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<document id="E5BD0E3B36A4C5CBB5CEC055DCEC6E66" ID-CLB-Dataset="21966" ID-DOI="10.5852/ejt.2016.195" ID-GBIF-Dataset="39071d6f-d900-4c81-ace1-dd03875018c5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837846" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590071140602" checkinUser="valdenar" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2016" docId="03EB5A00FFE37D77ACE7D63DFE1E990C" docLanguage="en" docName="ejt-2016-195.pdf.imf" docOrigin="European Journal of Taxonomy 195" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tylopus punctus Likhitrakarn &amp; Golovatch &amp; Panha 2016, sp. nov." docType="treatment" docUuid="35585ACA-8490-49EE-8DA5-73EB3EB04A5C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="32" masterDocId="FFD22278FFFF7D68AE62D451FFD19824" masterDocTitle="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina" masterLastPageNumber="47" masterPageNumber="1" pageNumber="29" updateTime="1698840626989" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CFD3C770A91AE24A0F31E897EBFA716E">The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina</mods:title>
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<mods:namePart id="0F1DAA61D86FF980AF09FC18EECBDD3A">Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation id="25CFAFC300C006554725452D9AF726DB">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand. &amp; Email: kongerrrr @ hotmail. com &amp; urn: lsid: zoobank. org: author: 442 E 2 E 6 E- 1 CB 3 - 47 B 2 - 9896 - 1 A 922 CE 509 E 1</mods:affiliation>
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<mods:namePart id="788568545B501FB88CEBBD29024ACC12">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="A69CEA2A7063C1B5AAF6436B168A5A19">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia. &amp; Corresponding author: sgolovatch @ yandex. ru &amp; urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:namePart id="99E41BD2A408E567CC8D9CE36556C349">Panha, Somsak</mods:namePart>
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<mods:affiliation id="E161B4A05DCAC932D6512687463A0BD4">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Corresponding author: somsak. pan @ chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:date id="C52B63A654A4C396C0C2411C1B02554F">2016</mods:date>
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<treatment id="03EB5A00FFE37D77ACE7D63DFE1E990C" ID-DOI="http://doi.org/10.5281/zenodo.3852406" ID-GBIF-Taxon="164246093" ID-Zenodo-Dep="3852406" LSID="urn:lsid:zoobank.org:act:35585ACA-8490-49EE-8DA5-73EB3EB04A5C" httpUri="http://treatment.plazi.org/id/03EB5A00FFE37D77ACE7D63DFE1E990C" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29">
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<taxonomicName id="4C429095FFE37D74ACE7D63DFC9A9AA3" ID-CoL="7CZPX" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[645,843,620,647]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="punctus" status="sp. nov.">
<emphasis id="B9363704FFE37D74ACE7D63DFC9A9AA3" bold="true" box="[645,843,620,647]" italics="true" pageId="28" pageNumber="29">Tylopus punctus</emphasis>
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<taxonomicNameLabel id="A2058A7FFFE37D74AD31D63CFC7F9AA3" box="[851,942,621,647]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C358B89DFFE37D74ACA0D6E2FCA09AE9" box="[706,881,691,717]" pageId="28" pageNumber="29" type="description">
<paragraph id="8BFDEB16FFE37D74ACA0D6E2FCA09AE9" blockId="28.[360,1228,620,717]" box="[706,881,691,717]" pageId="28" pageNumber="29">
