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<document id="98DD6D3D866C774AB1E0F02DC365DDB8" ID-CLB-Dataset="21966" ID-DOI="10.5852/ejt.2016.195" ID-GBIF-Dataset="39071d6f-d900-4c81-ace1-dd03875018c5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837846" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590071140602" checkinUser="valdenar" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2016" docId="03EB5A00FFE07D49AF9DD56EFDAE9AF7" docLanguage="en" docName="ejt-2016-195.pdf.imf" docOrigin="European Journal of Taxonomy 195" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tylopus baenzigeri Golovatch &amp; Enghoff 1993" docType="treatment" docVersion="7" lastPageNumber="34" masterDocId="FFD22278FFFF7D68AE62D451FFD19824" masterDocTitle="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina" masterLastPageNumber="47" masterPageNumber="1" pageNumber="32" updateTime="1698840626989" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D652BBF07C5A5D0A2BDFA24CD6D79ECF">The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina</mods:title>
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<mods:namePart id="9C5EE73C7D7B803A23BA3B84737E88B0">Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation id="655A0135E3BF3AE0322AA6D1AE7674F8">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand. &amp; Email: kongerrrr @ hotmail. com &amp; urn: lsid: zoobank. org: author: 442 E 2 E 6 E- 1 CB 3 - 47 B 2 - 9896 - 1 A 922 CE 509 E 1</mods:affiliation>
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<mods:namePart id="24813CF66E1D43F964141BCC711C54DB">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="453A2EFB2F686CE8C280AFD561F5703C">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia. &amp; Corresponding author: sgolovatch @ yandex. ru &amp; urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
<mods:nameIdentifier id="FA8342F57E75B70046A019A0A9B72276" type="email">sgolovatch@yandex.ru&amp;sgolovatch@yandex.ru&amp;somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:namePart id="14C0B9A31952DCA772DCBCABA449A4EF">Panha, Somsak</mods:namePart>
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<mods:affiliation id="3E3C0AAADC5B492DCB4453AC83037DC5">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Corresponding author: somsak. pan @ chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
<mods:nameIdentifier id="1A51BD4D6F194AD8766262516DF671C7" type="email">somsak.pan@chula.ac.th&amp;sgolovatch@yandex.ru&amp;somsak.pan@chula.ac.th</mods:nameIdentifier>
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<heading id="D0B55C7AFFE07D77AF9DD56EFBE2997E" box="[511,1075,319,346]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="32" reason="2">
<taxonomicName id="4C429095FFE07D77AF9DD56EFBE2997E" ID-CoL="7CZMK" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[511,1075,319,346]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="baenzigeri">
<emphasis id="B9363704FFE07D77AF9DD56EFD33997E" bold="true" box="[511,738,319,346]" italics="true" pageId="31" pageNumber="32">Tylopus baenzigeri</emphasis>
<bibRefCitation id="EFD396E7FFE07D77AC88D56EFBE2997E" author="Golovatch S. I. &amp; Enghoff H." box="[746,1075,319,346]" pageId="31" pageNumber="32" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
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<figureCitation id="1379F793FFE07D77ACB5D532FC8B9958" box="[727,858,354,381]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="31.[189,232,1894,1920]" captionStartId-1="32.[189,232,1930,1956]" captionStartId-2="33.[187,230,1930,1956]" captionStartId-3="38.[189,232,1821,1847]" captionTargetBox-0="[189,1398,599,1848]" captionTargetBox-1="[369,1231,950,1895]" captionTargetBox-2="[198,1389,743,1904]" captionTargetBox-3="[266,1310,1040,1771]" captionTargetId-0="figure@31.[189,1398,599,1848]" captionTargetId-1="figure@32.[359,1236,943,1899]" captionTargetId-2="figure@33.[198,1389,743,1904]" captionTargetId-3="figure@38.[266,1310,1040,1771]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionTargetPageId-3="38" captionText-0="Fig. 23. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." captionText-1="Fig. 24. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A–B. Right gonopod, lateral and mesal views, respectively. C. Leg 8. D. Legs of segment 11. Scale bars = 0.5 mm. Abbreviations are explained in the text." captionText-2="Fig. 25. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ, right gonopod. A–B. Mesal and lateral views, respectively. C–F. Distal part, submesal, sublateral, subcaudal and suboral views, respectively. Scale bar = 0.2 mm." captionText-3="Fig. 26. Distribution of the new eight species of Tylopus. Filled triangle. T. punctus sp. nov. Filled circle. T. acuminatus sp. nov. Crossed circle. T. acuminatus sp. nov., T. dorsalis sp. nov. Crossed square. T. acuminatus sp. nov. T. retusus sp. nov. Open circle. T. retusus sp. nov. Open triangle. T. baenzigeri Golovatch &amp; Enghoff, 1993. Filled square. T. hongkhraiensis sp. nov. Open diamond. T. thunghaihin sp. nov. Inverted filled triangle. T. flavolineatus sp. nov. Open square. T. moniliformis sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.3837921" figureDoi-1="http://doi.org/10.5281/zenodo.3837923" figureDoi-2="http://doi.org/10.5281/zenodo.3837925" figureDoi-3="http://doi.org/10.5281/zenodo.3837927" httpUri-0="https://zenodo.org/record/3837921/files/figure.png" httpUri-1="https://zenodo.org/record/3837923/files/figure.png" httpUri-2="https://zenodo.org/record/3837925/files/figure.png" httpUri-3="https://zenodo.org/record/3837927/files/figure.png" pageId="31" pageNumber="32">Figs 23–26</figureCitation>
</paragraph>
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<treatmentCitationGroup id="AB52CC38FFE07D77AED9D5F5FED299C6" pageId="31" pageNumber="32">
<treatmentCitation id="0AE3CD07FFE07D77AED9D5F5FCE4999A" author="Golovatch S. I. &amp; Enghoff H." box="[187,821,420,446]" page="101" pageId="31" pageNumber="32" year="1993">
<taxonomicName id="4C429095FFE07D77AED9D5F5FCE4999A" ID-CoL="7CZMK" authority="Golovatch &amp; Enghoff, 1993: 101" authorityName="Golovatch &amp; Enghoff" authorityPageNumber="101" authorityYear="1993" box="[187,821,420,446]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="baenzigeri">
<emphasis id="B9363704FFE07D77AED9D5F5FE4D999A" box="[187,412,420,446]" italics="true" pageId="31" pageNumber="32">Tylopus baenzigeri</emphasis>
<bibRefCitation id="EFD396E7FFE07D77AFC7D5F5FCE4999A" author="Golovatch S. I. &amp; Enghoff H." box="[421,821,420,446]" pageId="31" pageNumber="32" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993: 101</bibRefCitation>
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, figs 50–51 (
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locality, Doi Suthep National Park).
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</paragraph>
<paragraph id="8BFDEB16FFE07D77AED9D658FB569A00" blockId="31.[187,1159,521,548]" box="[187,1159,521,548]" pageId="31" pageNumber="32">
<treatmentCitationGroup id="AB52CC38FFE07D77AED9D658FB569A00" box="[187,1159,521,548]" pageId="31" pageNumber="32">
<taxonomicName id="4C429095FFE07D77AED9D658FE489A07" ID-CoL="7CZMK" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[187,409,521,547]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="baenzigeri">
<emphasis id="B9363704FFE07D77AED9D658FE489A07" box="[187,409,521,547]" italics="true" pageId="31" pageNumber="32">Tylopus baenzigeri</emphasis>
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–
<treatmentCitation id="0AE3CD07FFE07D77AFD5D658FD589A07" author="Enghoff H." box="[439,649,521,548]" page="99" pageId="31" pageNumber="32" year="2005">
<bibRefCitation id="EFD396E7FFE07D77AFD5D658FD589A07" author="Enghoff H." box="[439,649,521,548]" pageId="31" pageNumber="32" pagination="87 - 103" refId="ref19494" refString="Enghoff H. 2005. The millipedes of Thailand (Diplopoda). Steenstrupia 29 (1): 87 - 103." type="journal article" year="2005">Enghoff 2005: 99</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AE3CD07FFE07D77ACDAD658FC2E9A00" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Prateepasen R. &amp; Panha S." box="[696,1023,521,548]" page="25" pageId="31" pageNumber="32" year="2010">
<bibRefCitation id="EFD396E7FFE07D77ACDAD658FC2E9A00" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Prateepasen R. &amp; Panha S." box="[696,1023,521,548]" pageId="31" pageNumber="32" pagination="23 - 68" refId="ref20037" refString="Likhitrakarn N., Golovatch S. I., Prateepasen R. &amp; Panha S. 2010. Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 72: 23 - 68. http: // dx. doi. org / 10.3897 / zookeys. 72.744" type="journal article" year="2010">
Likhitrakarn
<emphasis id="B9363704FFE07D77AD30D65BFC579A07" box="[850,902,521,547]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
. 2010: 25
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE3CD07FFE07D77AA69D658FB529A00" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Panha S." box="[1035,1155,521,548]" page="65" pageId="31" pageNumber="32" year="2014">
<bibRefCitation id="EFD396E7FFE07D77AA69D658FB529A00" author="Likhitrakarn N. &amp; Golovatch S. I. &amp; Panha S." box="[1035,1155,521,548]" pageId="31" pageNumber="32" pagination="63 - 91" refId="ref19845" refString="Likhitrakarn N., Golovatch S. I. &amp; Panha S. 2014 a. Three new species of the millipede genus Tylopus Jeekel, 1968 from Thailand, with additional notes on the species described by Attems (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 435: 63 - 91. http: // dx. doi. org / 10.3897 / zookeys. 435.8286" type="journal article" year="2014">2014a: 65</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<caption id="DF3DBB9EFFE07D77AEDFD337FC399FCE" ID-DOI="http://doi.org/10.5281/zenodo.3837921" ID-Zenodo-Dep="3837921" httpUri="https://zenodo.org/record/3837921/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,232,1894,1920]" targetBox="[189,1398,599,1848]" targetPageId="31">
<paragraph id="8BFDEB16FFE07D77AEDFD337FC399FCE" blockId="31.[189,1399,1894,2027]" pageId="31" pageNumber="32">
<emphasis id="B9363704FFE07D77AEDFD337FECB9FA4" bold="true" box="[189,282,1894,1920]" pageId="31" pageNumber="32">Fig. 23.</emphasis>
<taxonomicName id="4C429095FFE07D77AF43D337FC859FA5" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[289,852,1894,1921]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="baenzigeri">
<emphasis id="B9363704FFE07D77AF43D337FDD19FA4" box="[289,512,1894,1920]" italics="true" pageId="31" pageNumber="32">Tylopus baenzigeri</emphasis>
<bibRefCitation id="EFD396E7FFE07D77AC65D337FC859FA5" author="Golovatch S. I. &amp; Enghoff H." box="[519,852,1894,1921]" pageId="31" pageNumber="32" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
, Ƌ.
<emphasis id="B9363704FFE07D77ADE4D337FC4A9FA4" bold="true" box="[902,923,1894,1920]" pageId="31" pageNumber="32">A</emphasis>
. Habitus, live coloration.
<emphasis id="B9363704FFE07D77AAACD337FAD49FA4" bold="true" box="[1230,1285,1894,1920]" pageId="31" pageNumber="32">B–C</emphasis>
. Anterior part of body, dorsal and lateral views, respectively.
<emphasis id="B9363704FFE07D77AD7AD3DBFC9E9F80" bold="true" box="[792,847,1930,1956]" pageId="31" pageNumber="32">D–E</emphasis>
. Segments 10 and 11, dorsal and lateral views, respectively.
<emphasis id="B9363704FFE07D77AF38D3FCFE409FE3" bold="true" box="[346,401,1965,1991]" pageId="31" pageNumber="32">F–H</emphasis>
. Posterior part of body, dorsal, ventral and lateral views, respectively.
<emphasis id="B9363704FFE07D77AA8BD3FCFAC39FE3" bold="true" box="[1257,1298,1965,1991]" pageId="31" pageNumber="32">I–J</emphasis>
. Sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
<subSubSection id="C358B89DFFDF7D49AEDFD55AFAA7996D" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="materials_examined">
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<heading id="D0B55C7AFFDF7D48AEDFD55AFE7B9901" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B9363704FFDF7D48AEDFD55AFE7B9901" bold="true" box="[189,426,267,293]" pageId="32" pageNumber="33">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFDF7D48AEDFD56BFC7C9953" blockId="32.[189,1398,313,375]" pageId="32" pageNumber="33">
<collectingCountry id="F355AB86FFDF7D48AEDFD56BFE889970" box="[189,345,314,340]" name="Thailand" pageId="32" pageNumber="33">THAILAND</collectingCountry>
: Ƌ, Royal Agricultural Station Angkhang, Fang,
<collectingRegion id="498625F4FFDF7D48ADCAD568FBE99977" box="[936,1080,313,339]" country="Thailand" name="Chiang Mai" pageId="32" pageNumber="33">Chiang Mai</collectingRegion>
,
<geoCoordinate id="EE768DD1FFDF7D48AA25D568FB089970" box="[1095,1241,313,340]" degrees="19" direction="north" minutes="54" orientation="latitude" pageId="32" pageNumber="33" precision="15" seconds="05" value="19.901388">19°54’05”N</geoCoordinate>
,
<geoCoordinate id="EE768DD1FFDF7D48AA85D56BFAA09970" box="[1255,1393,313,340]" degrees="99" direction="east" minutes="2" orientation="longitude" pageId="32" pageNumber="33" precision="15" seconds="25" value="99.040276">99° 2’25”E</geoCoordinate>
, ca
<quantity id="4CBA46F3FFDF7D48AEBCD50CFEE69953" box="[222,311,349,375]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="32" pageNumber="33" unit="m" value="1420.0">1420 m</quantity>
a.s.l.,
<date id="FFFCCDD6FFDF7D48AFE6D50CFD909953" box="[388,577,349,375]" pageId="32" pageNumber="33" value="2014-08-12">12 August 2014</date>
, leg. N. Likhitrakarn (
<collectionCode id="ED5373D3FFDF7D48AD24D50CFC719953" box="[838,928,349,375]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="32" pageNumber="33">CUMZ</collectionCode>
).
</paragraph>
<paragraph id="8BFDEB16FFDF7D48AEDFD5C7FE429995" blockId="32.[189,403,406,433]" box="[189,403,406,433]" pageId="32" pageNumber="33">
<heading id="D0B55C7AFFDF7D48AEDFD5C7FE429995" bold="true" box="[189,403,406,433]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B9363704FFDF7D48AEDFD5C7FE429995" bold="true" box="[189,403,406,433]" pageId="32" pageNumber="33">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFDF7D48AEDFD597FB799A01" blockId="32.[189,1398,453,550]" pageId="32" pageNumber="33">
MEASUREMENTS AND COLOUR. Length
<quantity id="4CBA46F3FFDF7D48AC01D594FD1F99C4" box="[611,718,453,480]" metricMagnitude="-2" metricUnit="m" metricValue="4.38" pageId="32" pageNumber="33" unit="mm" value="43.8">43.8 mm</quantity>
, width of midbody pro- and metazonae 3.51 and
<quantity id="4CBA46F3FFDF7D48AB65D594FAA399C4" box="[1287,1394,453,480]" metricMagnitude="-3" metricUnit="m" metricValue="5.04" pageId="32" pageNumber="33" unit="mm" value="5.04">5.04 mm</quantity>
, respectively. Coloration of live animal, as well as of alcohol material black-brown (
<figureCitation id="1379F793FFDF7D48AAFAD5B9FB3C9A26" box="[1176,1261,488,514]" captionStart="Fig" captionStartId="31.[189,232,1894,1920]" captionTargetBox="[189,1398,599,1848]" captionTargetId="figure@31.[189,1398,599,1848]" captionTargetPageId="31" captionText="Fig. 23. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837921" httpUri="https://zenodo.org/record/3837921/files/figure.png" pageId="32" pageNumber="33">Fig. 23</figureCitation>
A–G), head and venter dark to light brown, legs and antennae yellowish brown to light yellowish.
</paragraph>
<paragraph id="8BFDEB16FFDF7D49AEDFD617FAA7996D" blockId="32.[189,1399,581,889]" lastBlockId="33.[189,1399,267,329]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<collectionCode id="ED5373D3FFDF7D48AEDFD617FF2C9A79" box="[189,253,582,608]" pageId="32" pageNumber="33">BODY</collectionCode>
. Antennae rather short (
<figureCitation id="1379F793FFDF7D48AC7AD614FD539A7B" box="[536,642,581,607]" captionStart="Fig" captionStartId="31.[189,232,1894,1920]" captionTargetBox="[189,1398,599,1848]" captionTargetId="figure@31.[189,1398,599,1848]" captionTargetPageId="31" captionText="Fig. 23. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837921" httpUri="https://zenodo.org/record/3837921/files/figure.png" pageId="32" pageNumber="33">Fig. 23B</figureCitation>
), extending behind body segment 3 (Ƌ) when stretched dorsally. In width, segment 3 &lt;4 &lt;5 &lt;collum &lt;segment 2 &lt;6–17; thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 4+4 anterior, 1+1 intermediate and 4+4 posterior; a very faint incision laterally in posterior 1/3; caudal corner of paraterga very broadly rounded, not drawn behind rear tergal margin (
<figureCitation id="1379F793FFDF7D48AFFFD683FDF69AC8" box="[413,551,722,748]" captionStart="Fig" captionStartId="31.[189,232,1894,1920]" captionTargetBox="[189,1398,599,1848]" captionTargetId="figure@31.[189,1398,599,1848]" captionTargetPageId="31" captionText="Fig. 23. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837921" httpUri="https://zenodo.org/record/3837921/files/figure.png" pageId="32" pageNumber="33">
Fig. 23B
<emphasis id="B9363704FFDF7D48AC66D683FDC29AC8" bold="true" box="[516,531,722,748]" pageId="32" pageNumber="33">–</emphasis>
C
</figureCitation>
). Paraterga 2 well-developed (
<figureCitation id="1379F793FFDF7D48ADEFD683FBC69AC8" box="[909,1047,722,748]" captionStart="Fig" captionStartId="31.[189,232,1894,1920]" captionTargetBox="[189,1398,599,1848]" captionTargetId="figure@31.[189,1398,599,1848]" captionTargetPageId="31" captionText="Fig. 23. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837921" httpUri="https://zenodo.org/record/3837921/files/figure.png" pageId="32" pageNumber="33">
Fig. 23B
<emphasis id="B9363704FFDF7D48AD96D683FBD29AC8" bold="true" box="[1012,1027,722,748]" pageId="32" pageNumber="33">–</emphasis>
C
</figureCitation>
), horizontal, lateral edge with two minute incisions in anterior half, caudal corner very narrowly rounded. Paraterga 3 and 4 each nearly pointed, expanded laterally, with two minute incisions in the middle. Sterna densely setose and finely microtuberculate (
<figureCitation id="1379F793FFDF7D48AFBED76DFD9B9B72" box="[476,586,828,854]" captionStart="Fig" captionStartId="32.[189,232,1930,1956]" captionTargetBox="[369,1231,950,1895]" captionTargetId="figure@32.[359,1236,943,1899]" captionTargetPageId="32" captionText="Fig. 24. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A–B. Right gonopod, lateral and mesal views, respectively. C. Leg 8. D. Legs of segment 11. Scale bars = 0.5 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837923" httpUri="https://zenodo.org/record/3837923/files/figure.png" pageId="32" pageNumber="33">Fig. 24D</figureCitation>
); with a large, subtrapeziform, ventrally slightly concave, sternal lobe between Ƌ coxae 4 (
<figureCitation id="1379F793FFDF7D48AFD3D70EFDFA9B5D" box="[433,555,863,889]" captionStart="Fig" captionStartId="31.[189,232,1894,1920]" captionTargetBox="[189,1398,599,1848]" captionTargetId="figure@31.[189,1398,599,1848]" captionTargetPageId="31" captionText="Fig. 23. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837921" httpUri="https://zenodo.org/record/3837921/files/figure.png" pageId="32" pageNumber="33">
Fig. 23I
<emphasis id="B9363704FFDF7D48AC6DD70EFDCC9B5D" bold="true" box="[527,541,863,889]" pageId="32" pageNumber="33">–</emphasis>
J
</figureCitation>
). Legs rather long and slender, midbody ones ca 1.2–1.3 (Ƌ) as long as body height; Ƌ legs without tarsal brushes; Ƌ postfemora and tibiae each with an adenostyle (tubercle) on segments 8–17; Ƌ coxae, prefemora, femora and postfemora microtuberculate ventrally (
<figureCitation id="1379F793FFDE7D49AABFD57FFAB9996D" box="[1245,1384,302,329]" captionStart="Fig" captionStartId="32.[189,232,1930,1956]" captionTargetBox="[369,1231,950,1895]" captionTargetId="figure@32.[359,1236,943,1899]" captionTargetPageId="32" captionText="Fig. 24. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A–B. Right gonopod, lateral and mesal views, respectively. C. Leg 8. D. Legs of segment 11. Scale bars = 0.5 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837923" httpUri="https://zenodo.org/record/3837923/files/figure.png" pageId="33" pageNumber="34">
Fig. 24C
<emphasis id="B9363704FFDE7D49AB20D57EFA81996D" bold="true" box="[1346,1360,303,329]" pageId="33" pageNumber="34">–</emphasis>
D
</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3DBB9EFFDF7D48AEDFD3DBFEEB9FCF" ID-DOI="http://doi.org/10.5281/zenodo.3837923" ID-Zenodo-Dep="3837923" httpUri="https://zenodo.org/record/3837923/files/figure.png" pageId="32" pageNumber="33" startId="32.[189,232,1930,1956]" targetBox="[369,1231,950,1895]" targetPageId="32">
<paragraph id="8BFDEB16FFDF7D48AEDFD3DBFEEB9FCF" blockId="32.[189,1398,1930,2027]" pageId="32" pageNumber="33">
<emphasis id="B9363704FFDF7D48AEDFD3DBFECC9F80" bold="true" box="[189,285,1930,1956]" pageId="32" pageNumber="33">Fig. 24.</emphasis>
<taxonomicName id="4C429095FFDF7D48AF44D3DBFCB79F80" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[294,870,1930,1956]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="baenzigeri">
<emphasis id="B9363704FFDF7D48AF44D3DBFDD69F80" box="[294,519,1930,1956]" italics="true" pageId="32" pageNumber="33">Tylopus baenzigeri</emphasis>
<bibRefCitation id="EFD396E7FFDF7D48AC70D3DBFCB79F80" author="Golovatch S. I. &amp; Enghoff H." box="[530,870,1930,1956]" pageId="32" pageNumber="33" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
</taxonomicName>
, Ƌ.
<emphasis id="B9363704FFDF7D48ADFFD3DBFC059F80" bold="true" box="[925,980,1930,1956]" pageId="32" pageNumber="33">A–B</emphasis>
. Right gonopod, lateral and mesal views, respectively.
<emphasis id="B9363704FFDF7D48AFCBD3FCFE6F9FE3" bold="true" box="[425,446,1965,1991]" pageId="32" pageNumber="33">C</emphasis>
. Leg 8.
<emphasis id="B9363704FFDF7D48AC7CD3FFFDE29FEC" bold="true" box="[542,563,1966,1992]" pageId="32" pageNumber="33">D</emphasis>
. Legs of segment 11. Scale bars = 0.5 mm. Abbreviations are explained in the text.
</paragraph>
</caption>
<subSubSection id="C358B89DFFDE7D49AEDFD538FDAE9AF7" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BFDEB16FFDE7D49AEDFD538FEFE99A7" blockId="33.[189,303,361,387]" box="[189,303,361,387]" pageId="33" pageNumber="34">
<heading id="D0B55C7AFFDE7D49AEDFD538FEFE99A7" bold="true" box="[189,303,361,387]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B9363704FFDE7D49AEDFD538FEFE99A7" bold="true" box="[189,303,361,387]" pageId="33" pageNumber="34">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFDE7D49AEDFD5C6FCA59A1A" blockId="33.[189,1399,407,575]" pageId="33" pageNumber="34">
This new specimen agrees in most details with the original description (
<bibRefCitation id="EFD396E7FFDE7D49AA7ED5C6FAB89995" author="Golovatch S. I. &amp; Enghoff H." box="[1052,1385,407,433]" pageId="33" pageNumber="34" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff 1993</bibRefCitation>
), especially in gonopod structure which shows a huge spine
<emphasis id="B9363704FFDE7D49ADEED5EBFC4899F0" bold="true" box="[908,921,442,468]" pageId="33" pageNumber="34">z</emphasis>
serrate on both sides (
<figureCitation id="1379F793FFDE7D49AAD3D5EBFA9699F1" box="[1201,1351,442,469]" captionStart="Fig" captionStartId="32.[189,232,1930,1956]" captionTargetBox="[369,1231,950,1895]" captionTargetId="figure@32.[359,1236,943,1899]" captionTargetPageId="32" captionText="Fig. 24. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A–B. Right gonopod, lateral and mesal views, respectively. C. Leg 8. D. Legs of segment 11. Scale bars = 0.5 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837923" httpUri="https://zenodo.org/record/3837923/files/figure.png" pageId="33" pageNumber="34">
Figs 24A
<emphasis id="B9363704FFDE7D49AB40D5EBFAE099F0" bold="true" box="[1314,1329,442,468]" pageId="33" pageNumber="34">–</emphasis>
B
</figureCitation>
,
<figureCitation id="1379F793FFDE7D49AB35D5EBFAB699F0" box="[1367,1383,442,468]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[234,1349,804,1890]" captionTargetId="figure@5.[234,1349,804,1890]" captionTargetPageId="5" captionText="Fig. 3. Tylopus flavolineatus sp. nov., holotype, Ƌ, right gonopod. A–B. Mesal and lateral views, respectively. C–F. Distal part, subcaudal, subcaudal, suboral and sublateral views, respectively. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3837865" httpUri="https://zenodo.org/record/3837865/files/figure.png" pageId="33" pageNumber="34">3</figureCitation>
), and in a large body. Thus, to document its identity the new specimen has been illustrated in detail for comparative purposes (
<figureCitation id="1379F793FFDE7D49AFB1D651FD8E9A3F" box="[467,607,512,539]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="31.[189,232,1894,1920]" captionStartId-1="32.[189,232,1930,1956]" captionStartId-2="33.[187,230,1930,1956]" captionTargetBox-0="[189,1398,599,1848]" captionTargetBox-1="[369,1231,950,1895]" captionTargetBox-2="[198,1389,743,1904]" captionTargetId-0="figure@31.[189,1398,599,1848]" captionTargetId-1="figure@32.[359,1236,943,1899]" captionTargetId-2="figure@33.[198,1389,743,1904]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionText-0="Fig. 23. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively." captionText-1="Fig. 24. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ. A–B. Right gonopod, lateral and mesal views, respectively. C. Leg 8. D. Legs of segment 11. Scale bars = 0.5 mm. Abbreviations are explained in the text." captionText-2="Fig. 25. Tylopus baenzigeri Golovatch &amp; Enghoff, 1993, Ƌ, right gonopod. A–B. Mesal and lateral views, respectively. C–F. Distal part, submesal, sublateral, subcaudal and suboral views, respectively. Scale bar = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3837921" figureDoi-1="http://doi.org/10.5281/zenodo.3837923" figureDoi-2="http://doi.org/10.5281/zenodo.3837925" httpUri-0="https://zenodo.org/record/3837921/files/figure.png" httpUri-1="https://zenodo.org/record/3837923/files/figure.png" httpUri-2="https://zenodo.org/record/3837925/files/figure.png" pageId="33" pageNumber="34">Figs 23–25</figureCitation>
). The
<typeStatus id="54F955B4FFDE7D49ACCFD650FD319A3F" box="[685,736,513,539]" pageId="33" pageNumber="34">type</typeStatus>
locality of
<taxonomicName id="4C429095FFDE7D49AD0CD650FB739A3E" authority=", Doi Suthep" authorityName="Doi Suthep" box="[878,1186,512,539]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="baenzigeri">
<emphasis id="B9363704FFDE7D49AD0CD650FBD89A3F" box="[878,1033,513,539]" italics="true" pageId="33" pageNumber="34">T. baenzigeri</emphasis>
, Doi Suthep
</taxonomicName>
National Park, is located quite far away (about
<quantity id="4CBA46F3FFDE7D49AC78D675FDA49A1B" box="[538,629,548,575]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="34" unit="km" value="120.0">120 km</quantity>
) from this new place.
</paragraph>
<paragraph id="8BFDEB16FFDE7D49AEDFD633FEE99A5B" blockId="33.[189,312,610,639]" box="[189,312,610,639]" pageId="33" pageNumber="34">
<emphasis id="B9363704FFDE7D49AEDFD633FEE99A5B" bold="true" box="[189,312,610,639]" italics="true" pageId="33" pageNumber="34">Checklist</emphasis>
</paragraph>
<paragraph id="8BFDEB16FFDE7D49AEDFD6C7FDAE9AF7" blockId="33.[189,1398,662,724]" pageId="33" pageNumber="34">
<tableCitation id="C6C0DEADFFDE7D49AEDFD6C7FEC59A94" box="[189,276,662,688]" captionStart="Table 1" captionStartId="34.[189,258,268,294]" captionText="Table 1. Records of Tylopus species." pageId="33" pageNumber="34">Table 1</tableCitation>
presents a checklist of the known or new
<taxonomicName id="4C429095FFDE7D49AC9DD6C7FC8B9A94" authorityName="Jeekel" authorityYear="1968" box="[767,858,662,688]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B9363704FFDE7D49AC9DD6C7FC8B9A94" box="[767,858,662,688]" italics="true" pageId="33" pageNumber="34">Tylopus</emphasis>
</taxonomicName>
species, all arranged in alphabetic order. Data on their distribution are also provided.
</paragraph>
</subSubSection>
</treatment>
</document>
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