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<document id="6733792D5B18A5FF8CD62054973E6DE5" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2021" docId="03B18797FFFD9D1F4BA5021FD8CF222C" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Retrorsia simplicissima Shear &amp; Marek 2021, n. sp." docType="treatment" docVersion="6" lastPageNumber="103" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="101" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8E658D20897144D4BF39F655C9FC0841" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4535B5810FC2E7A5D4A1C512A5E72AC4">
<mods:title id="9C3CAA4CCBF8BC688B6764B745D7B204">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
</mods:titleInfo>
<mods:name id="615A17544CA7099A330744DD70423424" type="personal">
<mods:role id="B5211C8DA3B3BC80338313DCD2F034F9">
<mods:roleTerm id="C6B6E06ABED2591ECE5F77390A820A01">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B34988D60D380932612BAFBD18CC6EAA">Shear, William A.</mods:namePart>
<mods:affiliation id="25CF3D8C0B151FC4C4B7F6289246A584">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
</mods:name>
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<mods:role id="1AA5E9AA043BB764245ADB1528CA4DB1">
<mods:roleTerm id="D685619266CFC38D8414C3BC31C11912">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E7714990603C4534741D5AB0B1D329B">Marek, Paul E.</mods:namePart>
<mods:nameIdentifier id="946032C53D23E6DE668D9D993C1B88F7" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation id="CA35BE7A51FBAEE3367CDD8B75787816">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
<mods:nameIdentifier id="3333DA8CF2AE0B51A9156B5D2F9C3D9F" type="email">pmarek@vt.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0EA8F8C58B23B522FA93FFE43EA198F4">text</mods:typeOfResource>
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<mods:title id="8C2373BE4CC642A32E4E1FD74E6FE4C1">Zootaxa</mods:title>
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<mods:part id="2CF650E51E0ED5E5E59D117CA22A5200">
<mods:date id="CF4BBB0BC0D1F5ED256E0DC05D3C0B57">2021</mods:date>
<mods:detail id="6CE654ADB1A340AA878C1BDABA89DAFC" type="pubDate">
<mods:number id="394550D8C69A628649326CC8FAAB5EAF">2021-05-24</mods:number>
</mods:detail>
<mods:detail id="A9F60E57C527C4FBDFB56566BD0C6306" type="volume">
<mods:number id="B9881762373B66AE20E20C6247349DBC">4975</mods:number>
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<mods:classification id="318F1177741584CF6A7D61019578FFB8">journal article</mods:classification>
<mods:identifier id="8843D2667755E0E513F4FB5675E2E3FC" type="CLB-Dataset">6194</mods:identifier>
<mods:identifier id="6A66D3BD6F419211A5C19E4E72C4D9F6" type="DOI">10.11646/zootaxa.4975.1.3</mods:identifier>
<mods:identifier id="F6BDE84484C31E9924AD04E3B81031A3" type="GBIF-Dataset">529d48db-4299-4e2d-9aaf-6c4178c08b18</mods:identifier>
<mods:identifier id="C7F204A44D1E12B5C5F65C9881EBE54E" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B18797FFFD9D1F4BA5021FD8CF222C" ID-DOI="http://doi.org/10.5281/zenodo.4805011" ID-GBIF-Taxon="182644889" ID-Zenodo-Dep="4805011" LSID="urn:lsid:plazi:treatment:03B18797FFFD9D1F4BA5021FD8CF222C" httpUri="http://treatment.plazi.org/id/03B18797FFFD9D1F4BA5021FD8CF222C" lastPageId="22" lastPageNumber="103" pageId="20" pageNumber="101">
<subSubSection id="C302650AFFFD9D1D4BA5021FDB8D2772" box="[151,718,1828,1855]" pageId="20" pageNumber="101" type="nomenclature">
<paragraph id="8BA73681FFFD9D1D4BA5021FDB8D2772" blockId="20.[151,718,1828,1892]" box="[151,718,1828,1855]" pageId="20" pageNumber="101">
<heading id="D0EF81EDFFFD9D1D4BA5021FDB8D2772" bold="true" box="[151,718,1828,1855]" fontSize="11" level="1" pageId="20" pageNumber="101" reason="1">
<emphasis id="B96CEA93FFFD9D1D4BA5021FDB8D2772" bold="true" box="[151,718,1828,1855]" pageId="20" pageNumber="101">
<taxonomicName id="4C184D02FFFD9D1D4BA5021FDB3E2772" authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2021" box="[151,637,1828,1855]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="20" pageNumber="101" phylum="Arthropoda" rank="species" species="simplicissima" status="sp. nov.">
<emphasis id="B96CEA93FFFD9D1D4BA5021FD8F12772" bold="true" box="[151,434,1828,1855]" italics="true" pageId="20" pageNumber="101">Retrorsia simplicissima</emphasis>
Shear &amp; Marek
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFFD9D1D49BE021FDB8D2772" box="[652,718,1829,1855]" pageId="20" pageNumber="101" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFFD9D1D4BA50273D80F272E" box="[151,332,1865,1892]" pageId="20" pageNumber="101" type="description">
<paragraph id="8BA73681FFFD9D1D4BA50273D80F272E" blockId="20.[151,718,1828,1892]" box="[151,332,1865,1892]" pageId="20" pageNumber="101">
<figureCitation id="13232A04FFFD9D1D4BA50273D8542729" box="[151,279,1865,1892]" captionStart="FIGURES 50–55" captionStartId="21.[151,264,1885,1909]" captionTargetBox="[165,1422,182,1859]" captionTargetId="figure-17@21.[165,1422,181,1862]" captionTargetPageId="21" captionText="FIGURES 50–55. Species of Retrorsia and Retrorsioides, n. gen. 50–52. Retrorsia simplicissima, n. sp. 50. Head and anterior rings, dorsal view. 51. Midbody ring, dorsal view. 52. Pygidium, lateral view. SEM stub WS33-11. 53–55. Retrorsioides castellum, n. sp. 53. Head and anterior rings, dorsal view. 53. Midbody ring, dorsal view. 55. Pygidium, lateral view. SEM stub WS32-17." figureDoi="http://doi.org/10.5281/zenodo.4804976" httpUri="https://zenodo.org/record/4804976/files/figure.png" pageId="20" pageNumber="101">Figs 50–52</figureCitation>
,
<figureCitation id="13232A04FFFD9D1D4A100273D80F272E" box="[290,332,1865,1891]" captionStart="FIGURES 121–126" captionStartId="40.[151,264,1408,1432]" captionTargetBox="[160,1431,185,1372]" captionTargetId="figure-24@40.[151,1436,181,1384]" captionTargetPageId="40" captionText="FIGURES 121–126. Drawings of gonopods of polydesmid species. 121. Left gonopod of Retrorisia leonardi, mesal view. 122. Left gonopod of Retrorsia benedictae, mesal view. 123. Left gonopod of Retrorsia richarti,n. sp., mesal view. 124. Right gonopod of Retrorsia gracilis, n. sp., lateral view. 125, Gonopod tip of Retrorsia gracilis, mesal view. 126. Right gonopod of Retrorsia simplicissima, n. sp., lateral view." figureDoi="http://doi.org/10.5281/zenodo.4805071" httpUri="https://zenodo.org/record/4805071/files/figure.png" pageId="20" pageNumber="101">126</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFFD9D1D4BA502ABDA1227BE" pageId="20" pageNumber="101" type="materials_examined">
<paragraph id="8BA73681FFFD9D1D4BA502ABDA1227BE" blockId="20.[151,1437,1937,2035]" pageId="20" pageNumber="101">
<emphasis id="B96CEA93FFFD9D1D4BA502ABD99A27E6" bold="true" box="[151,217,1937,1963]" pageId="20" pageNumber="101">Type.</emphasis>
<materialsCitation id="3B703CDCFFFD9D1D4BD102ABDA0E27BE" ID-GBIF-Occurrence="3128519325" collectingDate="2004-12-04" collectionCode="CAS" collectorName="W. Leonard &amp; C. Richart. The" county="Lewis Co." location="Stillman Basin" municipality="Male" pageId="20" pageNumber="101" specimenCount="2" stateProvince="Washington" typeStatus="holotype">
<collectingMunicipality id="6BC3ACFBFFFD9D1D4BD102ABD85E27E6" box="[227,285,1937,1963]" pageId="20" pageNumber="101">Male</collectingMunicipality>
<typeStatus id="54A38823FFFD9D1D4A1502ABD8CA27E6" box="[295,393,1937,1963]" pageId="20" pageNumber="101" type="holotype">holotype</typeStatus>
from
<location id="8EC7605AFFFD9D1D4AE602ABDBC327E6" LSID="urn:lsid:plazi:treatment:03B18797FFFD9D1F4BA5021FD8CF222C:8EC7605AFFFD9D1D4AE602ABDBC327E6" box="[468,640,1937,1963]" county="Lewis Co." municipality="Male" name="Stillman Basin" pageId="20" pageNumber="101" stateProvince="Washington">Stillman Basin</location>
,
<quantity id="4CE09B64FFFD9D1D49BC02ABDBBB27E6" box="[654,760,1937,1963]" metricMagnitude="3" metricUnit="m" metricValue="2.8968192" pageId="20" pageNumber="101" unit="mi" value="1.8">1.8 miles</quantity>
on
<location id="8EC7605AFFFD9D1D481502ABDABD27E6" LSID="urn:lsid:plazi:treatment:03B18797FFFD9D1F4BA5021FD8CF222C:8EC7605AFFFD9D1D481502ABDABD27E6" box="[807,1022,1937,1963]" county="Lewis Co." municipality="Male" name="Weyerhauser Road" pageId="20" pageNumber="101" stateProvince="Washington">Weyerhauser Road</location>
4200 from
<location id="8EC7605AFFFD9D1D4FB902ABDD8527E6" LSID="urn:lsid:plazi:treatment:03B18797FFFD9D1F4BA5021FD8CF222C:8EC7605AFFFD9D1D4FB902ABDD8527E6" box="[1163,1222,1937,1963]" county="Lewis Co." municipality="Male" name="Road" pageId="20" pageNumber="101" stateProvince="Washington">Road</location>
4000, 2116 ft asl, 46.4956°, -123.2134°,
<collectingCounty id="62C64E0DFFFD9D1D4AA5028FDB4E2782" box="[407,525,1973,1999]" pageId="20" pageNumber="101">Lewis Co.</collectingCounty>
,
<collectingRegion id="49DCF863FFFD9D1D492A028FDBE32782" box="[536,672,1973,1999]" country="United States of America" name="Washington" pageId="20" pageNumber="101">Washington</collectingRegion>
, collected
<date id="FFA61041FFFD9D1D482A028FDA9C2782" box="[792,991,1973,1999]" pageId="20" pageNumber="101" value="2004-12-04">
<collectingDate id="EFE2E9A9FFFD9D1D482A028FDA9C2782" box="[792,991,1973,1999]" pageId="20" pageNumber="101" value="2004-12-04">4 December 2004</collectingDate>
</date>
by
<collectorName id="26ED5357FFFD9D1D4F3B028FDDCF2782" box="[1033,1164,1973,1999]" pageId="20" pageNumber="101">W. Leonard</collectorName>
and
<collectorName id="26ED5357FFFD9D1D4FF1028FDC332782" box="[1219,1392,1973,1999]" pageId="20" pageNumber="101">C. Richart. The</collectorName>
<typeStatus id="54A38823FFFD9D1D4E45028FD99E27BE" pageId="20" pageNumber="101" type="holotype">holotype</typeStatus>
is mounted on SEM stub WS 33–11, deposited in
<collectionCode id="ED09AE44FFFD9D1D482102E3DA0E27BE" box="[787,845,2009,2035]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="101" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF676609FFFC9D1C4BA50267D9BC27AF" ID-DOI="http://doi.org/10.5281/zenodo.4804976" ID-Zenodo-Dep="4804976" httpUri="https://zenodo.org/record/4804976/files/figure.png" pageId="21" pageNumber="102" startId="21.[151,264,1885,1909]" targetBox="[165,1422,182,1859]" targetPageId="21">
<paragraph id="8BA73681FFFC9D1C4BA50267D9BC27AF" blockId="21.[151,1437,1885,2018]" pageId="21" pageNumber="102">
<emphasis id="B96CEA93FFFC9D1C4BA50267D80C273B" bold="true" box="[151,335,1885,1910]" pageId="21" pageNumber="102">FIGURES 50–55</emphasis>
. Species of
<taxonomicName id="4C184D02FFFC9D1C4AFC0264DB6E2738" authorityName="Shelley" authorityYear="2003" box="[462,557,1886,1909]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEA93FFFC9D1C4AFC0264DB6E2738" box="[462,557,1886,1909]" italics="true" pageId="21" pageNumber="102">Retrorsia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C184D02FFFC9D1C49520267DBA82738" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[608,747,1885,1909]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="102" phylum="Arthropoda" rank="genus" status="gen. nov.">Retrorsioides</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFFC9D1C49CA0264DA032738" box="[760,832,1886,1910]" pageId="21" pageNumber="102" rank="genus">
<emphasis id="B96CEA93FFFC9D1C49CA0264DA79273B" bold="true" box="[760,826,1886,1910]" pageId="21" pageNumber="102">n. gen</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B96CEA93FFFC9D1C487A0264DAC4273B" bold="true" box="[840,903,1886,1910]" pageId="21" pageNumber="102">50–52</emphasis>
.
<emphasis id="B96CEA93FFFC9D1C48A70264DDBA273B" bold="true" box="[917,1273,1885,1910]" pageId="21" pageNumber="102">
<emphasis id="B96CEA93FFFC9D1C48A70264DDD32738" bold="true" box="[917,1168,1885,1910]" italics="true" pageId="21" pageNumber="102">
<taxonomicName id="4C184D02FFFC9D1C48A70264DDC82738" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[917,1163,1885,1910]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="102" phylum="Arthropoda" rank="species" species="simplicissima" status="sp. nov.">Retrorsia simplicissima</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25F57E8FFFC9D1C4FAA0264DD91273B" box="[1176,1234,1886,1910]" pageId="21" pageNumber="102" rank="species">n. sp.</taxonomicNameLabel>
50.
</emphasis>
Head and anterior rings, dorsal view.
<emphasis id="B96CEA93FFFC9D1C4AB502B8D8E327D7" bold="true" box="[391,416,1922,1946]" pageId="21" pageNumber="102">51</emphasis>
. Midbody ring, dorsal view.
<emphasis id="B96CEA93FFFC9D1C49FF02B8DBA527D7" bold="true" box="[717,742,1922,1946]" pageId="21" pageNumber="102">52</emphasis>
. Pygidium, lateral view. SEM stub WS33-11.
<emphasis id="B96CEA93FFFC9D1C4FF702B8DC4727D7" bold="true" box="[1221,1284,1922,1946]" pageId="21" pageNumber="102">53–55</emphasis>
.
<emphasis id="B96CEA93FFFC9D1C4E2002BBD81C27F3" bold="true" pageId="21" pageNumber="102">
<emphasis id="B96CEA93FFFC9D1C4E2002BBD84227F0" bold="true" italics="true" pageId="21" pageNumber="102">
<taxonomicName id="4C184D02FFFC9D1C4E2002BBD9BF27F0" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="102" phylum="Arthropoda" rank="species" species="castellum" status="sp. nov.">Retrorsioides castellum</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25F57E8FFFC9D1C4A35029CD80327F3" box="[263,320,1958,1982]" pageId="21" pageNumber="102" rank="species">n. sp.</taxonomicNameLabel>
53
</emphasis>
. Head and anterior rings, dorsal view.
<emphasis id="B96CEA93FFFC9D1C49D7029CDBBD27F3" bold="true" box="[741,766,1958,1982]" pageId="21" pageNumber="102">53</emphasis>
. Midbody ring, dorsal view.
<emphasis id="B96CEA93FFFC9D1C4F10029CDD7827F3" bold="true" box="[1058,1083,1958,1982]" pageId="21" pageNumber="102">55</emphasis>
. Pygidium, lateral view. SEM stub WS32-17.
</paragraph>
</caption>
<subSubSection id="C302650AFFFF9D1F4BF505ACDB3E20B4" pageId="22" pageNumber="103" type="diagnosis">
<paragraph id="8BA73681FFFF9D1F4BF505ACDB3E20B4" blockId="22.[151,1437,150,609]" pageId="22" pageNumber="103">
<emphasis id="B96CEA93FFFF9D1F4BF505ACD80120FD" bold="true" box="[199,322,150,176]" pageId="22" pageNumber="103">Diagnosis.</emphasis>
The gonopod (
<figureCitation id="13232A04FFFF9D1F4AD905ADDB0020FF" box="[491,579,151,178]" captionStart="FIGURES 121–126" captionStartId="40.[151,264,1408,1432]" captionTargetBox="[160,1431,185,1372]" captionTargetId="figure-24@40.[151,1436,181,1384]" captionTargetPageId="40" captionText="FIGURES 121–126. Drawings of gonopods of polydesmid species. 121. Left gonopod of Retrorisia leonardi, mesal view. 122. Left gonopod of Retrorsia benedictae, mesal view. 123. Left gonopod of Retrorsia richarti,n. sp., mesal view. 124. Right gonopod of Retrorsia gracilis, n. sp., lateral view. 125, Gonopod tip of Retrorsia gracilis, mesal view. 126. Right gonopod of Retrorsia simplicissima, n. sp., lateral view." figureDoi="http://doi.org/10.5281/zenodo.4805071" httpUri="https://zenodo.org/record/4805071/files/figure.png" pageId="22" pageNumber="103">Fig 126</figureCitation>
) is extremely simple, with the pulvillus about midway in its length, and only a hint of a pulvillar notch. The terminal zone tip is undivided and slightly retrorse. The anterior marginal setal row of the collum consists of 22 setae (
<figureCitation id="13232A04FFFF9D1F492E05E5DB3320B4" box="[540,624,222,249]" captionStart="FIGURES 50–55" captionStartId="21.[151,264,1885,1909]" captionTargetBox="[165,1422,182,1859]" captionTargetId="figure-17@21.[165,1422,181,1862]" captionTargetPageId="21" captionText="FIGURES 50–55. Species of Retrorsia and Retrorsioides, n. gen. 50–52. Retrorsia simplicissima, n. sp. 50. Head and anterior rings, dorsal view. 51. Midbody ring, dorsal view. 52. Pygidium, lateral view. SEM stub WS33-11. 53–55. Retrorsioides castellum, n. sp. 53. Head and anterior rings, dorsal view. 53. Midbody ring, dorsal view. 55. Pygidium, lateral view. SEM stub WS32-17." figureDoi="http://doi.org/10.5281/zenodo.4804976" httpUri="https://zenodo.org/record/4804976/files/figure.png" pageId="22" pageNumber="103">Fig. 50</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFFF9D1F4BF50439DD522150" box="[199,1041,259,285]" pageId="22" pageNumber="103" type="etymology">
<paragraph id="8BA73681FFFF9D1F4BF50439DD522150" blockId="22.[151,1437,150,609]" box="[199,1041,259,285]" pageId="22" pageNumber="103">
<emphasis id="B96CEA93FFFF9D1F4BF50439D80D2150" bold="true" box="[199,334,259,285]" pageId="22" pageNumber="103">Etymology.</emphasis>
The species name, an adjective, refers to the simple gonopods.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFFF9D1F4BF5041CDDD8219C" pageId="22" pageNumber="103" type="description">
<paragraph id="8BA73681FFFF9D1F4BF5041CDDD8219C" blockId="22.[151,1437,150,609]" pageId="22" pageNumber="103">
<emphasis id="B96CEA93FFFF9D1F4BF5041CD81A210D" bold="true" box="[199,345,294,320]" pageId="22" pageNumber="103">Description.</emphasis>
Length about 5.0 mm, greatest width
<quantity id="4CE09B64FFFF9D1F4833041DDA2B210C" box="[769,872,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="22" pageNumber="103" unit="mm" value="0.46">0.46 mm</quantity>
. Anterior metazonites (
<figureCitation id="13232A04FFFF9D1F4F58041DDDAA210C" box="[1130,1257,294,321]" captionStart="FIGURES 50–55" captionStartId="21.[151,264,1885,1909]" captionTargetBox="[165,1422,182,1859]" captionTargetId="figure-17@21.[165,1422,181,1862]" captionTargetPageId="21" captionText="FIGURES 50–55. Species of Retrorsia and Retrorsioides, n. gen. 50–52. Retrorsia simplicissima, n. sp. 50. Head and anterior rings, dorsal view. 51. Midbody ring, dorsal view. 52. Pygidium, lateral view. SEM stub WS33-11. 53–55. Retrorsioides castellum, n. sp. 53. Head and anterior rings, dorsal view. 53. Midbody ring, dorsal view. 55. Pygidium, lateral view. SEM stub WS32-17." figureDoi="http://doi.org/10.5281/zenodo.4804976" httpUri="https://zenodo.org/record/4804976/files/figure.png" pageId="22" pageNumber="103">Figs 50, 51</figureCitation>
) with four rows of setae, a few posterior segments may appear to have five rows due to intercalated setae. Epiproct not swollen, short. Anterior legs crassate, tarsi with sphaerotrichomes. Gonopod (
<figureCitation id="13232A04FFFF9D1F48AD0455DABC21C7" box="[927,1023,367,394]" captionStart="FIGURES 121–126" captionStartId="40.[151,264,1408,1432]" captionTargetBox="[160,1431,185,1372]" captionTargetId="figure-24@40.[151,1436,181,1384]" captionTargetPageId="40" captionText="FIGURES 121–126. Drawings of gonopods of polydesmid species. 121. Left gonopod of Retrorisia leonardi, mesal view. 122. Left gonopod of Retrorsia benedictae, mesal view. 123. Left gonopod of Retrorsia richarti,n. sp., mesal view. 124. Right gonopod of Retrorsia gracilis, n. sp., lateral view. 125, Gonopod tip of Retrorsia gracilis, mesal view. 126. Right gonopod of Retrorsia simplicissima, n. sp., lateral view." figureDoi="http://doi.org/10.5281/zenodo.4805071" httpUri="https://zenodo.org/record/4805071/files/figure.png" pageId="22" pageNumber="103">Fig. 126</figureCitation>
) with small prefemorite. Acropodite relatively short, stout, pulvillus near midlength of acropodite, pulvillar notch small, shallow, indicated by slight swelling immediately distal. Terminal zone simple, only slightly retrorse. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFFF9D1F4BF504E0D8CF222C" pageId="22" pageNumber="103" type="distribution">
<paragraph id="8BA73681FFFF9D1F4BF504E0DBB221B8" blockId="22.[151,1437,150,609]" box="[199,753,474,501]" pageId="22" pageNumber="103">
<emphasis id="B96CEA93FFFF9D1F4BF504E0D82221B9" bold="true" box="[199,353,474,500]" pageId="22" pageNumber="103">Distribution.</emphasis>
Only known from the
<typeStatus id="54A38823FFFF9D1F495204E1DBD321B8" box="[608,656,475,501]" pageId="22" pageNumber="103">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BA73681FFFF9D1F4BF504C5D8CF222C" blockId="22.[151,1437,150,609]" pageId="22" pageNumber="103">
<emphasis id="B96CEA93FFFF9D1F4BF504C5D8522254" bold="true" box="[199,273,511,537]" pageId="22" pageNumber="103">Notes.</emphasis>
This species seems marginal to our concept of
<emphasis id="B96CEA93FFFF9D1F481304C5DAD22255" box="[801,913,511,536]" italics="true" pageId="22" pageNumber="103">
<taxonomicName id="4C184D02FFFF9D1F481304C5DACE2255" authorityName="Shelley" authorityYear="2003" box="[801,909,511,536]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="genus">Retrorsia</taxonomicName>
,
</emphasis>
though the general appearance of the gonopod in comparison to the other species suggests it belongs here. The gonopod is among the simplest to be found in the family
<taxonomicName id="4C184D02FFFF9D1F4BD5077DD8CB222C" authorityName="Leach" authorityYear="1815" box="[231,392,583,609]" class="Diplopoda" family="Polydesmidae" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="family">Polydesmidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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