpUri="https://zenodo.org/record/205112/files/figure.png" pageId="2" pageNumber="40">Fig. 4</figureCitation>
|
||
. For comparison those of
|
||
<taxonomicName id="4C1E6BB0221FFF97FE80F9B10FBAF9E6" box="[301,443,1592,1616]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221FFF97FE80F9B10FBAF9E6" box="[301,443,1592,1616]" italics="true" pageId="2" pageNumber="40">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
are illustrated in
|
||
<figureCitation id="13250CB6221FFF97FDD2F9B00D0DF9E6" box="[639,780,1592,1616]" captionStart="FIGURES 3 – 6" captionStartId="4.[151,264,1902,1925]" captionTargetBox="[253,1338,558,1857]" captionTargetId="figure@4.[239,1338,553,1865]" captionTargetPageId="4" captionText="FIGURES 3 – 6. Median lobe of aedeagus in ventral and lateral view and left paramere of: (3) and (4) Hydroporus esseri sp. n.; (5) and (6) Hydroporus umbrosus (specimen from Berlin, Germany)." httpUri="https://zenodo.org/record/205112/files/figure.png" pageId="2" pageNumber="40">Figs 5 and 6</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA11033221FFF97FF6AF9D40FA5F9C2" blockId="2.[151,1437,151,2012]" box="[199,420,1628,1652]" pageId="2" pageNumber="40">
|
||
<emphasis id="B96ACC21221FFF97FF6AF9D40F26F9C2" bold="true" box="[199,295,1628,1652]" pageId="2" pageNumber="40">Female:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8BA11033221FFF97FF6AF9F70B45F822" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="40">
|
||
<emphasis id="B96ACC21221FFF97FF6AF9F70FCFF92E" bold="true" box="[199,462,1663,1688]" pageId="2" pageNumber="40">Differential diagnosis.</emphasis>
|
||
As stated in the Taxonomy section,
|
||
<taxonomicName id="4C1E6BB0221FFF97FCC4F9090DCCF92E" box="[873,973,1665,1688]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC21221FFF97FCC4F9090DCCF92E" box="[873,973,1665,1688]" italics="true" pageId="2" pageNumber="40">H. esseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96ACC21221FFF97FC78F9080A14F92E" bold="true" box="[981,1045,1664,1688]" pageId="2" pageNumber="40">
|
||
<taxonomicNameLabel id="A259715A221FFF97FC78F9080A14F92E" box="[981,1045,1664,1688]" pageId="2" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
belongs to the
|
||
<taxonomicName id="4C1E6BB0221FFF97FB6FF9080B33F92E" box="[1218,1330,1664,1688]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="species" species="puberulus">
|
||
<emphasis id="B96ACC21221FFF97FB6FF9080B33F92E" box="[1218,1330,1664,1688]" italics="true" pageId="2" pageNumber="40">puberulus</emphasis>
|
||
</taxonomicName>
|
||
-group of the genus. It is distinctly smaller than
|
||
<taxonomicName id="4C1E6BB0221FFF97FDEDF92D0CFAF90A" box="[576,763,1700,1724]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="species" species="glabriusculus">
|
||
<emphasis id="B96ACC21221FFF97FDEDF92D0CFAF90A" box="[576,763,1700,1724]" italics="true" pageId="2" pageNumber="40">H. glabriusculus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CE6BDD6221FFF97FCA4F92C0D97F90A" box="[777,918,1700,1724]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="2" pageNumber="40" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8–3.2 mm</quantity>
|
||
),
|
||
<taxonomicName id="4C1E6BB0221FFF97FC04F92D0A3AF90A" box="[937,1083,1700,1724]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="species" species="puberulus">
|
||
<emphasis id="B96ACC21221FFF97FC04F92D0A3AF90A" box="[937,1083,1700,1724]" italics="true" pageId="2" pageNumber="40">H. puberulus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CE6BDD6221FFF97FBE7F92C0AD2F90A" box="[1098,1235,1700,1724]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="2" pageNumber="40" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7–3.1 mm</quantity>
|
||
) and
|
||
<taxonomicName id="4C1E6BB0221FFF97FABFF92D0B9DF90A" box="[1298,1436,1700,1724]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="species" species="rufinasus">
|
||
<emphasis id="B96ACC21221FFF97FABFF92D0B9DF90A" box="[1298,1436,1700,1724]" italics="true" pageId="2" pageNumber="40">H. rufinasus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CE6BDD6221FFF97FF33F9400F29F956" box="[158,296,1736,1760]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="2" pageNumber="40" unit="mm" value="3.05" valueMax="3.2" valueMin="2.9">2.9–3.2 mm</quantity>
|
||
) (cf.
|
||
<bibRefCitation id="EF8F6DC2221FFF97FEC9F9400C76F956" author="Nilsson" box="[356,631,1736,1760]" pageId="2" pageNumber="40" refString="Nilsson, A. N. & Holmen, M. (1995) The aquatic Adephaga (Coleoptera) of Fennoscandia and Denmark. II. Dytiscidae. Fauna Entomologica Scandinavica, 32, 1 - 192." type="journal article" year="1995">Nilsson & Holmen 1995</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF8F6DC2221FFF97FD01F9410D7DF956" author="Larson" box="[684,892,1736,1760]" pageId="2" pageNumber="40" refString="Larson, D. J., Alarie, Y. & Roughley, R. E. (2000) Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. - Ottawa: NRC Research Press, 982 pp." type="book" year="2000">
|
||
Larson
|
||
<emphasis id="B96ACC21221FFF97FCACF9410D33F956" box="[769,818,1736,1760]" italics="true" pageId="2" pageNumber="40">et al</emphasis>
|
||
. 2000
|
||
</bibRefCitation>
|
||
), and also other characters separate the new species from these three.
|
||
<taxonomicName id="4C1E6BB0221FFF97FE39F9650C22F8B2" box="[404,547,1772,1796]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221FFF97FE39F9650C22F8B2" box="[404,547,1772,1796]" italics="true" pageId="2" pageNumber="40">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
is the only species of the group which comes closer to the new one. Size and colouration are the most obvious differences between them: total length:
|
||
<quantity id="4CE6BDD6221FFF97FC77F8980A33F89E" box="[986,1074,1808,1832]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="40" unit="mm" value="2.4">2.4 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C1E6BB0221FFF97FBF5F8990ABCF89E" box="[1112,1213,1809,1832]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC21221FFF97FBF5F8990ABCF89E" box="[1112,1213,1809,1832]" italics="true" pageId="2" pageNumber="40">H. esseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96ACC21221FFF97FB6BF8980B06F89E" bold="true" box="[1222,1287,1808,1832]" pageId="2" pageNumber="40">
|
||
<taxonomicNameLabel id="A259715A221FFF97FB6BF8980B06F89E" box="[1222,1287,1808,1832]" pageId="2" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<quantity id="4CE6BDD6221FFF97FAE5F8980EC2F8FA" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="2" pageNumber="40" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C1E6BB0221FFF97FF48F8BD0C9FF8FA" authority="Nilsson & Holmen 1995" authorityName="Nilsson & Holmen" authorityYear="1995" box="[229,670,1844,1868]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221FFF97FF48F8BD0F73F8FA" box="[229,370,1844,1868]" italics="true" pageId="2" pageNumber="40">H. umbrosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8F6DC2221FFF97FE2DF8BC0C97F8FA" author="Nilsson" box="[384,662,1844,1868]" pageId="2" pageNumber="40" refString="Nilsson, A. N. & Holmen, M. (1995) The aquatic Adephaga (Coleoptera) of Fennoscandia and Denmark. II. Dytiscidae. Fauna Entomologica Scandinavica, 32, 1 - 192." type="journal article" year="1995">Nilsson & Holmen 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; upper side generally darker and more uniform in the new species, in
|
||
<taxonomicName id="4C1E6BB0221FFF97FF3AF8D10F29F8C6" box="[151,296,1880,1904]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="40" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221FFF97FF3AF8D10F29F8C6" box="[151,296,1880,1904]" italics="true" pageId="2" pageNumber="40">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
large parts of head, sides of pronotum and elytra reddish brown, and elytra usually with a poorly delimited, but distinct blackish spot in posterior two thirds and a dark stripe along suture (cf.
|
||
<figureCitation id="13250CB6221FFF97FB07F8F40B37F822" box="[1194,1334,1916,1940]" captionStart="FIGURES 1 – 2" captionStartId="3.[151,264,1354,1377]" captionTargetBox="[276,1307,483,1331]" captionTargetId="figure@3.[276,1310,483,1331]" captionTargetPageId="3" captionText="FIGURES 1 – 2. Habitus of: (1) Hydroporus esseri sp. n., (2) Hydroporus umbrosus (specimen from Berlin, Germany)." httpUri="https://zenodo.org/record/205111/files/figure.png" pageId="2" pageNumber="40">Figs 1 and 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA11033221FFF96FF68F8290B7DFF41" blockId="2.[151,1437,151,2012]" lastBlockId="3.[151,1436,151,428]" lastPageId="3" lastPageNumber="41" pageId="2" pageNumber="40">
|
||
However, usually size and colour are not accepted as "good" characters, and because we could study only a single specimen, we do not know to which extent the new species varies with respect to these characters. Additionally,
|
||
<taxonomicName id="4C1E6BB0221EFF96FF3AFF110F27FF06" box="[151,294,152,176]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221EFF96FF3AFF110F27FF06" box="[151,294,152,176]" italics="true" pageId="3" pageNumber="41">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
is known sometimes to lack the blackish spot on the elytra and is then much more similar to the new species. That's why someone might argue that the description of the new species cannot be justified with the necessary certainty, and that the specimen from southern
|
||
<collectingCountry id="F30950A3221EFF96FD77FF570D2BFF41" box="[730,810,223,247]" name="Turkey" pageId="3" pageNumber="41">Turkey</collectingCountry>
|
||
might be nothing else than a variety of
|
||
<taxonomicName id="4C1E6BB0221EFF96FB4AFF690B74FF4E" box="[1255,1397,224,248]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221EFF96FB4AFF690B74FF4E" box="[1255,1397,224,248]" italics="true" pageId="3" pageNumber="41">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA11033221EFF96FF68FE8D0AE5FE1A" blockId="3.[151,1436,151,428]" pageId="3" pageNumber="41">
|
||
We wish to address these concerns, and point to the following facts: During the last years we have seen about 1,200 specimens of
|
||
<taxonomicName id="4C1E6BB0221EFF96FEDAFEA10C04FEF6" box="[375,517,296,320]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221EFF96FEDAFEA10C04FEF6" box="[375,517,296,320]" italics="true" pageId="3" pageNumber="41">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
, coming chiefly from Central Europe, but also many from Scandinavia, and some from Buryatia and Kamchatka. We have not been able to recognise a considerable variability in the specimens studied. In particular we noted that among all
|
||
<taxonomicName id="4C1E6BB0221EFF96FDD8FEF90D05FE3E" box="[629,772,368,392]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221EFF96FDD8FEF90D05FE3E" box="[629,772,368,392]" italics="true" pageId="3" pageNumber="41">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
studied there was none with a total length of
|
||
<quantity id="4CE6BDD6221EFF96FAA0FEE70B67FE31" box="[1293,1382,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="41" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, and only a few of
|
||
<quantity id="4CE6BDD6221EFF96FE9FFE1C0F8BFE1A" box="[306,394,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="41" unit="mm" value="2.6">2.6 mm</quantity>
|
||
, and that the species lacks that blackish spot on elytra only extremely rarely.
|
||
</paragraph>
|
||
<caption id="DF6140BB221EFF96FF3AFAC20B4CFAD4" box="[151,1357,1354,1378]" httpUri="https://zenodo.org/record/205111/files/figure.png" pageId="3" pageNumber="41" targetBox="[276,1307,483,1331]" targetPageId="3">
|
||
<paragraph id="8BA11033221EFF96FF3AFAC20B4CFAD4" blockId="3.[151,1357,1354,1378]" box="[151,1357,1354,1378]" pageId="3" pageNumber="41">
|
||
<emphasis id="B96ACC21221EFF96FF3AFAC20F3BFAD6" bold="true" box="[151,314,1354,1377]" pageId="3" pageNumber="41">FIGURES 1–2.</emphasis>
|
||
Habitus of: (1)
|
||
<taxonomicName id="4C1E6BB0221EFF96FE71FAC30C97FAD7" box="[476,662,1355,1377]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC21221EFF96FE71FAC30C97FAD7" box="[476,662,1355,1377]" italics="true" pageId="3" pageNumber="41">Hydroporus esseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96ACC21221EFF96FD31FAC30CD4FAD6" bold="true" box="[668,725,1355,1376]" pageId="3" pageNumber="41">
|
||
<taxonomicNameLabel id="A259715A221EFF96FD31FAC30CD4FAD6" box="[668,725,1355,1376]" pageId="3" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, (2)
|
||
<taxonomicName id="4C1E6BB0221EFF96FCABFAC30DE9FAD7" box="[774,1000,1355,1377]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221EFF96FCABFAC30DE9FAD7" box="[774,1000,1355,1377]" italics="true" pageId="3" pageNumber="41">Hydroporus umbrosus</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Berlin, Germany).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA11033221EFF96FF68FA010D0DFA72" blockId="3.[151,1436,1416,1476]" pageId="3" pageNumber="41">In addition, we have observed differences in several other characters, the combination of which leaves no doubt about the existence of two well separated species:</paragraph>
|
||
<paragraph id="8BA11033221EFF96FF3AFA7D0B67F986" blockId="3.[151,1436,1524,2017]" pageId="3" pageNumber="41">
|
||
- prosternal process before indistinct transverse protuberance with only very fine transverse carinae in
|
||
<taxonomicName id="4C1E6BB0221EFF96FA95FA7E0B9DF9BB" box="[1336,1436,1526,1549]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC21221EFF96FA95FA7E0B9DF9BB" box="[1336,1436,1526,1549]" italics="true" pageId="3" pageNumber="41">H. esseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96ACC21221EFF96FF68F9910F05F987" bold="true" box="[197,260,1561,1585]" pageId="3" pageNumber="41">
|
||
<taxonomicNameLabel id="A259715A221EFF96FF68F9910F05F987" box="[197,260,1561,1585]" pageId="3" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; in
|
||
<taxonomicName id="4C1E6BB0221EFF96FE82F9920FBFF987" box="[303,446,1561,1585]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221EFF96FE82F9920FBFF987" box="[303,446,1561,1585]" italics="true" pageId="3" pageNumber="41">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
transverse protuberance often more distinct and transverse carina before less weak;
|
||
</paragraph>
|
||
<paragraph id="8BA11033221EFF96FF3AF9B50A40F92A" blockId="3.[151,1436,1524,2017]" pageId="3" pageNumber="41">
|
||
- lanceolate part of prosternal process (blade) less vaulted and covered with several coarse punctures in
|
||
<taxonomicName id="4C1E6BB0221EFF96FA94F9B60B9AF9E3" box="[1337,1435,1598,1621]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC21221EFF96FA94F9B60B9AF9E3" box="[1337,1435,1598,1621]" italics="true" pageId="3" pageNumber="41">H. esseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A259715A221EFF96FF68F9E90EFDF9CE" box="[197,252,1633,1656]" pageId="3" pageNumber="41" rank="species">sp.n.</taxonomicNameLabel>
|
||
, sides of blade roughly punctured, but rest without such punctures in almost all
|
||
<taxonomicName id="4C1E6BB0221EFF96FBDCF9EA0AFEF9CF" box="[1137,1279,1633,1657]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221EFF96FBDCF9EA0AFEF9CF" box="[1137,1279,1633,1657]" italics="true" pageId="3" pageNumber="41">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
studied; sides of blade not flattened in
|
||
<taxonomicName id="4C1E6BB0221EFF96FE7AF90E0C3AF92B" box="[471,571,1670,1693]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC21221EFF96FE7AF90E0C3AF92B" box="[471,571,1670,1693]" italics="true" pageId="3" pageNumber="41">H. esseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A259715A221EFF96FDEFF90D0C80F92B" box="[578,641,1669,1693]" pageId="3" pageNumber="41" rank="species">
|
||
<emphasis id="B96ACC21221EFF96FDEFF90D0C80F92B" bold="true" box="[578,641,1669,1693]" pageId="3" pageNumber="41">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, but narrowly flattened in
|
||
<taxonomicName id="4C1E6BB0221EFF96FC06F90E0A38F92B" box="[939,1081,1669,1693]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221EFF96FC06F90E0A38F92B" box="[939,1081,1669,1693]" italics="true" pageId="3" pageNumber="41">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
</paragraph>
|
||
<paragraph id="8BA11033221EFF96FF3AF9210F7DF8BE" blockId="3.[151,1436,1524,2017]" pageId="3" pageNumber="41">
|
||
- midline of metacoxal processes very deeply impressed over its entire length in
|
||
<taxonomicName id="4C1E6BB0221EFF96FBF8F9220ABBF977" box="[1109,1210,1706,1729]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC21221EFF96FBF8F9220ABBF977" box="[1109,1210,1706,1729]" italics="true" pageId="3" pageNumber="41">H. esseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96ACC21221EFF96FB69F9210B04F977" bold="true" box="[1220,1285,1705,1729]" pageId="3" pageNumber="41">
|
||
<taxonomicNameLabel id="A259715A221EFF96FB69F9210B04F977" box="[1220,1285,1705,1729]" pageId="3" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; distinct, but less impressed in
|
||
<taxonomicName id="4C1E6BB0221EFF96FE23F9460C1CF953" box="[398,541,1741,1765]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221EFF96FE23F9460C1CF953" box="[398,541,1741,1765]" italics="true" pageId="3" pageNumber="41">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
; punctured grooves inside and next to metacoxal line also deeper impressed in the new species;
|
||
</paragraph>
|
||
<paragraph id="8BA11033221EFF96FF3AF89D0C26F8C2" blockId="3.[151,1436,1524,2017]" pageId="3" pageNumber="41">
|
||
- hind margin of each metacoxal process next to end of midline without distinct pointed prolongations in
|
||
<taxonomicName id="4C1E6BB0221EFF96FA2CF89E0F07F8E7" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC21221EFF96FA2CF89E0F07F8E7" italics="true" pageId="3" pageNumber="41">H. esseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96ACC21221EFF96FEA2F8B10F50F8E7" bold="true" box="[271,337,1849,1873]" pageId="3" pageNumber="41">
|
||
<taxonomicNameLabel id="A259715A221EFF96FEA2F8B10F50F8E7" box="[271,337,1849,1873]" pageId="3" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; at posterior end of midline each process with a very small but distinct pointed prolongation in almost all
|
||
<taxonomicName id="4C1E6BB0221EFF96FE94F8D60FC6F8C3" box="[313,455,1885,1909]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC21221EFF96FE94F8D60FC6F8C3" box="[313,455,1885,1909]" italics="true" pageId="3" pageNumber="41">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
studied;
|
||
</paragraph>
|
||
<paragraph id="8BA11033221EFF96FF3AF8090F52F80A" blockId="3.[151,1436,1524,2017]" pageId="3" pageNumber="41">- punctation on metaventrite, metacoxal plates and first and second abdominal ventrites coarser and denser in the new species;</paragraph>
|
||
<paragraph id="8BA11033221EFF91FF3AF8410AEFFF62" blockId="3.[151,1436,1524,2017]" lastBlockId="4.[151,1436,151,284]" lastPageId="4" lastPageNumber="42" pageId="3" pageNumber="41">
|
||
- second and third abdominal ventrites without traces of a suture between them in
|
||
<taxonomicName id="4C1E6BB0221EFF96FBCAF8420ACDF857" box="[1127,1228,1994,2017]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC21221EFF96FBCAF8420ACDF857" box="[1127,1228,1994,2017]" italics="true" pageId="3" pageNumber="41">H. esseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96ACC21221EFF96FB78F8410B16F857" bold="true" box="[1237,1303,1993,2017]" pageId="3" pageNumber="41">
|
||
<taxonomicNameLabel id="A259715A221EFF96FB78F8410B16F857" box="[1237,1303,1993,2017]" pageId="3" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, only a flat impression perceptible if illuminated adequately; among all
|
||
<taxonomicName id="4C1E6BB02219FF91FCD1FF110A0FFF06" box="[892,1038,152,176]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC212219FF91FCD1FF110A0FFF06" box="[892,1038,152,176]" italics="true" pageId="4" pageNumber="42">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
studied only a few with complete suture and several ones with incomplete suture, but not a single one without any traces of such;
|
||
</paragraph>
|
||
<paragraph id="8BA110332219FF91FF3AFF680ADDFEAA" blockId="4.[151,1436,151,284]" pageId="4" pageNumber="42">
|
||
- median lobe in the new species in ventral view narrower and more converging to tip, in lateral view more curved near basal part and dorsal side more straight between base and apex (cf.