<figureCitation id="1379F793FFE37D74ACA0D6E2FC999AE9" box="[706,840,691,717]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="29.[189,232,1894,1920]" captionStartId-1="30.[189,232,1966,1992]" captionTargetBox-0="[189,1398,911,1851]" captionTargetBox-1="[311,1244,1002,1916]" captionTargetId-0="figure@29.[189,1398,911,1851]" captionTargetId-1="figure@30.[304,1249,1000,1919]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." captionText-1="Fig. 22. Tylopus punctus sp. nov., holotype, Ƌ. AD. Left gonopod, lateral, mesal, subcaudal and suboral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text." figureDoi-0="http://doi.org/10.5281/zenodo.3837917" figureDoi-1="http://doi.org/10.5281/zenodo.3837919" httpUri-0="https://zenodo.org/record/3837917/files/figure.png" httpUri-1="https://zenodo.org/record/3837919/files/figure.png" pageId="28" pageNumber="29">Figs 2122</figureCitation>
,
<figureCitation id="1379F793FFE37D74AD36D6E2FCA09AE9" box="[852,881,691,717]" captionStart="Fig" captionStartId="38.[189,232,1821,1847]" captionTargetBox="[266,1310,1040,1771]" captionTargetId="figure@38.[266,1310,1040,1771]" captionTargetPageId="38" captionText="Fig. 26. Distribution of the new eight species of Tylopus. Filled triangle. T. punctus sp. nov. Filled circle. T. acuminatus sp. nov. Crossed circle. T. acuminatus sp. nov., T. dorsalis sp. nov. Crossed square. T. acuminatus sp. nov. T. retusus sp. nov. Open circle. T. retusus sp. nov. Open triangle. T. baenzigeri Golovatch &amp; Enghoff, 1993. Filled square. T. hongkhraiensis sp. nov. Open diamond. T. thunghaihin sp. nov. Inverted filled triangle. T. flavolineatus sp. nov. Open square. T. moniliformis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3837927" httpUri="https://zenodo.org/record/3837927/files/figure.png" pageId="28" pageNumber="29">26</figureCitation>
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<paragraph id="8BFDEB16FFE37D74AEDFD6A5FEE79B2A" blockId="28.[189,310,756,782]" box="[189,310,756,782]" pageId="28" pageNumber="29">
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<emphasis id="B9363704FFE37D74AEDFD6A5FEE79B2A" bold="true" box="[189,310,756,782]" pageId="28" pageNumber="29">Diagnosis</emphasis>
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Very similar to
<taxonomicName id="4C429095FFE37D74AF17D775FD679B1A" authority="(Pocock, 1895)" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[373,694,803,830]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="silvestris">
<emphasis id="B9363704FFE37D74AF17D775FE2B9B19" box="[373,506,803,829]" italics="true" pageId="28" pageNumber="29">T. silvestris</emphasis>
(
<bibRefCitation id="EFD396E7FFE37D74AC69D772FD7F9B1A" author="Pocock R. I." box="[523,686,803,830]" pageId="28" pageNumber="29" pagination="346 - 368" refId="ref20188" refString="Pocock R. I. 1895. Report upon the Chilopoda and Diplopoda obtained by P. W. Bassett-Smith, Esq., Surgeon R. N., and J. J. Walker, Esq., R. N., during the cruise in the Chinese Seas of H. M. S. ' Penguin', Commander W. U. Moore commanding. Annals and Magazine of Natural History series 6, 15: 346 - 368. http: // biodiversitylibrary. org / page / 19265532" type="journal article" year="1895">Pocock, 1895</bibRefCitation>
)
</taxonomicName>
, but differs in the smaller size, the width of midbody being
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(versus 3.0 mm); the caudal corner of midbody paraterga very narrowly rounded and not drawn behind the rear tergal margin (versus nearly pointed and produced behind the rear tergal margin), and gonopod process
<emphasis id="B9363704FFE37D74AFDED7DCFE1D9B83" bold="true" box="[444,460,909,935]" pageId="28" pageNumber="29">h</emphasis>
small and pointed (versus absent).
</paragraph>
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<emphasis id="B9363704FFE37D74AEDFD79FFE959BCC" bold="true" box="[189,324,974,1000]" pageId="28" pageNumber="29">Etymology</emphasis>
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To emphasize gonopod process
<emphasis id="B9363704FFE37D74AC51D7ACFD929C33" bold="true" box="[563,579,1021,1047]" pageId="28" pageNumber="29">h</emphasis>
being small and pointed; adjective.
</paragraph>
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<emphasis id="B9363704FFE37D74AEDFD06FFE7B9C7C" bold="true" box="[189,426,1086,1112]" pageId="28" pageNumber="29">Material examined</emphasis>
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<collectingCountry id="F355AB86FFE37D74AEDFD0C1FE8C9C8E" box="[189,349,1168,1194]" name="Myanmar" pageId="28" pageNumber="29">MYANMAR</collectingCountry>
: Ƌ,
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<location id="8E9DBDCDFFE37D74AFF0D0C1FDAA9C8E" LSID="urn:lsid:plazi:treatment:03EB5A00FFE37D77ACE7D63DFE1E990C:8E9DBDCDFFE37D74AFF0D0C1FDAA9C8E" box="[402,635,1168,1194]" country="Myanmar" county="Chan State" latitude="20.922222" longLatPrecision="21" longitude="96.56667" municipality="Aungban" name="Mintaingbin Forest" pageId="28" pageNumber="29">Mintaingbin Forest</location>
Camp
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, ca
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north of
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,
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,
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,
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, ca
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a.s.l.
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,
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</date>
, leg.
<collectorName id="26B78EC0FFE37D74AD2ED0E5FBCD9CE9" box="[844,1052,1203,1230]" pageId="28" pageNumber="29">H. Schillhammer</collectorName>
&amp; H.
<collectionCode id="ED5373D3FFE37D74AA00D0E5FBA89CEA" box="[1122,1145,1204,1230]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="28" pageNumber="29" type="Museum">V</collectionCode>
.
<collectorName id="26B78EC0FFE37D74AAEAD0E2FB289CE9" box="[1160,1273,1203,1229]" pageId="28" pageNumber="29">Shaverdo</collectorName>
(NHMW- 8248).
</paragraph>
</materialsCitation>
<paragraph id="8BFDEB16FFE37D74AE88D149FE8D9D16" blockId="28.[189,914,1304,1366]" box="[234,348,1304,1330]" pageId="28" pageNumber="29">
<typeStatus id="54F955B4FFE37D74AE88D149FE8D9D16" box="[234,348,1304,1330]" pageId="28" pageNumber="29" type="paratype">
<emphasis id="B9363704FFE37D74AE88D149FE8D9D16" bold="true" box="[234,348,1304,1330]" pageId="28" pageNumber="29">Paratype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BFDEB16FFE37D74AEDFD16AFC439D71" blockId="28.[189,914,1304,1366]" box="[189,914,1339,1366]" pageId="28" pageNumber="29">
<collectingCountry id="F355AB86FFE37D74AEDFD16AFE8A9D71" box="[189,347,1339,1365]" name="Myanmar" pageId="28" pageNumber="29">MYANMAR</collectingCountry>
:
<specimenCount id="9D44209FFFE37D74AF04D16AFE439D72" box="[358,402,1339,1366]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
, same data as for
<typeStatus id="54F955B4FFE37D74AC07D16AFD1D9D71" box="[613,716,1339,1365]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
(NHMW-8249).
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFE37D76AEDFD12DFBC69A5F" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="description">
<paragraph id="8BFDEB16FFE37D74AEDFD12DFE9F9DB2" blockId="28.[189,334,1404,1430]" box="[189,334,1404,1430]" pageId="28" pageNumber="29">
<heading id="D0B55C7AFFE37D74AEDFD12DFE9F9DB2" bold="true" box="[189,334,1404,1430]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B9363704FFE37D74AEDFD12DFE9F9DB2" bold="true" box="[189,334,1404,1430]" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFE37D74AEDFD1FAFE5E9E0B" blockId="28.[189,1399,1451,1583]" pageId="28" pageNumber="29">
MEASUREMENTS AND COLOUR. Length 27.5 (Ƌ) or
<quantity id="4CBA46F3FFE37D74AC8FD1FAFCEE9DE1" box="[749,831,1451,1477]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="28" pageNumber="29" unit="mm" value="29.0">29 mm</quantity>
(
<specimenCount id="9D44209FFFE37D74AD33D1FAFCB59DE1" box="[849,868,1451,1477]" pageId="28" pageNumber="29" type="female"></specimenCount>
), width of midbody pro- and metazonae 1.56 and
<quantity id="4CBA46F3FFE37D74AE92D19FFE9D9DCD" box="[240,332,1486,1513]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="28" pageNumber="29" unit="mm" value="2.1">2.1 mm</quantity>
(Ƌ) or 1.98 and
<quantity id="4CBA46F3FFE37D74AC75D19FFDA39DCD" box="[535,626,1486,1513]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="29" unit="mm" value="2.5">2.5 mm</quantity>
(
<specimenCount id="9D44209FFFE37D74ACE4D19FFD489DCC" box="[646,665,1486,1512]" pageId="28" pageNumber="29" type="female"></specimenCount>
), respectively. Coloration of alcohol material upon long-term preservation rather uniformly light brown to brown, with light yellow paraterga, antennae, venter and legs (
<figureCitation id="1379F793FFE37D74AE9ED245FE819E0B" box="[252,336,1556,1583]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
AI).
</paragraph>
<paragraph id="8BFDEB16FFE37D74AEDFD207FDEF9F04" blockId="28.[189,1399,1622,1824]" pageId="28" pageNumber="29">
HEAD. Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae rather short (
<figureCitation id="1379F793FFE37D74AEA4D228FEE19EB7" box="[198,304,1657,1683]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="28" pageNumber="29">Fig. 21B</figureCitation>
), reaching only in front of body segment 3 (Ƌ,
<specimenCount id="9D44209FFFE37D74AD38D22BFCBF9EB0" box="[858,878,1658,1684]" pageId="28" pageNumber="29" type="female"></specimenCount>
) when stretched dorsally. In width, segment 3 &lt;4 &lt;head &lt;collum &lt;segment 2 &lt;515; thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 4+4 anterior, 2+2 intermediate and 3+3 posterior; an evident lateral incision at about anterior 1/3; caudal corner of paraterga very broadly rounded, declined ventrad, not drawn behind rear tergal margin.
</paragraph>
<paragraph id="8BFDEB16FFE37D76AEDFD319FAA7996D" blockId="28.[189,1399,1863,1996]" lastBlockId="30.[189,1399,267,329]" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
BODY. Tegument smooth and shining, prozonae very finely shagreened, metaterga rather smooth and leathery, with posterior halves faintly rugulose, surface below paraterga finely microgranulate (
<figureCitation id="1379F793FFE37D74AEA4D3DFFECC9F8C" box="[198,285,1934,1960]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
AF). Postcollum metaterga with two transverse rows of setae: 2+
<quantity id="4CBA46F3FFE37D74AA2AD3DFFBA89F8C" box="[1096,1145,1934,1960]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="28" pageNumber="29" unit="in" value="2.0">2 in</quantity>
anterior (pre-sulcus) row, mostly abraded, and 2+
<quantity id="4CBA46F3FFE37D74AC6AD3E0FDE59FEF" box="[520,564,1969,1995]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="28" pageNumber="29" unit="in" value="2.0">2 in</quantity>
posterior (post-sulcus) row, mostly traceable as insertion points, except for segments 18 and 19, each latter with 4+4 setae in posterior row. Tergal setae simple, slender, about 1/3 of metatergal length (
<figureCitation id="1379F793FFE27D75AF83D57FFDA3996D" box="[481,626,302,329]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="29" pageNumber="30">Fig. 21A, F</figureCitation>
). Axial line visible both on pro- and metazonae. Paraterga welldeveloped (
<figureCitation id="1379F793FFE27D75AF27D503FE469948" box="[325,407,338,364]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="29" pageNumber="30">Fig. 21</figureCitation>
AG), especially so in Ƌ, set high (at upper ¼ of body height), anterior edge rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded, segments 2 and 3 slightly extending behind rear tergal margin, thereafter not surpassing rear tergal margin until segment 15 (
<figureCitation id="1379F793FFE27D75AEA4D5EAFEC899F1" box="[198,281,443,469]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="29" pageNumber="30">Fig. 21</figureCitation>
AD). Paratega 2 broad, anterior edge angular, lateral edge with two small, but evident incisions in anterior half. Following paraterga with an evident incision in anterior 1/3 of lateral edge (
<figureCitation id="1379F793FFE27D75AB61D58FFF249A38" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="29" pageNumber="30">Fig. 21A, C, F</figureCitation>
). Calluses on paraterga delimited by a sulcus only dorsally, rather narrow, a little wider on porebearing segments. Ozopores distinct, lying at about ¼ of metatergal length in front of posterior edge of metaterga. Transverse sulcus usually distinct (
<figureCitation id="1379F793FFE27D75AD67D619FC6B9A47" box="[773,954,584,611]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="29" pageNumber="30">Fig. 21A, D, F</figureCitation>
), slightly incomplete on segment 19, complete on segments 518, narrow, line-shaped, rather deep, reaching bases of paraterga, beaded at bottom. Stricture between pro- and metazonae evident, rather wide and deep, beaded at bottom down to base of paraterga (
<figureCitation id="1379F793FFE27D75AFD9D6E3FDF79AE8" box="[443,550,690,716]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="29" pageNumber="30">Fig. 21A</figureCitation>
, CF). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2 and 3, thereafter increasingly reduced and remaining a sharp caudal tooth until segment 9 (Ƌ) or 6 (
<specimenCount id="9D44209FFFE27D75AF24D6A9FE889B36" box="[326,345,760,786]" pageId="29" pageNumber="30" type="female"></specimenCount>
), retained as a small caudal tooth and increasingly reduced until segment 17, absent from segment 18 (Ƌ) or 7 (
<specimenCount id="9D44209FFFE27D75AFAED74AFE319B11" box="[460,480,795,821]" pageId="29" pageNumber="30" type="female"></specimenCount>
). Epiproct (
<figureCitation id="1379F793FFE27D75AC16D74AFD189B11" box="[628,713,795,821]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="29" pageNumber="30">Fig. 21</figureCitation>
EG) conical, flattened dorsoventrally, with two evident, rather large, rounded, apical papillae; tip subtruncate; lateral pre-apical papillae clear, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge small and well-separated (
<figureCitation id="1379F793FFE17D76AA99D57FFAB8996D" box="[1275,1385,302,329]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="30" pageNumber="31">Fig. 21G</figureCitation>
).
</paragraph>
<caption id="DF3DBB9EFFE27D75AEDFD337FD9E9FCE" ID-DOI="http://doi.org/10.5281/zenodo.3837917" ID-Zenodo-Dep="3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1894,1920]" targetBox="[189,1398,911,1851]" targetPageId="29">
<paragraph id="8BFDEB16FFE27D75AEDFD337FD9E9FCE" blockId="29.[189,1399,1894,2026]" pageId="29" pageNumber="30">
<emphasis id="B9363704FFE27D75AEDFD337FECD9FA4" bold="true" box="[189,284,1894,1920]" pageId="29" pageNumber="30">Fig. 21.</emphasis>
<taxonomicName id="4C429095FFE27D75AF47D337FE349FA4" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[293,485,1894,1920]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="punctus" status="sp. nov.">
<emphasis id="B9363704FFE27D75AF47D337FE349FA4" box="[293,485,1894,1920]" italics="true" pageId="29" pageNumber="30">Tylopus punctus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFE27D75AF8DD336FD9F9FA5" box="[495,590,1895,1921]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ.
<emphasis id="B9363704FFE27D75AC99D337FCE39FA4" bold="true" box="[763,818,1894,1920]" pageId="29" pageNumber="30">AB</emphasis>
. Anterior part of body, dorsal and lateral views, respectively.
<emphasis id="B9363704FFE27D75AF35D3D8FE419F87" bold="true" box="[343,400,1929,1955]" pageId="29" pageNumber="30">CD</emphasis>
. Segments 1112, dorsal and lateral views, respectively.
<emphasis id="B9363704FFE27D75AA48D3D8FBB29F87" bold="true" box="[1066,1123,1929,1955]" pageId="29" pageNumber="30">EG</emphasis>
. Posterior part of body, lateral, subdorsal and subventral views, respectively.
<emphasis id="B9363704FFE27D75AD5ED3FCFCBC9FE3" bold="true" box="[828,877,1965,1991]" pageId="29" pageNumber="30">HI</emphasis>
. Sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFE17D76AEDFD521FA9D99F4" blockId="30.[189,1399,368,465]" pageId="30" pageNumber="31">
STERNA. Sparsely setose, without modifications, but with a pair of small, blunt, fully separated cones between Ƌ coxae 4 (
<figureCitation id="1379F793FFE17D76AFCED5C2FDFE998A" box="[428,559,403,430]" captionStart="Fig" captionStartId="29.[189,232,1894,1920]" captionTargetBox="[189,1398,911,1851]" captionTargetId="figure@29.[189,1398,911,1851]" captionTargetPageId="29" captionText="Fig. 21. Tylopus punctus sp. nov., holotype, Ƌ. AB. Anterior part of body, dorsal and lateral views, respectively. CD. Segments 1112, dorsal and lateral views, respectively. EG. Posterior part of body, lateral, subdorsal and subventral views, respectively. HI. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837917" httpUri="https://zenodo.org/record/3837917/files/figure.png" pageId="30" pageNumber="31">Fig. 21H, I</figureCitation>
). Legs rather short and slender, midbody ones ca 1.11.2 (Ƌ) or 0.80.9 (
<specimenCount id="9D44209FFFE17D76AEAAD5E7FF0A99F4" box="[200,219,438,464]" pageId="30" pageNumber="31" type="female"></specimenCount>
) as long as body height, Ƌ prefemora not swollen, Ƌ tarsal brushes present only on legs 1 and 2.
</paragraph>
<paragraph id="8BFDEB16FFE17D76AEDFD5A9FBC69A5F" blockId="30.[189,1399,504,636]" pageId="30" pageNumber="31">
GONOPOD. Rather simple (
<figureCitation id="1379F793FFE17D76AF8AD5A9FDEE9A36" box="[488,575,504,530]" captionStart="Fig" captionStartId="30.[189,232,1966,1992]" captionTargetBox="[311,1244,1002,1916]" captionTargetId="figure@30.[304,1249,1000,1919]" captionTargetPageId="30" captionText="Fig. 22. Tylopus punctus sp. nov., holotype, Ƌ. AD. Left gonopod, lateral, mesal, subcaudal and suboral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837919" httpUri="https://zenodo.org/record/3837919/files/figure.png" pageId="30" pageNumber="31">Fig. 22</figureCitation>
); coxa slightly curved caudad, sparsely setose distoventrally. Prefemur sparsely setose, about 1/3 as long as femorite + postfemoral part. Femorite curved and stout, with an evident mesal groove and a strong distolateral sulcus demarcating a postfemoral part; process
<emphasis id="B9363704FFE17D76AB75D66FFAF69A7C" bold="true" box="[1303,1319,574,600]" pageId="30" pageNumber="31">h</emphasis>
small, slender and pointed; solenophore (
<emphasis id="B9363704FFE17D76AC31D633FDAE9A58" bold="true" box="[595,639,610,636]" pageId="30" pageNumber="31">sph</emphasis>
) clearly coiled, long and flattened.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFE17D77AEDFD6F2FE1E990C" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BFDEB16FFE17D76AEDFD6F2FEF59A99" blockId="30.[189,292,675,701]" box="[189,292,675,701]" pageId="30" pageNumber="31">
<heading id="D0B55C7AFFE17D76AEDFD6F2FEF59A99" bold="true" box="[189,292,675,701]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B9363704FFE17D76AEDFD6F2FEF59A99" bold="true" box="[189,292,675,701]" pageId="30" pageNumber="31">Remark</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFE17D76AEDFD680FB9E9B2B" blockId="30.[189,1399,721,783]" pageId="30" pageNumber="31">
<bibRefCitation id="EFD396E7FFE17D76AEDFD680FEBA9AC8" author="Pocock R. I." box="[189,363,721,748]" pageId="30" pageNumber="31" pagination="346 - 368" refId="ref20188" refString="Pocock R. I. 1895. Report upon the Chilopoda and Diplopoda obtained by P. W. Bassett-Smith, Esq., Surgeon R. N., and J. J. Walker, Esq., R. N., during the cruise in the Chinese Seas of H. M. S. ' Penguin', Commander W. U. Moore commanding. Annals and Magazine of Natural History series 6, 15: 346 - 368. http: // biodiversitylibrary. org / page / 19265532" type="journal article" year="1895">Pocock (1895)</bibRefCitation>
described only
<taxonomicName id="4C429095FFE17D76AC49D683FC849AC8" authority="(Pocock, 1895)" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[555,853,721,748]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="B9363704FFE17D76AC49D683FD469ACF" box="[555,663,721,747]" italics="true" pageId="30" pageNumber="31">T. doriae</emphasis>
(
<bibRefCitation id="EFD396E7FFE17D76ACCAD680FC9C9AC8" author="Pocock R. I." box="[680,845,721,748]" pageId="30" pageNumber="31" pagination="346 - 368" refId="ref20188" refString="Pocock R. I. 1895. Report upon the Chilopoda and Diplopoda obtained by P. W. Bassett-Smith, Esq., Surgeon R. N., and J. J. Walker, Esq., R. N., during the cruise in the Chinese Seas of H. M. S. ' Penguin', Commander W. U. Moore commanding. Annals and Magazine of Natural History series 6, 15: 346 - 368. http: // biodiversitylibrary. org / page / 19265532" type="journal article" year="1895">Pocock, 1895</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C429095FFE17D76ADF0D683FBC89ACF" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[914,1049,721,747]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="silvestris">
<emphasis id="B9363704FFE17D76ADF0D683FBC89ACF" box="[914,1049,721,747]" italics="true" pageId="30" pageNumber="31">T. silvestris</emphasis>
</taxonomicName>
from northern
<collectingCountry id="F355AB86FFE17D76AAB3D683FA949AC8" box="[1233,1349,722,748]" name="Myanmar" pageId="30" pageNumber="31">Myanmar</collectingCountry>
, the above being a third species of this genus also occurring in northern
<collectingCountry id="F355AB86FFE17D76ADB4D6A4FB9B9B2B" box="[982,1098,757,783]" name="Myanmar" pageId="30" pageNumber="31">Myanmar</collectingCountry>
.
</paragraph>
<caption id="DF3DBB9EFFE17D76AEDFD3FFFBBD9FC8" ID-DOI="http://doi.org/10.5281/zenodo.3837919" ID-Zenodo-Dep="3837919" httpUri="https://zenodo.org/record/3837919/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1966,1992]" targetBox="[311,1244,1002,1916]" targetPageId="30">
<paragraph id="8BFDEB16FFE17D76AEDFD3FFFBBD9FC8" blockId="30.[189,1398,1966,2028]" pageId="30" pageNumber="31">
<emphasis id="B9363704FFE17D76AEDFD3FFFEC69FEC" bold="true" box="[189,279,1966,1992]" pageId="30" pageNumber="31">Fig. 22.</emphasis>
<taxonomicName id="4C429095FFE17D76AF7FD3FFFE089FEC" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[285,473,1966,1992]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="punctus" status="sp. nov.">
<emphasis id="B9363704FFE17D76AF7FD3FFFE089FEC" box="[285,473,1966,1992]" italics="true" pageId="30" pageNumber="31">Tylopus punctus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFE17D76AFBDD3FFFDEA9FEC" box="[479,571,1966,1992]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ.
<emphasis id="B9363704FFE17D76ACBED3FFFCC49FEC" bold="true" box="[732,789,1966,1992]" pageId="30" pageNumber="31">AD</emphasis>
. Left gonopod, lateral, mesal, subcaudal and suboral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFE07D77AED9D55DFE1E990C" blockId="31.[187,463,267,296]" box="[187,463,267,296]" pageId="31" pageNumber="32">
<emphasis id="B9363704FFE07D77AED9D55DFE1E990C" bold="true" box="[187,463,267,296]" italics="true" pageId="31" pageNumber="32">New faunistic record</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>