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<document id="B8C19DEF5E1E4B558A5E027D162A969C" ID="10.11646/zootaxa.4238.3.1SLASH10400" ID-CLB-Dataset="36345" ID-DOI="10.11646/zootaxa.4238.3.1" ID-GBIF-Dataset="030203e2-e6a1-4b59-bfa0-0689bf817b08" ID-ISSN="1175-5326" ID-Zenodo-Dep="345756" ID-ZooBank="D123950C-DF2B-45EE-9966-813339A83491" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488787785636" checkinUser="plazi" docAuthor="Krikken, J. &amp; Huijbregts, J." docDate="2017" docId="03E5BB4DE837FFC6AAE4FF0FFDA5FC12" docLanguage="en" docName="zootaxa.4238.3.1SLASH10400.pdf" docOrigin="Zootaxa 4238 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus holosericus Harold 1877" docType="treatment" docVersion="7" lastPageNumber="321" masterDocId="FFDCC335E824FFD2AA73FF98FFDAFFDA" masterDocTitle="Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="334" masterPageNumber="301" pageNumber="320" updateTime="1698437609924" updateUser="ExternalLinkService">
<mods:mods id="AE85D106A414F9438F4AC24CB7FD5C16" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C1C8C083C1A88F1AB5901FD65D94BB0E">
<mods:title id="EBF0B66CA4EA04549A006126168621B7">Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="4E776DD39B328C95C7FD385CF6AE8139" type="personal">
<mods:role id="6DEAE598576D5708AD041F56DB86ADF0">
<mods:roleTerm id="EF439102DF2F8239A1F95E5EFB9EB708">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2496154061C2F38BF4529378F5867C48">Krikken, J.</mods:namePart>
</mods:name>
<mods:name id="A229DA2862B9CAAE4969DB459DD725EE" type="personal">
<mods:role id="498B73A48FCC1A9BEE992F9D5E3BE7EE">
<mods:roleTerm id="B43F50B00FED6B02A8DB2B4A9196A6F1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC032BE517F6463D5F192597EBA5AAEE">Huijbregts, J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="6ACE70E11759C1218A648394A4D48840">text</mods:typeOfResource>
<mods:relatedItem id="B115290FDD9B7B48AFC33037D504CD77" type="host">
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<mods:title id="10F2EBD262134EC05C9D667AACBA322F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A6854F7B343EE426E5541449E3773315">
<mods:date id="9A2087F555663C03416E7DB3D00D2CA8">2017</mods:date>
<mods:detail id="5E28D9A5B459945B9B1BC14B2D0DAF6A" type="volume">
<mods:number id="83C21E78F7A328D7E2A31FA28DCDCEDF">4238</mods:number>
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<mods:detail id="F6017E8542DF5042F3D40533C01547EF" type="issue">
<mods:number id="17DC696DBBDA343A0A88398084E4DAD2">3</mods:number>
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<mods:classification id="B3C7AD56DACA4DB179D6EC5531A175F1">journal article</mods:classification>
<mods:identifier id="9276517D8EEAF518E5E70DCA354CB538" type="CLB-Dataset">36345</mods:identifier>
<mods:identifier id="EECC5E3F00661923898BB9F7EAE2B477" type="DOI">10.11646/zootaxa.4238.3.1</mods:identifier>
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<mods:identifier id="5B8CFA9E7D9632AE634356EDC5CC2360" type="Zenodo-Dep">345756</mods:identifier>
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<treatment id="03E5BB4DE837FFC6AAE4FF0FFDA5FC12" ID-DOI="http://doi.org/10.5281/zenodo.6030188" ID-GBIF-Taxon="127719080" ID-Zenodo-Dep="6030188" LSID="urn:lsid:plazi:treatment:03E5BB4DE837FFC6AAE4FF0FFDA5FC12" httpUri="http://treatment.plazi.org/id/03E5BB4DE837FFC6AAE4FF0FFDA5FC12" lastPageId="20" lastPageNumber="321" pageId="19" pageNumber="320">
<subSubSection id="C35659D0E837FFC1AAE4FF0FFEAEFF09" pageId="19" pageNumber="320" type="nomenclature">
<paragraph id="8BF30A5BE837FFC1AAE4FF0FFDADFF68" blockId="19.[151,631,151,211]" box="[151,631,151,178]" pageId="19" pageNumber="320">
<heading id="D0BBBD37E837FFC1AAE4FF0FFDADFF68" bold="true" box="[151,631,151,178]" fontSize="11" level="1" pageId="19" pageNumber="320" reason="1">
<taxonomicName id="4C4C71D8E837FFC1AAE4FF0FFDADFF68" ID-CoL="7XN45" authority="Harold, 1877" authorityName="Harold" authorityYear="1877" box="[151,631,151,178]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="holosericus">
<emphasis id="B938D649E837FFC1AAE4FF0FFDADFF68" bold="true" box="[151,631,151,178]" pageId="19" pageNumber="320">
<emphasis id="B938D649E837FFC1AAE4FF0FFE17FF6B" bold="true" box="[151,461,151,177]" italics="true" pageId="19" pageNumber="320">Onthophagus holosericus</emphasis>
<bibRefCitation id="EFDD77AAE837FFC1ABA7FF00FDADFF68" author="Harold" box="[468,631,152,178]" pageId="19" pageNumber="320" refString="Harold, E. de (1877) Enumeration des Lamellicornes Coprophages rapportes de l'Archipel Malais, de la Nouvelle Guinee et de l'Australie boreale par M. M. J. Doria, O. Beccari et L. M. D'Albertis. Annali del Museo Civico di Storia Naurale di Genova, 10, 38 - 73." type="journal article" year="1877">Harold, 1877</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FF22FEAEFF09" blockId="19.[151,631,151,211]" box="[151,372,186,211]" pageId="19" pageNumber="320">
(
<figureCitation id="137716DEE837FFC1AAECFF22FF37FF09" box="[159,237,186,211]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[207,1396,233,1891]" captionTargetId="figure@5.[177,1413,195,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Sulawesi Onthophagus species, oblique habitus. 1, O. bisscrutator, holotype male from Toraut. 2, O. scrutator, male from Moramo. 3, O. bongkudai, holotype male from Moajat. 4, O. sangirensis, male from Toraut. 5, O. holosericus, male from Marena, 6, female from Marena. 7, O. sopu, holotype male from Sopu. 8, O. manguliensis, male from Mangoli. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345757/files/figure.png" pageId="19" pageNumber="320">Figs. 5</figureCitation>
,
<figureCitation id="137716DEE837FFC1AA89FF23FE99FF09" box="[250,323,187,211]" captionStart="FIGURES 19 – 24" captionStartId="8.[151,264,1879,1902]" captionTargetBox="[171,1436,193,1838]" captionTargetId="figure@8.[151,1436,193,1858]" captionTargetPageId="8" captionText="FIGURES 19 – 24. Sulawesi Onthophagus species, dorsal, oblique (20 only) forebody. 19, O. sangirensis, male from Manado. 20, O. holosericus, female from Kendari, oblique. 21, O. holosericus, male from Marena. 22, O. holosericus, female from Marena. 23, O. begoniophilus, holotype male major from Ambang. 24, O. begoniophilus, paratype male medium from Ambang. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345760/files/figure.png" pageId="19" pageNumber="320">20–22</figureCitation>
,
<figureCitation id="137716DEE837FFC1AB3DFF22FEB1FF09" box="[334,363,186,211]" captionStart="FIGURES 39 – 46" captionStartId="15.[151,264,1581,1604]" captionTargetBox="[151,1415,230,1560]" captionTargetId="figure@15.[151,1436,193,1560]" captionTargetPageId="15" captionText="FIGURES 39 – 46. Sulawesi Onthophagus body parts. 39, O. bongkudai, head full-face, holotype male from Moajat. 40, O. bongkudai, left elytron dorsal, holotype male from Moajat. 41, O. bongkudai, protibial upper side, holotype male from Moajat. 42, O. bongkudai, metatibial upperside, holotype male from Moajat. 43, O. bongkudai, aedeagus lateral, holotype male from Moajat. 44, O. bongkudai, head full-face, paratype female. 45, O. sangirensis, aedeagus lateral, male from Toraut. 46, O. holosericus, aedeagus lateral, male from Marena. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345763/files/figure.png" pageId="19" pageNumber="320">46</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E837FFC1AAE4FE9AFD19FB3C" pageId="19" pageNumber="320" type="materials_examined">
<paragraph id="8BF30A5BE837FFC1AAE4FE9AFBEDFEE5" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
<materialsCitation id="3B240006E837FFC1AAE4FE9AFBEDFEE5" ID-GBIF-Occurrence="1438340114" collectingDate="1974-07" collectionCode="MSNG" collectorName="O. Beccari &amp; Onth. &amp; Har." country="Indonesia" location="Celebes" pageId="19" pageNumber="320" specimenCount="2" stateProvince="Celebes" typeStatus="lectotype">
<emphasis id="B938D649E837FFC1AAE4FE9AFE54FEC1" bold="true" box="[151,398,258,283]" pageId="19" pageNumber="320">Material examined.</emphasis>
<typeStatus id="54F7B4F9E837FFC1ABD1FE9BFDC8FEC1" box="[418,530,259,283]" pageId="19" pageNumber="320" type="lectotype">Lectotype</typeStatus>
male,
<emphasis id="B938D649E837FFC1A80BFE9AFC9BFEC1" bold="true" box="[632,833,258,283]" pageId="19" pageNumber="320">here designated</emphasis>
(
<collectionCode id="ED5D929EE837FFC1A92EFE9BFC6EFEC1" box="[861,948,259,283]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="19" pageNumber="320">MSNG</collectionCode>
), from
<collectingCountry id="F35B4ACBE837FFC1AE6DFE9AFB54FEC1" box="[1054,1166,258,283]" name="Indonesia" pageId="19" pageNumber="320">Indonesia</collectingCountry>
, labelled: &quot;
<collectingRegion id="4988C4B9E837FFC1AF57FE9AFA5AFEC1" box="[1316,1408,258,283]" country="Indonesia" name="Sulawesi Utara" pageId="19" pageNumber="320">Celebes</collectingRegion>
\ Kandari.
<collectingDate id="EFB6D573E837FFC1AA8AFEBEFE98FEE5" box="[249,322,294,319]" pageId="19" pageNumber="320" value="1974-07">
<date id="FFF22C9BE837FFC1AA8AFEBEFE98FEE5" box="[249,322,294,319]" pageId="19" pageNumber="320" value="1974-07">VII.74</date>
</collectingDate>
\
<collectorName id="26B96F8DE837FFC1AB2BFEBEFE14FEE5" box="[344,462,294,319]" pageId="19" pageNumber="320">O.Beccari</collectorName>
&quot;, &quot;
<typeStatus id="54F7B4F9E837FFC1AB99FEBEFDE8FEE5" box="[490,562,294,319]" pageId="19" pageNumber="320">Typus</typeStatus>
&quot;(red print), &quot;
<collectorName id="26B96F8DE837FFC1A8B5FEBEFCD2FEE5" box="[710,776,294,319]" pageId="19" pageNumber="320">Onth.</collectorName>
\
<taxonomicName id="4C4C71D8E837FFC1A96EFEBEFC46FEE5" box="[797,924,294,319]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="holosericus">holosericus</taxonomicName>
\
<collectorName id="26B96F8DE837FFC1A9C1FEBFFC39FEE4" box="[946,995,295,318]" pageId="19" pageNumber="320">Har.</collectorName>
typo!&quot;.
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAB4FED2FD6AFEB9" blockId="19.[151,1436,258,2011]" box="[199,688,330,355]" pageId="19" pageNumber="320">
The
<specimenCount id="9D4AC1D2E837FFC1AA8AFED3FE4BFEB9" box="[249,401,330,355]" pageId="19" pageNumber="320" type="generic">28 specimens</specimenCount>
examined are as follows:
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FEF7FD65FE5D" blockId="19.[151,1436,258,2011]" box="[151,703,367,391]" pageId="19" pageNumber="320">
<materialsCitation id="3B240006E837FFC1AAE4FEF7FD65FE5D" ID-GBIF-Occurrence="1438340103" box="[151,703,367,391]" location="CENTRAL SULAWESI" pageId="19" pageNumber="320" specimenCount="1" stateProvince="CENTRAL SULAWESI">
<collectingRegion id="4988C4B9E837FFC1AAE4FEF7FE75FE5D" box="[151,431,367,391]" country="Indonesia" name="Sulawesi Tengah" pageId="19" pageNumber="320">CENTRAL SULAWESI</collectingRegion>
: BANGGAI DISTRICT
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FE0BFDB3FE14" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
- Matanyo Forest, N of Kayutanyo,
<quantity id="4CB4A7BEE837FFC1A811FE0BFD6AFE70" box="[610,688,403,427]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="19" pageNumber="320" unit="m" value="170.0">170 m</quantity>
,
<date id="FFF22C9BE837FFC1A8B2FE0AFCB6FE71" box="[705,876,402,427]" pageId="19" pageNumber="320" value="1989-10-30" valueMax="1989-11-04" valueMin="1989-10-30">30.x–4.xi.1989</date>
, Krikken &amp; Van der Blom, sw18b, multistratal evergreen forest, 2 fish traps,
<quantity id="4CB4A7BEE837FFC1AB97FE2FFDD5FE15" box="[484,527,438,463]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="19" pageNumber="320" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE837FFC1A866FE2FFDB3FE14" box="[533,617,439,462]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FE43FCD2FDCC" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
- Matanyo Forest, N of Kayutanyo,
<quantity id="4CB4A7BEE837FFC1A816FE43FD6EFE28" box="[613,692,475,499]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="320" unit="m" value="120.0">120 m</quantity>
,
<date id="FFF22C9BE837FFC1A8B5FE43FCB3FE29" box="[710,873,475,499]" pageId="19" pageNumber="320" value="1989-10-30" valueMax="1989-10-04" valueMin="1989-10-30">30.x–4.x.1989</date>
, Krikken &amp; Van der Blom, sw16a, multistratal evergreen forest, 2 human excrement traps,
<quantity id="4CB4A7BEE837FFC1A8F1FE67FD77FDCD" box="[642,685,510,535]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="19" pageNumber="320" unit="in" value="2.0">2 in</quantity>
<collectionCode id="ED5D929EE837FFC1A8C7FE67FCD2FDCC" box="[692,776,511,534]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FDBBFCD2FD84" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
- Matanyo Forest, N of Kayutanyo,
<quantity id="4CB4A7BEE837FFC1A816FDBBFD6EFDE0" box="[613,692,547,571]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="320" unit="m" value="120.0">120 m</quantity>
,
<date id="FFF22C9BE837FFC1A8B5FDBBFCB3FDE1" box="[710,873,547,571]" pageId="19" pageNumber="320" value="1989-10-30" valueMax="1989-10-04" valueMin="1989-10-30">30.x–4.x.1989</date>
, Krikken &amp; Van der Blom, sw17a, multistratal evergreen forest, 2 human excrement traps,
<quantity id="4CB4A7BEE837FFC1A8F1FDDFFD77FD85" box="[642,685,582,607]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="19" pageNumber="320" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE837FFC1A8C7FDDFFCD2FD84" box="[692,776,583,606]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FDF3FC9EFD59" blockId="19.[151,1436,258,2011]" box="[151,836,619,643]" pageId="19" pageNumber="320">
<collectingRegion id="4988C4B9E837FFC1AAE4FDF3FE75FD59" box="[151,431,619,643]" country="Indonesia" name="Sulawesi Tengah" pageId="19" pageNumber="320">
<collectionCode id="ED5D929EE837FFC1AAE4FDF3FE75FD59" box="[151,431,619,643]" pageId="19" pageNumber="320">CENTRAL SULAWESI</collectionCode>
</collectingRegion>
:
<collectionCode id="ED5D929EE837FFC1ABCAFDF3FC9EFD59" box="[441,836,619,643]" pageId="19" pageNumber="320">LORE LINDU NATIONAL PARK</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FD17FED2FD10" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
- Marena Shelter,
<quantity id="4CB4A7BEE837FFC1ABF3FD16FE13FD7C" box="[384,457,654,679]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="320" unit="m" value="600.0">600 m</quantity>
,
<date id="FFF22C9BE837FFC1ABA6FD16FD5BFD7D" box="[469,641,654,679]" pageId="19" pageNumber="320" value="1985-12-14" valueMax="1985-12-17" valueMin="1985-12-14">14–17.xii.1985</date>
, J. Krikken, pw65a, second-growth forest, 2 human excrement traps,
<quantity id="4CB4A7BEE837FFC1AFFDFD17FF77FD11" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="19" pageNumber="320" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE837FFC1AAC7FD2BFED2FD10" box="[180,264,691,714]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FD4FFED2FCC8" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
- Marena Shelter,
<quantity id="4CB4A7BEE837FFC1ABF3FD4EFE13FD34" box="[384,457,726,751]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="320" unit="m" value="600.0">600 m</quantity>
,
<date id="FFF22C9BE837FFC1ABA6FD4EFD9BFD35" box="[469,577,726,751]" pageId="19" pageNumber="320" value="2014-12-17">14–17.xii</date>
,1985, J. Krikken, pw65b, second-growth forest, 2 human excrement traps,
<quantity id="4CB4A7BEE837FFC1AFFDFD4FFF77FCC9" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="19" pageNumber="320" unit="in" value="3.0">3 in</quantity>
<collectionCode id="ED5D929EE837FFC1AAC7FD63FED2FCC8" box="[180,264,763,786]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FC87FE07FC80" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
- Marena forest (hill crest),
<quantity id="4CB4A7BEE837FFC1AB86FC86FD9AFCEC" box="[501,576,798,823]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="320" unit="m" value="650.0">650 m</quantity>
,
<date id="FFF22C9BE837FFC1A83DFC86FD20FCED" box="[590,762,798,823]" pageId="19" pageNumber="320" value="1985-12-14" valueMax="1985-12-17" valueMin="1985-12-14">14–17.xii.1985</date>
, J. Krikken, pw63a, multistratal evergreen forest, 2 human excrement traps,
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<collectionCode id="ED5D929EE837FFC1ABFAFCDBFE07FC80" box="[393,477,835,858]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FCFFFEBBFC78" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
- Marena forest (hill crest),
<quantity id="4CB4A7BEE837FFC1AB88FCFEFD9DFCA4" box="[507,583,870,895]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="320" unit="m" value="650.0">650 m</quantity>
,
<date id="FFF22C9BE837FFC1A824FCFEFCD4FCA5" box="[599,782,870,895]" pageId="19" pageNumber="320" value="1985-12-14" valueMax="1985-12-17" valueMin="1985-12-14">14–17.xii. 1985</date>
, J. Krikken, pw63b, multistratal evergreen forest, 2 fish traps,
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<collectionCode id="ED5D929EE837FFC1AB7EFC13FEBBFC78" box="[269,353,907,930]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FC37FE07FC30" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
- Marena forest (near river),
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,
<date id="FFF22C9BE837FFC1A827FC36FCDAFC1D" box="[596,768,942,967]" pageId="19" pageNumber="320" value="1985-12-14" valueMax="1985-12-17" valueMin="1985-12-14">14–17.xii.1985</date>
, J. Krikken, pw64a, multistratal evergreen forest, 6 human excrement traps,
<quantity id="4CB4A7BEE837FFC1AB24FC4AFE58FC31" box="[343,386,978,1003]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="19" pageNumber="320" unit="in" value="5.0">5 in</quantity>
<collectionCode id="ED5D929EE837FFC1ABFAFC4BFE07FC30" box="[393,477,979,1002]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FC6FFEC7FBE8" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
- Marena forest (nr river),
<quantity id="4CB4A7BEE837FFC1AB93FC6EFDF0FBD4" box="[480,554,1014,1039]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="320" unit="m" value="600.0">600 m</quantity>
,
<date id="FFF22C9BE837FFC1A845FC6EFD38FBD5" box="[566,738,1014,1039]" pageId="19" pageNumber="320" value="1985-12-14" valueMax="1985-12-17" valueMin="1985-12-14">14–17.xii.1985</date>
, J. Krikken, pw64b, multistratal evergreen forest, 6 fish traps,
<quantity id="4CB4A7BEE837FFC1AAE4FB82FF18FBE9" box="[151,194,1050,1075]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="19" pageNumber="320" unit="in" value="5.0">5 in</quantity>
<collectionCode id="ED5D929EE837FFC1AABAFB83FEC7FBE8" box="[201,285,1051,1074]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FBA7FD04FB8D" blockId="19.[151,1436,258,2011]" box="[151,734,1087,1111]" pageId="19" pageNumber="320">
<materialsCitation id="3B240006E837FFC1AAE4FBA7FD04FB8D" ID-GBIF-Occurrence="1438340111" box="[151,734,1087,1111]" location="SOUTHEAST SULAWESI" pageId="19" pageNumber="320" specimenCount="1" stateProvince="SOUTHEAST SULAWESI">
<collectingRegion id="4988C4B9E837FFC1AAE4FBA7FE0BFB8D" box="[151,465,1087,1111]" country="Indonesia" name="Sulawesi Tenggara" pageId="19" pageNumber="320">SOUTHEAST SULAWESI</collectingRegion>
: KENDARI DISTRICT
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FBFBFCD8FBA1" blockId="19.[151,1436,258,2011]" box="[151,770,1122,1147]" pageId="19" pageNumber="320">
- Kendari,
<date id="FFF22C9BE837FFC1AB5CFBFAFE54FBA1" box="[303,398,1122,1147]" pageId="19" pageNumber="320" value="1874-07">vii.1874</date>
, O. Beccari, lectotype, in MSNG
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FB1FFC2EFB44" blockId="19.[151,1436,258,2011]" box="[151,1012,1158,1183]" pageId="19" pageNumber="320">
- Kendari,
<quantity id="4CB4A7BEE837FFC1AB5CFB1FFEACFB45" box="[303,374,1158,1183]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="320" unit="km" value="40.0">40 km</quantity>
W of Kendari,
<date id="FFF22C9BE837FFC1A856FB1EFD75FB45" box="[549,687,1158,1183]" pageId="19" pageNumber="320" value="1980-08-12">12.viii.1980</date>
, P.R. Deeleman,
<quantity id="4CB4A7BEE837FFC1A91DFB1FFC43FB45" box="[878,921,1158,1183]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="19" pageNumber="320" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE837FFC1A9D3FB1FFC2EFB44" box="[928,1012,1159,1182]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAE4FB33FD19FB3C" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
- Mekara,
<quantity id="4CB4A7BEE837FFC1AB5EFB33FEB3FB19" box="[301,361,1194,1219]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="320" unit="km" value="7.0">7 km</quantity>
S of Lambuya,
<quantity id="4CB4A7BEE837FFC1A859FB33FDB2FB18" box="[554,616,1195,1219]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="320" unit="m" value="70.0">70 m</quantity>
,
<date id="FFF22C9BE837FFC1A804FB32FCC1FB19" box="[631,795,1194,1219]" pageId="19" pageNumber="320" value="1989-11-15" valueMax="1989-11-18" valueMin="1989-11-15">15–18.xi.1989</date>
, Krikken &amp; Van der Blom, sw34a,
<emphasis id="B938D649E837FFC1AEC8FB33FA88FB18" box="[1211,1362,1195,1218]" italics="true" pageId="19" pageNumber="320">
<taxonomicName id="4C4C71D8E837FFC1AEC8FB33FB21FB18" box="[1211,1275,1195,1218]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="19" pageNumber="320" phylum="Tracheophyta" rank="genus">Pinus</taxonomicName>
-Acacia
</emphasis>
forest plantation, 2 human excrement traps,
<quantity id="4CB4A7BEE837FFC1A84EFB57FDB2FB3D" box="[573,616,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="19" pageNumber="320" unit="in" value="2.0">2 in</quantity>
<collectionCode id="ED5D929EE837FFC1A81CFB57FD19FB3C" box="[623,707,1231,1254]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="320">RMNH</collectionCode>
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E837FFC6AAB4FB6AFDA5FC12" lastPageId="20" lastPageNumber="321" pageId="19" pageNumber="320" type="description">
<paragraph id="8BF30A5BE837FFC1AAB4FB6AFB43FA89" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
<emphasis id="B938D649E837FFC1AAB4FB6AFE83FAD1" bold="true" box="[199,345,1266,1291]" pageId="19" pageNumber="320">Description.</emphasis>
Body length
<quantity id="4CB4A7BEE837FFC1A875FB6AFD42FAD0" box="[518,664,1266,1291]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" metricValueMax="9.8" metricValueMin="6.6" pageId="19" pageNumber="320" unit="mm" value="8.2" valueMax="9.8" valueMin="6.6">6.6–9.8 mm</quantity>
. Head and pronotum usually metallic green or cupreous; elytra patterned yellow(-brown) and black, mainly scattered (partly subsymmetric), matt, largely due to microreticulation (
<date id="FFF22C9BE837FFC1AAECFAA3FF14FA89" box="[159,206,1339,1363]" pageId="19" pageNumber="320" value="1980-10">x80</date>
); elytral colour pattern quite variable. Pilosity pale yellow, dorsally long, conspicuous.
</paragraph>
<paragraph id="8BF30A5BE837FFC1AAB4FAC6FC4DF9A9" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">Cephalic margin slightly reflexed; clypeal border generally round; clypeus anteromedially with robust spatuliform horn on reflexed apex (major males), clypeus posterior surface slightly concave, clypeogenal transition virtually continuous at border; genal border widely round; clypeofrontal ridge absent in males; clypeal surface at horn base smooth, impunctate, shiny; posterolateral part of clypeus and anteromedian part of frons rugulatepunctate. Posterior part of frons and vertex impunctate, microreticulate; genal surface with several vague punctures, microreticulate; paramedian protrusions between anterior part of eyes moderately distinct. Vertexal base slightly transversely elevated. Eyes with approximately 8 facet rows across widest point. Minor males have their ornamentation reduced, and may lack clypeal protrusion completely.</paragraph>
<paragraph id="8BF30A5BE837FFC1AAB4F9E7FC77F891" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">Pronotum strongly convex, discal midline not impressed; anterior section of lateral border distinctly sinuate, anterolateral corner slightly reflexed, angle rounded off, posterior section distinctly sinuate; pronotal base medially at most indistinctly marginate; anterior declivity medially topped by sharp triangular, anteriorly directed lobe (dorsal view), anterior side with upright keel; this protuberance reduced in minor males. Anterior declivity of pronotum (largely) impunctate, rest of pronotal derm coarsely (isodiametrically) punctate throughout, punctural setae abundant, semierect, long (many times their punctural diameter).</paragraph>
<paragraph id="8BF30A5BE837FFC1AAB4F8CEFE1BF801" blockId="19.[151,1436,258,2011]" pageId="19" pageNumber="320">
Elytral striae discally well defined, fine, with indistinct punctures; separated by many times their diameter, scarcely crenulate on interstriae; interstriae almost flat; interstriae with irregular rows of seta-bearing punctures, and microreticulation (
<date id="FFF22C9BE837FFC1ABEDF807FE16F86D" box="[414,460,1951,1975]" pageId="19" pageNumber="320" value="1980-10">x80</date>
), matt; many punctures (especially on anterior section of interstriae) anteriorly with contiguous microgranules.
</paragraph>
<paragraph id="8BF30A5BE830FFC6AAB4FF0FFDE4FEC7" blockId="20.[151,1437,151,968]" pageId="20" pageNumber="321">Antennal club brown. Metaventrite medially impunctate; sublaterally with abundant, coarse, mainly setabearing punctures; lateral wings with abundant, seta-bearing, subocellate punctures. Visible abdominal ventrites 1– 6 with transverse row of seta-bearing punctures, setae long. Pygidium matt, black, with sparse, seta-bearing punctures; base with transverse ridge.</paragraph>
<paragraph id="8BF30A5BE830FFC6AAB4FEB0FD66FE52" blockId="20.[151,1437,151,968]" pageId="20" pageNumber="321">Legs black-brown, generally robust. Protibia with 3+(1) larger denticles, spur elongate-acuminate. Mesotibiae and metatibiae and spurs unmodified. Femoral undersides with sparse seta-bearing punctures. Metatarsomere 1 almost as long as metatarsomeres 2–5 combined.</paragraph>
<paragraph id="8BF30A5BE830FFC6AAB4FE0CFE40FE77" blockId="20.[151,1437,151,968]" box="[199,410,404,429]" pageId="20" pageNumber="321">
Aedeagus,
<figureCitation id="137716DEE830FFC6AB30FE0CFE4CFE77" box="[323,406,404,429]" captionStart="FIGURES 39 – 46" captionStartId="15.[151,264,1581,1604]" captionTargetBox="[151,1415,230,1560]" captionTargetId="figure@15.[151,1436,193,1560]" captionTargetPageId="15" captionText="FIGURES 39 – 46. Sulawesi Onthophagus body parts. 39, O. bongkudai, head full-face, holotype male from Moajat. 40, O. bongkudai, left elytron dorsal, holotype male from Moajat. 41, O. bongkudai, protibial upper side, holotype male from Moajat. 42, O. bongkudai, metatibial upperside, holotype male from Moajat. 43, O. bongkudai, aedeagus lateral, holotype male from Moajat. 44, O. bongkudai, head full-face, paratype female. 45, O. sangirensis, aedeagus lateral, male from Toraut. 46, O. holosericus, aedeagus lateral, male from Marena. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345763/files/figure.png" pageId="20" pageNumber="321">Fig. 46</figureCitation>
.
</paragraph>
<paragraph id="8BF30A5BE830FFC6AAB4FE2FFD43FDC2" blockId="20.[151,1437,151,968]" pageId="20" pageNumber="321">
<emphasis id="B938D649E830FFC6AAB4FE2FFDA9FE0A" bold="true" box="[199,627,439,464]" pageId="20" pageNumber="321">Variation and sexual dimorphism.</emphasis>
Variation in colour, size, and secondary sexual characters may be considerable. Minor males, without clypeal projection, may look like females, but lack, apart from other reductions, the transverse clypeofrontal ridge.
</paragraph>
<paragraph id="8BF30A5BE830FFC6AAB4FDBCFC99FD16" blockId="20.[151,1437,151,968]" pageId="20" pageNumber="321">
<emphasis id="B938D649E830FFC6AAB4FDBCFEC6FDE6" box="[199,284,548,572]" italics="true" pageId="20" pageNumber="321">Female</emphasis>
with clypeus entirely densely rugulate, posteriorly to paramedian frontal protrusions; clypeal horn absent; clypeofrontal ridge generally straight, virtually reaching clypeogenal border on either side; frons between anterior part of eyes with two low but distinct paramedian protrusions. Anterior declivity of pronotum with pair of smooth concavities, separated by short, median longitudinal ridge, declivity posteriorly topped by sharp, transverse, variably arcuate crest. Pygidium strongly convex.
</paragraph>
<paragraph id="8BF30A5BE830FFC6AAB4FD4FFD26FCE2" blockId="20.[151,1437,151,968]" pageId="20" pageNumber="321">
<emphasis id="B938D649E830FFC6AAB4FD4FFCE7FD2A" bold="true" box="[199,829,727,752]" pageId="20" pageNumber="321">
<typeStatus id="54F7B4F9E830FFC6AAB4FD4FFEE7FD2A" box="[199,317,727,752]" pageId="20" pageNumber="321" type="lectotype">Lectotype</typeStatus>
designation and spelling of species epithet
</emphasis>
. Harold’s description (1877: 69) is not explicit about the number of specimens he saw, and therefore it seems prudent to stabilize the nomenclature by designating the specimen we studied from the MSNG as the
<typeStatus id="54F7B4F9E830FFC6A8FEFC87FD22FCE2" box="[653,760,799,824]" pageId="20" pageNumber="321" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BF30A5BE830FFC6AAB4FCDDFDA5FC12" blockId="20.[151,1437,151,968]" pageId="20" pageNumber="321">
In his article
<bibRefCitation id="EFDD77AAE830FFC6AB13FCDCFDD8FC86" author="Harold" box="[352,514,836,861]" pageId="20" pageNumber="321" refString="Harold, E. de (1877) Enumeration des Lamellicornes Coprophages rapportes de l'Archipel Malais, de la Nouvelle Guinee et de l'Australie boreale par M. M. J. Doria, O. Beccari et L. M. D'Albertis. Annali del Museo Civico di Storia Naurale di Genova, 10, 38 - 73." type="journal article" year="1877">Harold (1877)</bibRefCitation>
twice spelled the species-group name as “
<taxonomicName id="4C4C71D8E830FFC6A99EFCDCFBB4FC86" box="[1005,1134,836,860]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="holosericus">
<emphasis id="B938D649E830FFC6A99EFCDCFBB4FC86" box="[1005,1134,836,860]" italics="true" pageId="20" pageNumber="321">holosericus</emphasis>
</taxonomicName>
”. This is also the spelling that Harold used on the original
<typeStatus id="54F7B4F9E830FFC6A878FCF0FDE1FC5A" box="[523,571,872,896]" pageId="20" pageNumber="321">type</typeStatus>
label.
<bibRefCitation id="EFDD77AAE830FFC6A8FAFCFFFCA7FC5A" author="Lansberge" box="[649,893,871,896]" pageId="20" pageNumber="321" refString="Lansberge, J. W. van (1883) Revision des Onthophagus de l'archipel indo-neerlandaise, avec description des especes nouvelles. Notes Leyden Museum, 5, 41 - 82." type="journal article" year="1883">Lansberge (1883: 75)</bibRefCitation>
seems to have been the first to use the spelling “
<emphasis id="B938D649E830FFC6AAD0FC14FEEAFC7E" box="[163,304,908,932]" italics="true" pageId="20" pageNumber="321">holosericeus</emphasis>
”, without explicit justification; subsequent authors (including
<bibRefCitation id="EFDD77AAE830FFC6A982FC15FB69FC7E" author="Boucomont" box="[1009,1203,908,932]" pageId="20" pageNumber="321" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel malais (Coleopt.). Annales de la Societe Entomologique de France, 73, 238 - 350." type="journal article" year="1914">Boucomont 1914</bibRefCitation>
and
<bibRefCitation id="EFDD77AAE830FFC6AE99FC14FA4EFC7F" author="Balthasar" box="[1258,1428,908,933]" pageId="20" pageNumber="321" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region (Coleoptera Lamellicornia). Uol. 2. Tschechoslowakische Akademie der Wissenschaften, Prague, 627 pp." type="book" year="1963">Balthasar 1963</bibRefCitation>
) also used this spelling without justification.
</paragraph>
</subSubSection>
</treatment>
</document>
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