138 lines
20 KiB
XML
138 lines
20 KiB
XML
<document id="FC288C9FDB481EC9C6AF46FC8D549E95" ID-CLB-Dataset="3091" ID-DOI="10.1093/zoolinnean/zlaa180" ID-GBIF-Dataset="90f6f9e0-4cdd-4e1b-82b0-5c976e702317" ID-ISSN="0024-4082" ID-Zenodo-Dep="5752961" ID-ZooBank="529D72D0-2863-41BC-AEB0-D421B716CFFD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638521496927" checkinUser="plazi" docAuthor="Szawaryn, Karol" docDate="2021" docId="03EDD1467E48FFD5FC44FA9BA02D3CE7" docLanguage="en" docName="zlaa180.pdf" docOrigin="Zoological Journal of the Linnean Society 193" docSource="https://doi.org/10.1093/zoolinnean/zlaa180" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Baltosidis damzeni Szawaryn 2021, SP. NOV." docType="treatment" docUuid="58C958C5-6656-4749-94C1-69E7666B58EC" docUuidSource="ZooBank" docVersion="10" lastPageNumber="1303" masterDocId="FFD4A93E7E4EFFDCFFFEFF95A2273D15" masterDocTitle="The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles" masterLastPageNumber="1309" masterPageNumber="1294" pageNumber="1300" updateTime="1699471844068" updateUser="ExternalLinkService">
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<mods:title id="C1854D46E6484526745C32BCA558A0AE">The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles</mods:title>
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<mods:namePart id="F6E502214723351B914D701CBEB52FB7">Szawaryn, Karol</mods:namePart>
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<mods:title id="7FBBD2AADE3A585F4F1432B19F22E802">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="D17F8BC0FD2353F2B9C78F52B1459355">2021</mods:date>
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<mods:number id="DE7E9D397478023A9FBC5952D1C790CD">2021-01-21</mods:number>
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<mods:number id="4DC5768FC25D2209265CA2607A3539D8">193</mods:number>
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<mods:url id="30ACE6DAAE257911868D5856A14222AE">https://doi.org/10.1093/zoolinnean/zlaa180</mods:url>
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<treatment id="03EDD1467E48FFD5FC44FA9BA02D3CE7" ID-DOI="http://doi.org/10.5281/zenodo.5753016" ID-GBIF-Taxon="191189738" ID-Zenodo-Dep="5753016" LSID="urn:lsid:zoobank.org:act:58C958C5-6656-4749-94C1-69E7666B58EC" httpUri="http://treatment.plazi.org/id/03EDD1467E48FFD5FC44FA9BA02D3CE7" lastPageId="9" lastPageNumber="1303" pageId="6" pageNumber="1300">
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<subSubSection id="C35E33DB7E48FFDAFC44FA9BA7043833" box="[954,1315,1294,1318]" pageId="6" pageNumber="1300" type="nomenclature">
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<paragraph id="8BFB60507E48FFDAFC44FA9BA7043833" blockId="6.[954,1315,1294,1318]" box="[954,1315,1294,1318]" pageId="6" pageNumber="1300">
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<heading id="D0B3D73C7E48FFDAFC44FA9BA7043833" box="[954,1315,1294,1318]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="1300" reason="2">
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<taxonomicName id="4C441BD37E48FFDAFC44FA9BA69D3830" authority="Szawaryn, 2021" authorityName="Szawaryn" authorityYear="2021" box="[954,1210,1294,1317]" class="Insecta" family="Coccinellidae" genus="Baltosidis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1298" phylum="Arthropoda" rank="genus" status="gen. nov.">BALTOSIDIS DAMZENI</taxonomicName>
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<emphasis id="B930BC427E48FFDAFB3EFA84A7043833" bold="true" box="[1216,1315,1294,1318]" pageId="6" pageNumber="1300">
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<taxonomicNameLabel id="A20301397E48FFDAFB3EFA84A7043833" box="[1216,1315,1294,1318]" pageId="6" pageNumber="1300" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35E33DB7E48FFD5FBD4FAAFA02D3CE7" lastPageId="9" lastPageNumber="1303" pageId="6" pageNumber="1300" type="description">
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<paragraph id="8BFB60507E48FFDAFBD4FAAFA6943846" blockId="6.[1066,1203,1338,1363]" box="[1066,1203,1338,1363]" pageId="6" pageNumber="1300">
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(
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<figureCitation id="137F7CD57E48FFDAFBCDFAAFA68C3846" box="[1075,1195,1338,1363]" captionStart="Figure 3" captionStartId="8.[164,244,1520,1542]" captionTargetBox="[323,1283,195,1481]" captionTargetId="figure-154@8.[323,1283,195,1481]" captionTargetPageId="8" captionText="Figure 3. Baltosidis damzeni. A, habitus ventral. B, head, prosternum, mesoventrite. C, pronotum. D, tip of elytra with sutural lines. E, habitus lateral. F, details of maxillary palp. G, postcoxal lines on ventrite 1. Holotype, MAIG 5988 (A–E); paratype AWL 154 (F); paratype, MAIG 5967 (G)." figureDoi="http://doi.org/10.5281/zenodo.5752969" httpUri="https://zenodo.org/record/5752969/files/figure.png" pageId="6" pageNumber="1300">FIG. 3A–G</figureCitation>
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)
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</paragraph>
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<paragraph id="8BFB60507E48FFDAFCC5FAF6A7B23882" blockId="6.[827,1443,1378,1431]" pageId="6" pageNumber="1300">
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<emphasis id="B930BC427E48FFDAFCC5FAF6A6A0386D" box="[827,1159,1378,1400]" italics="true" pageId="6" pageNumber="1300">Z o o b a n k r e g i s t r a t i o n:</emphasis>
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u r n: l s i d: z o o b a n k. org:act:
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<uuid id="FFE25A857E48FFDAFC77FA14A7B23882" box="[905,1429,1409,1431]" pageId="6" pageNumber="1300">A2F18452-6E52-41FB-86BB-3FB99A29D35B</uuid>
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</paragraph>
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<paragraph id="8BFB60507E48FFDAFCC5FA2BA1C33BFD" blockId="6.[827,1443,1470,1768]" pageId="6" pageNumber="1300">
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<emphasis id="B930BC427E48FFDAFCC5FA2BA61138C6" box="[827,1078,1470,1491]" italics="true" pageId="6" pageNumber="1300">Material examined:</emphasis>
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<materialsCitation id="3B2C6A0D7E48FFDAFBAFFA2BA6203B07" ID-GBIF-Occurrence="3415081304" pageId="6" pageNumber="1300" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus id="54FFDEF27E48FFDAFBAFFA2BA6E138C1" box="[1105,1222,1470,1492]" pageId="6" pageNumber="1300" type="holotype">Holotype</typeStatus>
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. MAIG no. 5988, donated by Jonas Damzen, FT-IR 9694 (
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<figureCitation id="137F7CD57E48FFDAFB00FA48A71A38E6" box="[1278,1341,1501,1523]" captionStart="Figure 5" captionStartId="10.[163,244,898,920]" captionTargetBox="[164,784,196,858]" captionTargetId="figure-427@10.[163,784,195,859]" captionTargetPageId="10" captionText="Figure 5. FT-IR absorbance spectra of analysed amber pieces with the indication of museum inventory numbers above each spectrum." figureDoi="http://doi.org/10.5281/zenodo.5752974" httpUri="https://zenodo.org/record/5752974/files/figure.png" pageId="6" pageNumber="1300">Fig. 5</figureCitation>
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),
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<specimenCount id="9D42ABD97E48FFDAFAAFFA48A7B838E6" box="[1361,1439,1501,1523]" pageId="6" pageNumber="1300" type="female">female</specimenCount>
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, no syn-inclusions.
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</materialsCitation>
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<typeStatus id="54FFDEF27E48FFDAFBF3FA69A6A53B04" box="[1037,1154,1532,1553]" pageId="6" pageNumber="1300" type="paratype">Paratypes</typeStatus>
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. MAIG no. 5967, donated by Jonas Damzen, sex unknown, no syn-inclusions; MAIG no. 3687, donated by Tomasz Dębowski & Maciej Szulimowicz,
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<date id="FFFA46907E48FFDAFC28F9CDA6743B78" box="[982,1107,1624,1645]" pageId="6" pageNumber="1300" value="2000-09-09">09.09.2000</date>
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, sex unknown, syninclusions:
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<taxonomicName id="4C441BD37E48FFDAFCC5F9E3A1FE3B99" authorityName="Imhoff" authorityYear="1856" box="[827,985,1654,1676]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1300" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
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1,
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<taxonomicName id="4C441BD37E48FFDAFBFBF9E2A6883B99" box="[1029,1199,1655,1676]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="1300" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
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1; AWI-154, FT-IR 14954 (
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<figureCitation id="137F7CD57E48FFDAFC65F900A1C33BBE" box="[923,996,1685,1707]" captionStart="Figure 5" captionStartId="10.[163,244,898,920]" captionTargetBox="[164,784,196,858]" captionTargetId="figure-427@10.[163,784,195,859]" captionTargetPageId="10" captionText="Figure 5. FT-IR absorbance spectra of analysed amber pieces with the indication of museum inventory numbers above each spectrum." figureDoi="http://doi.org/10.5281/zenodo.5752974" httpUri="https://zenodo.org/record/5752974/files/figure.png" pageId="6" pageNumber="1300">Fig. 5</figureCitation>
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), sex unknown, no syn-inclusions. Other material. MAIG no. 5985, sex unknown, no syn-inclusions.
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</paragraph>
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<paragraph id="8BFB60507E48FFDBFCC5F887A0343BCE" blockId="6.[827,1442,1809,1862]" lastBlockId="7.[145,760,1671,1755]" lastPageId="7" lastPageNumber="1301" pageId="6" pageNumber="1300">
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<emphasis id="B930BC427E48FFDAFCC5F887A68D3A33" box="[827,1194,1809,1831]" italics="true" pageId="6" pageNumber="1300">
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Diagnosis:
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<taxonomicName id="4C441BD37E48FFDAFC37F884A6193A33" authority="Szawaryn, 2021" authorityName="Szawaryn" authorityYear="2021" box="[969,1086,1809,1830]" class="Insecta" family="Coccinellidae" genus="Baltosidis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1298" phylum="Arthropoda" rank="genus" status="gen. nov.">Baltosidis</taxonomicName>
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<taxonomicName id="4C441BD37E48FFDAFBB9F884A68D3A33" authority="Szawaryn, 2021" authorityName="Szawaryn" authorityYear="2021" box="[1095,1194,1809,1830]" pageId="6" pageNumber="1300" rank="species" species="damzeni" status="sp. nov.">damzeni</taxonomicName>
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</emphasis>
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can be distinguished from other
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<taxonomicName id="4C441BD37E48FFDAFC3FF8A5A6B33A53" box="[961,1172,1840,1862]" class="Insecta" family="Coccinellidae" genus="Baltosidis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1300" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B930BC427E48FFDAFC3FF8A5A61F3A50" box="[961,1080,1840,1861]" italics="true" pageId="6" pageNumber="1300">Baltosidis</emphasis>
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species
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</taxonomicName>
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by glabrous pronotum and elytra, abdominal postcoxal lines almost fused with accessory lines, but not forming distinctly closed v-shape. Ventral surface glabrous.
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</paragraph>
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<paragraph id="8BFB60507E49FFD5FF6FF894A0DD3C0D" blockId="7.[145,762,1793,1907]" lastBlockId="9.[145,762,197,280]" lastPageId="9" lastPageNumber="1303" pageId="7" pageNumber="1301">
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<emphasis id="B930BC427E49FFDBFF6FF894A3003A03" box="[145,295,1793,1814]" italics="true" pageId="7" pageNumber="1301">Description:</emphasis>
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Total length =
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<quantity id="4CBCCDB57E49FFDBFE03F894A0973A03" box="[509,688,1793,1814]" metricMagnitude="-3" metricUnit="m" metricValue="1.255" metricValueMax="1.28" metricValueMin="1.23" pageId="7" pageNumber="1301" unit="mm" value="1.255" valueMax="1.28" valueMin="1.23">1.23–1.28 mm</quantity>
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, total width =
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<quantity id="4CBCCDB57E49FFDBFF0DF88AA3BC3A20" box="[243,411,1823,1845]" metricMagnitude="-4" metricUnit="m" metricValue="8.3" metricValueMax="8.6" metricValueMin="8.0" pageId="7" pageNumber="1301" unit="mm" value="0.83" valueMax="0.86" valueMin="0.8">0.80–0.86 mm</quantity>
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. Body elongate oval (
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<figureCitation id="137F7CD57E49FFDBFD68F88AA0CE3A20" box="[662,745,1823,1845]" captionStart="Figure 3" captionStartId="8.[164,244,1520,1542]" captionTargetBox="[323,1283,195,1481]" captionTargetId="figure-154@8.[323,1283,195,1481]" captionTargetPageId="8" captionText="Figure 3. Baltosidis damzeni. A, habitus ventral. B, head, prosternum, mesoventrite. C, pronotum. D, tip of elytra with sutural lines. E, habitus lateral. F, details of maxillary palp. G, postcoxal lines on ventrite 1. Holotype, MAIG 5988 (A–E); paratype AWL 154 (F); paratype, MAIG 5967 (G)." figureDoi="http://doi.org/10.5281/zenodo.5752969" httpUri="https://zenodo.org/record/5752969/files/figure.png" pageId="7" pageNumber="1301">Fig. 3A</figureCitation>
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). Head, pronotum and elytra glabrous (
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<figureCitation id="137F7CD57E49FFDBFD92F8ABA0CF3A46" box="[620,744,1854,1876]" captionStart="Figure 3" captionStartId="8.[164,244,1520,1542]" captionTargetBox="[323,1283,195,1481]" captionTargetId="figure-154@8.[323,1283,195,1481]" captionTargetPageId="8" captionText="Figure 3. Baltosidis damzeni. A, habitus ventral. B, head, prosternum, mesoventrite. C, pronotum. D, tip of elytra with sutural lines. E, habitus lateral. F, details of maxillary palp. G, postcoxal lines on ventrite 1. Holotype, MAIG 5988 (A–E); paratype AWL 154 (F); paratype, MAIG 5967 (G)." figureDoi="http://doi.org/10.5281/zenodo.5752969" httpUri="https://zenodo.org/record/5752969/files/figure.png" pageId="7" pageNumber="1301">Fig. 3C, G</figureCitation>
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). Pronotum covered with medium-sized, irregular punctures about same size as those on elytra. Metaventrite covered with small punctures separated by a distance of about 3–4× its diameter (
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<figureCitation id="137F7CD57E49FFDBFAD4F950A7A63BCF" box="[1322,1409,1733,1755]" captionStart="Figure 3" captionStartId="8.[164,244,1520,1542]" captionTargetBox="[323,1283,195,1481]" captionTargetId="figure-154@8.[323,1283,195,1481]" captionTargetPageId="8" captionText="Figure 3. Baltosidis damzeni. A, habitus ventral. B, head, prosternum, mesoventrite. C, pronotum. D, tip of elytra with sutural lines. E, habitus lateral. F, details of maxillary palp. G, postcoxal lines on ventrite 1. Holotype, MAIG 5988 (A–E); paratype AWL 154 (F); paratype, MAIG 5967 (G)." figureDoi="http://doi.org/10.5281/zenodo.5752969" httpUri="https://zenodo.org/record/5752969/files/figure.png" pageId="7" pageNumber="1301">Fig. 3A</figureCitation>
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), punctures not associated with setae, without additional rows of large punctures anteriorly to hind coxae, metaventral postcoxal lines complete laterally, straight, discrimen visible in posterior half. Elytra covered with dense punctures of similar size as those on pronotum, irregularly distributed, punctures not associated with setae, sutural line present in apical third (
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<figureCitation id="137F7CD57E46FFD4FEC2F930A3B73BAF" box="[316,400,1701,1723]" captionStart="Figure 3" captionStartId="8.[164,244,1520,1542]" captionTargetBox="[323,1283,195,1481]" captionTargetId="figure-154@8.[323,1283,195,1481]" captionTargetPageId="8" captionText="Figure 3. Baltosidis damzeni. A, habitus ventral. B, head, prosternum, mesoventrite. C, pronotum. D, tip of elytra with sutural lines. E, habitus lateral. F, details of maxillary palp. G, postcoxal lines on ventrite 1. Holotype, MAIG 5988 (A–E); paratype AWL 154 (F); paratype, MAIG 5967 (G)." figureDoi="http://doi.org/10.5281/zenodo.5752969" httpUri="https://zenodo.org/record/5752969/files/figure.png" pageId="8" pageNumber="1302">Fig. 3D</figureCitation>
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), elytral margin narrow. Elytral epipleuron punctate, glabrous, incomplete, reaching to posterior border of ventrite 3 (
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<figureCitation id="137F7CD57E46FFD4FDDDF977A0523BE2" box="[547,629,1762,1784]" captionStart="Figure 3" captionStartId="8.[164,244,1520,1542]" captionTargetBox="[323,1283,195,1481]" captionTargetId="figure-154@8.[323,1283,195,1481]" captionTargetPageId="8" captionText="Figure 3. Baltosidis damzeni. A, habitus ventral. B, head, prosternum, mesoventrite. C, pronotum. D, tip of elytra with sutural lines. E, habitus lateral. F, details of maxillary palp. G, postcoxal lines on ventrite 1. Holotype, MAIG 5988 (A–E); paratype AWL 154 (F); paratype, MAIG 5967 (G)." figureDoi="http://doi.org/10.5281/zenodo.5752969" httpUri="https://zenodo.org/record/5752969/files/figure.png" pageId="8" pageNumber="1302">Fig. 3A</figureCitation>
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). Wings well developed. Ventrite 1 as long as 2 and 3 combined, surface covered with punctures of smaller size than those on metaventrite, abdominal postcoxal lines reaching one-seventh of length of ventrite 1 at level of hind coxa, incomplete, accessory line distinctly curved, long, separated from postcoxal line only in apical part, not forming v-shape pattern (
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<figureCitation id="137F7CD57E46FFD4FB29F956A70B3BCC" box="[1239,1324,1731,1753]" captionStart="Figure 3" captionStartId="8.[164,244,1520,1542]" captionTargetBox="[323,1283,195,1481]" captionTargetId="figure-154@8.[323,1283,195,1481]" captionTargetPageId="8" captionText="Figure 3. Baltosidis damzeni. A, habitus ventral. B, head, prosternum, mesoventrite. C, pronotum. D, tip of elytra with sutural lines. E, habitus lateral. F, details of maxillary palp. G, postcoxal lines on ventrite 1. Holotype, MAIG 5988 (A–E); paratype AWL 154 (F); paratype, MAIG 5967 (G)." figureDoi="http://doi.org/10.5281/zenodo.5752969" httpUri="https://zenodo.org/record/5752969/files/figure.png" pageId="8" pageNumber="1302">Fig. 3G</figureCitation>
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). Ventrite 2 slightly longer than ventrite 3, ventrites 3 and 4 subequal in length, ventrite 5 large, rounded, about as long as ventrite 2. Ventrites 2–5 glabrous. Femora swollen, tibiae extended apically, tarsi tetramerous. Coxites of female genitalia can be observed in
|
||
<typeStatus id="54FFDEF27E47FFD5FD69FF50A0DE3DCE" box="[663,761,197,219]" pageId="9" pageNumber="1303" type="holotype">holotype</typeStatus>
|
||
(MAIG 5988) specimen: they are narrow, elongate, with one to two long setae attached to stylus (
|
||
<figureCitation id="137F7CD57E47FFD5FD67FE97A0CD3C0D" box="[665,746,258,280]" captionStart="Figure 3" captionStartId="8.[164,244,1520,1542]" captionTargetBox="[323,1283,195,1481]" captionTargetId="figure-154@8.[323,1283,195,1481]" captionTargetPageId="8" captionText="Figure 3. Baltosidis damzeni. A, habitus ventral. B, head, prosternum, mesoventrite. C, pronotum. D, tip of elytra with sutural lines. E, habitus lateral. F, details of maxillary palp. G, postcoxal lines on ventrite 1. Holotype, MAIG 5988 (A–E); paratype AWL 154 (F); paratype, MAIG 5967 (G)." figureDoi="http://doi.org/10.5281/zenodo.5752969" httpUri="https://zenodo.org/record/5752969/files/figure.png" pageId="9" pageNumber="1303">Fig. 3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFB60507E47FFD5FF6FFED4A0943C81" blockId="9.[145,760,321,404]" pageId="9" pageNumber="1303">
|
||
<emphasis id="B930BC427E47FFD5FF6FFED4A3333C43" box="[145,276,321,342]" italics="true" pageId="9" pageNumber="1303">Etymology:</emphasis>
|
||
The specific epithet is dedicated to my friend Jonas Damzen (
|
||
<collectingRegion id="4980AEB27E47FFD5FE5DFEF5A3DB3C63" box="[419,508,352,374]" country="Lithuania" name="Vilniaus" pageId="9" pageNumber="1303">Vilnius</collectingRegion>
|
||
,
|
||
<collectingCountry id="F35320C07E47FFD5FDF4FEF5A0A43C63" box="[522,643,352,374]" name="Lithuania" pageId="9" pageNumber="1303">Lithuania</collectingCountry>
|
||
) collector and donator of the
|
||
<typeStatus id="54FFDEF27E47FFD5FE97FEEAA3BD3C81" box="[361,410,383,404]" pageId="9" pageNumber="1303">type</typeStatus>
|
||
specimens to the MAIG.
|
||
</paragraph>
|
||
<paragraph id="8BFB60507E47FFD5FF6FFE28A02D3CE7" blockId="9.[145,761,445,498]" pageId="9" pageNumber="1303">
|
||
<emphasis id="B930BC427E47FFD5FF6FFE28A3AB3CC7" box="[145,396,445,466]" italics="true" pageId="9" pageNumber="1303">Locality and horizon:</emphasis>
|
||
Eocene, Baltic amber, Gulf of Gdańsk area (secondary deposit).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |