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<document id="F93BC1D703F513296049063E5DDFC2C5" ID-CLB-Dataset="298914" ID-DOI="10.11646/zootaxa.5474.1.3" ID-GBIF-Dataset="9a12ec35-d848-4b69-8cee-d67b7bcd5716" ID-ISSN="1175-5326" ID-Zenodo-Dep="12526909" ID-ZooBank="EE52D68D-7082-43D2-B326-026621AE36C5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719300134533" checkinUser="plazi" docAuthor="Moreno, Carlos &amp; Zilberman, Bruno" docDate="2024" docId="03B187855C46FFFFABACD787FDA5609A" docLanguage="en" docName="zootaxa.5474.1.3.pdf" docOrigin="Zootaxa 5474 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5474.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratoxenus inflatus Moreno &amp; Zilberman 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="42" masterDocId="FF88FFFD5C40FFF8AB3BD348FF996720" masterDocTitle="Revalidation of the termitophilous genus Ceratoxenus Mann and description of two new species from the Neotropical region (Coleoptera: Staphylinidae: Aleocharinae: Corotocini)" masterLastPageNumber="49" masterPageNumber="35" pageNumber="41" updateTime="1719412577716" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="20A789A707F4D90D01984E8591BE3D30" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DD255D45FA747CF6D80A8A995C4F7FD6">
<mods:title id="618D3903BA7D2B15BD374901D5F5036A">Revalidation of the termitophilous genus Ceratoxenus Mann and description of two new species from the Neotropical region (Coleoptera: Staphylinidae: Aleocharinae: Corotocini)</mods:title>
</mods:titleInfo>
<mods:name id="E4EF91D7DA8E2F925533B22BA719B13F" type="personal">
<mods:role id="333BFA0A06DB189B9D90833DAB2116FD">
<mods:roleTerm id="881E17D036D0F128745755408649053B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="99940B6339931616A1795CBFB83A5A2F">Moreno, Carlos</mods:namePart>
<mods:nameIdentifier id="33CDC96AA3930065ED97694C4F78E6E7" type="ORCID">0000-0002-6195-8648</mods:nameIdentifier>
<mods:affiliation id="8C77FDA1215D20C79924AF8D041FF2D7">Programa de Pós-Graduação em Sistemática, Taxonomia Animal e Biodiversidade, LaC-Laboratório de Coleoptera, Museu de Zoologia da Universidade de São Paulo, 04263 - 000, São Paulo, Brazil.</mods:affiliation>
<mods:nameIdentifier id="7F7BC12EBA3791BD1747D3CD2A75DEA0" type="email">cmpiresilva@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="3BB8CF0B9B51F70347F9BE22A050CC5F" type="personal">
<mods:role id="8268E86C9A7C59F98D49FCA5925600EF">
<mods:roleTerm id="8018CA17BDB5344EFFBCDB63B88EE6E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="72736AFABB052321B639E4E473E23914">Zilberman, Bruno</mods:namePart>
<mods:nameIdentifier id="C2473B9D926EEE4F8DF83186085DE04F" type="ORCID">0000-0003-2613-4827</mods:nameIdentifier>
<mods:affiliation id="F57CC51919F9A8B61F58C7060A6D7C17">Programa de Pós-Graduação em Sistemática, Taxonomia Animal e Biodiversidade, LaC-Laboratório de Coleoptera, Museu de Zoologia da Universidade de São Paulo, 04263 - 000, São Paulo, Brazil.</mods:affiliation>
<mods:nameIdentifier id="E0EA55BC9214AF9094A84CE8F76A64BF" type="email">brunozilberman@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="CE4760931ECF383EE83485A8DC103177">text</mods:typeOfResource>
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<mods:title id="60FE38D50D1D53E3C474F46B3171A09A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="32AF27A4B128CF16EF7710E4CBD960AA">
<mods:date id="49C9B04413FA8B0B078AC10C9CB2D029">2024</mods:date>
<mods:detail id="DD9A0970D01B313822703E00557945ED" type="pubDate">
<mods:number id="F2DCFBE5276547F3A73C154A40CBFD10">2024-06-25</mods:number>
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<mods:detail id="074C8DADE35E425DF40A5B34554E0CD0" type="volume">
<mods:number id="D656EE2C5C3AD1279E27F364A3731390">5474</mods:number>
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<mods:number id="ABB252F11DFE4D1993FAABDE09A9FC33">1</mods:number>
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<mods:identifier id="3F868E27584A2368A61263399A607189" type="CLB-Dataset">298914</mods:identifier>
<mods:identifier id="6872457DA8BF48888C9156E19DB3D0DE" type="DOI">10.11646/zootaxa.5474.1.3</mods:identifier>
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<treatment id="03B187855C46FFFFABACD787FDA5609A" ID-DOI="http://doi.org/10.5281/zenodo.12526977" ID-GBIF-Taxon="233791022" ID-Zenodo-Dep="12526977" LSID="urn:lsid:plazi:treatment:03B187855C46FFFFABACD787FDA5609A" httpUri="http://treatment.plazi.org/id/03B187855C46FFFFABACD787FDA5609A" lastPageId="7" lastPageNumber="42" pageId="6" pageNumber="41">
<subSubSection id="C30265185C46FFFEABACD787FC8963C9" box="[151,784,1230,1257]" pageId="6" pageNumber="41" type="nomenclature">
<paragraph id="8BA736935C46FFFEABACD787FC8963C9" blockId="6.[151,784,1230,1293]" box="[151,784,1230,1257]" pageId="6" pageNumber="41">
<heading id="D0EF81FF5C46FFFEABACD787FC8963C9" bold="true" box="[151,784,1230,1257]" fontSize="11" level="1" pageId="6" pageNumber="41" reason="1">
<emphasis id="B96CEA815C46FFFEABACD787FC8963C9" bold="true" box="[151,784,1230,1257]" pageId="6" pageNumber="41">
<taxonomicName id="4C184D105C46FFFEABACD787FD3563C8" authority="Moreno &amp; Zilberman" authorityName="Moreno &amp; Zilberman" authorityYear="2024" box="[151,684,1230,1257]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="inflatus" status="sp. nov.">
<emphasis id="B96CEA815C46FFFEABACD787FE0C63C9" bold="true" box="[151,405,1230,1257]" italics="true" pageId="6" pageNumber="41">Ceratoxenus inflatus</emphasis>
Moreno &amp; Zilberman
</taxonomicName>
<taxonomicNameLabel id="A25F57FA5C46FFFEA988D787FC8963C9" box="[691,784,1231,1257]" pageId="6" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30265185C46FFFEABACD7BBFEF9622D" box="[151,352,1266,1293]" pageId="6" pageNumber="41" type="description">
<paragraph id="8BA736935C46FFFEABACD7BBFEF9622D" blockId="6.[151,784,1230,1293]" box="[151,352,1266,1293]" pageId="6" pageNumber="41">
(
<figureCitation id="13232A165C46FFFEABA4D7BBFF76622D" box="[159,239,1267,1293]" captionStart="FIGURE 4" captionStartId="7.[151,250,1306,1330]" captionTargetBox="[151,1436,181,1282]" captionTargetId="figure-17@7.[151,1436,181,1282]" captionTargetPageId="7" captionText="FIGURE 4. Ceratoxenus inflatus Moreno and Zilberman sp. nov. Holotype. Habitus lateral (A), dorso-lateral (B), dorsal (C), and labels (D)." figureDoi="http://doi.org/10.5281/zenodo.12526929" httpUri="https://zenodo.org/record/12526929/files/figure.png" pageId="6" pageNumber="41">Figs. 4</figureCitation>
,
<figureCitation id="13232A165C46FFFEABC0D7BAFE92622D" box="[251,267,1266,1293]" captionStart="FIGURE 5" captionStartId="8.[151,250,1659,1683]" captionTargetBox="[151,1436,684,1636]" captionTargetId="figure-340@8.[151,1436,684,1636]" captionTargetPageId="8" captionText="FIGURE 5. Ceratoxenus inflatus Moreno and Zilberman sp. nov. Holotype.Habitus ventral, head and thorax (A), and abdominal sternites II–V (B)." figureDoi="http://doi.org/10.5281/zenodo.12526933" httpUri="https://zenodo.org/record/12526933/files/figure.png" pageId="6" pageNumber="41">5</figureCitation>
,
<figureCitation id="13232A165C46FFFEAA2CD7BBFECE622D" box="[279,343,1267,1293]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,885,1824]" captionTargetId="figure-305@9.[151,1436,885,1824]" captionTargetPageId="9" captionText="FIGURE 6. Comparative external morphology of head and thorax among Ceratoxenus species: C. tricornis Mann, 1923 (A, D), C. inflatus sp. nov. (C, E), and C. akerata sp. nov. (C, F). White arrows point to the lateral horns, yellow arrows point to the gena with a distinctive carina in C. inflatus and C. akerata, and uncarinated in C. tricornis, while cyan arrows point to the medial projection between lateral horns." figureDoi="http://doi.org/10.5281/zenodo.12526937" httpUri="https://zenodo.org/record/12526937/files/figure.png" pageId="6" pageNumber="41">6B, E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30265185C46FFFEABACD672FD666099" pageId="6" pageNumber="41" type="diagnosis">
<paragraph id="8BA736935C46FFFEABACD672FD8362E1" blockId="6.[151,1437,1338,1977]" pageId="6" pageNumber="41">
<emphasis id="B96CEA815C46FFFEABACD672FE936274" bold="true" box="[151,266,1338,1364]" pageId="6" pageNumber="41">Diagnosis</emphasis>
: The species is recognized by the larger and closely spaced lateral projections at frons; by the rounded triangular medial projection, overtaking the level of frontal elevations; gena elevated, broadly rounded, forming a longitudinal carina behind eyes; pronotum with anterior impression shorter and noticebly narrower than posterior, and with anterior margin arcuated.
</paragraph>
<paragraph id="8BA736935C46FFFEABFCD683FD91619D" blockId="6.[151,1437,1338,1977]" pageId="6" pageNumber="41">
<emphasis id="B96CEA815C46FFFEABFCD683FE9762C5" bold="true" box="[199,270,1483,1509]" pageId="6" pageNumber="41">Head.</emphasis>
Gena somewhat smooth, broadly elevated behind eyes; head surface finely punctuate; lateral projections at frons larger and closely spaced; medial process roundely projected. Long bristles absent on vertex (
<figureCitation id="13232A165C46FFFEAE08D6A7FA016129" box="[1331,1432,1519,1545]" captionStart="FIGURE 4" captionStartId="7.[151,250,1306,1330]" captionTargetBox="[151,1436,181,1282]" captionTargetId="figure-17@7.[151,1436,181,1282]" captionTargetPageId="7" captionText="FIGURE 4. Ceratoxenus inflatus Moreno and Zilberman sp. nov. Holotype. Habitus lateral (A), dorso-lateral (B), dorsal (C), and labels (D)." figureDoi="http://doi.org/10.5281/zenodo.12526929" httpUri="https://zenodo.org/record/12526929/files/figure.png" pageId="6" pageNumber="41">Figs. 4C</figureCitation>
,
<figureCitation id="13232A165C46FFFEABACD55BFF4A610D" box="[151,211,1555,1581]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,885,1824]" captionTargetId="figure-305@9.[151,1436,885,1824]" captionTargetPageId="9" captionText="FIGURE 6. Comparative external morphology of head and thorax among Ceratoxenus species: C. tricornis Mann, 1923 (A, D), C. inflatus sp. nov. (C, E), and C. akerata sp. nov. (C, F). White arrows point to the lateral horns, yellow arrows point to the gena with a distinctive carina in C. inflatus and C. akerata, and uncarinated in C. tricornis, while cyan arrows point to the medial projection between lateral horns." figureDoi="http://doi.org/10.5281/zenodo.12526937" httpUri="https://zenodo.org/record/12526937/files/figure.png" pageId="6" pageNumber="41">6B,E</figureCitation>
). Antenna with antennomere 1 cylindrical, with almost same length of antennomeres 2–4 combined; 2–4 longer than wide, subequal in length; 5–8 shorter than previous, slightly longer than wide, all subequal in length; 9– 10 slightly shorter than previous, subquadrate; 11 conical. Second maxilar palpomere broadly rounded and inflated at apex. Postmentum longer than wide occupying more than half of total head length,, slightly longer than gula, entirely covered by long bristles.
</paragraph>
<paragraph id="8BA736935C46FFFEABFCD58FFADB60B5" blockId="6.[151,1437,1338,1977]" pageId="6" pageNumber="41">
<emphasis id="B96CEA815C46FFFEABFCD58FFEBE61C1" bold="true" box="[199,295,1735,1761]" pageId="6" pageNumber="41">Thorax.</emphasis>
Pronotum slightly broader on posterior region; pronotum light-brown with lateral margins darker; anterior impression shorter and noticeably narrower than posterior, and with anterior margin strongly arcuate, bearing long bristles marginally; lateral margins parallel with long bristles sparsely distributed (
<figureCitation id="13232A165C46FFFEAF8CD447FA816009" box="[1207,1304,1807,1833]" captionStart="FIGURE 4" captionStartId="7.[151,250,1306,1330]" captionTargetBox="[151,1436,181,1282]" captionTargetId="figure-17@7.[151,1436,181,1282]" captionTargetPageId="7" captionText="FIGURE 4. Ceratoxenus inflatus Moreno and Zilberman sp. nov. Holotype. Habitus lateral (A), dorso-lateral (B), dorsal (C), and labels (D)." figureDoi="http://doi.org/10.5281/zenodo.12526929" httpUri="https://zenodo.org/record/12526929/files/figure.png" pageId="6" pageNumber="41">Figs. 4C</figureCitation>
,
<figureCitation id="13232A165C46FFFEAE18D447FADC6009" box="[1315,1349,1807,1833]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,885,1824]" captionTargetId="figure-305@9.[151,1436,885,1824]" captionTargetPageId="9" captionText="FIGURE 6. Comparative external morphology of head and thorax among Ceratoxenus species: C. tricornis Mann, 1923 (A, D), C. inflatus sp. nov. (C, E), and C. akerata sp. nov. (C, F). White arrows point to the lateral horns, yellow arrows point to the gena with a distinctive carina in C. inflatus and C. akerata, and uncarinated in C. tricornis, while cyan arrows point to the medial projection between lateral horns." figureDoi="http://doi.org/10.5281/zenodo.12526937" httpUri="https://zenodo.org/record/12526937/files/figure.png" pageId="6" pageNumber="41">6E</figureCitation>
). Elytra elongate, wider on posterior region; light-brown, outer margin darker. Legs with pro- and mesocoxae transverse; metacoxa triangular. Tibia slightly longer than femur, covered with long and thick bristles along entire length; outer margins of tibia slightly convex, and inner margin straight; pro-, meso-, and metatarsi increasing in length.
</paragraph>
<paragraph id="8BA736935C46FFFEABFCD4D6FD666099" blockId="6.[151,1437,1338,1977]" box="[199,767,1950,1977]" pageId="6" pageNumber="41">
<emphasis id="B96CEA815C46FFFEABFCD4D6FEDA6098" bold="true" box="[199,323,1950,1976]" pageId="6" pageNumber="41">Abdomen.</emphasis>
As described to genus. Male unknown.
</paragraph>
</subSubSection>
<caption id="DF67661B5C47FFFFABACD652FEB46276" ID-DOI="http://doi.org/10.5281/zenodo.12526929" ID-Zenodo-Dep="12526929" httpUri="https://zenodo.org/record/12526929/files/figure.png" pageId="7" pageNumber="42" startId="7.[151,250,1306,1330]" targetBox="[151,1436,181,1282]" targetPageId="7" targetType="figure">
<paragraph id="8BA736935C47FFFFABACD652FEB46276" blockId="7.[151,1436,1306,1366]" pageId="7" pageNumber="42">
<emphasis id="B96CEA815C47FFFFABACD652FE8D6212" bold="true" box="[151,276,1306,1330]" pageId="7" pageNumber="42">FIGURE 4.</emphasis>
<taxonomicName id="4C184D105C47FFFFAA20D653FD7B6212" authority="Moreno and Zilberman" authorityName="Moreno and Zilberman" authorityYear="2024" box="[283,738,1306,1330]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="inflatus" status="sp. nov.">
<emphasis id="B96CEA815C47FFFFAA20D653FE776212" box="[283,494,1306,1330]" italics="true" pageId="7" pageNumber="42">Ceratoxenus inflatus</emphasis>
Moreno and Zilberman
</taxonomicName>
<emphasis id="B96CEA815C47FFFFA9D3D652FCA06212" bold="true" box="[744,825,1306,1330]" pageId="7" pageNumber="42">
<taxonomicNameLabel id="A25F57FA5C47FFFFA9D3D652FCA06212" box="[744,825,1306,1330]" pageId="7" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype. Habitus lateral (A), dorso-lateral (B), dorsal (C), and labels (D).
</paragraph>
</caption>
<subSubSection id="C30265185C47FFFFABFCD6E0FA0B62C7" pageId="7" pageNumber="42" type="materials_examined">
<paragraph id="8BA736935C47FFFFABFCD6E0FA0B62C7" blockId="7.[151,1437,1448,1978]" pageId="7" pageNumber="42">
<emphasis id="B96CEA815C47FFFFABFCD6E0FE3462E2" bold="true" box="[199,429,1448,1474]" pageId="7" pageNumber="42">Material examined</emphasis>
.
<materialsCitation id="3B703CCE5C47FFFFAA84D6E0FA0B62C7" ID-GBIF-Occurrence="4899569303" collectorName="A. Emerson" country="Guyana" location="Field Museum of Natural History" pageId="7" pageNumber="42" specimenCount="2" typeStatus="holotype">
<emphasis id="B96CEA815C47FFFFAA84D6E0FDC762E2" bold="true" box="[447,606,1448,1474]" pageId="7" pageNumber="42">
<typeStatus id="54A388315C47FFFFAA84D6E0FDC762E2" box="[447,606,1448,1474]" pageId="7" pageNumber="42" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
. Female.
<collectingCountry id="F30F76035C47FFFFA9EED6E0FCD662E2" box="[725,847,1448,1474]" name="Guyana" pageId="7" pageNumber="42">GUYANA</collectingCountry>
. “TOPOTYPE”; “Kartabo, British Guiana, 13: III: 1924”; “Coll. by,
<collectorName id="26ED53455C47FFFFAA67D685FE4462C7" box="[348,477,1485,1511]" pageId="7" pageNumber="42">A.Emerson</collectorName>
, #24.50 [field number]”; “FMNHINS [
<location id="8EC760485C47FFFFA8A9D684FA9662C6" LSID="urn:lsid:plazi:treatment:03B187855C46FFFFABACD787FDA5609A:8EC760485C47FFFFA8A9D684FA9662C6" box="[914,1295,1484,1511]" country="Guyana" name="Field Museum of Natural History" pageId="7" pageNumber="42">Field Museum of Natural History</location>
] 4518561”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30265185C47FFFFABFCD6B8FB52610E" pageId="7" pageNumber="42" type="etymology">
<paragraph id="8BA736935C47FFFFABFCD6B8FB52610E" blockId="7.[151,1437,1448,1978]" pageId="7" pageNumber="42">
<emphasis id="B96CEA815C47FFFFABFCD6B8FED7612A" bold="true" box="[199,334,1520,1546]" pageId="7" pageNumber="42">Etymology.</emphasis>
The epithet is derived from Latin,
<taxonomicName id="4C184D105C47FFFFA9EFD6B8FCB1612A" authorityName="Moreno and Zilberman" authorityYear="2024" box="[724,808,1520,1546]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="inflatus">
<emphasis id="B96CEA815C47FFFFA9EFD6B8FCB1612A" box="[724,808,1520,1546]" italics="true" pageId="7" pageNumber="42">inflatus</emphasis>
</taxonomicName>
, essentially meaning “inflate”, in allusion to the bloated elevated gena, and to the conspicuously rounded lateral projections of head. Noun in masculine.
</paragraph>
</subSubSection>
<subSubSection id="C30265185C47FFFFABFCD570FE1A6052" pageId="7" pageNumber="42" type="discussion">
<paragraph id="8BA736935C47FFFFABFCD570FE1A6052" blockId="7.[151,1437,1448,1978]" pageId="7" pageNumber="42">
<emphasis id="B96CEA815C47FFFFABFCD570FEA26172" bold="true" box="[199,315,1592,1618]" pageId="7" pageNumber="42">Remarks.</emphasis>
The specimen studied contains the Emerson’s label “24.50” dated “
<date id="FFA610535C47FFFFAF62D570FB426172" box="[1113,1243,1592,1618]" pageId="7" pageNumber="42" value="1924-03-13">13.III.1924</date>
”, and seemingly several termitophiles were collected by him in this date in different nests of
<taxonomicName id="4C184D105C47FFFFA8CDD515FB686156" baseAuthorityName="Holmgren" baseAuthorityYear="1910" box="[1014,1265,1629,1654]" class="Insecta" family="Termitidae" genus="Nasutitermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="guyanae">
<emphasis id="B96CEA815C47FFFFA8CDD515FB686156" box="[1014,1265,1629,1654]" italics="true" pageId="7" pageNumber="42">Nasutitermes guyanae</emphasis>
</taxonomicName>
. The specimen here studied was not referenced in Emerson's synonyms paper, but he mentioned
<specimenCount id="9D1EFD1A5C47FFFFAF16D5C9FB5E61BA" box="[1069,1223,1664,1691]" count="1" pageId="7" pageNumber="42" type="generic">one specimen</specimenCount>
labeled as &quot;24.61&quot; as &quot;
<taxonomicName id="4C184D105C47FFFFABFBD5ECFE16619E" authorityName="Mann" authorityYear="1923" box="[192,399,1700,1726]" class="Insecta" family="Staphylinidae" genus="Xenopelta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B96CEA815C47FFFFABFBD5ECFE16619E" box="[192,399,1700,1726]" italics="true" pageId="7" pageNumber="42">Xenopelta cornuta</emphasis>
</taxonomicName>
&quot; from this same date and locatity. Probably, these
<specimenCount id="9D1EFD1A5C47FFFFA8F2D5EDFBF7619E" box="[969,1134,1700,1727]" count="2" pageId="7" pageNumber="42" type="generic">two specimens</specimenCount>
belong to the same species (
<taxonomicName id="4C184D105C47FFFFAB9AD581FE8C61C2" authorityName="Moreno and Zilberman" authorityYear="2024" box="[161,277,1736,1763]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="inflatus">
<emphasis id="B96CEA815C47FFFFAB9AD581FE8C61C2" box="[161,277,1736,1763]" italics="true" pageId="7" pageNumber="42">C. inflatus</emphasis>
</taxonomicName>
), and based on these data we deduce that the host of
<taxonomicName id="4C184D105C47FFFFA858D581FC4E61C2" authorityName="Moreno and Zilberman" authorityYear="2024" box="[867,983,1736,1763]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="inflatus">
<emphasis id="B96CEA815C47FFFFA858D581FC4E61C2" box="[867,983,1736,1763]" italics="true" pageId="7" pageNumber="42">C. inflatus</emphasis>
</taxonomicName>
is
<taxonomicName id="4C184D105C47FFFFA8CCD581FB6861C2" baseAuthorityName="Holmgren" baseAuthorityYear="1910" box="[1015,1265,1737,1762]" class="Insecta" family="Termitidae" genus="Nasutitermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="guyanae">
<emphasis id="B96CEA815C47FFFFA8CCD581FB6861C2" box="[1015,1265,1737,1762]" italics="true" pageId="7" pageNumber="42">Nasutitermes guyanae</emphasis>
</taxonomicName>
, as this species was the only termite with termitophiles
<bibRefCitation id="EF894B625C47FFFFA96CD5A5FC956027" author="Emerson, A. E." box="[599,780,1772,1799]" pageId="7" pageNumber="42" pagination="369 - 395" refId="ref7276" refString="Emerson, A. E. (1935) Termitophile distribution and quantitative characters as indicators of physiological speciation in British Guiana termites (Isoptera). Annals of the Entomological Society of America, 28, 369 - 395. https: // doi. org / 10.1093 / aesa / 28.3.369" type="journal article" year="1935">Emerson (1935)</bibRefCitation>
recorded at that particular place and date. It is noteworthy that the author erroneous synonym would stand along with any likely output, since there is no possibility of any of the abovementioned specimens belong to the genus
<emphasis id="B96CEA815C47FFFFA9D3D47CFCF8606E" box="[744,865,1844,1870]" italics="true" pageId="7" pageNumber="42">
<taxonomicName id="4C184D105C47FFFFA9D3D47CFCC4606E" box="[744,861,1844,1870]" class="Insecta" family="Staphylinidae" genus="Xenopelta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="genus">Xenopelta</taxonomicName>
.
</emphasis>
For comparison with other species, please see the other remarks above.
</paragraph>
</subSubSection>
<subSubSection id="C30265185C47FFFFABFCD434FC0760B6" box="[199,926,1916,1942]" pageId="7" pageNumber="42" type="biology_ecology">
<paragraph id="8BA736935C47FFFFABFCD434FC0760B6" blockId="7.[151,1437,1448,1978]" box="[199,926,1916,1942]" pageId="7" pageNumber="42">
<emphasis id="B96CEA815C47FFFFABFCD434FEBE60B6" bold="true" box="[199,295,1916,1942]" pageId="7" pageNumber="42">Biology.</emphasis>
Probably
<taxonomicName id="4C184D105C47FFFFAAA1D435FD3B60B6" authorityName="Mann" authorityYear="1923" baseAuthorityName="Holmgren" baseAuthorityYear="1910" box="[410,674,1917,1942]" class="Insecta" family="Termitidae" genus="Nasutitermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="guayanae">
<emphasis id="B96CEA815C47FFFFAAA1D435FD3B60B6" box="[410,674,1917,1942]" italics="true" pageId="7" pageNumber="42">Nasutitermes guayanae</emphasis>
</taxonomicName>
(see remarks section).
</paragraph>
</subSubSection>
<subSubSection id="C30265185C47FFFFABFCD4E8FDA5609A" box="[199,572,1952,1978]" pageId="7" pageNumber="42" type="distribution">
<paragraph id="8BA736935C47FFFFABFCD4E8FDA5609A" blockId="7.[151,1437,1448,1978]" box="[199,572,1952,1978]" pageId="7" pageNumber="42">
<emphasis id="B96CEA815C47FFFFABFCD4E8FEF8609A" bold="true" box="[199,353,1952,1978]" pageId="7" pageNumber="42">Distribution.</emphasis>
<collectingCountry id="F30F76035C47FFFFAA5CD4E8FE26609A" box="[359,447,1952,1978]" name="Guyana" pageId="7" pageNumber="42">Guyana</collectingCountry>
—Kartabo.
</paragraph>
</subSubSection>
</treatment>
</document>
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