|
||
<figureCitation id="13250CB62219FF91FBEFFE8C0ACFFEAA" box="[1090,1230,260,284]" captionStart="FIGURES 3 – 6" captionStartId="4.[151,264,1902,1925]" captionTargetBox="[253,1338,558,1857]" captionTargetId="figure@4.[239,1338,553,1865]" captionTargetPageId="4" captionText="FIGURES 3 – 6. Median lobe of aedeagus in ventral and lateral view and left paramere of: (3) and (4) Hydroporus esseri sp. n.; (5) and (6) Hydroporus umbrosus (specimen from Berlin, Germany)." httpUri="https://zenodo.org/record/205112/files/figure.png" pageId="4" pageNumber="42">Figs 3 and 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA110332219FF91FF6AFEC40A48FE42" blockId="4.[151,1436,332,501]" pageId="4" pageNumber="42">
|
||
Finally we want to point to the big gap between the geographical distribution area of
|
||
<taxonomicName id="4C1E6BB02219FF91FB21FEC50B1AFED2" box="[1164,1307,332,356]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC212219FF91FB21FEC50B1AFED2" box="[1164,1307,332,356]" italics="true" pageId="4" pageNumber="42">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
and that of the new species. According to
|
||
<bibRefCitation id="EF8F6DC22219FF91FE46FEE70D3AFE31" author="Nilsson" box="[491,827,367,391]" pageId="4" pageNumber="42" refString="Nilsson, A. N. & Holmen, M. (1995) The aquatic Adephaga (Coleoptera) of Fennoscandia and Denmark. II. Dytiscidae. Fauna Entomologica Scandinavica, 32, 1 - 192." type="journal article" year="1995">Nilsson & Holmen (1995: 63)</bibRefCitation>
|
||
, the distribution of
|
||
<taxonomicName id="4C1E6BB02219FF91FBB9FEF90AA0FE3E" box="[1044,1185,368,392]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC212219FF91FBB9FEF90AA0FE3E" box="[1044,1185,368,392]" italics="true" pageId="4" pageNumber="42">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
covers most of Northern, Central and Eastern Europe, plus Asian
|
||
<collectingCountry id="F30950A32219FF91FD2AFE1C0CD3FE1A" box="[647,722,404,428]" name="Russia" pageId="4" pageNumber="42">Russia</collectingCountry>
|
||
(from Western Siberia to Far East). The
|
||
<collectingCountry id="F30950A32219FF91FB35FE1C0AF3FE1A" box="[1176,1266,404,428]" name="Ukraine" pageId="4" pageNumber="42">Ukraine</collectingCountry>
|
||
and the Southern Territories of
|
||
<collectingCountry id="F30950A32219FF91FEF3FE3F0FA8FE79" box="[350,425,439,463]" name="Russia" pageId="4" pageNumber="42">Russia</collectingCountry>
|
||
are additionally given in
|
||
<bibRefCitation id="EF8F6DC22219FF91FD61FE3F0D77FE79" author="Nilsson" box="[716,886,439,463]" pageId="4" pageNumber="42" refString="Nilsson, A. N. (2011) Catalogue of Palearctic Dytiscidae (Coleoptera). Internet version 2011 - 01 - 01. Available from http: // www 2. emg. umu. se / projects / biginst / andersn / Cat _ main. htm (accessed in March 2011)." type="journal article" year="2011">Nilsson (2011)</bibRefCitation>
|
||
; these are the most southern areas so far known, and both more than
|
||
<quantity id="4CE6BDD62219FF91FEDAFE540FE0FE42" box="[375,481,476,500]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="42" unit="km" value="1000.0">1,000 km</quantity>
|
||
apart from the
|
||
<typeStatus id="54A5AE912219FF91FD20FE550CBCFE43" box="[653,701,477,501]" pageId="4" pageNumber="42">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C1E6BB02219FF91FC90FE550DA1FE42" box="[829,928,477,500]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC212219FF91FC90FE550DA1FE42" box="[829,928,477,500]" italics="true" pageId="4" pageNumber="42">H. esseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96ACC212219FF91FC05FE540DE9FE42" bold="true" box="[936,1000,476,500]" pageId="4" pageNumber="42">
|
||
<taxonomicNameLabel id="A259715A2219FF91FC05FE540DE9FE42" box="[936,1000,476,500]" pageId="4" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13250CB62219FF91FC5BFE540A3AFE42" box="[1014,1083,476,500]" captionStart="FIGURE 7" captionStartId="5.[151,250,1880,1903]" captionTargetBox="[265,1323,1133,1858]" captionTargetId="figure@5.[264,1323,1132,1860]" captionTargetPageId="5" captionText="FIGURE 7. Distribution of Hydroporus esseri sp. n. (black star) and western part of that of Hydroporus umbrosus (dark greyish area)." httpUri="https://zenodo.org/record/205113/files/figure.png" pageId="4" pageNumber="42">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6140BB2219FF91FF3AF8E60D50F812" httpUri="https://zenodo.org/record/205112/files/figure.png" pageId="4" pageNumber="42" targetBox="[253,1338,558,1857]" targetPageId="4">
|
||
<paragraph id="8BA110332219FF91FF3AF8E60D50F812" blockId="4.[151,1436,1902,1956]" pageId="4" pageNumber="42">
|
||
<emphasis id="B96ACC212219FF91FF3AF8E60F3BF832" bold="true" box="[151,314,1902,1925]" pageId="4" pageNumber="42">FIGURES 3–6.</emphasis>
|
||
Median lobe of aedeagus in ventral and lateral view and left paramere of: (3) and (4)
|
||
<taxonomicName id="4C1E6BB02219FF91FB31F8E70B57F833" box="[1180,1366,1903,1925]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC212219FF91FB31F8E70B57F833" box="[1180,1366,1903,1925]" italics="true" pageId="4" pageNumber="42">Hydroporus esseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96ACC212219FF91FAF1F8E70B94F832" bold="true" box="[1372,1429,1903,1924]" pageId="4" pageNumber="42">
|
||
<taxonomicNameLabel id="A259715A2219FF91FAF1F8E70B94F832" box="[1372,1429,1903,1924]" pageId="4" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; (5) and (6)
|
||
<taxonomicName id="4C1E6BB02219FF91FEA4F8060FEAF812" box="[265,491,1934,1956]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC212219FF91FEA4F8060FEAF812" box="[265,491,1934,1956]" italics="true" pageId="4" pageNumber="42">Hydroporus umbrosus</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Berlin, Germany).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA110332218FF90FF6AFF1F0F88FEAA" blockId="5.[151,1437,151,1076]" pageId="5" pageNumber="43">
|
||
<emphasis id="B96ACC212218FF90FF6AFF1F0F2BFF06" bold="true" box="[199,298,151,176]" pageId="5" pageNumber="43">Habitat.</emphasis>
|
||
The single specimen was collected in very shallow water among gravel and pebbles of a backwater pool near the banks of the Köprülü River. The water was clear and without any submerged vegetation (
|
||
<figureCitation id="13250CB62218FF90FA89FF340EA6FF41" captionStart="FIGURES 8 – 9" captionStartId="6.[151,264,1641,1664]" captionTargetBox="[264,1322,193,1620]" captionTargetId="figure@6.[264,1323,193,1620]" captionTargetPageId="6" captionText="FIGURES 8 – 9. The Köprülü River in southern Turkey; arrow: small backwater pool, locus typicus of Hydroporus esseri sp. n." httpUri="https://zenodo.org/record/205114/files/figure.png" pageId="5" pageNumber="43">Figs 8 and 9</figureCitation>
|
||
). Together with the
|
||
<typeStatus id="54A5AE912218FF90FE23FF570FF1FF41" box="[398,496,223,247]" pageId="5" pageNumber="43" type="holotype">holotype</typeStatus>
|
||
few specimens of
|
||
<taxonomicName id="4C1E6BB02218FF90FD65FF680A2EFF41" authority="Brulle, 1832" authorityName="Brulle" authorityYear="1832" box="[712,1071,223,248]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="dilatatus">
|
||
<emphasis id="B96ACC212218FF90FD65FF680D87FF4E" box="[712,902,224,248]" italics="true" pageId="5" pageNumber="43">Agabus dilatatus</emphasis>
|
||
(Brullé, 1832)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1E6BB02218FF90FBC5FF680ED9FEAA" authority="Forster, 1771" authorityName="Forster" authorityYear="1771" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="nebulosus">
|
||
<emphasis id="B96ACC212218FF90FBC5FF680B34FF4E" box="[1128,1333,224,248]" italics="true" pageId="5" pageNumber="43">Agabus nebulosus</emphasis>
|
||
(Forster, 1771)
|
||
</taxonomicName>
|
||
were collected.
|
||
</paragraph>
|
||
<paragraph id="8BA110332218FF90FF6AFEAF0A9FFE42" blockId="5.[151,1437,151,1076]" pageId="5" pageNumber="43">
|
||
The closely related
|
||
<taxonomicName id="4C1E6BB02218FF90FE09FEA10C32FEF6" box="[420,563,296,320]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="umbrosus">
|
||
<emphasis id="B96ACC212218FF90FE09FEA10C32FEF6" box="[420,563,296,320]" italics="true" pageId="5" pageNumber="43">H. umbrosus</emphasis>
|
||
</taxonomicName>
|
||
occurs all over its range in shallow and well vegetated standing water bodies e.g. exposed fens, bogs, shallow pools, marshland and well vegetated lake shores (
|
||
<bibRefCitation id="EF8F6DC22218FF90FB81FEC40ABFFED2" author="Horion" box="[1068,1214,332,356]" pageId="5" pageNumber="43" refString="Horion, A. (1941) Faunistik der deutschen Kafer, I. Adephaga - Caraboidea. Krefeld: Goecke Verlag, 463 pp." type="book" year="1941">Horion 1941</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8F6DC22218FF90FB64FEC40ED0FE31" author="Nilsson" pageId="5" pageNumber="43" refString="Nilsson, A. N. & Holmen, M. (1995) The aquatic Adephaga (Coleoptera) of Fennoscandia and Denmark. II. Dytiscidae. Fauna Entomologica Scandinavica, 32, 1 - 192." type="journal article" year="1995">Nilsson & Holmen 1995</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8F6DC22218FF90FF71FEF80F95FE31" author="Hess" box="[220,404,367,392]" pageId="5" pageNumber="43" refString="Hess, M., Spitzenberg, D., Bellstedt, R., Heckes, U., Hendrich, L. & Sondermann, W. (1999) Artenbestand und Gefahrdungssituation der Wasserkafer Deutschlands (Coleoptera: Hydradephaga, Hydrophiloidea part., Dryopoidea part.; Microsporidae, Hydraenidae, Scirtidae). Naturschutz und Landschaftsplanung, 31 (7), 197 - 211." type="journal article" year="1999">
|
||
Hess
|
||
<emphasis id="B96ACC212218FF90FEB7FEF90F4AFE3E" box="[282,331,368,392]" italics="true" pageId="5" pageNumber="43">et al</emphasis>
|
||
. 1999
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8F6DC22218FF90FE33FEE70C4BFE31" author="Hendrich" box="[414,586,367,391]" pageId="5" pageNumber="43" refString="Hendrich, L. (2003) Die Wasserkafer von Berlin. Struktur der aquatischen Kaferfauna (Hydradephaga, Hydrophiloidea, Dryopoidea [partim] und Staphylinoidea [partim]) in anthropogen beeinflussten Gewassern von Berlin - Taxonomische, raumliche, faunistische und okologische Aspekte. Dissertation, Fakultat VII, Institut fur Biologie und Okologie der Technischen Universitat Berlin. Berlin: dissertation. de - Verlag im Internet, 563 pp." type="book" year="2003">Hendrich 2003</bibRefCitation>
|
||
). If it is assumed that
|
||
<taxonomicName id="4C1E6BB02218FF90FC93FEF90DA0FE3E" box="[830,929,369,392]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="esseri" status="sp. nov.">
|
||
<emphasis id="B96ACC212218FF90FC93FEF90DA0FE3E" box="[830,929,369,392]" italics="true" pageId="5" pageNumber="43">H. esseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96ACC212218FF90FC05FEF80DE9FE3E" bold="true" box="[936,1000,368,392]" pageId="5" pageNumber="43">
|
||
<taxonomicNameLabel id="A259715A2218FF90FC05FEF80DE9FE3E" box="[936,1000,368,392]" pageId="5" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
prefers similar habitats, then we might conclude that the single specimen found was flushed away by heavy rain from its original habitat further upstream or from a much smaller tributary of the Köprülü River, and that the collecting site does not represent its preferred habitat. This would also explain why only one and not more specimens have been collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30443B82218FF90FF6AFE770ABBFD8A" pageId="5" pageNumber="43" type="etymology">
|
||
<paragraph id="8BA110332218FF90FF6AFE770ABBFD8A" blockId="5.[151,1437,151,1076]" pageId="5" pageNumber="43">
|
||
<emphasis id="B96ACC212218FF90FF6AFE770FA6FDAE" bold="true" box="[199,423,511,536]" pageId="5" pageNumber="43">Derivatio nominis.</emphasis>
|
||
This species is dedicated to the coleopterist Jens Esser (Berlin,
|
||
<collectingCountry id="F30950A32218FF90FB3FFE770AFEFDA1" box="[1170,1279,511,535]" name="Germany" pageId="5" pageNumber="43">Germany</collectingCountry>
|
||
), specialist in
|
||
<taxonomicName id="4C1E6BB02218FF90FF3AFDAC0F4CFD8A" box="[151,333,548,572]" class="Insecta" family="Cryptophagidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="family">Cryptophagidae</taxonomicName>
|
||
, who collected the
|
||
<typeStatus id="54A5AE912218FF90FD89FDAC0C88FD8A" box="[548,649,548,572]" pageId="5" pageNumber="43" type="holotype">holotype</typeStatus>
|
||
; the specific epithet is a noun in the genitive case.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30443B82218FF90FF6AFDCF0F1AFB82" pageId="5" pageNumber="43" type="distribution">
|
||
<paragraph id="8BA110332218FF90FF6AFDCF0B88FDE9" blockId="5.[151,1437,151,1076]" box="[199,1417,583,608]" pageId="5" pageNumber="43">
|
||
<emphasis id="B96ACC212218FF90FF6AFDCF0F61FDD6" bold="true" box="[199,352,583,608]" pageId="5" pageNumber="43">Distribution.</emphasis>
|
||
Only known from the
|
||
<typeStatus id="54A5AE912218FF90FDCDFDC00C91FDD6" box="[608,656,584,608]" pageId="5" pageNumber="43">type</typeStatus>
|
||
locality, but probably more widespread in southern
|
||
<collectingCountry id="F30950A32218FF90FB75FDCF0B29FDE9" box="[1240,1320,583,607]" name="Turkey" pageId="5" pageNumber="43">Turkey</collectingCountry>
|
||
(
|
||
<figureCitation id="13250CB62218FF90FA9AFDCF0B7AFDE9" box="[1335,1403,583,607]" captionStart="FIGURE 7" captionStartId="5.[151,250,1880,1903]" captionTargetBox="[265,1323,1133,1858]" captionTargetId="figure@5.[264,1323,1132,1860]" captionTargetPageId="5" captionText="FIGURE 7. Distribution of Hydroporus esseri sp. n. (black star) and western part of that of Hydroporus umbrosus (dark greyish area)." httpUri="https://zenodo.org/record/205113/files/figure.png" pageId="5" pageNumber="43">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA110332218FF90FF6AFDE30F1AFB82" blockId="5.[151,1437,151,1076]" pageId="5" pageNumber="43">
|
||
<emphasis id="B96ACC212218FF90FF6AFDE30F50FD32" bold="true" box="[199,337,619,644]" pageId="5" pageNumber="43">Study area.</emphasis>
|
||
The Köprülü Canyon National Park is located in the western part of the Taurus Mountains in southern
|
||
<collectingCountry id="F30950A32218FF90FF50FD070F4CFD11" box="[253,333,655,679]" name="Turkey" pageId="5" pageNumber="43">Turkey</collectingCountry>
|
||
between Antalya and Manavgat at elevations of
|
||
<quantity id="4CE6BDD62218FF90FCC6FD070DB0FD11" box="[875,945,655,679]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="43" unit="m" value="110.0">110 m</quantity>
|
||
to
|
||
<quantity id="4CE6BDD62218FF90FC7EFD070A2FFD11" box="[979,1070,655,679]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="43" unit="m" value="2500.0">2,500 m</quantity>
|
||
(summit of the Bozburun mountain range) on very heterogeneous geomorphologic structure. It encompasses 37,000 ha, lying at a distance of
|
||
<quantity id="4CE6BDD62218FF90FA2DFD3C0EBAFD59" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="43" unit="km" value="90.0">90 km</quantity>
|
||
northeast from the province of Antalya. The reserve is separated by the Köprülü River canyon in a western and eastern part. The eastern part is dominated by hills up to
|
||
<quantity id="4CE6BDD62218FF90FCB7FD740D60FCA2" box="[794,865,764,788]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="43" unit="m" value="850.0">850 m</quantity>
|
||
above sea level with several deep canyons joining to the Köprülü Canyon. The western portion consists of higher altitudes often above
|
||
<quantity id="4CE6BDD62218FF90FBEAFC970AA4FC8E" box="[1095,1189,799,824]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="43" unit="m" value="1000.0">1,000 m</quantity>
|
||
, with Bozburun range raising over
|
||
<quantity id="4CE6BDD62218FF90FE8CFCCC0F7EFCEA" box="[289,383,836,860]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="43" unit="m" value="2000.0">2,000 m</quantity>
|
||
, being the western part boundary. The park has a rich flora with 900 to 1,000 species including stands of cypress forest and many endemics. The area comprises the whole range of vegetation zones from Mediterranean to alpine environments. The area has a Mediterranean climate with high precipitation (
|
||
<quantity id="4CE6BDD62218FF90FAB7FC040B95FC12" box="[1306,1428,908,932]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="43" unit="mm" value="1700.0">1,700 mm</quantity>
|
||
) throughout the winter months. Nevertheless, owing to the karst, water availability during summer months is restricted. Average annual temperature is 18.2 C°. The environment of the park still faces many threats. Overgrazing and illegal cutting of trees and shrubs have caused degradation of many parts of the natural vegetation (
|
||
<bibRefCitation id="EF8F6DC22218FF90FAF9FC7F0F11FB82" author="Cetinkaya" pageId="5" pageNumber="43" refString="Cetinkaya, G. (2002) The progress report regarding " Research for the establishment of Koprulu Kanyon National park as a biosphere reserve ". Saint-Petersburg State University of Architecture and Civil Engineering, 80 pp. (unpublished report)." type="book" year="2002">Cetinkaya 2002</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |