366 lines
68 KiB
XML
366 lines
68 KiB
XML
<document id="E7130E8A052003939A744DB0B2D3B571" ID-DOI="10.1515/aemnp-2017-0099" ID-ISSN="0374-1036" ID-Zenodo-Dep="5317730" ID-ZooBank="27FC96A5-11A0-478C-B9C2-1514BDB0D45E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630128735338" checkinUser="marcus" docAuthor="Tkoč, Michal, Ježek, Jan & Pont, François Le" docDate="2017" docId="03AA74003B3DFFE3FE0B098BFD06C18C" docLanguage="en" docName="ActaEntMusNatPra.2017.57.2.765-774.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 57 (2)" docSource="http://dx.doi.org/10.1515/aemnp-2017-0099" docStyle="DocumentStyle:60E99616ED1F2F5EE637DA5E5309A4D2.7:ActaEntMusNatPra.2017.journal_article" docStyleId="60E99616ED1F2F5EE637DA5E5309A4D2" docStyleName="ActaEntMusNatPra.2017.journal_article" docStyleVersion="7" docTitle="Alepia bifida Tkoč & Ježek & Pont 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="772" masterDocId="FF930C783B3CFFE4FFEE0D78FFC6C35F" masterDocTitle="Alepia bifida, a new species of moth flies from Central America (Diptera: Psychodidae: Psychodinae)" masterLastPageNumber="774" masterPageNumber="765" pageNumber="766" updateTime="1699161950608" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D08494CC9AD2071598BFD8A3C10323A0">Alepia bifida, a new species of moth flies from Central America (Diptera: Psychodidae: Psychodinae)</mods:title>
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<mods:namePart id="394A703BE3469C8EC944BA3C5287FDA0">Tkoč, Michal</mods:namePart>
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<mods:affiliation id="5FC96952F568A0A4BBE7D8B1746D1A72">1,2,4) &) &) National Museum, Department of Entomology, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic; e-mails: michaltkoc @ gmail. com, jan. jezek @ o 2 active. cz & 1,2,4) &) Charles University in Prague, Faculty of Science, Department of Zoology, Viničná 7, CZ- 128 44 Praha 2, Czech Republic, & 1,2,4) &) Corresponding author</mods:affiliation>
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<mods:namePart id="2B6294FC5EA36B9EE76D05603597A63E">Ježek, Jan</mods:namePart>
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<mods:affiliation id="FBB2BAC0096D0799289B4C339CC55293">1,2,4) &) &) National Museum, Department of Entomology, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic; e-mails: michaltkoc @ gmail. com, jan. jezek @ o 2 active. cz</mods:affiliation>
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<mods:namePart id="FAB853C58AEA1FB52B98A83F4BA6FA70">Pont, François Le</mods:namePart>
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<mods:title id="7FBC4EC3E97ADD5DEADF487B1CBFBBD9">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<taxonomicName id="4C03BE953B3DFFE5FE0B098BFDB7C654" ID-CoL="9J3NJ" authority="Tkoč & Ježek & Pont, 2017" authorityName="Tkoč & Ježek & Pont" authorityYear="2017" box="[485,625,1267,1291]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="766" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
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<emphasis id="B97719043B3DFFE5FE0B098BFDB7C654" bold="true" box="[485,625,1267,1291]" italics="true" pageId="1" pageNumber="766">Alepia bifida</emphasis>
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(
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<figureCitation id="1338D9933B3DFFE5FDC3086AFD42C67A" box="[557,644,1298,1317]" captionStart-0="Figs 1–6" captionStart-1="Figs 7–10" captionStart-2="Figs 11–18" captionStart-3="Figs 19–27" captionStartId-0="2.[151,188,1539,1558]" captionStartId-1="3.[113,150,1501,1520]" captionStartId-2="4.[151,188,1518,1537]" captionStartId-3="5.[113,150,1514,1533]" captionTargetBox-0="[165,1121,215,1505]" captionTargetBox-1="[123,1087,231,1463]" captionTargetBox-2="[169,1107,217,1488]" captionTargetBox-3="[138,1079,217,1492]" captionTargetId-0="figure-10@2.[159,1125,208,1540]" captionTargetId-1="figure-14@3.[91,1097,215,1483]" captionTargetId-2="figure-10@4.[151,1119,211,1500]" captionTargetId-3="figure-14@5.[115,1088,193,1499]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Figs 1–6. Alepia bifida sp. nov., male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture." captionText-1="Figs 7–10. Alepia bifida sp. nov., female. 7–8 – last abdominal segments and genitalia (7 – dorsal view, 8 – lateral view); 9–10 – genital chamber and subgenital plate in detail (9 – lateral view, 10 – caudal view)." captionText-2="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." captionText-3="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi-0="http://doi.org/10.5281/zenodo.5317734" figureDoi-1="http://doi.org/10.5281/zenodo.5317736" figureDoi-2="http://doi.org/10.5281/zenodo.5317740" figureDoi-3="http://doi.org/10.5281/zenodo.5317744" httpUri-0="https://zenodo.org/record/5317734/files/figure.png" httpUri-1="https://zenodo.org/record/5317736/files/figure.png" httpUri-2="https://zenodo.org/record/5317740/files/figure.png" httpUri-3="https://zenodo.org/record/5317744/files/figure.png" pageId="1" pageNumber="766">Figs 1–27</figureCitation>
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)
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leg. (
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|
||
22614, dissected).
|
||
<typeStatus id="54B87BB43B3DFFE5FD7108F4FD36C6C2" box="[671,752,1420,1439]" pageId="1" pageNumber="766" type="allotype">ALLOTYPE</typeStatus>
|
||
:
|
||
<specimenCount id="9D050E9F3B3DFFE5FD1008F4FCE6C6C0" box="[766,800,1420,1439]" pageId="1" pageNumber="766" type="female">1 ♀</specimenCount>
|
||
, same data as holotype (
|
||
<collectionCode id="ED125DD33B3DFFE5FC1308F4FBF8C6C0" box="[1021,1086,1420,1439]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="766">NMPC</collectionCode>
|
||
: slides
|
||
<collectorName id="26F6A0C03B3DFFE5FF4908DFFF29C6E5" box="[167,239,1447,1466]" pageId="1" pageNumber="766">Cat. No.</collectorName>
|
||
34706,
|
||
<collectorName id="26F6A0C03B3DFFE5FEDC08DFFEBEC6E5" box="[306,376,1447,1466]" pageId="1" pageNumber="766">Inv. No.</collectorName>
|
||
22618, dissected).
|
||
<typeStatus id="54B87BB43B3DFFE5FDF508DFFDB5C6E7" box="[539,627,1447,1466]" pageId="1" pageNumber="766" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<specimenCount id="9D050E9F3B3DFFE5FD9308DFFD6AC6E5" box="[637,684,1447,1466]" pageId="1" pageNumber="766" type="male">4 ♂♂</specimenCount>
|
||
<specimenCount id="9D050E9F3B3DFFE5FD5F08DFFD17C6E5" box="[689,721,1447,1466]" pageId="1" pageNumber="766" type="female">1 ♀</specimenCount>
|
||
, same data as
|
||
<typeStatus id="54B87BB43B3DFFE5FCA408DFFC53C6E5" box="[842,917,1447,1466]" pageId="1" pageNumber="766" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED125DD33B3DFFE5FC4E08DFFC27C6E5" box="[928,993,1447,1466]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="766">NMPC</collectionCode>
|
||
: slides
|
||
<collectorName id="26F6A0C03B3DFFE5FBF008DFFF54C68B" pageId="1" pageNumber="766">Cat. No.</collectorName>
|
||
34703–34705, 34707,
|
||
<collectorName id="26F6A0C03B3DFFE5FEB808B9FE5BC68B" box="[342,413,1473,1492]" pageId="1" pageNumber="766">Inv. No.</collectorName>
|
||
22615–22617, 22619, mostly dissected; number 34705/22617 = slide with one shallow depression and two non-dissected specimens)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319969D3B3DFFE3FF9F0881FD65C10C" lastPageId="7" lastPageNumber="772" pageId="1" pageNumber="766" type="description">
|
||
<paragraph id="8BBCC5163B3DFFE5FF9F0881FB87C50F" blockId="1.[113,1090,1340,1616]" pageId="1" pageNumber="766">
|
||
<emphasis id="B97719043B3DFFE5FF9F0881FE85C54F" bold="true" box="[113,323,1528,1552]" pageId="1" pageNumber="766">
|
||
Description.
|
||
<emphasis id="B97719043B3DFFE5FEEB0880FE85C54F" bold="true" box="[261,323,1528,1552]" italics="true" pageId="1" pageNumber="766">Male.</emphasis>
|
||
</emphasis>
|
||
Eyes separated (
|
||
<figureCitation id="1338D9933B3DFFE5FE110881FD88C54F" box="[511,590,1529,1552]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="1" pageNumber="766">Fig. 11</figureCitation>
|
||
), frontal suture largely U-shaped with central triangular ligament with concave sides. Eye bridge formed by three facet rows, the number of facets in the apices (not pointed as in female) is more reduced. Number of antennomeres 16.
|
||
</paragraph>
|
||
<caption id="DF7C959E3B3EFFE6FF790B7BFDE3C514" ID-DOI="http://doi.org/10.5281/zenodo.5317734" ID-Zenodo-Dep="5317734" httpUri="https://zenodo.org/record/5317734/files/figure.png" pageId="2" pageNumber="767" startId="2.[151,188,1539,1558]" targetBox="[165,1121,215,1505]" targetPageId="2">
|
||
<paragraph id="8BBCC5163B3EFFE6FF790B7BFDE3C514" blockId="2.[151,1126,1539,1611]" pageId="2" pageNumber="767">
|
||
Figs 1–6.
|
||
<taxonomicName id="4C03BE953B3EFFE6FF040B7BFE90C549" authority="Tkoč & Ježek & Pont, 2017" authorityName="Tkoč & Ježek & Pont" authorityYear="2017" box="[234,342,1539,1558]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="767" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
|
||
<emphasis id="B97719043B3EFFE6FF040B7BFE90C549" box="[234,342,1539,1558]" italics="true" pageId="2" pageNumber="767">Alepia bifida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244A47F3B3EFFE6FEB50B7BFE58C549" box="[347,414,1539,1558]" pageId="2" pageNumber="767" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7C959E3B3FFFE7FF9F08A5FC6FC554" ID-DOI="http://doi.org/10.5281/zenodo.5317736" ID-Zenodo-Dep="5317736" httpUri="https://zenodo.org/record/5317736/files/figure.png" pageId="3" pageNumber="768" startId="3.[113,150,1501,1520]" targetBox="[123,1087,231,1463]" targetPageId="3">
|
||
<paragraph id="8BBCC5163B3FFFE7FF9F08A5FC6FC554" blockId="3.[113,1088,1501,1547]" pageId="3" pageNumber="768">
|
||
Figs 7–10.
|
||
<taxonomicName id="4C03BE953B3FFFE7FF3F08A5FEF9C6AF" authority="Tkoč & Ježek & Pont, 2017" authorityName="Tkoč & Ježek & Pont" authorityYear="2017" box="[209,319,1501,1520]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="768" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
|
||
<emphasis id="B97719043B3FFFE7FF3F08A5FEF9C6AF" box="[209,319,1501,1520]" italics="true" pageId="3" pageNumber="768">Alepia bifida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244A47F3B3FFFE7FEAB08A5FE4CC6AF" box="[325,394,1501,1520]" pageId="3" pageNumber="768" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, female. 7–8 – last abdominal segments and genitalia (7 – dorsal view, 8 – lateral view); 9–10 – genital chamber and subgenital plate in detail (9 – lateral view, 10 – caudal view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7C959E3B38FFE0FF790896FEF2C50E" ID-DOI="http://doi.org/10.5281/zenodo.5317740" ID-Zenodo-Dep="5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="4" pageNumber="769" startId="4.[151,188,1518,1537]" targetBox="[169,1107,217,1488]" targetPageId="4">
|
||
<paragraph id="8BBCC5163B38FFE0FF790896FEF2C50E" blockId="4.[151,1127,1518,1617]" pageId="4" pageNumber="769">
|
||
Figs 11–18.
|
||
<taxonomicName id="4C03BE953B38FFE0FEED0896FEB4C55E" authority="Tkoč & Ježek & Pont, 2017" authorityName="Tkoč & Ježek & Pont" authorityYear="2017" box="[259,370,1518,1537]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="769" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
|
||
<emphasis id="B97719043B38FFE0FEED0896FEB4C55E" box="[259,370,1518,1537]" italics="true" pageId="4" pageNumber="769">Alepia bifida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244A47F3B38FFE0FE970896FE79C55E" box="[377,447,1518,1537]" pageId="4" pageNumber="769" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7C959E3B39FFE1FF9F0892FED2C512" ID-DOI="http://doi.org/10.5281/zenodo.5317744" ID-Zenodo-Dep="5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="5" pageNumber="770" startId="5.[113,150,1514,1533]" targetBox="[138,1079,217,1492]" targetPageId="5">
|
||
<paragraph id="8BBCC5163B39FFE1FF9F0892FED2C512" blockId="5.[113,1089,1514,1613]" pageId="5" pageNumber="770">
|
||
Figs 19–27.
|
||
<taxonomicName id="4C03BE953B39FFE1FF320892FE8CC6A2" authority="Tkoč & Ježek & Pont, 2017" authorityName="Tkoč & Ježek & Pont" authorityYear="2017" box="[220,330,1514,1533]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="770" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
|
||
<emphasis id="B97719043B39FFE1FF320892FE8CC6A2" box="[220,330,1514,1533]" italics="true" pageId="5" pageNumber="770">Alepia bifida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244A47F3B39FFE1FEBE0892FE53C6A2" box="[336,405,1514,1533]" pageId="5" pageNumber="770" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P
|
||
<subScript id="1787C7533B39FFE1FF2D0B6AFF0FC542" attach="left" box="[195,201,1554,1565]" fontSize="5" pageId="5" pageNumber="770">1</subScript>
|
||
, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBCC5163B3AFFE2FF790DA4FE5BC66C" blockId="6.[151,1127,220,1587]" pageId="6" pageNumber="771">
|
||
Scape 2.4 times as long as pedicel, cylindrical, narrower basally, pedicel almost globular, pressed in longitudinal axis (
|
||
<figureCitation id="1338D9933B3AFFE2FE260D84FDD1C24C" box="[456,535,252,275]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">Fig. 12</figureCitation>
|
||
). Flagellomeres bowl shaped, apical ones missing. Length ratio of maxillary palpomeres 1.0: 1.9: 2.2: 2.2, palpomere 4 not annulate (
|
||
<figureCitation id="1338D9933B3AFFE2FBE70C64FB9FC26C" box="[1033,1113,284,307]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">Fig. 15</figureCitation>
|
||
). Terminal lobes of labium, cibarium and epipharynx as in
|
||
<figureCitation id="1338D9933B3AFFE2FD1A0C44FCF3C20C" box="[756,821,316,339]" captionStart="Figs 1–6" captionStartId="2.[151,188,1539,1558]" captionTargetBox="[165,1121,215,1505]" captionTargetId="figure-10@2.[159,1125,208,1540]" captionTargetPageId="2" captionText="Figs 1–6. Alepia bifida sp. nov., male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture." figureDoi="http://doi.org/10.5281/zenodo.5317734" httpUri="https://zenodo.org/record/5317734/files/figure.png" pageId="6" pageNumber="771">Figs 2</figureCitation>
|
||
and
|
||
<figureCitation id="1338D9933B3AFFE2FC860C44FC40C20C" box="[872,902,316,339]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">16</figureCitation>
|
||
. Thoracic sclerites as in
|
||
<figureCitation id="1338D9933B3AFFE2FF5C0C24FF39C22C" box="[178,255,348,371]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">Fig. 19</figureCitation>
|
||
. Wings sharply pointed, maculated, spots on veins as well as wing membrane very dark and conspicuous (
|
||
<figureCitation id="1338D9933B3AFFE2FE670C04FE0EC2CC" box="[393,456,380,403]" captionStart="Figs 1–6" captionStartId="2.[151,188,1539,1558]" captionTargetBox="[165,1121,215,1505]" captionTargetId="figure-10@2.[159,1125,208,1540]" captionTargetPageId="2" captionText="Figs 1–6. Alepia bifida sp. nov., male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture." figureDoi="http://doi.org/10.5281/zenodo.5317734" httpUri="https://zenodo.org/record/5317734/files/figure.png" pageId="6" pageNumber="771">Fig. 3</figureCitation>
|
||
),
|
||
<quantity id="4CFB68F33B3AFFE2FE320C04FD9DC2CC" box="[476,603,380,403]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="6" pageNumber="771" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.1–2.5 mm</quantity>
|
||
long in
|
||
<typeStatus id="54B87BB43B3AFFE2FD5E0C04FCCBC2CC" box="[688,781,380,403]" pageId="6" pageNumber="771" type="holotype">holotype</typeStatus>
|
||
and male
|
||
<typeStatus id="54B87BB43B3AFFE2FC960C04FC24C2CC" box="[888,994,380,403]" pageId="6" pageNumber="771" type="paratype">paratypes</typeStatus>
|
||
. Completely strengthened veins: Sc, R 1, CuA 1, CuA 2 strenghtened basally. Radial, medial and cubital veins black spotted at the end, as well as radial and medial forks, basal cell and basis of M 3 and CuA 1. Basal costal nodes well visible, Sc uninterrupted, M 3, CuA 1 and CuA 2 not touching at basis of wing. R 5 extends in the apex of the wing. Radial and medial forks complete, cross veins missing. The wing margin reaching CuA 2 is a little distad of medial fork, medial fork conspicuously distad of radial fork. Medial wing angle 222° (BCD). Wing indices: AB: AC: AD = 3.7: 3.6: 4.5; BC: CD: BD = 1.0: 1.0: 1.9; maximum wing length equal to 2.7 times its maximum width. Halteres clubbed (
|
||
<figureCitation id="1338D9933B3AFFE2FDD90F04FD41C1CC" box="[567,647,636,659]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">Fig. 20</figureCitation>
|
||
), with a maximum length equal hardly to 4.1 times its maximum width. Ratios of lengths of femora, tibiae and first tarsomeres: P 1 = 1.8: 1.7: 1.0; P 2 = 2.0: 2.4: 1.2; P 3 = 2.0: 2.7: 1.3; fore claws bent and pointed distad (
|
||
<figureCitation id="1338D9933B3AFFE2FBE50FC4FB9CC18C" box="[1035,1114,700,723]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">Fig. 21</figureCitation>
|
||
). Aedeagal complex as in
|
||
<figureCitation id="1338D9933B3AFFE2FE720FA4FDD0C1AC" box="[412,534,732,755]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">Figs 17, 18</figureCitation>
|
||
,
|
||
<figureCitation id="1338D9933B3AFFE2FDCE0FA4FDFBC1AC" box="[544,573,732,755]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">22</figureCitation>
|
||
, basiphallus almost straight and narrow from lateral view, however, with a U-shaped loop proximally before apex (
|
||
<figureCitation id="1338D9933B3AFFE2FCC70F84FCBDC04C" box="[809,891,764,787]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">Figs 18</figureCitation>
|
||
,
|
||
<figureCitation id="1338D9933B3AFFE2FC6B0F84FC65C04C" box="[901,931,764,787]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">22</figureCitation>
|
||
). Basiphallus very widened and pressed distally, S-shaped, with small blunt teeth, backward oriented. Distiphallus prolonged by bent bifid arm, spanner- or bottle-opener shaped (
|
||
<figureCitation id="1338D9933B3AFFE2FCBF0E44FC61C00C" box="[849,935,828,851]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">Figs 18</figureCitation>
|
||
,
|
||
<figureCitation id="1338D9933B3AFFE2FC5B0E44FC15C00C" box="[949,979,828,851]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">22</figureCitation>
|
||
), from which one protuberance is blunt in contrast to very pointed hooked tip. Aedeagal complex overlaid basally by a large hyaline bell-shaped tunica (hypandrium), distinctly narrowed in the middle (lateral view), quite bare (
|
||
<figureCitation id="1338D9933B3AFFE2FE5C0EE4FDF7C0EC" box="[434,561,924,947]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">Figs 17, 18</figureCitation>
|
||
,
|
||
<figureCitation id="1338D9933B3AFFE2FDD30EE4FD9DC0EC" box="[573,603,924,947]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">22</figureCitation>
|
||
). Tunica is articulated proximally to basiphallus and protuberances of gonocoxites. Gonocoxites almost cylindrical, only slightly expanded medially, 1.3 times shorter than gonostyli (
|
||
<figureCitation id="1338D9933B3AFFE2FDB00EA4FD1FC0AC" box="[606,729,988,1011]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">Figs 17, 18</figureCitation>
|
||
,
|
||
<figureCitation id="1338D9933B3AFFE2FD0A0EA4FCC7C0AC" box="[740,769,988,1011]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">23</figureCitation>
|
||
), both parts with setae. Gonostyli hardly S-shaped in dorsal view, sticky, sickle-shaped from lateral view, subapically inconspicuously inflated and tapered to a narrow, blunt tip. Epandrium bare, with an inconspicuous operculum in dorsal view (
|
||
<figureCitation id="1338D9933B3AFFE2FE5B0944FDC3C70C" box="[437,517,1084,1107]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">Fig. 24</figureCitation>
|
||
), well visible from lateral view (
|
||
<figureCitation id="1338D9933B3AFFE2FC8E0944FC76C70C" box="[864,944,1084,1107]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">Fig. 25</figureCitation>
|
||
). Ventral plate of epandrium reduced to only two sclerotized converging ribs jointed with surstyli (
|
||
<figureCitation id="1338D9933B3AFFE2FBE70924FB9CC72C" box="[1033,1114,1116,1139]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">Fig. 24</figureCitation>
|
||
). Hypandrium hyaline, lateral margin of 9th sternite grows together or connected with proximal border of epandrium and proximal protuberance of gonocoxite. Epiproct transversal as a fold, hardly visible, hypoproct shortly tongue-shaped, both parts with microsetae (
|
||
<figureCitation id="1338D9933B3AFFE2FC5809C4FBC5C78C" box="[950,1027,1212,1235]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">Fig. 24</figureCitation>
|
||
). Surstyli almost circular basally, gradually tapering caudally (
|
||
<figureCitation id="1338D9933B3AFFE2FD2609A4FC82C7AC" box="[712,836,1244,1267]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">Figs 24, 25</figureCitation>
|
||
), with numerous accessory tenacula in a dark elliptical area near the base of surstyli (
|
||
<figureCitation id="1338D9933B3AFFE2FD120984FCB1C64C" box="[764,887,1276,1299]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">Figs 24–27</figureCitation>
|
||
), subapically with one tenaculum (
|
||
<figureCitation id="1338D9933B3AFFE2FEFA0864FE56C66C" box="[276,400,1308,1331]" captionStart="Figs 19–27" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[138,1079,217,1492]" captionTargetId="figure-14@5.[115,1088,193,1499]" captionTargetPageId="5" captionText="Figs 19–27. Alepia bifida sp. nov., male. 19 – thoracic sclerites, lateral view; 20 – haltere, lateral view; 21 – tarsal claw of P1, lateral view; 22 – aedeagal complex, lateral view; 23 – gonopod, lateral view; 24–25 – epandrium and surstyli (24 – dorsal view, 25 – lateral view); 26–27 – hooded tip of accessory tenaculum in detail (26 – dorsal view, 27 – ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5317744" httpUri="https://zenodo.org/record/5317744/files/figure.png" pageId="6" pageNumber="771">Figs 24, 25</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBCC5163B3AFFE3FF5B0843FD8DC28C" blockId="6.[151,1127,220,1587]" lastBlockId="7.[113,1089,220,723]" lastPageId="7" lastPageNumber="772" pageId="6" pageNumber="771">
|
||
<emphasis id="B97719043B3AFFE2FF5B0843FEC9C60C" bold="true" box="[181,271,1339,1363]" italics="true" pageId="6" pageNumber="771">Female.</emphasis>
|
||
Eyes separated (
|
||
<figureCitation id="1338D9933B3AFFE2FE2A0844FDC4C60C" box="[452,514,1340,1363]" captionStart="Figs 1–6" captionStartId="2.[151,188,1539,1558]" captionTargetBox="[165,1121,215,1505]" captionTargetId="figure-10@2.[159,1125,208,1540]" captionTargetPageId="2" captionText="Figs 1–6. Alepia bifida sp. nov., male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture." figureDoi="http://doi.org/10.5281/zenodo.5317734" httpUri="https://zenodo.org/record/5317734/files/figure.png" pageId="6" pageNumber="771">Fig. 1</figureCitation>
|
||
), frontal suture triangular, frons bare, frontoclypeus with insertions of setae arranged in two oval areas touched on the level of antennal bows and divergent near tentorial pits. The minimum distance between eyes corresponds approximately to the diameter of two facets. Eye bridge formed by three facet rows, the number of facets in the pointed apices being more reduced. Vertex rounded, occipital lobe small, with a shallow indentation apically and a sclerotized rim. Scape twice as long as pedicel, cylindrical, gradually narrower basally, pedicel globular (
|
||
<figureCitation id="1338D9933B3AFFE2FDE30884FD9DC54C" box="[525,603,1532,1555]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">Fig. 13</figureCitation>
|
||
). Flagellomeres spindle-shaped, narrow (
|
||
<figureCitation id="1338D9933B3AFFE2FBFE0884FF72C56C" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">Figs 13, 14</figureCitation>
|
||
), apical ones missing. Ascoids simple, paired, needle-shaped (
|
||
<figureCitation id="1338D9933B3AFFE2FCA40B64FC5FC56C" box="[842,921,1564,1587]" captionStart="Figs 11–18" captionStartId="4.[151,188,1518,1537]" captionTargetBox="[169,1107,217,1488]" captionTargetId="figure-10@4.[151,1119,211,1500]" captionTargetPageId="4" captionText="Figs 11–18. Alepia bifida sp. nov., male and female. 11 – detail of frons, male; 12–13 – basal antennomeres (12 – male, 13 – female); 14 – medial antennomeres, female; 15 – maxilla and palpus maxillaris, male; 16 – labium, labial lobes and epipharynx, dorsal view, male; 17–18 – aedeagal complex and gonopods, male (17 – dorsal view; 18 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5317740" httpUri="https://zenodo.org/record/5317740/files/figure.png" pageId="6" pageNumber="771">Fig. 14</figureCitation>
|
||
). Terminal lobes of labium, cibarium and epipharynx as in
|
||
<figureCitation id="1338D9933B3BFFE3FDF60DA4FD9EC3AC" box="[536,600,220,243]" captionStart="Figs 1–6" captionStartId="2.[151,188,1539,1558]" captionTargetBox="[165,1121,215,1505]" captionTargetId="figure-10@2.[159,1125,208,1540]" captionTargetPageId="2" captionText="Figs 1–6. Alepia bifida sp. nov., male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture." figureDoi="http://doi.org/10.5281/zenodo.5317734" httpUri="https://zenodo.org/record/5317734/files/figure.png" pageId="7" pageNumber="772">Fig. 1</figureCitation>
|
||
. Wing shape and maculation similar to male, length
|
||
<quantity id="4CFB68F33B3BFFE3FF520D84FEFBC24C" box="[188,317,252,275]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="7" pageNumber="772" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
|
||
(
|
||
<typeStatus id="54B87BB43B3BFFE3FEA30D84FE63C24C" box="[333,421,252,275]" pageId="7" pageNumber="772" type="allotype">allotype</typeStatus>
|
||
and female
|
||
<typeStatus id="54B87BB43B3BFFE3FDC20D84FD4BC24C" box="[556,653,252,275]" pageId="7" pageNumber="772" type="paratype">paratype</typeStatus>
|
||
). Ratios of lengths of femora, tibiae and first tarsomeres: P 1 = 2.0: 1.8: 1.0; P 2 = 2.2: 2.6: 1.3; P 3 = 2.3: 3.1: 1.4. Genitalia as figured (
|
||
<figureCitation id="1338D9933B3BFFE3FF970C44FF1FC20C" box="[121,217,316,339]" captionStart="Figs 1–6" captionStartId="2.[151,188,1539,1558]" captionTargetBox="[165,1121,215,1505]" captionTargetId="figure-10@2.[159,1125,208,1540]" captionTargetPageId="2" captionText="Figs 1–6. Alepia bifida sp. nov., male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture." figureDoi="http://doi.org/10.5281/zenodo.5317734" httpUri="https://zenodo.org/record/5317734/files/figure.png" pageId="7" pageNumber="772">Figs 4, 5</figureCitation>
|
||
,
|
||
<figureCitation id="1338D9933B3BFFE3FF0A0C44FED8C20C" box="[228,286,316,339]" captionStart="Figs 7–10" captionStartId="3.[113,150,1501,1520]" captionTargetBox="[123,1087,231,1463]" captionTargetId="figure-14@3.[91,1097,215,1483]" captionTargetPageId="3" captionText="Figs 7–10. Alepia bifida sp. nov., female. 7–8 – last abdominal segments and genitalia (7 – dorsal view, 8 – lateral view); 9–10 – genital chamber and subgenital plate in detail (9 – lateral view, 10 – caudal view)." figureDoi="http://doi.org/10.5281/zenodo.5317736" httpUri="https://zenodo.org/record/5317736/files/figure.png" pageId="7" pageNumber="772">7–10</figureCitation>
|
||
). Subgenital plate (
|
||
<figureCitation id="1338D9933B3BFFE3FE050C44FDF6C20C" box="[491,560,316,339]" captionStart="Figs 1–6" captionStartId="2.[151,188,1539,1558]" captionTargetBox="[165,1121,215,1505]" captionTargetId="figure-10@2.[159,1125,208,1540]" captionTargetPageId="2" captionText="Figs 1–6. Alepia bifida sp. nov., male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture." figureDoi="http://doi.org/10.5281/zenodo.5317734" httpUri="https://zenodo.org/record/5317734/files/figure.png" pageId="7" pageNumber="772">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="1338D9933B3BFFE3FDD20C44FDA3C20C" box="[572,613,316,339]" captionStart="Figs 7–10" captionStartId="3.[113,150,1501,1520]" captionTargetBox="[123,1087,231,1463]" captionTargetId="figure-14@3.[91,1097,215,1483]" captionTargetPageId="3" captionText="Figs 7–10. Alepia bifida sp. nov., female. 7–8 – last abdominal segments and genitalia (7 – dorsal view, 8 – lateral view); 9–10 – genital chamber and subgenital plate in detail (9 – lateral view, 10 – caudal view)." figureDoi="http://doi.org/10.5281/zenodo.5317736" httpUri="https://zenodo.org/record/5317736/files/figure.png" pageId="7" pageNumber="772">7–9</figureCitation>
|
||
) bilobed, with a shallow concavity caudally, with densely spaced microtrichia and sparsely covered by long setae. Genital chamber (
|
||
<figureCitation id="1338D9933B3BFFE3FBFF0C24FF6EC2CC" captionStart="Figs 7–10" captionStartId="3.[113,150,1501,1520]" captionTargetBox="[123,1087,231,1463]" captionTargetId="figure-14@3.[91,1097,215,1483]" captionTargetPageId="3" captionText="Figs 7–10. Alepia bifida sp. nov., female. 7–8 – last abdominal segments and genitalia (7 – dorsal view, 8 – lateral view); 9–10 – genital chamber and subgenital plate in detail (9 – lateral view, 10 – caudal view)." figureDoi="http://doi.org/10.5281/zenodo.5317736" httpUri="https://zenodo.org/record/5317736/files/figure.png" pageId="7" pageNumber="772">Figs 7–10</figureCitation>
|
||
) with simple structures, without net- or wart-like decoration. Cerci almost triangular and slightly bent, three times longer than its bases (
|
||
<figureCitation id="1338D9933B3BFFE3FD710CE4FD23C2EC" box="[671,741,412,435]" captionStart="Figs 1–6" captionStartId="2.[151,188,1539,1558]" captionTargetBox="[165,1121,215,1505]" captionTargetId="figure-10@2.[159,1125,208,1540]" captionTargetPageId="2" captionText="Figs 1–6. Alepia bifida sp. nov., male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture." figureDoi="http://doi.org/10.5281/zenodo.5317734" httpUri="https://zenodo.org/record/5317734/files/figure.png" pageId="7" pageNumber="772">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="1338D9933B3BFFE3FD1C0CE4FCDAC2EC" box="[754,796,412,435]" captionStart="Figs 7–10" captionStartId="3.[113,150,1501,1520]" captionTargetBox="[123,1087,231,1463]" captionTargetId="figure-14@3.[91,1097,215,1483]" captionTargetPageId="3" captionText="Figs 7–10. Alepia bifida sp. nov., female. 7–8 – last abdominal segments and genitalia (7 – dorsal view, 8 – lateral view); 9–10 – genital chamber and subgenital plate in detail (9 – lateral view, 10 – caudal view)." figureDoi="http://doi.org/10.5281/zenodo.5317736" httpUri="https://zenodo.org/record/5317736/files/figure.png" pageId="7" pageNumber="772">7, 8</figureCitation>
|
||
), rounded caudally, setose, connected by a wrinkled membrane (
|
||
<figureCitation id="1338D9933B3BFFE3FE130CC4FDFBC28C" box="[509,573,444,467]" captionStart="Figs 1–6" captionStartId="2.[151,188,1539,1558]" captionTargetBox="[165,1121,215,1505]" captionTargetId="figure-10@2.[159,1125,208,1540]" captionTargetPageId="2" captionText="Figs 1–6. Alepia bifida sp. nov., male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture." figureDoi="http://doi.org/10.5281/zenodo.5317734" httpUri="https://zenodo.org/record/5317734/files/figure.png" pageId="7" pageNumber="772">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBCC5163B3BFFE3FF7E0CA4FD65C10C" blockId="7.[113,1089,220,723]" pageId="7" pageNumber="772">
|
||
<emphasis id="B97719043B3BFFE3FF7E0CA4FF07C2AC" bold="true" box="[144,193,476,499]" italics="true" pageId="7" pageNumber="772">Egg.</emphasis>
|
||
Three times as long asmaximum width, with characteristic wrinkled structures (
|
||
<figureCitation id="1338D9933B3BFFE3FC330CA4FBDBC2AC" box="[989,1053,476,499]" captionStart="Figs 1–6" captionStartId="2.[151,188,1539,1558]" captionTargetBox="[165,1121,215,1505]" captionTargetId="figure-10@2.[159,1125,208,1540]" captionTargetPageId="2" captionText="Figs 1–6. Alepia bifida sp. nov., male and female.1 – head, frontal view, female; 2 – cibarium, epipharynx and labial lobes, dorsolateral view, male; 3 – wing, male; 4 – subgenital plate, dorso-subventral view, female; 5 – cerci, dorsal view, female; 6 – egg, with detail of sculpture." figureDoi="http://doi.org/10.5281/zenodo.5317734" httpUri="https://zenodo.org/record/5317734/files/figure.png" pageId="7" pageNumber="772">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="1338D9933B3BFFE3FBCB0CA4FBF2C2AC" box="[1061,1076,476,499]" captionStart="Figs 7–10" captionStartId="3.[113,150,1501,1520]" captionTargetBox="[123,1087,231,1463]" captionTargetId="figure-14@3.[91,1097,215,1483]" captionTargetPageId="3" captionText="Figs 7–10. Alepia bifida sp. nov., female. 7–8 – last abdominal segments and genitalia (7 – dorsal view, 8 – lateral view); 9–10 – genital chamber and subgenital plate in detail (9 – lateral view, 10 – caudal view)." figureDoi="http://doi.org/10.5281/zenodo.5317736" httpUri="https://zenodo.org/record/5317736/files/figure.png" pageId="7" pageNumber="772">7</figureCitation>
|
||
).
|
||
<emphasis id="B97719043B3BFFE3FF9F0C84FEACC14C" bold="true" box="[113,362,508,531]" pageId="7" pageNumber="772">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C03BE953B3BFFE3FE810C84FE30C14C" authority="Tkoč & Ježek & Pont, 2017" authorityName="Tkoč & Ježek & Pont" authorityYear="2017" box="[367,502,508,532]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="772" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
|
||
<emphasis id="B97719043B3BFFE3FE810C84FE30C14C" box="[367,502,508,532]" italics="true" pageId="7" pageNumber="772">Alepia bifida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244A47F3B3BFFE3FE150C84FD8AC14C" box="[507,588,508,531]" pageId="7" pageNumber="772" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is morphologically similar to
|
||
<taxonomicName id="4C03BE953B3BFFE3FC680C84FE90C16C" authority="Bravo, Lago & Castro, 2004" authorityName="Bravo, Lago & Castro" authorityYear="2004" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="772" phylum="Arthropoda" rank="species" species="recurva">
|
||
<emphasis id="B97719043B3BFFE3FC680C84FC32C14C" box="[902,1012,508,531]" italics="true" pageId="7" pageNumber="772">A. recurva</emphasis>
|
||
<bibRefCitation id="EF92B8E73B3BFFE3FC170C84FE90C16C" author="BRAVO F. & LAGO A. & CASTRO I." pageId="7" pageNumber="772" pagination="589 - 599" refId="ref3454" refString="BRAVO F., LAGO A. & CASTRO I. 2004: Dez especies novas de Alepia Enderlein (Diptera: Psychodidae) do Brasil. Neotropical Entomology 33: 589 - 599." type="journal article" year="2004">Bravo, Lago & Castro, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F31485863B3BFFE3FE740F64FE19C16C" box="[410,479,540,563]" name="Brazil" pageId="7" pageNumber="772">Brazil</collectingCountry>
|
||
. However, wing characters, basiphallus, distiphallus and gonostyli show distinct differences (see the
|
||
<tableCitation id="C681F0AD3B3BFFE3FDAA0F44FD55C10C" box="[580,659,572,595]" captionStart="Table 1" captionStartId="7.[113,160,777,796]" captionText="Table 1. Comparison of diagnostic characters of males of Alepia bifida sp. nov. and A. recurva Bravo, Lago & Castro, 2004. For figures of A. recurva see BRAVO et al. (2004)." pageId="7" pageNumber="772">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319969D3B3BFFE3FF9F0F24FB87C12C" box="[113,1089,604,628]" pageId="7" pageNumber="772" type="etymology">
|
||
<paragraph id="8BBCC5163B3BFFE3FF9F0F24FB87C12C" blockId="7.[113,1089,220,723]" box="[113,1089,604,628]" pageId="7" pageNumber="772">
|
||
<emphasis id="B97719043B3BFFE3FF9F0F24FF37C12C" bold="true" box="[113,241,604,627]" pageId="7" pageNumber="772">Etymology.</emphasis>
|
||
Named after the shape of the bifurcated distiphallus (Lat.
|
||
<emphasis id="B97719043B3BFFE3FCA50F24FC57C12C" box="[843,913,604,628]" italics="true" pageId="7" pageNumber="772">bifidus</emphasis>
|
||
,
|
||
<emphasis id="B97719043B3BFFE3FC730F24FC75C12C" box="[925,947,604,627]" italics="true" pageId="7" pageNumber="772">-a</emphasis>
|
||
,
|
||
<emphasis id="B97719043B3BFFE3FC500F24FC21C12C" box="[958,999,604,627]" italics="true" pageId="7" pageNumber="772">-um</emphasis>
|
||
= bifid).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319969D3B3BFFE3FF9F0F04FF2FC1EC" pageId="7" pageNumber="772" type="biology_ecology">
|
||
<paragraph id="8BBCC5163B3BFFE3FF9F0F04FF2FC1EC" blockId="7.[113,1089,220,723]" pageId="7" pageNumber="772">
|
||
<emphasis id="B97719043B3BFFE3FF9F0F04FF0BC1CC" bold="true" box="[113,205,636,659]" pageId="7" pageNumber="772">Biology.</emphasis>
|
||
Unknown. Adults were collected by light trap in the forest on the slopes of Cerro Musún Mt.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319969D3B3BFFE3FF9F0FC4FD06C18C" box="[113,704,700,723]" pageId="7" pageNumber="772" type="distribution">
|
||
<paragraph id="8BBCC5163B3BFFE3FF9F0FC4FD06C18C" blockId="7.[113,1089,220,723]" box="[113,704,700,723]" pageId="7" pageNumber="772">
|
||
<emphasis id="B97719043B3BFFE3FF9F0FC4FEC2C18C" bold="true" box="[113,260,700,723]" pageId="7" pageNumber="772">Distribution.</emphasis>
|
||
Currently recorded only from
|
||
<collectingCountry id="F31485863B3BFFE3FDA50FC4FD7AC18C" box="[587,700,700,723]" name="Nicaragua" pageId="7" pageNumber="772">Nicaragua</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |